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Ver. 24.11
November 5th, 2024

041.0000 Family Polypodiaceae J. & C.Presl; Delic. Prag. 159 (1822)
67 genera, ca. 1700 spp. Phylogeny: Wei & Zhang 2022
Species listed in database: 1735; hybrids: 46
= Platyceriaceae Ching; Acta Phytotax. Sin. 16(3): 18 (1978)
= Drynariaceae Ching; Acta Phytotax. Sin. 16: 1-37 (1978)
= Grammitidaceae Newman; Hist. Brit. Ferns: 7 (1840) (Grammitaceae auct., missp.)
= Gymnogrammitidaceae Ching; Acta Phytotax. Sin. 11: 12 (1966)
= Loxogrammaceae Ching ex Pic.Serm.; Webbia 29: 11 (1975)
= Platyceriaceae Ching; Sunyatsenia 5: ? (1940)
= Pleurosoriopsidaceae Sa.Kurata & Ikebe ex Ching; Acta Phytotax. Sin. 16(4): 17 (1978)
041.3501 Genus Lepisorus (J.Sm.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 47 (1933), nom. cons. prop.
Distribution: China (many), E- and tropical Asia, 1 Hawaii, few Africa, Madagascar and W-Pacific. Phylogeny and broad circumscription see Zhao et al. (2019), confirmed by PPG II. There is considerable taxonomic confusion in China because of previous oversplitting.
Species listed in database: 95
= Belvisia Mirb.; Hist. Nat. Vég. [Lamarck & Mirbel] 4: 65 (1803)
= Caobangia A.R.Sm. & X.C.Zhang; Novon 12(4): 546 (2002)
= Cheiropteris Christ; Bull. Herb. Boissier, sér. 1, 6: 876 (1898), not Chiropteris J. G. Kurr ex H. G. Bronn (1858)
= Drymotaenium Makino; Bot. Mag. (Tokyo) 15: 120 (1901) (nom. rej. prop. fide Fraser-Jenkins 2008)
= Ellipinema Li Bing Zhang & Liang Zhang; Molec. Phylogen. Evol. 148: 6 (2020)
= Hyalolepis Kunze; Linnaea 23: 258 (1850)
= Hymenolepis Kaulf.; Enum. Filic. [Kaulfuss] 146 (1824) (nec Cassini (1817))
= Lemmaphyllum C.Presl; Abh. Königl. Böhm. Ges. Wiss., ser. 5, 6: 157 (1851)
= Lepidogrammitis Ching; Sunyatsenia 5: 258 (1940)
= Lepidomicrosorium Ching & K.H.Shing; Bot. Res. Acad. Sin. 1983(1): 1 (1983)
= Macroplethus C.Presl; Abh. Königl. Böhm. Ges. Wiss., ser. 5, 6: 141 (1851), nom. rej.
= Neocheiropteris Christ; Mém. Soc. Bot. France 1 [Hist. Acrostich.]: 21 (1905) s. str.
= Neolepisorus Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 11 (1940)
= Paragramma (Blume) T.Moore; Index Fil. [T. Moore] 1: 32 (1857)
= Platygyria Ching & S.K.Wu; Acta Bot. Yunnan. 2(1): 67 (1980)
= Pleopeltis sect.Lepisorus J.Sm.; Compan. Bot. Mag. 13 (1846)
= Tricholepidium Ching; Acta Phytotax. Geobot. 29(1-5): 41 (1978)
= Weatherbya Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 191, t. 6 (1947)
= xEllipisorus Li Bing Zhang & Liang Zhang; Phytotaxa 464(4): 299 (2020)
= xLepinema Li Bing Zhang & Liang Zhang; Phytotaxa 455(4): 264 (2020)
species :
Lepisorus abbreviatus (Fée) Li Wang; Bot. J. Linn. Soc. 162(1): 35 (2010)
Distribution: Vietnam
41 VIE
Remarks: sect. 13 Belvisia
Synonyms: hidden
= Belvisia abbreviata (Fée) Hovenkamp & Franken; Blumea 37(2): 519 (1993)
= Belvisia hymenolepioides (Christ) Ching; Acta Phytotax. Sin. 10: 301 (1965)
= Belvisia hymenolepioides (Christ) V.N.Tu; Bot. Zhurn. (Moscow & Leningrad) 64(11): 1671 (1979), isonym
= Drymoglossum abbreviatum Fée; Mém. Foug. 7 (Icon. Esp. Nouv.): 26, t. 10, f. 2 (1857)
= Hymenolepis abbreviata Fée; Mém. Foug. 10 (Icon. Esp. Nouv.): 12 (1865)
= Hymenolepis hymenolepioides (Christ) Ching ex C.Chr. & Tardieu; Notul. Syst. (Paris) 8: 185 (1939)
= Lemmaphyllum abbreviatum Kuhn & C.Chr.; Dansk Bot. Ark. 6(3): 50 (1929)
= Macroplethus abbreviatus (Fée) Tagawa; Acta Phytotax. Geobot. 11: 234 (1942)
= Polypodium abbreviatum (Fée) C.Chr.; Index Filic., Suppl. Prelim. 25 (1917)
= Polypodium hymenolepioides Christ; J. Bot. (Morot) 19: 75 (1905)
= Polypodium lineare var. abbreviatum Christ; Bull. Acad. Int. Géogr. Bot. sér. 3, 11(153-154): 208 (1902)
species :
Lepisorus accedens (Blume) Hosok.; Trans. Nat. Hist. Soc. Formosa 31: 477 (1941)
Distribution: peninsular Thailand; peninsular Malaysia (Terengganu, Perak, Pahang, Selangor, Negeri Sembilan, Johor); Sumatra; Borneo (Mt. Kinabalu, Sabah); Sulawesi; Philippines; Java; Lesser Sunda Isl. (Lombok, Bali); Moluccas (Seram, etc.); New Guinea (Irian Jaya, Papua New Guinea); Bismarck Arch. (Admiralty Isl., Admiralty Isl., New Ireland, New Britain); Solomon Isl. (Bougainville, Choiseul, Guadalcanal, Kolombangara, Makira, Santa Ysabel); Australia (NE-Queensland); Vanuatu; Fiji (Viti Levu, Vanua Levu, Taveuni); American Samoa (Tutuila, Ofu, Olosega, Ta´u); [Western] Samoa (Savai´i, ´Upolu); Fed. States of Micronesia (Pohnpei, Kosrae)
41 THA 42 BOR JAW LSI MLY MOL NWG PHI SUL SUM 43 BIS SOL 50 AUS 60 ASA FIJ SAM VAN 62 MIC
Remarks: sect. 11 Weatherbya
Synonyms: hidden
= Craspedaria cuspidiflora (Reinw.) Fée; Mém. Foug. 5 (Gen. Filic.): 264 (1852)
= Craspedaria javanensis Fée; Mém. Foug. 7 (Icon. Esp. Nouv.): 65 (1857)
= Drynaria acuminata Brack.; U.S. Expl. Exped., Filic. 16: 47 (1854) (non p. 41)
= Drynaria cuspidiflora (Reinw.) J.Sm.; J. Bot. (Hooker) 3: 397 (1841)
= Goniophlebium javanense (Fée) T.Moore; Index Fil. [T. Moore] 390 (1862)
= Lemmaphyllum accedens (Blume) Donk; Reinwardtia 2: 409 (1954)
= Lepidogrammitis accedens (Blume) Saiki; J. Phytogeogr. Taxon. 35(2): 80 (1987)
= Lepidogrammitis damunensis (Rosenst.) Saiki; J. Phytogeogr. Taxon. 35(2): 80 (1987) (without basionym page)
= Microsorum accedens (Blume) Copel.; Univ. Calif. Publ. Bot. 16: 111-116 (1929)
= Phymatopsis accedens (Blume) J.Sm.; Hist. Fil. 104 (1875)
= Pleopeltis accedens (Blume) T.Moore; Index Fil. [T. Moore] 77 (1857)
= Pleopeltis acuminata (Brack.) T.Moore; Index Fil. [T. Moore] 344 (1862)
= Pleopeltis damunensis Alderw.; Bull. Dépt. Agric. Indes Néerl. 27: 4 (1909)
= Pleuridium cuspidiflorum (Reinw.) J.Sm.; Bonplandia (Hannover) 9: 261 (1861)
= Polypodium accedens Blume; Enum. Pl. Javae 2: 121 (1828)
= Polypodium cuspidiflorum Reinw.
