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Ver. 25.03
Mar. 6th, 2025

030.0000 Family Pteridaceae E.D.M.Kirchn.; Schul-Bot. 109 (1831)
48 genera, ca. 1350 spp. The family has been split extensively in smaller subgroups, but Smith et al. (2006) and PPG-I argue for keeping them together.
Species listed in database: 1402; hybrids: 68
= Acrostichaceae Mett. ex A.B.Frank; Syn. Pflanzenk., ed. 2, 3: 1458 (1877), nom. rej.
= Actiniopteridaceae Pic.Serm.; Webbia 17: 5 (1962)
= Adiantaceae Newman; Hist. Brit. Ferns 5 (1840) nom. cons. over Parkeriaceae (1825)
= Anopteraceae Doweld; Tent. Syst. Pl. Vasc. (Tracheophyta) 51 (2001)
= Antrophyaceae (Link) Ching; Acta Phytotax. Sin. 16: 11 (1978)
= Ceratopteridaceae Underw.; Native Ferns, ed. 6: 65, 78 (1900)
= Cheilanthaceae B.K.Nayar; Taxon 19: 233 (1970)
= Cryptogrammaceae Pic.Serm.; Webbia 17: 299 (1963)
= Hemionitidaceae Pic.Serm.; Webbia 21: 487 (1966)
= Negripteridaceae Pic.Serm.; Nuovo Giorn. Bot. Ital., n. s. 53: 160 (1946)
= Parkeriaceae Hook.; Exot. Fl. 2: t. 147 (1825), nom. rej. in favor of Adiantaceae
= Platyzomataceae Nakai; Bull. Natl. Sci. Mus., Tokyo, 29: 4 (1950)
= Pteridaceae Reichenb.; Handb. Nat. Pfl.-Syst. 138 (1837)
= Sinopteridaceae Koidz.; Acta Phytotax. Geobot. 3: 50 (1934)
= Taenitidaceae (C.Presl) Pic.Serm.; Webbia 29: 1 (1974)
= Vittariaceae Ching; Sunyatsenia 5: 232 (1940)
030.6017 Genus Oeosporangium Vis.; Atti Reale Ist. Veneto Sci. Lett. Arti, Ser. 3, 12: 656 (1867)
Distribution: Old World. This is the correct name for Cheilosoria and one Old World Cheilanthes clade fide Fraser-Jenkins et al. (2017) and after rejection of Allosorus (2019). The current circumscription is probably inhomogenous and needs further refinement.
Species listed in database: 29; hybrids: 10
= Cheilosoria Trevis.; Atti Reale Ist. Veneto Sci. Lett. Arti, Ser. 5, 3: 579 (1877)
= Parallosorus Keyserl.; Polyp. Herb. Bunge 5 (1873)
species :
Oeosporangium acrosticum (Balb.) L.Sáez & Aymerich; Orsis 31: 33 (2017)
Distribution: Portugal; Spain; Andorra; Gibraltar; Baleares; Croatia; Greece (widespread, but not North, incl. Sporades, Kiklades); Crete (widespread); Morocco; Algeria; Tunisia; Libya; Cape Verde Isl. (Fogo Isl.); ?Ethiopia; Turkmenistan; Tajikistan; East Aegaean Isl. (widespread, incl. Rhodos); Afghanistan; Pakistan (Baluchistan, NW-Pakistan, Chitral, Murree, Hazara); India (Himachal Pradesh, Jammu & Kashmir)
12 ADO BAL GIB POR SPA 13 CRO GRC KRI 20 ALG LBY MOR TUN 21 CVI 32 TKM TZK 34 AFG EAI 40 IND PAK
Remarks: allotetraploid between O. pteridioides and O. persicum, therefore best to be recognized at species level. For a long time known as Cheilanthes pteridioides, while Cheilanthes maderensis in the previous sense is todays O. pteridioides
Synonyms: hidden
= Adiantum odorum (Sw.) DC.; Fl. Franç. [de Candolle & Lamarck], éd. 3, 6: 237 (1815)
= Allosorus acrosticus (Balb.) Christenh.; Willdenowia 42(2): 284 (2012)
= Allosorus pusillus Bernh.; Neues J. Bot. 12: 36 (1806), nom. superfl.
= Cheilanthes acrostica (Balb.) Tod.; Giorn. Sci. Nat. Econ. Palermo 1: 215 (seors 10) (1866)
= Cheilanthes fragrans var. acrostica (Balb.) Fiori; Fl. Ital. Crypt. 5: 267 (1943)
= Cheilanthes fragrans var. paleacea Pamp.; Pl. Tripolitanae 11 (1914)
= Cheilanthes odora Sw.; Syn. Fil. [Swartz] 127, 327 (1806), nom. superfl.
= Cheilanthes pteridioides f. paleacea (Pamp.) Maire & Weiller; Encycl. Biol. 33: 71 (1952)
= Cheilanthes pteridioides subsp. acrostica (Balb.) O.Bolòs, Vigo, Masalles & Ninot; Fl. Man. Paisos Catalans 1213 (1990)
= Hemionitis acrostica (Balb.) Mosyakin; Phytotaxa 373(2): 164-168 (2018)
= Hemionitis atreyu Christenh.; Global Fl. 1(4): 10 (2018), nom. nov. superfl. for Cheilanthes acrostica Tod.
