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Ver. 24.10
October 9th, 2024

030.0000 Family Pteridaceae E.D.M.Kirchn.; Schul-Bot. 109 (1831)
48 genera, ca. 1350 spp. The family has been split extensively in smaller subgroups, but Smith et al. (2006) and PPG-I argue for keeping them together.
Species listed in database: 1399; hybrids: 67
= Acrostichaceae Mett. ex A.B.Frank; Syn. Pflanzenkd. (Leunis) ed. 2, 3: 1458 (1877), nom. rej.
= Actiniopteridaceae Pic.Serm.; Webbia 17: 5 (1962)
= Adiantaceae Newman; Hist. Brit. Ferns: 5 (1840) nom. cons. over Parkeriaceae (1825)
= Anopteraceae Doweld; Tent. Syst. Pl. Vasc.: 51 (2001)
= Antrophyaceae (Link) Ching; Acta Phytotax. Sin. 16: 11 (1978)
= Ceratopteridaceae Underw.; Our nat. Ferns ed. 6: 65, 78 (1900)
= Cheilanthaceae B.K.Nayar; Taxon 19: 233 (1970)
= Cryptogrammaceae Pic.Serm.; Webbia 17: 299 (1963)
= Hemionitidaceae Pic.Serm.; Webbia 21: 487 (1966)
= Negripteridaceae Pic.Serm.; Nuovo Giorn. Bot. Ital., ser. 2, 53: 160 (1946)
= Parkeriaceae Hook.; Exot. Fl. 2: t. 147 (1825), nom. rej. in favor of Adiantaceae
= Platyzomataceae Nakai; Bull. Natl. Sci. Mus., Tokyo, 29: 4 (1950)
= Pteridaceae Reichenb.; Handb. Nat. Pfl.-Syst. 138 (1837)
= Sinopteridaceae Koidz.; Acta Phytotax. Geobot. 3: 50 (1934)
= Taenitidaceae (C.Presl) Pic.Serm.; Webbia 29: 1 (1974)
= Vittariaceae Ching; Sunyatsenia 5: 232 (1940)
030.6029 Genus Lytoneuron (Klotzsch) Yesilyurt; Phytotaxa 221(2): 116 (2015)
Distribution: South America (mostly Brazil). Separation from Doryopteris based on molecular results (Yesilyurt et al. 2015)
Species listed in database: 17
= Doryopteris sect.Lytoneuron Klotzsch; Linnaea 20(3): 343 (1847)
species :
Lytoneuron acutilobum (Prantl) Yesilyurt; Phytotaxa 221(2): 117 (2015)
Distribution: Brazil (Paraná, Rio de Janeiro, Santa Catarina, São Paulo)
84 BRA
Synonyms: hidden
= Doryopteris acutiloba (Prantl) Diels; Nat. Pflanzenfam. [Engler & Prantl] I, 4: 269 (1899)
= Hemionitis acutiloba (Prantl) Christenh.; Global Fl. 1(4): 7 (2018)
= Pellaea acutiloba Prantl; Bot. Jahrb. Syst. 3: 425 (1882)
species :
Lytoneuron bradei (Rosenst.) Yesilyurt; Phytotaxa 376(3): 140 (2018), comb. Inval.
Distribution: Brazil (São Paulo)
84 BRA
Synonyms: hidden
= Doryopteris bradei Rosenst.; Hedwigia 66: 360 (1915)
species :
Lytoneuron columbinum (Hook.) Yesilyurt; Phytotaxa 376(3): 146 (2018)
Distribution: Brazil (Rio de Janeiro, Minas Gerais, Espírito Santo)
84 BRA
Synonyms: hidden
= Doryopteris columbina (Hook.) Diels; Nat. Pflanzenfam. [Engler & Prantl] I, 4: 269 (1899)
= Hemionitis columbina (Hook.) Christenh.; Global Fl. 1(4): 12 (2018)
= Lytoneuron columbinum (Hook.) Smith-Braga & Schwartsburd; Brittonia 72(4): 364 (2020), isonym
= Pellaea columbina (Hook.) Hook.; Syn. Fil. [W. J. Hooker & Baker] 146 (1867)
= Pellaea lomariacea var. columbina Hook.; Sp. Fil. [W. J. Hooker] 2: 133, t. 1 12 A (1858)
variant :
var. caparaoense Brade ex Smith-Braga & Schwartsburd; Brittonia 72(4): 5 (2020)
Distribution: Brazil (Minas Gerais, Espírito Santo)
84 BRA
species :
Lytoneuron crenulans (Fée) Yesilyurt; Phytotaxa 221(2): 117 (2015)