= Polypodium damunense Rosenst.; Repert. Spec. Nov. Regni Veg. 6: 42 (1908)
= Polypodium lyciifolium Bory; Voy. Monde, Crypt. 1: 260 (1828)
= Polypodium redimiens Brause; Bot. Jahrb. Syst. 56(2): 193 (1920)
= Weatherbya accedens (Blume) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 191, t. 6 (1947)
= Weatherbya damunensis (Rosenst.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 191 (1947)
species :
Lepisorus affinis Ching; Acta Phytotax. Sin. 8: 169, t. 22(28) (1959)
Distribution: China (Hainan)
36 CHI
Remarks: sect. 10 Lepisorus
Synonyms: hidden
= Lepisorus longifolius Ching; Acta Phytotax. Sin. 8: 170 (1959) (non (Blume) Holttum)
= Lepisorus longus Ching; Acta Phytotax. Sin. 10: 192 (1965), nom. nov. for L. longifolius Ching
species :
Lepisorus alberti (Regel) Ching; Acta Bot. Yunnan. 5(1): 20 (1983)
Distribution: China (Gansu, Hebei, Henan, Nei Mongol, Qinghai, Shaanxi, Shanxi, Sichuan, Xinjiang, Yunnan); Tibet
36 CHI TIB
Remarks: sect. 9 Hymenophyton. Records for Taiwan refer to L. clathratus fide Knapp (Ferns of Taiwan, 2011 ff.)
Synonyms: hidden
= Lepisorus clathratus var. papakensis (Masam.) Tagawa; Acta Phytotax. Geobot. 5(2): 111 (1936)
= Lepisorus papakensis (Masam.) Ching & Y.X.Lin; Acta Bot. Yunnan. 5(1): 7 (1983)
= Lepisorus pumilus Ching & S.K.Wu; Acta Bot. Yunnan. 5(1): 16 (1983)
= Polypodium alberti Regel; Trudy Imp. S.-Peterburgsk. Bot. Sada 7: 622 (1881)
= Polypodium lineare var. alberti (Regel) C.Chr.; Index Fil. [T. Moore] 507, 540 (1906)
= Polypodium papakense Masam.; J. Soc. Trop. Agric. 6: 569 (1934)
species :
Lepisorus amaurolepidus (Sledge) Bir & Trikha; Bir & Vasudeva, J. Bombay Nat. Hist. Soc. 68: 192 (1971)
Distribution: S-India; Sri Lanka
40 IND SRL
Remarks: sect. 17 Pseudovittaria. Included in L. contortus by Kandel & Fraser-Jenkins (2020)
Synonyms: hidden
= Lepisorus amaurolepidus (Sledge) B.K.Nayar & S.Kaur; Companion Ferns Brit. India 84 (1974), isonym
= Pleopeltis amaurolepida Sledge; Bull. Brit. Mus. (Nat. Hist.), Bot. 2: 136 (1960)
species :
Lepisorus angustus Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 86 (1933)
Distribution: China (Anhui, Chongqing, Gansu, Guangxi, Henan, Hubei, Hunan, Shaanxi, Sichuan, Yunnan, S-Zhejiang); Tibet; ?Japan
36 CHI TIB
Remarks: sect. 17 Pseudovittaria. Included in L. thunbergianus by Fraser-Jenkins et al. 2015 but kept separate on genetic basis by Zhao et al. 2019
Synonyms: hidden
= Lepisorus nanchuanensis Ching; Bull. Bot. Res., Harbin 3(4): 4 (1983)
= Lepisorus niger Ching; Fl. Xizangica 1: 312 (1983) (fide Fraser-Jenkins et al. 2015)
= Lepisorus pseudoangustus Ching; Fl. Xizangica 1: 310 (1983)
= Lepisorus thunbergianus var. angustus (Ching) Sa.Kurata; Sci. Rep. Yokosuka City Mus. 11: 39 (1965)
= Pleopeltis caudatoattenuata (Takeda) Panigrahi & Patnaik; Curr. Sci. 34: 127 (1965)
= Polypodium caudatoattenuatum (Takeda) C.Chr.; Index Filic., Suppl. Tert.: 146 (1934)
= Polypodium lineare f. caudatoattenuatum Takeda; Notes Roy. Bot. Gard. Edinburgh 8(39): 269-270 (1915)
species :
Lepisorus annamensis (C.Chr.) Li Wang; Bot. J. Linn. Soc. 162(1): 35 (2010)
Distribution: China (Hainan); Thailand; Laos; Vietnam; peninsular Malaysia (Perak); Borneo (Sabah); Sulawesi
36 CHI 41 LAO THA VIE 42 BOR MLY SUL
Remarks: sect. 13 Belvisia
Synonyms: hidden
= Belvisia annamensis (C.Chr.) Tagawa; Acta Phytotax. Geobot. 22: 107 (1967)
= Belvisia annamensis (C.Chr.) V.N.Tu; Bot. Zhurn. (Moscow & Leningrad) 64(11): 1670 (1979), isonym
= Hymenolepis annamensis C.Chr.; Dansk Bot. Ark. 6(3): 54-70 (1929)
= Hymenolepis callifolia var. paltonioides C.Chr.; (1929)
= Macroplethus annamensis (C.Chr.) Tagawa; Acta Phytotax. Geobot. 11: 234 (1942)
species :
Lepisorus annuifrons (Makino) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 70 (1933)
Distribution: South Korea (incl. Cheju Isl.); Japan
38 JAP KOR
Remarks: sect. 12 Ussurienses
Synonyms: hidden
= Pleopeltis annuifrons (Makino) Nakai; (1933)
= Polypodium annuifrons Makino; Bot. Mag. (Tokyo) 13: 48 (1899)
= Polypodium japonense Makino; Bot. Mag. (Tokyo) 12: 165 (1898), nomen
species :
Lepisorus asterolepis (Baker) Ching; Fl. Jiangsuensis 1: 74 (1977)
Distribution: China (Anhui, Chongqing, Fujian, Guangxi, Guizhou, Hubei, Hunan, Jiangsu, Jiangxi, Shaanxi, Sichuan, Yunnan, Zhejiang); Tibet; ?Bhutan; NE-India (Arunachal Pradesh); Myanmar (Kachin, etc.); Vietnam
36 CHI TIB 40 IND 41 MYA VIE
Remarks: sect. 10 Lepisorus. Included in L. macrosphaerus by Fraser-Jenkins et al. 2015 but a separate species fide Zhao et al. 2019 on genetic grounds
Synonyms: hidden
= Lepisorus longipes Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(4): 25 (1984)
= Lepisorus macrosphaerus var. asterolepis (Baker) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4(3): 74 (1933)
= Lepisorus macrosphaerus var. asterolepis (Baker) R.D.Dixit; R. D. Dixit, Census Indian Pteridophytes 41 (1984), isonym
= Pleopeltis asterolepis (C.Chr.) Panigrahi; Pl. Sci. Res. (India), Abstr. 23rd. Ann. Conf. Khurda: 36 (1998)
= Pleopeltis macrosphaera var. asterolepis (Baker) Panigrahi & Patnaik; Proc. Nat. Acad. Sci. India, B, 34: 482, f. 1-5 (1964)
= Polypodium asterolepis Baker; J. Bot. 18: 230 (1880) (non Polypodium astrolepis Liebm.)