= Oeosporangium pteridioides subsp. acrosticum (Balb.) Fraser-Jenk. & Pariyar; Annot. Checkl. Ind. Pterid. 1: 259 (2016)
= Pteris acrostica Balb.; Elenco 98 (1801)
species :
Oeosporangium belangeri (Bory) Fraser-Jenk.; Annot. Checkl. Ind. Pterid. 1: 255 (2016)
Name: Belangers spleenwort
Distribution: China (Hainan); Nepal; India (Assam State, Meghalaya, Mizoram, Tripura, West Bengal); Bangladesh; Myanmar; Thailand; Laos; Cambodia; Vietnam; peninsular Malaysia; Borneo (Mt. Kinabalu, Sabah); Philippines (Luzon)
36 CHI 40 BAN IND NEP 41 CBD LAO MYA THA VIE 42 BOR MLY PHI
Synonyms: hidden
= Allosorus caespitosus C.Presl; Tent. Pterid. 152 (1836)
= Allosorus cambodiensis (Baker) Kuntze; Revis. Gen. Pl. 2: 806 (1891)
= Cheilanthes belangeri (Bory) C.Chr.; Index Filic. 172 (1905)
= Cheilanthes griffithiana Fée; Mém. Foug. 5 (Gen. Filic.) 157 (1852)
= Cheilanthes malaccensis Fée; Mém. Foug. 5 (Gen. Filic.) 167 (1852)
= Cheilanthes varians Wall. ex Hook.; Sp. Fil. [W.J.Hooker] 2: 89, t. 103 A (1852)
= Cheilosoria belangeri (Bory) Ching & K.H.Shing; Glossary Lycopods Ferns 39 (1982)
= Hemionitis belangeri (Bory) Christenh.; Global Fl. 1(4): 10 (2018)
= Pellaea cambodiensis Baker; Ann. Bot. (Oxford) 5: 213 (1891)
= Pteridella belangeri (Bory) Mett. ex Kuhn; Reisen Ost-Afr. [Decken] 3(3): 16 (1879)
= Pteris belangeri Bory; Voy. Indes Or. [Bélanger] 2: 44 (1833)
= Pteris caespitosa Wall.; Numer. List [Wallich] no. 90 (1829), nomen
= Pteris varians Wall.; Numer. List [Wallich] no. 86 (1829), nomen
species :
Oeosporangium chinense (Baker) Fraser-Jenk.; Annot. Checkl. Ind. Pterid. 2: 451 (2018)
Distribution: China (Chongqing, N-Guangxi, NE-Guizhou, W-Hubei, Sichuan)
36 CHI
Synonyms: hidden
= Cheilanthes chinensis (Baker) Domin; Biblioth. Bot. 86: 133 nota (1913)
= Hemionitis chinensis (Baker) Christenh.; Global Fl. 1(4): 11 (2018)
= Notholaena chinensis Baker; Gard. Chron., Ser. 2, 14: 494 (1880)
species :
Oeosporangium chusanum (Hook.) Fraser-Jenk.; Annot. Checkl. Ind. Pterid. 1: 256 (2016)
Distribution: China (Anhui, Chongqing, Fujian, Gansu, Guangdong, Guangxi, Guizhou, Henan, Hongkong, Hubei, Hunan, Jiangsu, Jiangxi, Shaanxi, Sichuan, Zhejiang); South Korea (Cheju Isl.); Japan; Taiwan; Laos; Vietnam; Sulawesi; Philippines (Luzon)
36 CHI 38 JAP KOR TAI 41 LAO VIE 42 PHI SUL
Synonyms: hidden
= Adiantopsis fordii (Baker) C.Chr.; Index Filic. 22 (1905)
= Cheilanthes bockii Diels; Bot. Jahrb. Syst. 29: 199 (1900)
= Cheilanthes boltonii Copel.; Fragm. Fl. Philipp. 192 (1905)
= Cheilanthes chusana Hook.; Sp. Fil. [W.J.Hooker] 2: 95, t. 106 B (1852)
= Cheilanthes fordii Baker; J. Bot. 17: 304 (1879)
= Cheilanthes mysurensis var. chusana (Hook.) C.Chr.; Index Filic. 293 (1901)
= Cheilanthes mysurensis var. giraldii Christ; Notul. Syst. (Paris) 1(2): 51 (1909)
= Cheilanthes tenuifolia Hook.; Five Months Yang-Tsze [Blakiston] 362 (1862) (non Sw.)
= Cheilosoria chusana (Hook.) Ching; J. Arnold Arbor. 64(1): 19 (1983)
= Hemionitis chusana (Hook.) Christenh.; Global Fl. 1(4): 11 (2018)
species :
Oeosporangium coriaceum (Decne.) Fraser-Jenk. & Pariyar; Annot. Checkl. Ind. Pterid. 1: 264 (2016)
Distribution: SE-Egypt; Niger; Chad; Sudan; Ethiopia; Eritrea; Djibouti; Somalia; Tanzania; SW-Iran; Saudi Arabia; Oman; Yemen (N-Yemen, S-Yemen)
20 EGY 22 NGR 24 CHA DJI ERI ETH SOM SUD 25 TAN 34 IRN 35 OMA SAU YEM
Synonyms: hidden
= Allosorus coriaceus (Decne.) Christenh.; Willdenowia 42(2): 284 (2012)
= Cheilanthes arabica (Decne.) Buchinger; Flora (Regensburg) 303 (1846)
= Cheilanthes coriacea Decne.; Arch. Mus. Hist. Nat. 2: 190 (1841)
= Cheilanthes decaisnei Kunze; Index Sem. (Leipzig) 1850: 23, 307 (1850)
= Hemionitis morla Christenh.; Global Fl. 1(4): 18 (2018), nom. nov. for Cheilanthes coriacea Decne.
= Pellaea abyssinica Fée; Mém. Foug. 6 (Icon. Esp. Nouv.): sub t. 3 (1853)
= Pellaea arabica Fée; Mém. Foug. 5 (Gen. Filic.) 130 (1852)
= Pellaea lomarioides Baker; Bull. Misc. Inform. Kew 1895: 229 (1895)
species :
Oeosporangium delicatulum (Tagawa & K.Iwats.) Fraser-Jenk. & Pariyar; Annot. Checkl. Ind. Pterid. 1: 265 (2016)
Distribution: N-Thailand (Chiang Mai)
41 THA
Conservation Status: DD in Thailand
Synonyms: hidden
= Cheilanthes delicatula Tagawa & K.Iwats.; Acta Phytotax. Geobot. 25: 19 (1971)
= Hemionitis delicatessa Christenh.; Global Fl. 1(4): 12 (2018), nom. nov.