Distribution: Peru; Bolivia (Chuquisaca, Cochabamba, La Paz, Santa Cruz); Brazil (Bahia, Espírito Santo, Minas Gerais, Paraná, Rio de Janeiro, Rio Grande do Sul, Santa Catarina, São Paulo); Paraguay; Argentina (Jujuy, Salta)
83 BOL PER 84 BRA 85 ARG PAR
Synonyms: hidden
= Doryopteris actinophylla (Baker) Rosenst.; Hedwigia 46: 85 (1906)
= Doryopteris crenulans (Fée) Christ; Schwacke, Pl. Nov. Mineir. 2: 26 (1900)
= Doryopteris lomariacea var. actinophylla (Baker) Luetzelb.; Estud. Bot. Nordéste 3: 248 (1923)
= Hemionitis crenulans (Fée) Christenh.; Global Fl. 1(4): 12 (2018)
= Pellaea crenulans Fée; Crypt. Vasc. Brésil 2: 27, t. 87, f. 3 (1873)
= Pteris lomariacea var. actinophylla Baker; Fl. Bras. [Martius] 1(2): 406 (1870)
species :
Lytoneuron feei (Brade) Yesilyurt; Phytotaxa 221(2): 117 (2015)
Distribution: Brazil (Minas Gerais, Rio de Janeiro)
84 BRA
Synonyms: hidden
= Doryopteris feei Brade; Arch. Inst. Biol. Veg. 1: 226, f. 4, t. 5 (1935)
= Doryopteris feei f. major Brade; Arq. Inst. Biol. Veg. 1(3): 226 (1935)
= Hemionitis falkor Christenh.; Global Fl. 1(4): 13 (2018), nom. nov.
species :
Lytoneuron itatiaiense (Fée) Yesilyurt; Phytotaxa 221(2): 117 (2015)
Distribution: Brazil (São Paulo, Rio de Janeiro, Minas Gerais)
84 BRA
Synonyms: hidden
= Doryopteris itatiaiensis (Fée) Christ; Bull. Herb. Boissier, sér. 2, 2: 549 (1902)
= Doryopteris lomariacea var. itatiaiensis (Fée) Luetzelb.; Estud. Bot. Nordéste 3: 248 (1923)
= Hemionitis itatiaiensis (Fée) Christenh.; Global Fl. 1(4): 16 (2018)
= Pellaea itatiaiensis Fée; Crypt. Vasc. Brésil 2: 26, t. 88, f. 1 (1873)
= Pellaea lomariacea var. itatiaiensis (Fée) Prantl; Bot. Jahrb. Syst. 3: 425 (1882)
= Pteris itatiaiensis (Fée) Szyszyl.; Itin. Princip. Coburg. 2: 119 (1888)
species :
Lytoneuron lomariaceum (Kunze ex Klotzsch) Yesilyurt; Phytotaxa 221(2): 116 (2015)
Distribution: Venezuela (Bolívar); Guyana; Peru; Brazil (Bahia, Brasilia Distrito Federal, Espírito Santo, Goiás, Minas Gerais, Mato Grosso do Sul, Mato Grosso, Paraná, Rio de Janeiro, Rio Grande do Sul, Santa Catarina, São Paulo); Paraguay (Alto Paraná, Amambay, Caaguazu, Canindeyu, Cordillera, San Pedro); Uruguay (Rivera); Argentina (Misiones)
82 GUY VEN 83 PER 84 BRA 85 ARG PAR URU
Synonyms: hidden
= Allosorus lomariaceus (Kunze ex Klotzsch) Kuntze; Revis. Gen. Pl. 2: 806 (1891)
= Doryopteris lomariacea Kunze ex Klotzsch; Linnaea 20: 343 (1847)
= Hemionitis lomariacea (Kunze ex Klotzsch) Christenh.; Global Fl. 1(4): 17 (2018)
= Pellaea lomariacea (Kunze ex Klotzsch) Hook.; (1858)
= Pellaea lomariacea var. digitatopalmata Hook.; Sp. Fil. [W. J. Hooker] 2: 133 (1858)
= Pteris lomariacea (Kunze ex Klotzsch) Kunze; Linnaea 22: 576 (1849)
species :
Lytoneuron magdalenense (Brade) Yesilyurt; Fern Gaz. 22(1): 27 (2023)
Distribution: Brazil (Rio de Janeiro)
84 BRA
Synonyms: hidden
= Doryopteris subsimplex var. magdalenensis Brade; Arch. Inst. Biol. Veg. 2: 3 (1935)
= Hemionitis magdalenensis (Brade) Christenh.; Global Fl. 1(4): 17 (2018)
= Doryopteris magdalenensis (Brade) Brade; Arch. Jard. Bot. Rio de Janeiro 18: 40, f. 37-41 (1965)