= Polypodium excavatum var. asterolepis (Baker) C.Chr.; Index Filic. 9: 525 (1906)
= Polypodium macrosphaerum var. asterolepis (Baker) C.Chr.; (1924)
species :
Lepisorus balteiformis (Brause) Hovenkamp; Blumea 43: 110 (1998)
Distribution: New Guinea (Irian Jaya, Papua New Guinea)
42 NWG
Remarks: sect. 1 Paragramma
Synonyms: hidden
= Paragramma balteiformis (Brause) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 190 (1947)
= Polypodium balteiforme Brause; Bot. Jahrb. Syst. 66: 194 (1920)
species :
Lepisorus bicolor (Takeda) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 66 (1933)
Distribution: China (Guizhou, Sichuan, Yunnan); Tibet; Nepal; Bhutan; N-India (Himachal Pradesh, Uttarakhand, Darjeeling, Sikkim, Arunachal Pradesh); Myanmar (Kachin, Chin, etc.); Thailand
36 CHI TIB 40 BHU IND NEP 41 MYA THA
Remarks: sect. 8 Pachyphyllon
Synonyms: hidden
= Lepisorus kashyapii (Mehra) Mehra; Res. Bull. Punjab Univ., New Ser., Sci. 13: 24 (1962), comb. inval.
= Lepisorus kashyapii var. major Bir & Trikha; Amer. Fern J. 71(2): 55 (1981)
= Lepisorus kashyapii var. minor Bir & Trikha; Amer. Fern J. 71(2): 55 (1981)
= Pleopeltis bicolor (Takeda) Sledge; Bull. Brit. Mus. (Nat. Hist.), Bot. 2: 138 (1960)
= Pleopeltis kashyapii (Mehra) Alston & Bonner; Candollea 15: 208 (1956), nom. inval.
= Polypodium bicolor (Takeda) Alston ex Schelpe; Amer. Fern J. 44: 59 (1954) (non Mett.)
= Polypodium excavatum var. bicolor Takeda; Notes Roy. Bot. Gard. Edinburgh 8: 280 (1915)
= Polypodium kashyapii Mehra; Punjab Univ. Bot. Publ. [Ferns of Mussoorie]: 24 (1939), nom. nud. (fide Fraser-Jenkins et al. 2015)
species :
Lepisorus boninensis Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 76 (1933)
Distribution: Japan (Izu Isl.); Ryukyu Isl.; Bonin Isl. (Chichijima, Anijima, Otohtojima, Hahajima); Volcano Isl. (Kita-Iwojima, Minami-Iwojima)
38 JAP KZN NNS OGA
Remarks: sect. 15 Obtusi
Conservation Status: IUCN R. NT in Japan
Synonyms: hidden
= Pleopeltis boninensis (Christ) H.Ohba; Sci. Rep. Tohoku Imp. Univ., Ser. 4, Biol. 36: 120 (1971)
= Polypodium boninense Christ; Warb., Monsunia 1: 61 (1900)
species :
Lepisorus buergerianus (Miq.) C.F.Zhao, R.Wei & X.C.Zhang; Cladistics 36(3): 253 (2019)
Distribution: China (Anhui, Chongqing, Fujian, Gansu, Guangdong, Guangxi, Guizhou, Hongkong, Hubei, Hunan, Jiangxi, Sichuan, Yunnan, Zhejiang); Japan; Ryukyu Isl.; N-Vietnam
36 CHI 38 JAP NNS 41 VIE
Remarks: sect. 7 Lepidomicrosorium
Synonyms: hidden
= Lepidomicrosorium asarifolium Ching & K.H.Shing; Bot. Res. Acad. Sin. 1983(1): 11 (1983)
= Lepidomicrosorium brevipes Ching & K.H.Shing; Bot. Res. Acad. Sin. 1983(1): 13 (1983)
= Lepidomicrosorium buergerianum (Miq.) Ching & K.H.Shing; Fl. Jiangxi 1: 322 (1993)
= Lepidomicrosorium emeicola Ching & K.H.Shing; Bot. Res. Acad. Sin. 1983(1): 7 (1983)
= Lepidomicrosorium hederaceum (Christ) Ching; Bot. Res. Acad. Sin. 1983(1): 11 (1983)
= Lepidomicrosorium lanceolatum Ching & P.S.Wang; Bot. Res. Acad. Sin. 1983(1): 8 (1983)
= Lepidomicrosorium latibasis Ching & K.H.Shing; Bot. Res. Acad. Sin. 1983(1): 6 (1983)
= Lepidomicrosorium microsorioides (W.M.Chu) Ching & W.M.Chu; Bot. Res. Acad. Sin. 1983(1): 7 (1983)
= Lepidomicrosorium subhastatum (Baker) Ching; Bot. Res. Acad. Sin. 1983(1): 12 (1983)
= Lepidomicrosorium subsessile Ching & K.H.Shing; Bot. Res. Acad. Sin. 1983(1): 6 (1983)
= Lepidomicrosorium suijiangense Ching & W.M.Chu; Bot. Res. Acad. Sin. 1983(1): 13 (1983)
= Lepidomicrosorium yiliangense Ching & K.H.Shing; Bot. Res. Acad. Sin. 1983(1): 7 (1983)
= Leptochilus buergerianus (Miq.) Bosman; Leiden Bot. Ser. 14: 114 (1991)
= Microsorum buergerianum (Miq.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 295-311 (1933)
= Microsorum buergerianum var. ohwianum (Tagawa) Tagawa; Acta Phytotax. Geobot. 14: 9 (1949)
= Microsorum ohwianum Tagawa; J. Jap. Bot. 12: 752 (1936)
= Microsorum subhastatum (Baker) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 295-311 (1933)
= Microsorum subhastatum var. longifrons (Takeda) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 298 (1933)
= Neocheiropteris buergeriana (Miq.) Fraser-Jenk.; Indian J. Forest. 42(4): 314 (2019), isonym
= Neocheiropteris buergeriana (Miq.) Nakaike; Ill. Pterid. Japan 2: 518 (1981)
= Neocheiropteris subhastata (Baker) Tagawa; J. Jap. Bot. 27: 217 (1952)
= Neolepisorus microsorioides W.M.Zhu; Acta Bot. Yunnan. 1(2): 96 (1979)
= Polypodium buergerianum Miq.; Ann. Mus. Bot. Lugduno-Batavi 3: 170 (1867)
= Polypodium buergerianum var. stipitatum Takeda; Notes Roy. Bot. Gard. Edinburgh 8(39): 290-291 (1915)
= Polypodium hederaceum Christ; Bull. Acad. Int. Géogr. Bot. 215, c. fig. (1902)
= Polypodium subhastatum Baker; J. Bot. 27: 177 (1889)
= Polypodium subhastatum var. hederaceum (Christ) Takeda; Notes Roy. Bot. Gard. Edinburgh 8(39): 292 (1915)
= Polypodium subhastatum var. longifrons Takeda; Notes Roy. Bot. Gard. Edinburgh 8(39): 292 (1915)
= Polypodium superficiale var. chinense Rosenst.; Repert. Spec. Nov. Regni Veg. 13(355-358): 134 (1914)
= Tricholepidium buergerianum (Miq.) Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 69 (2008)
species :
Lepisorus carnosus (J.Sm.) C.F.Zhao, R.Wei & X.C.Zhang; Cladistics 36(3): 252 (2019)
Name: Penny fern
Distribution: China (Fujian, Gansu, Guangdong, Guangxi, Guizhou, Hubei, Hunan, Jiangxi, Shanxi, Sichuan, Yunnan, Zhejiang); Bhutan; NE-India (Sikkim, Meghalaya, Arunachal Pradesh); Myanmar (Kachin, etc.); Thailand; Vietnam; Philippines (Luzon)
36 CHI 40 BHU IND 41 MYA THA VIE 42 PHI
Remarks: not in Nepal or Taiwan. Sect. 4 Lemmaphyllum. Lemmaphyllum diversum and L. drymoglossoides are frequently kept separate
Synonyms: hidden
= Colysis diversa (Rosenst.) Nakai ex Nemoto; Fl. Japan. [Makino & Nemoto] Suppl. 20 (1936)
= Drymoglossum carnosum Hook.; Gen. Fil. [W. J. Hooker] t. 78A (1842)
= Drymoglossum carnosum var. minus Hook.; Sp. Fil. [W. J. Hooker] Fil. [W. J. Hooker] 5: 189 (1864)
= Drymoglossum carnosum Wall. ex J.Sm.; Gen. Fil. [W. J. Hooker] t. 78A (1841) (excl. var.)