species :
Oeosporangium duthiei (Baker) Fraser-Jenk. & Ph.Wangdi; Pterid. Bhutan 132 (2023)
Distribution: China (SW-Sichuan); Tibet; Nepal; Bhutan; India (Uttarakhand, Sikkim)
36 CHI TIB 40 BHU IND NEP
Conservation Status: IUCN CR for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Aleuritopteris duthiei (Baker) Ching; Bull. Fan Mem. Inst. Biol. Bot., Ser. 2, 1: 271 (1949)
= Cheilanthes duthiei Baker; Ann. Bot. (Oxford) 5: 210 (1891)
= Hemionitis duthiei (Baker) Christenh.; Global Fl. 1(4): 13 (2018)
= Leptolepidium duthiei (Baker) X.C.Zhang & G.M.Zhang; Acta Phytotax. Sin. 40: 286 (2002)
species :
Oeosporangium elegans (Poir.) Fraser-Jenk. & Pariyar; Annot. Checkl. Ind. Pterid. 1: 265 (2016)
Distribution: China (Fujian, Guangdong, Hainan, Zhejiang); India (Andhra Pradesh, Karnataka, Kerala, Odisha, Tamil Nadu); Sri Lanka; Myanmar; Thailand; Laos; Vietnam
36 CHI 40 IND SRL 41 LAO MYA THA VIE
Remarks: not in Taiwan fide Knapp (Ferns of Taiwan, 2011 ff.). E-Asian records for this species frequently refer to the closely related C. chusana
Synonyms: hidden
= Adiantum scandicinum Willd.; Syn. Fil. [Swartz] 326 (1806) (in obs.)
= Cheilanthes cadieri Christ; J. Bot. [Morot] 19: 70 (1905)
= Cheilanthes melanocoma Bory; Voy. Indes Or. [Bélanger] 2(1): 71 (1833)
= Cheilanthes mysurensis Wall. ex Hook.; Sp. Fil. [W.J.Hooker] 2: 94, 100A (1852) (non Asplenium mysurense B. Heyne ex Roth (1821))
= Cheilanthes opposita Kaulf.; Enum. Filic. [Kaulfuss] 211 (1824)
= Cheilanthes swartzii Webb & Berthel.; Hist. Nat. Iles Canaries (Phytogr.) 3: 453 (1849)
= Cheilosoria mysuriensis (Wall. ex Hook.) Ching & K.H.Shing; Fl. Fujianica 1: 84 (1982)
= Hemionitis opposita (Kaulf.) Christenh.; Global Fl. 1(4): 18 (2018)
= Pteris elegans Poir.; Encycl. [J. Lamarck et al.] 5: 718 (1804), non Sw. 1817, nec al., nec Cheilanthes elegans Desv. 1811.
species :
Oeosporangium fragile (Hook.) Fraser-Jenk.; Annot. Checkl. Ind. Pterid. 1: 265 (2016)
Distribution: China (S-Yunnan); Myanmar; N-Thailand
36 CHI 41 MYA THA
Synonyms: hidden
= Cheilanthes fragilis Hook.; Fil. Exot., t. 96 (1859)
= Cheilosoria fragilis (Hook.) Ching & K.H.Shing; Glossary Lycopods Ferns 39 (1982)
= Hemionitis fragilis (Hook.) Christenh.; Global Fl. 1(4): 14 (2018)
species :
Oeosporangium guanchicum (Bolle) Fraser-Jenk. & Pariyar; Annot. Checkl. Ind. Pterid. 1: 265 (2016)
Name: Guanchen-Schuppenfarn (DE); Helecho de roca, helecho chico (ES)
Distribution: Portugal; Spain; France; Corsica; Sardinia; Italy; Greece (only E-Peloponnisos); Morocco; Algeria; Tunisia; Madeira (Madeira Isl.); Canary Isl. (Fuerteventura, Gran Canaria, Tenerife, La Gomera, Hierro, La Palma Isl., Lanzarote); East Aegaean Isl. (Ikaria)
12 COR FRA POR SAR SPA 13 GRC ITA 20 ALG MOR TUN 21 CNY MDR 34 EAI
Synonyms: hidden
= Allosorus guanchicus (Bolle) Christenh.; Willdenowia 42(2): 284 (2012)
= Cheilanthes guanchica Bolle; Bonplandia (Hannover) 7: 107 (1859)
= Cheilanthes pteridioides subsp. guanchica (Bolle) O.Bolòs, Vigo, Masalles & Ninot; Fl. Man. Paisos Catalans 1213 (1990)
= Cheilanthes ×sventenii Benl; Nova Hedwigia 12: 147, t. 28 (1966)
= Hemionitis guanchica (Bolle) Christenh.; Global Fl. 1(4): 15 (2018)
species :
Oeosporangium hancockii (Baker) Fraser-Jenk.; Annot. Checkl. Ind. Pterid. 1: 257 (2016)
Distribution: China (Gansu, Yunnan, W-Guizhou, Sichuan); Tibet; Bhutan
36 CHI TIB 40 BHU
Remarks: not in India
Synonyms: hidden
= Cheilanthes bonatiana Brause; Hedwigia 64: 203, t. 4f (1914)
= Cheilanthes hancockii Baker; Bull. Misc. Inform. Kew 1895: 54 (1895)
= Cheilanthes taliensis Christ; Mém. Soc. Bot. France 1 [Hist. Acrostich.]: 58 (1905)
= Cheilanthes wilsonii Christ; Bull. Acad. Int. Géogr. Bot., Sér. 3, 16: 132 (1906)
= Cheilosoria hancockii (Baker) Ching & K.H.Shing; Glossary Lycopods Ferns 39 (1982)
= Hemionitis hancockii (Baker) Christenh.; Global Fl. 1(4): 15 (2018)
species :
Oeosporangium hispanicum (Mett.) Fraser-Jenk. & Pariyar; Annot. Checkl. Ind. Pterid. 1: 265 (2016)
Distribution: Portugal; Spain; France; Morocco; Algeria; ?Tunisia
12 FRA POR SPA 20 ALG MOR
Remarks: pteridioides agg.