species :
Lytoneuron microphyllum (Christ) Yesilyurt; Phytotaxa 376(3): 143 (2018)
Distribution: Brazil (Rio de Janeiro, Minas Gerais)
84 BRA
Synonyms: hidden
= Allosorus microphyllus Kuntze; Revis. Gen. Pl. 2: 806 (1891), nom. repl., non Liebm. 1849
= Cassebeera microphylla Christ; Schwacke, Pl. Nov. Mineir. 2: 25 (1900), nom. repl., non (Link) J. Sm. 1841
= Cheilanthes monticola Mart. ex Prantl; Bot. Jahrb. Syst. 3: 422 (1882), nom. nud., non Gardner ex Hook. 1842
= Doryopteris microphylla Christ; Bull. Herb. Boissier, sér. 2, 2: 546 (1902), nom. repl.
= Hemionitis pyornkrachzarkii Christenh.; Global Fl. 1(4): 20 (2018), nom. nov. for Pellaea microphylla Fée
= Lytoneuron microphyllum (Christ) Smith-Braga & Schwartsb.; Brittonia 72(4): 371 (2020), isonym
= Pellaea glaziovii var. minor Baker; Fl. Bras. [Martius] 1(2): 595 (1870)
= Pellaea microphylla Fée; Crypt. Vasc. Brésil 1: 43, t. 4, f. 2 (1869) (non Mett. (1869))
species :
Lytoneuron ornithopus (Mett. ex Hook. & Baker) Yesilyurt; Phytotaxa 221(2): 117 (2015)
Distribution: Bolivia (Santa Cruz); Brazil (Pará, ?Tocantins, Bahia, ?Maranhão, Brasilia Distrito Federal, Goiás, Mato Grosso, Mato Grosso do Sul, Espírito Santo, Minas Gerais, Rio de Janeiro, São Paulo, Paraná)
83 BOL 84 BRA
Synonyms: hidden
= Doryopteris alcicornis (Prantl) Diels; Nat. Pflanzenfam. [Engler & Prantl] I, 4: 184: 270 (1899)
= Doryopteris ornithopus (Mett. ex Hook. & Baker) J.Sm.; Hist. Fil. 289 (1875)
= Doryopteris trilobata J.Prado; Kew Bull. 48(1): 13 (1993)
= Hemionitis ornithopus (Mett. ex Hook. & Baker) Christenh.; Global Fl. 1(4): 18 (2018)
= Hemionitis trilobata (J.Prado) Christenh.; Global Fl. 1(4): 22 (2018)
= Pellaea alcicornis Prantl; Bot. Jahrb. Syst. 3: 429 (1882)
= Pteris cheirophylla Kunze ex Baker; Fl. Bras. [Martius] 1(2): 406 (1870), in syn.
= Pteris lomariacea var. digitato-palmata Hook.; Sp. Fil. [W. J. Hooker] 2: 133 (1858)
= Pteris ornithopus Mett. ex Hook. & Baker; Syn. Fil. [W. J. Hooker & Baker] 166 (1867)
variant :
var. pygmaeum (Brade) Smith-Braga & Schwartsburd; Brittonia 72(4): 11 (2020)
Distribution: Brazil (Bahia, Espírito Santo, Goias, Minas Gerais)
84 BRA
Synonyms: hidden
= Doryopteris ornithopus var. pygmaea Brade; Bol. Mus. Nac. Rio de Janeiro 5(3): 95 (1929)
species :
Lytoneuron paradoxum (Fée) Yesilyurt; Phytotaxa 221(2): 117 (2015)
Distribution: Brazil (Minas Gerais, Rio de Janeiro, São Paulo)
84 BRA
Synonyms: hidden
= Cassebeera paradoxa Fée; Mém. Foug. 7 (Icon. Esp. Nouv.): 30, t. 20, f. 2 (1857) (non Hook. (1858))
= Doryopteris paradoxa (Fée) Christ; Bull. Herb. Boissier, sér. 2, 2: 546 (1902)
= Doryopteris vestita (Hook. & Baker) Diels; Nat. Pflanzenfam. [Engler & Prantl] I, 4: 269 (1899)
= Hemionitis aconitifolia Christenh.; Global Fl. 1(4): 7 (2018), nom. nov.
= Pellaea columbina var. vestita Baker; Syn. Fil. [W. J. Hooker & Baker] 146 (1867)
= Pellaea paradoxa Fée; Crypt. Vasc. Brésil 1: 43 (1869), nom. illeg.
= Pellaea vestita (Baker) Prantl; Bot. Jahrb. Syst. 3: 423 (1882)
= Pteris paradoxa (Fée) Kuhn; Hedwigia 35: 160 (1896)