= Drymoglossum subcordatum Fée; Mém. Foug. 3 (Hist. Vittar.): 29 (1852)
= Elaphoglossum carnosum (Hook.) Keyserl.; Polyp. Herb. Bunge 36 (1873)
= Goniophlebium moupinense (Franch.) Bedd.; Suppl. Ferns Brit. Ind. 91 (1892)
= Lemmaphyllum adnascens Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 101 (1933)
= Lemmaphyllum carnosum (Hook.) C.Presl; Abh. Königl. Böhm. Ges. Wiss., ser. 5, 6: 158 (1851)
= Lemmaphyllum carnosum var. drymoglossoides (Baker) X.P.Wei; J. Syst. Evol. 51(4): 493 (2013)
= Lemmaphyllum carnosum var. longifolium S.Rahman & T.Sen; Indian Fern J. 15(1-2): 182 (1998)
= Lemmaphyllum christensenii Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 98 (1933) (non Polypodium christensenianum Maxon)
= Lemmaphyllum chui Saiki; J. Phytogeogr. Taxon. 35(2): 79 (1987)
= Lemmaphyllum diversum (Rosenst.) Tagawa; Acta Phytotax. Geobot. 14: 9 (1949)
= Lemmaphyllum drymoglossoides (Baker) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 100 (1933)
= Lemmaphyllum intermedium (Ching) Li Wang; Bot. J. Linn. Soc. 162(1): 36 (2010)
= Lepidogrammitis adnascens (Ching) Ching; Sunyatsenia 5: 258 (1940)
= Lepidogrammitis christensenii (Ching) Ching; Sunyatsenia 5: 258 (1940)
= Lepidogrammitis diversa (Rosenst.) Ching; Acta Bot. Yunnan. 1(1): 24 (1979), et Fl. Xizangica 1: 313 (1983)
= Lepidogrammitis drymoglossoides (Baker) Ching; Sunyatsenia 5: 258 (1940)
= Lepidogrammitis elongata Ching; Fl. Tsinling. 2: 232 (1974)
= Lepidogrammitis intermedia Ching; Fl. Tsinling. 2: 231 (1974)
= Lepidogrammitis kansuensis Ching; Fl. Tsinling. 2: 232 (1974)
= Notholaena carnosa Wall.; Numer. List. [Wallich] no. 138 (1829), nomen
= Oetosis carnosa (Wall. ex J.Sm.) Kuntze; Revis. Gen. Pl. 2: 817 (1891)
= Polypodium cyclophyllum Baker; Ann. Bot. (Oxford) 5: 473 (1891)
= Polypodium diversum Rosenst.; Hedwigia 66: 346 (1915)
= Polypodium drymoglossoides Baker; J. Bot. 25: 170 (1887)
= Polypodium moupinense Franch.; Nouv. Arch. Mus. Hist. Nat., sér. 2, 10: 121 (1887)
= Polypodium wangii C.Chr.; Index Filic., Suppl. Tert.: 161 (1934)
= Pteris piloselloides Thunb.; Fl. Jap. [Thunberg] 331 (1784) (non L. (1763))
= Taenitis carnosa (Hook.) Mett.; Fil. Hort. Bot. Lips. 28 (1856)
= Taenitis piloselloides (Thunb.) Wall.; Numer. List. [Wallich] no. 62 (1829), nomen
species :
Lepisorus clathratus (C.B.Clarke) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 71 (1933)
Distribution: Siberia (Altai); Kazakhstan; Kyrgyzstan; E-Afghanistan; China (Henan, Qinghai, Shaanxi, Guangxi, Sichuan, Yunnan); Tibet; Mongolia; Japan; Ryukyu Isl.; Taiwan; Pakistan (Kurram, Hazara); Nepal; Bhutan; India (Sikkim, Darjeeling Hills, Meghalaya, Arunachal Pradesh, Himachal Pradesh, Jammu & Kashmir)
30 SIB 32 KAZ KGZ 34 AFG 36 CHI TIB 37 MON 38 JAP NNS TAI 40 BHU IND NEP PAK
Remarks: sect. 9 Hymenophyton
Synonyms: hidden
= Lepisorus clathratus var. namegatae Sa.Kurata; Hokuriko J. Bot. 5: 17 (1956)
= Lepisorus henanensis Ching & S.K.Wu; Acta Bot. Yunnan. 5(1): 13 (1983)
= Lepisorus namegatae (Sa.Kurata) Ching & Y.X.Lin; Acta Bot. Yunnan. 5(1): 16 (1983)
= Lepisorus nepalensis K.Iwats.; Fl. E. Himal., 3rd. Rep.: 198 (1975) (fide Fraser-Jenkins 2008)
= Lepisorus petiolatus Ching & Y.X.Lin; Acta Phytotax. Sin. 31(5): 475 (1993)
= Lepisorus sinuatus (Ching & S.K.Wu) S.L.Yu; Acta Phytotax. Sin. 35: 343 (1997)
= Lepisorus thaipaiensis Ching & S.K.Wu; Acta Bot. Yunnan. 5(1): 8 (1983)
= Lepisorus variabilis Ching & S.K.Wu; Fl. Xizangica 1: 308 (1983) (fide Fraser-Jenkins 1997)
= Lepisorus venosus Ching & S.K.Wu; Acta Bot. Yunnan. 5(1): 15 (1983)
= Platygyria kongtingensis Ching & Y.X.Lin; Acta Bot. Yunnan. 5(1): 22 (1983)
= Platygyria muliensis Ching & S.K.Wu; Acta Bot. Yunnan. 5(1): 22 (1983)
= Platygyria sinuata Ching & S.K.Wu; Acta Bot. Yunnan. 2(1): 71 (1980)
= Platygyria variabilis Ching & S.K.Wu; Acta Bot. Yunnan. 5(1): 21 (1983) (fide Fraser-Jenkins 1997)
= Pleopeltis clathrata (C.B.Clarke) Bedd.; Ferns Brit. India 348 (1883)
= Pleopeltis clathrata (C.B.Clarke) Czerep.; Vasc. Pl. Russia & Adj. States (former Russia): 414 (1995), isonym
= Pleopeltis clathrata var. namegatae (Sa.Kurata) Ohwi; Fl. Jap. [Ohwi] 147 (1957)
= Pleopeltis variabilis (Ching & S.K.Wu) S.Singh & Panigrahi; J. Hill Res. 3: 18 (1990) (fide Fraser-Jenkins 1997)
= Polypodium clathratum C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 559, t. 82, f. 1 (1880)
species :
Lepisorus confluens W.M.Chu; Acta Bot. Yunnan., Suppl. 5: 55 (1992)
Distribution: China (Yunnan)
36 CHI
Remarks: sect. 17 Pseudovittaria
Synonyms: hidden
= Lepisorus iridescens Ching & Y.X.Lin; Fl. Reipubl. Popularis Sin. [Fl. China Ed. Committee] 6(2): 82, t. 16: 3-4 (2000)