Synonyms: hidden
= Allosorus hispanicus (Mett.) Christenh.; Willdenowia 42(2): 284 (2012)
= Cheilanthes hispanica Mett.; Abh. Senckenberg. Naturf. Ges. 3(1) [Aspl.]: 74 (1859)
= Hemionitis hispanica (Mett.) Christenh.; Global Fl. 1(4): 15 (2018)
species :
Oeosporangium insigne (Ching) Fraser-Jenk.; Annot. Checkl. Ind. Pterid. 1: 265 (2016)
Distribution: China (N-Sichuan); E-Tibet
36 CHI TIB
Synonyms: hidden
= Cheilanthes insignis Ching; Fl. Tsinling. 2: 208 (1974)
= Cheilanthes tenuifolia C.Chr.; (1924) (non Sw. 1806)
= Cheilosoria insignis (Ching) Ching & K.H.Shing; Fl. Xizang. 1: 91 (1983)
= Hemionitis insignis (Ching) Christenh.; Global Fl. 1(4): 16 (2018)
species :
Oeosporangium kuhnii (Milde) Fraser-Jenk.; Annot. Checkl. Ind. Pterid. 1: 261 (2016)
Distribution: Russian Far East (Primorye); China (Gansu, Hebei, Henan, Jilin, Liaoning, Nei Mongol, Shaanxi, Shanxi, Sichuan, Yunnan); Tibet; North Korea; South Korea (incl. Cheju Isl.); Japan
31 RFE 36 CHI TIB 38 JAP KOR NKO
Remarks: not in India
Synonyms: hidden
= Aleuritopteris caesia (Christ) Ching; Hong Kong Naturalist 10: 202 (1941)
= Aleuritopteris caesia var. efarinosa H.S.Kung, Li Bing Zhang & X.S.Guo; Acta Bot. Yunnan. 17(4): 422 (1995)
= Aleuritopteris kuhnii (Milde) Ching; Hong Kong Naturalist 10: 202 (1941)
= Aleuritopteris kuhnii f. efarinosa (Makino) Tagawa; Acta Phytotax. Geobot. 14(6): 191 (1952)
= Aleuritopteris kuhnii var. brandtii (Franch. & Sav.) Tagawa; Acta Phytotax. Geobot. 14(6): 191 (1952)
= Aleuritopteris tenella (Ching & S.K.Wu) Saiki; J. Phytogeogr. Taxon. 32(1): 11 (1984)
= Cheilanthes brandtii Franch. & Sav.; Enum. Pl. Jap. 2: 212, 620 (1877)
= Cheilanthes brandtii var. efarinosa Makino; J. Jap. Bot. 1(7): 26 (1917)
= Cheilanthes caesia Christ; Bull. Acad. Int. Géogr. Bot., Sér. 3, 16: 133 (1906)
= Cheilanthes farinosa var. brandtii (Franch. & Sav.) C.Chr.; Index Filic. 3: 172 (1905)
= Cheilanthes kuhnii f. efarinosa (Makino) Nakaike; J. Nippon Fernist Club 3(Suppl. 2): 40 (2004)
= Cheilanthes kuhnii Milde; Bot. Zeitung (Berlin) 25: 149 (1867)
= Cheilanthes kuhnii var. brandtii (Franch. & Sav.) Tagawa; J. Jap. Bot. 13(3): 184 (1934)
= Cheilanthes kuhnii var. caesia (Christ) C.Chr.; Acta Horti Gothob. 1: 90 (1924)
= Cheilanthes lanceolata C.Chr.; Bot. Gaz. 66: 334 (1913)
= Hemionitis brandtii (Franch. & Sav.) Christenh.; Global Fl. 1(4): 10 (2018)
= Hemionitis caesia (Christ) Christenh.; Global Fl. 1(4): 11 (2018)
= Hemionitis kuhnii (Milde) Christenh.; Global Fl. 1(4): 16 (2018)
= Leptolepidium caesium (Christ) K.H.Shing & S.K.Wu; Acta Bot. Yunnan. 1(1): 117 (1979)
= Leptolepidium kuhnii (Milde) K.H.Shing & S.K.Wu; Acta Bot. Yunnan. 1(1): 117 (1979)
= Leptolepidium kuhnii var. brandtii (Franch. & Savigny) K.H.Shing & S.K.Wu; Acta Bot. Yunnan. 1(1): 118 (1979)
= Leptolepidium tenellum Ching & S.K.Wu; Acta Bot. Yunnan. 1(1): 117 (1979)
species :
Oeosporangium mairei (Brause) Fraser-Jenk.; Annot. Checkl. Ind. Pterid. 1: 265 (2016)
Distribution: China (Guizhou, Hunan, Shaanxi, SW-Sichuan, Yunnan)
36 CHI
Synonyms: hidden
= Cheilanthes brausei Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 122 (2008), nom. nov.
= Hemionitis mairei (Brause) Christenh.; Global Fl. 1(4): 17 (2018)
= Mildella mairei (Brause) C.C.Hall & Lellinger; Amer. Fern J. 57: 133 (1967)
= Pellaea mairei Brause; Hedwigia 64: 201, t. 4 C (1914), non Cheilanthes mairei Brause
species :
Oeosporangium patulum (Baker) Fraser-Jenk.; Annot. Checkl. Ind. Pterid. 1: 265 (2016)
Distribution: China (Chongqing, Guangxi, Guizhou, W-Hubei)
36 CHI
Synonyms: hidden
= Cheilanthes patula Baker; J. Bot. 26: 225 (1888)
= Cheilosoria patula (Baker) P.S.Wang & X.Y.Wang; Pterid. Fl. Guizhou 188 (2001)
= Hemionitis patula (Baker) Christenh.; Global Fl. 1(4): 19 (2018)
= Pellaea patula (Baker) Ching; Acta Phytotax. Sin. 10: 302 (1965)
species :
Oeosporangium pauperculum (Christ) Fraser-Jenk. & Pariyar; Annot. Checkl. Ind. Pterid. 1: 265 (2016)
Distribution: China (W-Sichuan)
36 CHI
Synonyms: hidden
= Cheilanthes christii Fraser-Jenk. & Yatsk.; Indian Fern J. 27(1-2): 213 (2011), nom. nov.
= Cheilanthes paupercula (Christ) Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 122 (2008)
= Hemionitis christii (Fraser-Jenk. & Yatsk.) Christenh.; Global Fl. 1(4): 11 (2018)
= Mildella paupercula (Christ) C.C.Hall & Lellinger; Amer. Fern J. 57: 131 (1967)
= Pellaea paupercula (Christ) Ching; Bull. Fan Mem. Inst. Biol. Bot. 2: 203 (1931)
= Pteris paupercula Christ; Bull. Acad. Int. Géogr. Bot., Sér. 3, 16: 131 (1906)
species :
Oeosporangium persicum (Bory) Vis.; Mem. Reale Ist. Veneto Sci.16: 44 (1872)
Distribution: Italy; Croatia; Bosnia & Hercegovina; Montenegro; North Macedonia; Albania; Bulgaria; Greece (scattered mainland); Crete (rare); Crimea; ?Algeria; Kazakhstan; Turkmenistan; Uzbekistan; Kyrgyzstan; Tajikistan; Northern Caucasus; Georgia [Caucasus]; Armenia; Azerbaijan; East Aegaean Isl. (Lesvos, Chios, Ikaria, Samos, Simi); Turkey (Anatolia); NE-Iraq; Iran; Afghanistan; Pakistan (Chitral, Baluchistan, Murree, Gilgit, Baltistan); India (Himachal Pradesh, Jammu & Kashmir)
13 ALB BIH BUL CRO GRC ITA KRI MNT NMA 14 KRY 32 KAZ KGZ TKM TZK UZB 33 ARM AZE GEO NCS 34 AFG EAI IRN IRQ TUR 40 IND PAK
Remarks: pteridioides agg.