species :
Lytoneuron poseidonii Smith-Braga & Schwartsburd; Brittonia 72(4): 12 (2020)
Distribution: Brazil (Espírito Santo)
84 BRA
Remarks: possibly a synonym of Doryopteris varians fide Yesilyurt (2023)
species :
Lytoneuron quinquelobatum (Fée) Yesilyurt; Phytotaxa 221(2): 117 (2015)
Distribution: Brazil (Paraná, Rio de Janeiro)
84 BRA
Synonyms: hidden
= Doryopteris glaziovii (Baker) Diels; Nat. Pflanzenfam. [Engler & Prantl] I, 4: 269 (1899)
= Doryopteris quinquelobata (Fée) Diels; Nat. Pflanzenfam. [Engler & Prantl] I, 4: 269 (1899)
= Doryopteris quinquelobata var. septemlobata Brade; Bol. Mus. Nac. Rio de Janeiro 5: 93 (1929)
= Hemionitis quinquelobata (Fée) Christenh.; Global Fl. 1(4): 20 (2018)
= Pellaea glaziovii Baker; Fl. Bras. [Martius] 1(2): 595 (1870) (excl. var. minor)
= Pellaea quinquelobata Fée; Crypt. Vasc. Brésil 1: 42, t. 10, f. 1 (1869)
= Pteris quinquelobata (Fée) Baker; Ann. Bot. (Oxford) 5: 215 (1891)
species :
Lytoneuron rosenstockii (Brade) Yesilyurt; Phytotaxa 221(2): 117 (2015)
Distribution: Brazil (Minas Gerais, Rio de Janeiro)
84 BRA
Synonyms: hidden
= Doryopteris rosenstockii Brade; Bol. Mus. Nac. Rio de Janeiro 7(3): 143 (1931)
= Hemionitis rosenstockii (Brade) Christenh.; Global Fl. 1(4): 20 (2018)
species :
Lytoneuron rufum (Brade) Yesilyurt; Phytotaxa 221(2): 117 (2015)
Distribution: Brazil (Minas Gerais)
84 BRA
Synonyms: hidden
= Doryopteris rufa Brade; Bol. Mus. Rio de Janeiro 6: 94, t. 2 (1929)
= Hemionitis perelinae Christenh.; Global Fl. 1(4): 19 (2018), nom. nov.
species :
Lytoneuron subsimplex (Fée) Yesilyurt; Phytotaxa 221(2): 117 (2015)
Distribution: Brazil (Rio de Janeiro)
84 BRA
Synonyms: hidden
= Doryopteris subsimplex (Fée) Diels; Nat. Pflanzenfam. [Engler & Prantl] I, 4: 269 (1899)
= Hemionitis subsimplex (Fée) Christenh.; Global Fl. 1(4): 21 (2018)
= Pellaea subsimplex Fée; Crypt. Vasc. Brésil 1: 44, t. 4, f. 3 (1869)
= Pteris longula Mett.; Kuhn, Linnaea 36: 88 (1869)
= Pteris triphylla Baker; Fl. Bras. [Martius] 1(2): 596 (1870) (non J. Agardh (1839))
species :
Lytoneuron tijucanum (Brade) Yesilyurt; Phytotaxa 221(2): 117 (2015)
Distribution: Brazil (Rio de Janeiro)
84 BRA
Synonyms: hidden
= Doryopteris tijucana Brade & Rosenst.; Bol. Mus. Rio de Janeiro 7: 144, t. 9 (1931)
= Hemionitis tijucana (Brade & Rosenst.) Christenh.; Global Fl. 1(4): 22 (2018)
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A complete, synonymic checklist of the Higher Plants of the World


Version 24.10 (October 9th, 2024)

Covering 355,419 accepted species,
8,711 hybrids, 47,863 accepted infraspecific taxa and 1,000,150 synonyms,
in total 1,412,143* names

[*about 25,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus" are deliberately omitted]


With complete, detailed distribution data (countries and regions) and literature references

Arranged in linear phylogenetic sequence reflecting true relationships

Fully searchable and continously updated

With 38,209 pictures of 15,787 species

Assembled through 40 years independently from other databases

In cooperation with Catalogue of Life