species :
Lepisorus contortus (Christ) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 90 (1933)
Distribution: China (Anhui, Chongqing, ?Fujian, Gansu, Henan, Hubei, Jiangxi, Shaanxi, Sichuan, Yunnan, ?Zhejiang); Taiwan; Nepal; Bhutan; India (Himachal Pradesh, Uttarakhand, Sikkim, Arunachal Pradesh, S-India); Sri Lanka; Myanmar (Kachin, etc.); N-Thailand
36 CHI 38 TAI 40 BHU IND NEP SRL 41 MYA THA
Remarks: sect. 17 Pseudovittaria
Synonyms: hidden
= Lepisorus crassirhizoma Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 3(4): 3 (1983)
= Lepisorus jinfoshanensis Ching; Bull. Bot. Res., Harbin 3(4): 4 (1983)
= Lepisorus khullarii P.C.Pande & H.C.Pande; Illustr. Fern Fl. Kumaon Himalaya 1: 67 (2003) (fide Fraser-Jenkins 2008)
= Lepisorus nyalamensis Ching & S.K.Wu; Fl. Xizangica 1: 311 (1983) (fide Fraser-Jenkins et al. 2015)
= Pleopeltis contorta (Christ) Alston & Bonner; Candollea 15: 209 (1956)
= Polypodium contortum (Christ) Christ; Bot. Gaz. 61: 347 (1911)
= Polypodium lineare f. contortum (Christ) Takeda; Notes Roy. Bot. Gard. Edinburgh 8: 270 (1915)
= Polypodium lineare var. contortum Christ; Nuovo Giorn. Bot. Ital., n. s. 4: 98 (1897)
= Polypodium thunbergianum var. contortum (Christ) C.Chr.; Index Filic., Suppl. Tert.: 160 (1934)
species :
Lepisorus crassipes Ching & Y.X.Lin; Acta Bot. Yunnan. 5(1): 18 (1983)
Distribution: China (Gansu, Hebei, Hubei, Qinghai, Shaanxi, Sichuan)
36 CHI
Remarks: sect. 9 Hymenophyton. Often included in L. clathratus
Synonyms: hidden
= Lepisorus hsiawutaiensis Ching & S.K.Wu; Acta Bot. Yunnan. 5(1): 6 (1983)
= Lepisorus kansuensis Ching & S.K.Wu; Acta Bot. Yunnan. 5(1): 17 (1983)
= Lepisorus ligulatus Ching & S.K.Wu; Acta Bot. Yunnan. 5(1): 6 (1983)
= Lepisorus maowenensis Ching & S.K.Wu; Acta Bot. Yunnan. 5(1): 14 (1983)
= Lepisorus patungensis Ching & S.K.Wu; Acta Bot. Yunnan. 5(1): 11 (1983)
= Lepisorus shanyangensis Ching & Y.X.Lin; Acta Bot. Yunnan. 5(1): 19 (1983)
= Lepisorus sungpanensis Ching & Y.X.Lin; Acta Bot. Yunnan. 5(1): 16 (1983)
species :
Lepisorus distans (Makino) Ching; Acta Phytotax. Sin. 10: 302 (1965)
Distribution: Russian Far East (Kuril Isl.); China (Anhui, Jiangxi, Shandong, Zhejiang); Japan
31 RFE 36 CHI 38 JAP
Remarks: sect. 12 Ussurienses. Often included as variety in L. ussuriensis
Synonyms: hidden
= Lepisorus ussuriensis var. distans (Makino) Tagawa; Acta Phytotax. Geobot. 11: 236 (1942)
= Pleopeltis distans (Makino) Vorosch.; Florist. Issl. Razn. Raionakh SSSR [A. K. Skvortsov] 145 (1985) (non Kaulf. (1824), nec D. Don (1825))
= Pleopeltis ussuriensis var. distans (Makino) Okuyama
= Polypodium annuifrons var. distans (Makino) Nakai; Fl. Kor. 2: 414 (1911)
= Polypodium distans (Makino) Makino; Bot. Mag. (Tokyo) 20: 31 (1906), not Kaulfuss (1824), nor D. Don (1825), nor Raddi (1825)
= Polypodium lineare var. distans Makino; Bot. Mag. (Tokyo) 15: 60 (1901)
species :
Lepisorus eilophyllus (Diels) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 65 (1933)
Distribution: China (Gansu, Guizhou, Hubei, Sichuan, Yunnan); Tibet; NE-India (Arunachal Pradesh, Nagaland, Sikkim); Myanmar (Kachin, etc.); Thailand
36 CHI TIB 40 IND 41 MYA THA
Remarks: sect. 17 Pseudovittaria
Conservation Status: IUCN VU for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Lepisorus neolewisii K.H.Shing; Acta Phytotax. Sin. 31(6): 572 (1993)
= Lepisorus pseudolewisii K.H.Shing; Vasc. Pl. Hengduan Mount. 1: 166 (1993), without latin descr. or type
= Lepisorus subconfluens Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 85 (1933)
= Pleopeltis subconfluens (Ching) Panigrahi & Patnaik; Curr. Sci. 34: 127 (1965)
= Pleopeltis subconfluens (Ching) Tagawa & K.Iwats.; (1967), isonym
= Polypodium eilophyllum Diels; Bot. Jahrb. Syst. 29: 204 (1900), based on P. involutum Baker, J. Bot. 27: 177 (1889), not Desv. (1811), nor Mett. (1856)
= Polypodium involutum Baker; J. Bot. 27: 177 (1889) (non Desv. (1811) nec Mett. (1856))
= Polypodium lewisii Christ; (non Baker)
= Polypodium lineare C.Chr.; Index Filic. 540 (1906), nom. illeg. (non Thunb.)
= Polypodium subconfluens (Ching) C.Chr.; Index Filic., Suppl. Tert.: 159 (1934)
species :
Lepisorus elegans Ching & W.M.Chu; Acta Bot. Yunnan., Suppl. 5: 55 (1992)
Distribution: China (Yunnan)
36 CHI
Remarks: sect. 17 Pseudovittaria. Sometimes included in L. contortus
species :
Lepisorus ensatus (Thunb.) C.F.Zhao, R.Wei & X.C.Zhang; Cladistics 36(3): 251 (2019)
Distribution: China (Sichuan, Guizhou, Yunnan); South Korea (Cheju Isl.); Japan (incl. Izu Isl.); Ryukyu Isl.; Taiwan; NE-India (Manipur, Meghalaya); Vietnam; Philippines (Mountain Prov.)