Synonyms: hidden
= Acrostichum microphyllum Bertol.; Propag. Agric. 6: 343 (1856)
= Allosorus persicus (Bory) Christenh.; Willdenowia 42(2): 284 (2012)
= Cheilanthes fimbriata Vis.; Fl. Dalmat. 1: 42, t. 1, f. 1 (1842)
= Cheilanthes persica (Bory) Mett.; Fil. Afr. [Kuhn]: 73 (1868)
= Cheilanthes szovitsii Fisch. & C.A.Mey.; Bull. Soc. Imp. Naturalistes Moscou 6: 260 (1833), nom. nud.
= Hemionitis persica (Bory) Christenh.; Global Fl. 1(4): 19 (2018)
= Myriopteris szovitzii (Fisch. & C.A.Mey.) J.Sm.; Hist. Fil. 281 (1875)
= Notholaena microphylla Bolle; Sitzungsber. Ges. Naturf. Freunde Berlin (1858)
= Notholaena persica (Bory) Bory; Voy. Indes Or. [Bélanger] 2(1): 23 (1833)
= Notholaena szovitsii (Fisch. & C.A.Mey.) Keyserl.; Polyp. Herb. Bunge 28 (1873)
= Oeosporangium fimbriatum (Vis.) Vis.; (1867)
= Oeosporangium szovitsii (Fisch. & C.A.Mey.) Vis.; Atti Reale Ist. Veneto Sci. Lett. Arti, Ser. 3, 12: 656 (1867)
species :
Oeosporangium pteridioides (Reichard) Fraser-Jenk. & Pariyar; Annot. Checkl. Ind. Pterid. 1: 258 (2016)
Name: Madeira-Schuppenfarn (DE); Culantrillo de risco (ES)
Distribution: Portugal; Spain; France; +Monaco; Corsica; Sardinia; Italy; Sicily; Pantelleria; Croatia; Bosnia & Hercegovina; Montenegro; North Macedonia; Albania; Bulgaria; Greece (Kiklades, E-Peloponnisos); Crete; Morocco; S-Algeria (Hoggar); Azores (Pico); Madeira (Madeira Isl.); Canary Isl. (Lanzarote, Fuerteventura, Gran Canaria, Tenerife, La Gomera, Hierro, La Palma Isl.); N-Chad (Tibesti); Ethiopia; Djibouti; Turkmenistan; Northern Caucasus; Armenia; Azerbaijan; East Aegaean Isl. (Karpathos, Dodecanese, Lesvos); Turkey (Anatolia); Iraq; Iran; Cyprus; Lebanon; Syria; Israel; Palestine; Sinai peninsula; Jordania; Saudi Arabia; United Arab Emirates; Oman; Yemen (N-Yemen)
12 COR FRA MCO POR SAR SPA 13 ALB BIH BUL CRO GRC ITA KRI MNT NMA PTL SIC 20 ALG MOR 21 AZO CNY MDR 24 CHA DJI ETH 32 TKM 33 ARM AZE NCS 34 CYP EAI IRN IRQ ISR JOR LEB PAL SIN SYR TUR 35 OMA SAU UAE YEM
Remarks: for a long time known as Cheilanthes maderensis, while Cheilanthes pteridioides in the previous sense is todays O. acrosticum
Synonyms: hidden
= Adiantum fragrans (L.) L.f.; Suppl. Pl. 447 (1782), nom. illeg.
= Adiantum odoratum Poir.; Encycl. [J. Lamarck et al.] Suppl. 1: 142 (1810)
= Adiantum pusillum All. ex Pollini; Fl. Veron. 3: 293 (1824)
= Adiantum pusillum Willd. ex Bernh.; Neues J. Bot. 1(2): 36 (1806)
= Adiantum suaveolens Poir.; Encycl. [J. Lamarck et al.] Suppl. 1: 142 (1810)
= Allosorus fragrans (L.f.) Farw.; Amer. Midl. Naturalist 12: 284 (1931)
= Allosorus pteridioides (Reichard) Christenh.; Willdenowia 42(2): 284 (2012)
= Cheilanthes fragrans (L.) Sw.; Syn. Fil. [Swartz] 127 (1806) (nom. illeg.)
= Cheilanthes fragrans (L.) Webb ex Benth.; Hist. Nat. Iles Canaries (Phytogr.) 3: 452 (1847)
= Cheilanthes fragrans subsp. maderensis (Lowe) Benl; Mitt. Bot. Staatssamml. München 6: 33 (1966)
= Cheilanthes maderensis Lowe; J. Bot. [Hooker] 1: 26 (1834) (nomen), ex Trans. Cambridge Philos. Soc. 6: 528 (1838)
= Cheilanthes microphylla Buch; Abh. Königl. Akad. Wiss. Berlin 1816-1817: 360 (1819), nom. illeg.
= Cheilanthes odora proles maderensis (Lowe) Rouy; Fl. France [Rouy & Foucaud] 14: 389 (1913)
= Cheilanthes pteridiodes var. maderensis (Lowe) Trab.; Mém. Soc. Sci. Nat. Maroc. 8(1): 164 (1924)
= Cheilanthes pteridioides (Reichard) C.Chr.; Index Filic. 178 (1905)
= Cheilanthes pteridioides f. maderensis (Lowe) Maire & Weiller; Encycl. Biol. 33: 71 (1952)
= Cheilanthes pteridioides subsp. maderensis (Lowe) O.Bolòs, Vigo, Masalles & Ninot; Fl. Man. Paisos Catalans 1213 (1990)
= Cheilanthes pteridioides var. maderensis (Lowe) Trab.; Mém. Soc. Sci. Nat. Maroc 8(1): 164 (1924)
= Cheilanthes squamosa Heer & Regel; Linnaea 24: 174 (1851)
= Cheilanthes suaveolens Sw.; Syn. Fil. [Swartz] 127 (1806), nom. superfl.
= Hemionitis maderensis (Lowe) Christenh.; Global Fl. 1(4): 17 (2018)
= Hemionitis pteridioides (Reichard) Christenh.; Global Fl. 1(4): 19 (2018)
= Hemionitis quezelii (Tardieu) Christenh.; Global Fl. 1(4): 20 (2018)
= Negripteris quezelii Tardieu; Mission Bot. Tibesti 92, t. 1(1-5) (1958)
= Polypodium fragrans L.; Mant. Pl. 2: 307 (1771) (non L. 1753)
= Polypodium pteridioides Reichard; Syst. Pl., ed. 4 [Willdenow] 4: 424 (1780), nom. nov. pro Polypodium fragrans L. (1771), non L. (1753), nec P. pterioides Lam. (1779)
= Polypodium pteryoides Bory; Ess. Isl. Fortunées 311 (1803)
= Pteris fragrans (L.) Lag.; Anales Ci. Nat. 5: 158 (1802)
species :
Oeosporangium pulchellum (Bory ex Willd.) Fraser-Jenk. & Pariyar; Annot. Checkl. Ind. Pterid. 1: 264 (2016)
Name: Zierlicher Schuppenfarn (DE); Doradilla fina (ES)
Distribution: Madeira (Madeira Isl.); Canary Isl. (Gran Canaria, Tenerife, La Gomera, Hierro, La Palma Isl.)
21 CNY MDR
Synonyms: hidden
= Allosorus fragilis Christenh.; Willdenowia 42(2): 284 (2012), nom. nov.
= Allosorus pulchellus (Bory ex Willd.) C.Presl; Tent. Pterid. 152 (1836)
= Cheilanthes pulchella Bory ex Willd.; Sp. Pl., ed. 4 [Willdenow] 5(1): 456 (1810)
= Hemionitis pulchella (Bory ex Willd.) Christenh.; Global Fl. 1(4): 20 (2018)
species :
Oeosporangium smithii (C.Chr.) Fraser-Jenk. & Pariyar; Annot. Checkl. Ind. Pterid. 1: 265 (2016)
Distribution: China (W-Sichuan, NW-Yunnan)
36 CHI
Synonyms: hidden
= Cheilanthes smithii (C.Chr.) R.M.Tryon; Amer. Fern J. 76(4): 185 (1986)
= Hemionitis christensenii Christenh.; Global Fl. 1(4): 11 (2018), nom. nov. for Pellaea smithii C. Chr.
= Mildella smithii (C.Chr.) C.C.Hall & Lellinger; Amer. Fern J. 57: 132 (1967)
= Pellaea smithii C.Chr.; Acta Horti Gothob. 1: 84, t. 18 (1924)
species :
Oeosporangium stramineum (Ching) Fraser-Jenk.; Annot. Checkl. Ind. Pterid. 1: 260 (2016)
Distribution: China (Gansu, S-Qinghai, Xinjiang); Tibet; Bhutan; India (Sikkim)
36 CHI TIB 40 BHU IND
Synonyms: hidden
= Cheilanthes tibetica Fraser-Jenk. & Wangdi; Fern Gaz. 18(5): 218 (2009), nom. nov. for Pellaea straminea Ching
= Hemionitis straminea (Ching) Christenh.; Global Fl. 1(4): 21 (2018)
= Hemionitis tibetica (Fraser-Jenk. & Wangdi) Christenh.; Global Fl. 1(4): 22 (2018), nom. superfl.
= Mildella straminea (Ching) C.C.Hall & Lellinger; Amer. Fern J. 57: 130 (1967)
= Pellaea straminea Ching; Bull. Fan Mem. Inst. Biol. Bot. 2: 203, t. 17 (1931), non C. straminea Brause 1913
= Pellaea straminea var. tibetica Ching; Fl. Xizang. 1: 91 (1983)
species :
Oeosporangium subvillosum (Hook.) Fraser-Jenk. & Pariyar; Annot. Checkl. Ind. Pterid. 1: 260 (2016)
Distribution: China (Guizhou, Sichuan, Yunnan); Tibet; Pakistan (Hazara); Nepal; Bhutan; India (Jammu & Kashmir, Himachal Pradesh, Sikkim, Uttarakhand, West Bengal); Myanmar
36 CHI TIB 40 BHU IND NEP PAK 41 MYA
Synonyms: hidden
= Aleuritopteris subvillosa (Hook.) Ching; Hong Kong Naturalist 10: 203 (1941)
= Aleuritopteris subvillosa var. dilatata (Brause) H.S.Kung; Fl. Sichuan. 6: 256 (1988)
= Aleuritopteris subvillosa var. tibetica (Ching & S.K.Wu) H.S.Kung; Fl. Sichuan. 6: 258 (1988)
= Allosorus subvillosus (Hook.) Farw.; Amer. Midl. Naturalist 12: 284 (1931)
= Cheilanthes subvillosa Hook.; Sp. Fil. [W.J.Hooker] 2: 87, t. 98B (1852)
= Cheilanthes yunnanensis Brause; Hedwigia 54(2): 204 (1914)
= Cheilanthes yunnanensis var. dilatata Brause; Hedwigia 54(2): 205, t. 4, f. G (1913)
= Hemionitis subvillosa (Hook.) Christenh.; Global Fl. 1(4): 21 (2018)
= Leptolepidium subvillosum (Hook.) K.H.Shing & S.K.Wu; Acta Bot. Yunnan. 1(1): 116 (1979)
= Leptolepidium subvillosum var. dilatatum (Brause) K.H.Shing & S.K.Wu; Acta Bot. Yunnan. 1(1): 116 (1979)