36 CHI 38 JAP KOR NNS TAI 40 IND 41 VIE 42 PHI
Remarks: sect. 3 Neolepisorus
Conservation Status: IUCN EN for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Microsorum ensatum (Thunb.) H.Itô; J. Jap. Bot. 11: 96 (1935)
= Microsorum reticulatum Ching ex L.Shi; Acta Phytotax. Sin. 37(1): 100 (1999)
= Neocheiropteris ensata (Thunb.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 108-112 (1933)
= Neocheiropteris ensata f. izuensis (Sa.Kurata & Satake ex Sa.Kurata) Seriz.; J. Phytogeogr. Taxon. 29(1): 24 (1981)
= Neocheiropteris ensata f. undulatodentata Sugim.; J. Geobot. 24(3): 61 (1977)
= Neocheiropteris ensata var. izuensis Sa.Kurata & Satake; J. Geobot. 5(1): 5 (1962 (May 1962))
= Neolepisorus ensatus (Thunb.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 14 (1940)
= Neolepisorus ensatus var. izuensis (Sa.Kurata & Satake) Nakaike; Checkl. Lycophytes Ferns Japan: 36 (2013)
= Neolepisorus tenuipes Ching & K.H.Shing; Acta Phytotax. Sin. 21(3): 273 (1983) (fide Fraser-Jenkins 2008)
= Pleopeltis ensata (Thunb.) Bedd.; Suppl. Ferns Brit. Ind. 94 (1892), isonym
= Pleopeltis ensata (Thunb.) T.Moore; Index Fil. [T. Moore] 346 (1862)
= Polypodium ensatum Thunb.; Trans. Linn. Soc. 2: 341 (1794)
= Polypodium oligolepis Baker; Bull. Misc. Inform. Kew 1898: 231 (1898)
= Polypodium phyllitidis Thunb.; Fl. Jap. [Thunberg] 335 (1784) (non L. (1753))
variant :
var. platyphyllus (Tagawa) comb.ined.
Distribution: Japan; Taiwan; etc.
38 JAP TAI
Synonyms: hidden
= Neocheiropteris ensata var. platyphylla (Tagawa) Tagawa ex Ohwi; Fl. Jap. [Ohwi] 149 (1957)
= Neolepisorus platyphyllus (Tagawa) L.Y.Kuo; Taiwania 64(4): 379 (2019)
= Neolepisorus ensatus f. platyphyllus (Tagawa) Ching & K.H.Shing; Acta Phytotax. Sin. 21(3): 275 (1983)
= Neolepisorus ensatus var. platyphyllus Tagawa; J. Jap. Bot. 22(10-12): 162 (1948)
species :
Lepisorus excavatus (Bory ex Willd.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 68 (1939)
Distribution: Guinea; Sierra Leone; Liberia; Ivory Coast; Nigeria; Cameroon; Bioko Isl. [Fernando Poo]; São Tomé; Annobon Isl.; Equatorial Guinea (Rio Muni); Gabon; Congo [Brazzaville]; D.R.Congo [Zaire]; Rwanda; Burundi; Sudan; Ethiopia; Somalia; Uganda; Kenya; Tanzania; Angola; Zambia; Malawi; E-Zimbabwe; Mozambique; South Africa (Limpopo, Mpulamanga); Madagascar; Comores (Grande Comore, Mayotte); +Mauritius; La Réunion
22 GUI IVO LBR NGA SIE 23 ANN BIO BUR CMN CON EQG GAB RWA SAO ZAI 24 ETH SOM SUD 25 KEN TAN UGA 26 ANG MLW MOZ ZAM ZIM 27 SAF 29 COM MAU MDG REU
Remarks: sect. 8 Pachyphyllon. L. bampsii, L. vesiculari-paleaceus and L. schraderi are possibly separate species.
Synonyms: hidden
= Drynaria excavata (Bory ex Willd.) Fée; Mém. Foug. 5 (Gen. Filic.): 270 (1852)
= Drynaria phlebodes (Kunze ex Mett.) Fée; Mém. Foug. 5 (Gen. Filic.): 270 (1852)
= Lepisorus bampsii (Pic.Serm.) Zink; Syst. Fern Gen. Lepisorus 36 (1993)
= Lepisorus corradii Pic.Serm.; Nuovo Giorn. Bot. Ital., ser. 2, 52: 30, t. 3 (1946)
= Lepisorus mildbraedii (Hieron.) Pic.Serm.; Nuovo Giorn. Bot. Ital., ser. 2, 52: 33 (1946)
= Lepisorus phlebodes (Kunze ex Mett.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 72 (1933)
= Lepisorus preussii (Hieron.) Pic.Serm.; Nuovo Giorn. Bot. Ital., ser. 2, 52: 32 (1946)
= Lepisorus preussii var. angustifolius (Hieron.) Pic.Serm.; Nuovo Giorn. Bot. Ital., ser. 2, 52: 32 (1946)
= Lepisorus preussii var. angustipaleaceus (Hieron.) Pic.Serm.; Nuovo Giorn. Bot. Ital., ser. 2, 52: 32 (1946)
= Lepisorus preussii var. ledermannii (Hieron.) Pic.Serm.; Nuovo Giorn. Bot. Ital., ser. 2, 52: 32 (1946)
= Lepisorus preussii var. winkleri (Hieron.) Pic.Serm.; Nuovo Giorn. Bot. Ital., ser. 2, 52: 32 (1946)
= Lepisorus rotundus (Bonap.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 51 (1933)
= Lepisorus stolzii (Hieron.) Pic.Serm.; Nuovo Giorn. Bot. Ital., ser. 2, 52: 33 (1946)
= Lepisorus vesiculari-paleaceus (Hieron.) Pic.Serm.; Nuovo Giorn. Bot. Ital., ser. 2, 62: 33 (1946)
= Phymatodes excavata (Bory ex Willd.) C.Presl; Tent. Pterid. 196 (1836)
= Phymatodes simplex (Sw.) C.Presl; Tent. Pterid. 196 (1836), nom. illeg.
= Pleopeltis bampsii Pic.Serm.; Bull. Jard. Bot. Natl. Belg. 53(1-2): 200 (1983)
= Pleopeltis corradii (Pic.Serm.) Cufod.; Phyton (Horn) 4: 189 (1952)
= Pleopeltis excavata (Bory ex Willd.) Sledge; Bull. Brit. Mus. (Nat. Hist.), Bot. 2: 138 (1960), isonym
= Pleopeltis excavata (Bory ex Willd.) T.Moore; Index Fil. [T. Moore] 347 (1862)
= Pleopeltis grevilleana (Wall.) T.Moore; Index Fil. [T. Moore] 78 (1857)
= Pleopeltis mildbraedii (Hieron.) Pic.Serm.; Webbia 23: 189 (1968)
= Pleopeltis phlebodes (Kunze ex Mett.) Pic.Serm.; Webbia 23: 189 (1968)
= Pleopeltis preussii (Hieron.) Tardieu; Mém. Inst. Franc. Afrique Noire 28: 217, t. 44(6-8) (1953)
= Pleopeltis preussii var. angustifolia (Hieron.) Tardieu; Mém. Inst. Franc. Afrique Noire 28: 219 (1953), nom. nud.
= Pleopeltis preussii var. winkleri (Hieron.) Tardieu; Mém. Inst. Franc. Afrique Noire 28: 219 (1953), nom. nud.
= Pleopeltis rotunda (Bonap.) Tardieu; Mém. Inst. Franc. Afrique Noire 28: 219, t. 44(9-11) (1953)
= Pleopeltis simplex (C.Presl) Bedd.; Ferns Brit. India 347 (1883), nom. illeg.
= Pleopeltis stolzii (Hieron.) Pic.Serm.; Webbia 23: 189 (1968)
= Pleopeltis vesicularipaleacea (Hieron.) Pic.Serm.; Webbia 23: 190 (1968)
= Polypodium bullatum Baker; J. Linn. Soc., Bot. 15: 420 (1876)
= Polypodium excavatum Bory ex Willd.; Sp. Pl., ed. 4 [Willdenow] 5(1): 158 (1810)
= Polypodium excavatum var. concolor Takeda; Notes Roy. Bot. Gard. Edinburgh 8: 280 (1915), nom. inval.
= Polypodium excavatum var. phlebodes (Mett.) C.Chr.; Index Filic. 525 (1906)
= Polypodium excavatum var. vesiculari-paleaceum (Hieron.) C.Chr.; Notizbl. Bot. Gart. Berlin-Dahlem 9: 186 (1924)
= Polypodium grevilleanum Wall.; Numer. List. [Wallich] no. 5169 (1831), nom. nud.
= Polypodium lineare var. simplex (C.Presl) Baker; Syn. Fil. [W. J. Hooker & Baker], ed. 2: 354 (1874)
= Polypodium mildbraedii Hieron.; Bot. Jahrb. Syst. 46: 391 (1911)
= Polypodium phlebodes Kunze ex Mett.; Abh. Senckenberg. Naturf. Ges. 2(2) [Pheg. u. Asp.]: 92, no. 163 (1857)
= Polypodium preussii Hieron.; Bot. Jahrb. Syst. 46: 386 (1911)
= Polypodium preussii var. angustifolium Hieron.; Bot. Jahrb. Syst. 46(3): 388 (1911)
= Polypodium preussii var. angustipaleaceum Hieron.; Bot. Jahrb. Syst. 46(3): 388 (1911)
= Polypodium preussii var. ledermannii Hieron.; Bot. Jahrb. Syst. 46(3): 388 (1911)
= Polypodium preussii var. winkleri Hieron.; Bot. Jahrb. Syst. 46(3): 388 (1911)
= Polypodium rotundum Bonap.; Notes Ptéridol. 16: 47 (1924)
= Polypodium simplex Sw.; J. Bot. (Schrader) 1800(2): 19 (1801), non Burm.f. 1768
= Polypodium stolzii Hieron.; Bot. Jahrb. Syst. 46: 389 (1911)
= Polypodium vesicularipaleaceum Hieron.; Bot. Jahrb. Syst. 46: 390 (1911)
species :
Lepisorus fortuni (T.Moore) C.M.Kuo; Taiwania 30: 68 (1985)
Distribution: China (Anhui, Gansu, Guangxi, Guizhou, Hubei, Hunan, Macao, Shaanxi, Sichuan, Yunnan, Zhejiang); Tibet; Ryukyu Isl.; Bonin Isl. (Otohtojima, Hahajima); Taiwan; Bhutan; NE-India (Arunachal Pradesh, Nagaland); Myanmar; Vietnam; peninsular Malaysia (Perak)
36 CHI TIB 38 NNS OGA TAI 40 BHU IND 41 MYA VIE 42 MLY
Remarks: sect. 3 Neolepisorus. L. chinensis might be a separate species. There is confusion between the synonyms of L. fortun(e)i and L. ovatus (Neocheiropteris ovata)
Conservation Status: IUCN EN for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Drynaria fortuni T.Moore; Gard. Chron. 1855: 708 (1855) ex J. Sm. in Seem. (1857)
= Lepisorus undulatus Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(4): 23 (1984)
= Microsorum chinense (Mett. ex Kuhn) Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 70 (2008)
= Microsorum excelsum Ching & S.K.Wu; Fl. Xizangica 1: 327 (1983) (fide Fraser-Jenkins 2008)
= Microsorum fortuni (T.Moore) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 304 (1933)
= Microsorum henryi (Christ) C.M.Kuo; Taiwania 30: 67 (1985) (fide Fraser-Jenkins 2008)
= Microsorum takedae (Nakai) H.Itô; J. Jap. Bot. 11: 97 (1935)
= Neocheiropteris chinensis (Mett. ex Kuhn) Fraser-Jenk., Pariyar & Kandel; Ferns Fern Allies Nepal 1: 39 (2015)
= Neocheiropteris fortuni (T.Moore) Bosman ex Fraser-Jenk., Pariyar & Kandel; Ferns Fern Allies Nepal 1: 39 (2015)
= Neocheiropteris henryi (Christ) Fraser-Jenk.; New Species Syndr. Indian Pteridol. 179-180 (1997)
= Neolepisorus fortuni (T.Moore) Li Wang; Bot. J. Linn. Soc. 162(1): 36 (2010)
= Neolepisorus phyllomanes f. deltoideus (Baker) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 14 (1940), based on Polypodium deltoideum Baker, J. Bot. 26: 230 (1880), not Sw. (1788), nor Liebmann (1849)
= Phymatodes takedae Nakai; Bot. Mag. (Tokyo) 43: 2 (1929)
= Polypodium bandoi C.Chr.; Index Filic., Suppl. Tert.: 145 (1934)
= Polypodium chinense Mett. ex Kuhn; J. Bot. 6: 270 (1868)
= Polypodium deltoideum Baker; J. Bot. 26: 230 (1888) (non Sw. (1788) nec Liebm (1849))
= Polypodium fortuni (T.Moore) E.J.Lowe; Ferns 1: t. 42B (1856), not Kunze ex Mettenius (1856)
= Polypodium henryi Christ; Bull. Herb. Boissier, sér. 1, 6: 873 (1898) (non Diels (1899))
= Polypodium normale var. polysorum Baker; J. Bot. 13: 202 (1875)
= Polypodium rosthornii Diels; Bot. Jahrb. Syst. 29: 205 (1900)
= Polypodium sinense Christ; Mém. Soc. Bot. France 1 [Hist. Acrostich.]: 16 (1905)
= Polypodium takedae (Nakai) C.Chr.; Index Filic., Suppl. Tert.: 160 (1934)
species :
Lepisorus hachijoensis Sa.Kurata; Sci. Rep. Yokosuka City Mus. 11: 40 (1965)
Distribution: South Korea; Japan (Izu Isl.)
38 JAP KOR
Remarks: sect. 15 Obtusi
Synonyms: hidden
= Pleopeltis hachijoensis (Sa.Kurata) Á.Löve & D.Löve; Taxon 26(2-3): 324 (1977)
species :
Lepisorus henryi (Hieron. ex C.Chr.) Li Wang; Bot. J. Linn. Soc. 162(1): 35 (2010)
Distribution: China (Yunnan); Nepal; Bhutan; NE-India (Assam, Sikkim, Arunachal Pradesh, Darjeeling); Myanmar (Kachin, etc.); Thailand; Laos; Vietnam
36 CHI 40 BHU IND NEP 41 LAO MYA THA VIE
Remarks: sect. 13 Belvisia
Synonyms: hidden
= Belvisia henryi (Hieron. & C.Chr.) Raym.; Mém. Jard. Bot. Montréal 55: 32 (1962)
= Belvisia henryi (Hieron. ex C.Chr.) Tagawa; H. Hara, Fl. E. Himal. 490 (1960)
= Hymenolepis henryi Hieron. ex C.Chr.; Dansk Bot. Ark. 6(3): 67 (1929)
= Macroplethus henryi (Hieron. ex C.Chr.) Tagawa; Acta Phytotax. Geobot. 11: 234 (1942)
species :
Lepisorus heterolepis (Rosenst.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 86 (1933)
Distribution: China (Yunnan)
36 CHI
Remarks: sect. 17 Pseudovittaria. Included in L. contortus by Kandel & Fraser-Jenkins (2020) but kept separate on genetic grounds by Zhao et al. 2019
Synonyms: hidden
= Pleopeltis heterolepis (Rosenst.) Tagawa & K.Iwats.; Acta Phytotax. Geobot. 22: 100 (1967)
= Polypodium heterolepis (Rosenst.) C.Chr.; Index Filic., Suppl. Tert.: 150 (1934)
= Polypodium lineare var. heterolepis Rosenst.; Repert. Spec. Nov. Regni Veg. 12: 247 (1913)
= Polypodium loriforme var. heterolepis (Rosenst.) C.Chr.; Contr. U. S. Natl. Herb. 26(6): 318 (1931)
species :
Lepisorus jakonensis (Blanf.) Fraser-Jenk. & J.Krieg.; Indian J. Forest. 42(4): 310 (2019)
Distribution: China (Yunnan); Tibet; Nepal; Bhutan; India (Himachal Pradesh, Uttarakhand, Arunachal Pradesh, Sikkim, ?Darjeeling); Myanmar (Kachin)
36 CHI TIB 40 BHU IND NEP 41 MYA
Remarks: sect. 2 Pseudoclathrati
Synonyms: hidden
= Ellipinema jakonense (Blanf.) Li Bing Zhang & Liang Zhang; Molec. Phylogen. Evol. 148: 6 (2020)
= Lepisorus jakonensis (Blanf.) Ching; Acta Bot. Yunnan. 5(1): 5 (1983), comb. inval.