= Leptolepidium subvillosum var. tibeticum Ching & S.K.Wu; Acta Bot. Yunnan. 1(1): 116 (1979)
species :
Oeosporangium thwaitesii (Mett.) Fraser-Jenk.; Annot. Checkl. Ind. Pterid. 1: 262 (2016)
Distribution: S-India (Tamil Nadu); Sri Lanka
40 IND SRL
Conservation Status: IUCN CR for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Aleuritopteris thwaitesii (Mett.) Saiki; J. Phytogeogr. Taxon. 32(1): 11 (1984)
= Cheilanthes keralensis N.C.Nair & S.R.Ghosh; J. Indian Bot. Soc. 55 52 (1976)
= Cheilanthes laxa T.Moore; Index Fil. [T. Moore] 245 (1861), nomen
= Cheilanthes thwaitesii Mett.; Linnaea 36: 82 (1869)
= Hemionitis keralensis (N.C.Nair & S.R.Ghosh) Christenh.; Global Fl. 1(4): 16 (2018)
= Hemionitis thwaitesii (Mett.) Christenh.; Global Fl. 1(4): 22 (2018)
species :
Oeosporangium tinaei (Tod.) Fraser-Jenk.; Annot. Checkl. Ind. Pterid. 1: 265 (2016)
Name: Korsischer Schuppenfarn (DE); Helecho de risco (ES)
Distribution: Portugal; Spain; France; Corsica; Sardinia; Italy; Sicily; Morocco; Algeria; Tunisia; Madeira (Madeira Isl.); Canary Isl. (Tenerife); Turkey (Anatolia)
12 COR FRA POR SAR SPA 13 ITA SIC 20 ALG MOR TUN 21 CNY MDR 34 TUR
Synonyms: hidden
= Allosorus tinaei (Tod.) Christenh.; Willdenowia 42(2): 284 (2012)
= Cheilanthes corsica Reichst. & Vida; Candollea 28: 85 (1973)
= Cheilanthes pteridioides subsp. tinaei (Tod.) O.Bolòs, Vigo, Masalles & Ninot; Fl. Man. Paisos Catalans 1213 (1990)
= Cheilanthes tinaei Tod.; Giorn. Sci. Nat. Econ. Palermo 1: 217 (seors. 12) (1866)
= Hemionitis tinaei (Tod.) Christenh.; Global Fl. 1(4): 22 (2018)
species :
Oeosporangium tirajanae (T.S.Velázquez) comb.ined.
Name: Tirajana-Schuppenfarn (DE)
Distribution: Canary Isl. (Gran Canaria)
21 CNY
Synonyms: hidden
= Cheilanthes tirajanae T.S.Velázquez; Bot. Macaronés. 28: 22 (2013)
species :
Oeosporangium trichophyllum (Baker) Fraser-Jenk.; Annot. Checkl. Ind. Pterid. 1: 262 (2016)
Distribution: China (W-Yunnan, W-Sichuan); E-Tibet; ?India (?Sikkim)
36 CHI TIB
Synonyms: hidden
= Aleuritopteris delavayi (Baker) B.K.Nayar & Kaur; Companion Handb. Ferns Brit. India [Nayar & Kaur] 26 (1974)
= Cheilanthes delavayi Baker; Ann. Bot. (Oxford) 5: 211 (1891) (non Gymnogramma delavayi Baker)
= Cheilanthes trichophylla Baker; Ann. Bot. (Oxford) 5: 211 (1891)
= Cheilanthes undulata C.Hope & C.H.Wright; Gard. Chron., Ser. 3, 34: 397 (1903)
= Hemionitis trichophylla (Baker) Christenh.; Global Fl. 1(4): 22 (2018)
= Pellaea trichophylla (Baker) Ching; Acta Phytotax. Sin. 10: 302 (1965)
species :
Oeosporangium velutinum (Tardieu & C.Chr.) Fraser-Jenk.; ined.
Distribution: SW-Thailand; Vietnam
41 THA VIE
Synonyms: hidden
= Cheilanthes velutina (Tardieu & C.Chr.) Fraser-Jenk.; New Sp. Syndr. Indian Pteridol. 84 (1997)
= Notholaena velutina Tardieu & C.Chr.; Notul. Syst. (Paris) 6: 167, t. 1(5-7) (1938)
= Hemionitis velutina (Tardieu & C.Chr.) Christenh.; Global Fl. 1(4): 22 (2018)
species :
Oeosporangium yunnanense (Ching) Fraser-Jenk. & Pariyar; Annot. Checkl. Ind. Pterid. 1: 264 (2016)
Distribution: China (Sichuan, Yunnan); Bhutan; India (?Sikkim)
36 CHI 40 BHU IND
Conservation Status: IUCN CR for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Cheilanthes bhutanica Fraser-Jenk. & Wangdi; Fern Gaz. 18(5): 220 (2009), nom. nov.
= Hemionitis bhutanica (Fraser-Jenk. & Wangdi) Christenh.; Global Fl. 1(4): 10 (2018), nom. superfl.
= Hemionitis yunnanensis (Ching) Christenh.; Global Fl. 1(4): 23 (2018)
= Pellaea yunnanensis Ching; Acta Phytotax. Sin. 20(2): 235 (1982), non Brause (1913)
species :
Oeosporangium ×duriense (Mendonça & Vasc.) comb.ined.
Distribution: Portugal; Spain
12 POR SPA
Remarks: Oeosporangium hispanicum x pteridioides
Synonyms: hidden
= Allosorus ×prototinaei (Rasbach, Reichst. & Schneller) Rumsey; Fern Gaz. 19(7): 277 (2014)
= Cheilanthes ×duriensis Mendonça & Vasc.; Anales Inst. Vinho do Porto 15(4): 47 (1956)
= Cheilanthes ×prototinaei Rasbach, Reichst. & Schneller; Webbia 37(1): 45 (1983)
= Hemionitis ×prototinaei (Rasbach, Reichst. & Schneller) F.M.Vázquez; Folia Bot. Extremadur. 15: 149 (2021)
= Oeosporangium ×prototinaei (Rasbach, Reichst. & Schneller) Arana & Mor.-Saiz; Phytotaxa 433(3): 192 (2020)
species :
Oeosporangium ×ibericum (Rasbach & Reichst.) Arana & Mor.-Saiz; Phytotaxa 433(3): 191 (2020)
Distribution: Spain
12 SPA
Remarks: Oeosporangium hispanicum x tinaei
Synonyms: hidden
= Cheilanthes ×iberica Rasbach & Reichst.; Webbia 35(2): 264 (1982)
= Allosorus ×ibericus (Rasbach & Reichst.) Rumsey; Fern Gaz. 19(7): 277 (2014)
= Hemionitis ×iberica (Rasbach & Reichst.) F.M.Vázquez; Folia Bot. Extremadur. 15: 149 (2021)
species :
Oeosporangium ×insulare (Rasbach & Reichst.) Arana & Mor.-Saiz; Phytotaxa 433(3): 191 (2020)
Distribution: Spain; Corsica; Sardinia; Canary Isl. (Tenerife)
12 COR SAR SPA 21 CNY
Remarks: Oeosporangium guanchicum x tinaei
Synonyms: hidden
= Cheilanthes ×insularis Rasbach & Reichst.; Webbia 35(2): 266 (1982)
= Allosorus ×insularis (Rasbach & Reichst.) Rumsey; Fern Gaz. 19(7): 277 (2014)
= Hemionitis ×insularis (Rasbach & Reichst.) F.M.Vázquez; Folia Bot. Extremadur. 15: 149 (2021)
species :
Oeosporangium ×kochianum (Rasbach, Reichst. & Schneller) L.Sáez & Aymerich; Orsis 31: 33 (2017), as ´kochianus´
Distribution: Spain
12 SPA
Remarks: Oeosporangium pteridioides x tinaei
Synonyms: hidden
= Cheilanthes ×kochiana Rasbach, Reichst. & Schneller; Webbia 37(1): 48 (1983)
= Allosorus ×kochianus (Rasbach, Reichst. & Schneller) Rumsey; Fern Gaz. 19(7): 277 (2014)
= Allosurus ×kochianus (Rasbach, Reichst. & Schneller) L.Sáez & Aymerich; Orsis 29: 26 (2015), isonym
= Hemionitis ×kochiana (Rasbach, Reichst. & Schneller) F.M.Vázquez; Folia Bot. Extremadur. 15: 149 (2021)
species :
Oeosporangium ×kurdicum (Rasbach & Reichst.) Fraser-Jenk. & Pariyar; Annot. Checkl. Ind. Pterid. 1: 266 (2016)
Distribution: NE-Iraq; E-Afghanistan
34 AFG IRQ
Remarks: Oeosporangium acrosticum x persicum
Synonyms: hidden
= Allosorus ×kurdicus (Rasbach & Reichst.) Rumsey; Fern Gaz. 19(7): 277 (2014)
= Cheilanthes ×kurdica Rasbach & Reichst.; Webbia 37(1): 58 (1983)
species :
Oeosporangium ×malacitense (Rasbach & Reichst.) Arana & Mor.-Saiz; Phytotaxa 433(3): 191 (2020)
Distribution: Spain
12 SPA
Remarks: Oeosporangium acrostichum x guanchicum
Synonyms: hidden
= Cheilanthes ×malacitensis Rasbach & Reichst.; Webbia 35(2): 269 (1982)
= Allosorus ×malacitensis (Rasbach & Reichst.) Rumsey; Fern Gaz. 19(7): 277 (2014)
= Hemionitis ×malacitensis (Rasbach & Reichst.) F.M.Vázquez; Folia Bot. Extremadur. 15: 149 (2021)
species :
Oeosporangium ×marchettianum (Rasbach, Reichst. & Schneller) Arana & Mor.-Saiz; Phytotaxa 433(3): 191 (2020)
Distribution: Italy
13 ITA
Remarks: Oeosporangium acrosticum x pteridioides
Synonyms: hidden
= Allosorus ×marchettianus (Rasbach, Reichst. & Schneller) Rumsey; Fern Gaz. 19(7): 277 (2014)
= Cheilanthes ×marchettiana Rasbach, Reichst. & Schneller; Webbia 37(1): 55 (1983)
= Hemionitis ×marchettiana (Rasbach, Reichst. & Schneller) F.M.Vázquez; Folia Bot. Extremadur. 15: 149 (2021)
= Oeosporangium pteridioides subsp. reichsteinii Fraser-Jenk. & Pariyar; Annot. Checkl. Ind. Pterid. 1: 266 (2016) [nothosubsp.]
species :
Oeosporangium ×meridionale (F.M.Vázquez) comb.ined.
Distribution: Spain
12 SPA
Remarks: Oeosporangium acrostichum x tinaei
Synonyms: hidden
= Hemionitis ×meridionalis F.M.Vázquez; Folia Bot. Extremadur. 15: 150 (2021)
species :
Oeosporangium ×teneriffae (Rasbach & Reichst.) Arana & Mor.-Saiz; Phytotaxa 433(3): 192 (2020)
Distribution: Canary Isl. (Tenerife)
21 CNY
Remarks: Oeosporangium pteridioides x pulchellum
Synonyms: hidden
= Cheilanthes ×teneriffae Rasbach & Reichst.; Webbia 35(2): 262 (1982)
= Allosorus ×teneriffae (Rasbach & Reichst.) Rumsey; Fern Gaz. 19(7): 277 (2014)
species :
Oeosporangium ×tolocense (Rasbach, Reichst. & Schneller) Arana & Mor.-Saiz; Phytotaxa 433(3): 192 (2020)
Distribution: Spain
12 SPA
Remarks: Oeosporangium guanchicum x pteridioides
Synonyms: hidden
= Cheilanthes ×tolocensis Rasbach, Reichst. & Schneller; Webbia 37(1): 52 (1983)
= Hemionitis ×tolocensis (Rasbach, Reichst. & Schneller) F.M.Vázquez; Folia Bot. Extremadur. 15: 150 (2021)
= Allosorus ×tolocensis (Rasbach, Reichst. & Schneller) Rumsey; Fern Gaz. 19(7): 277 (2014)
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A complete, synonymic checklist of the Higher Plants of the World

Version 25.03 (March 6th, 2025)

Covering 356,112 accepted species,
8,776 hybrids, 48,085 accepted infraspecific taxa and 1,002,454 synonyms,
in total 1,415,427* names

[*about 25,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus" are deliberately omitted]

With complete, detailed distribution data (countries and regions) and literature references

Arranged in linear phylogenetic sequence reflecting true relationships

Fully searchable and continously updated

With 38,209 pictures of 15,787 species

Assembled through 40 years independently from other databases

In cooperation with Catalogue of Life