= Lepisorus pseudoclathratus Ching & S.K.Wu; Acta Bot. Yunnan. 5(1): 10 (1983) (tentative synonym fide Zhao et al. 2019
= Pleopeltis jakonensis (Blanf.) S.Singh & Panigrahi; J. Hill Res. 3: 17 (1990), with incorrect basionym ref.
= Polypodium clathratum var. jakonense Blanf.; J. Simla Naturalists´ Soc. 1(2): 14 (1886)
species :
Lepisorus kawakamii (Hayata) Tagawa; Acta Phytotax. Geobot. 5: 109 (1936)
Distribution: Taiwan
38 TAI
Remarks: sect. 10 Lepisorus
Synonyms: hidden
= Polypodium arisanense Rosenst.; Hedwigia 66: 347 (1915) (non Hayata (1914))
= Polypodium kawakamii Hayata; Bot. Mag. (Tokyo) 23: 77 (1909)
= Polypodium macrosorum Baker; J. Bot. 23: 106 (1885) (non Fée (1850))
species :
Lepisorus kolesnikovii (Tzvelev) Shmakov; Opred. Paporotn. Rossii (Key Ferns Russia) 32 (1999)
Distribution: Russian Far East (Khabarovsk, Primorye)
31 RFE
Synonyms: hidden
= Pleopeltis kolesnikovii Tzvelev; Novosti Sist. Vyssh. Rast. 26: 10 (1989)
species :
Lepisorus kuchenensis (Y.C.Wu) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 69 (1933)
Distribution: China (Guangxi, Guizhou, Yunnan); +Taiwan; Vietnam
36 CHI 38 TAI 41 VIE
Remarks: sect. 10 Lepisorus
Conservation Status: very rare or extinct in Taiwan, not seen there since the 1940s
Synonyms: hidden
= Pleopeltis kuchenensis (Y.C.Wu) Á.Löve & D.Löve; Taxon 26(2-3): 324 (1977)
= Polypodium kuchenense Y.C.Wu; Bull. Dept. Biol. Sun Yatsen Univ. 3 [Polyp. Yaoshan.]: 276, t. 129 (1932)
species :
Lepisorus kuratae T.Fujiw. & Seriz.; J. Pl. Res. 131(6). 956 (2018)
Distribution: Japan (Honshu, Shikoku)
38 JAP
Remarks: An allotetraploid species derived from L. thunbergianus and an ancestral ¿race of L. tosaensis
species :
Lepisorus lewisii (Baker) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 65 (1933)
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hunan, Jiangxi, Sichuan, Zhejiang)
36 CHI
Remarks: sect. 17 Pseudovittaria
Synonyms: hidden
= Polypodium lewisii Baker; J. Bot. 13: 201 (1875)
species :
Lepisorus likiangensis Ching & S.K.Wu; Acta Bot. Yunnan. 5(1): 12 (1983)
Distribution: China (Gansu, Guizhou, Qinghai, Shaanxi, Sichuan)
36 CHI
Remarks: sect. 9 Hymenophyton. Often included in L. clathratus s. l.
Synonyms: hidden
= Lepisorus coaetaneus Ching & Y.X.Lin; Acta Bot. Yunnan. 5(1): 12 (1983)
= Lepisorus shansiensis Ching & Y.X.Lin; Acta Bot. Yunnan. 5(1): 18 (1983)
= Lepisorus shensiensis Ching & S.K.Wu; Acta Bot. Yunnan. 5(1): 14 (1983)
species :
Lepisorus lineariformis Ching & S.K.Wu; Fl. Xizangica 1: 309 (1983)
Distribution: SE-Tibet (Medog)
36 TIB
Remarks: sect. 17 Pseudovittaria. Included in L. contortus by Kandel & Fraser-Jenkins 2020 but kept separate on genetic basis by Zhao et al. 2019
species :
Lepisorus longifolius (Blume) Holttum; Rev. Fl. Malaya 2: 151 (1954)
Distribution: Myanmar; peninsular Thailand; Cambodia; Vietnam; peninsular Malaysia (widespread); Singapore; Sumatra; Borneo (Mt. Kinabalu, Sabah, Sarawak, Kalimantan); Brunei; Sulawesi; Palawan; Philippines; Java
41 CBD MYA THA VIE 42 BOR BRN JAW MLY PHI PLW SNG SUL SUM
Remarks: not in India. Sect. 1 Paragramma
Conservation Status: endangered (Singapore)
Synonyms: hidden
= Drynaria revoluta J.Sm.; J. Bot. (Hooker) 3: 421 (1841), nom. nud.
= Grammitis decurrens Blume; Enum. Pl. Javae 2: 119 (1828), nom. illeg., non Wall. ex Hook. & Grev. 1827
= Grammitis longifolia Blume; Enum. Pl. Javae 2: 119 (1828)
= Microsorum longifolium (Blume) Copel.; Univ. Calif. Publ. Bot. 16: 112 (1929)
= Niphobolus longifolius (Blume) Keyserl.; Polyp. Herb. Bunge 38 (1873)
= Paragramma decurrens (Blume) T.Moore; Index Fil. [T. Moore] 32 (1857), nom. illeg.
= Paragramma longifolia (Blume) T.Moore; Index Fil. [T. Moore] 32 (1857)
= Phymatodes longifolia (Blume) J.Sm.; Cat. Cult. Ferns: 9 (1857)
= Phymatodes revoluta (Alderw.) Tardieu & C.Chr.; Fl. Indo-Chine [P. H. Lecomte et al.] 7(2): 468 (1941)
= Phymatosorus longifolius (Blume) Bosman & Fraser-Jenk.; New Species Syndr. Indian Pteridol. 196 (1997)
= Phymatosorus revolutus (J.Sm.) V.N.Tu; J. Biol. (Vietnam) 1(2): 20 (1979)
= Pleopeltis decurrens Blume; Enum. Pl. Javae 2: addend. (1828)
= Pleopeltis longifolia (Blume) Bedd.; Ferns Brit. India 349, t. 196 (1883)
= Pleopeltis revoluta (Alderw.) Alderw.; Bull. Dépt. Agric. Indes Néerl. 27: 5 (1909)
= Polypodium contiguum Hook.; Hooker´s Icon. Pl. 6: t. 987 (1854) (nomen, non Brack. (1854))
= Polypodium decurrens (Blume) Kunze; Bot. Zeitung (Berlin) 421 (1846), nom. illeg. (non Raddi (1819))
= Polypodium longifolium (Blume) Mett.; Abh. Senckenberg. Naturf. Ges. 2(2) [Pheg. u. Asp.]: 87, no. 153, t. 1, f. 6 (1857) (non Cav. (1802) nec C. Presl (1822))
= Polypodium productum Christ; Philipp. J. Sci. 2: 178 (1907)
= Polypodium revolutum (J.Sm.) C.Chr.; Index Filic. 331 (1905), 559 (1906)
= Polypodium revolutum Alderw.; Malayan Ferns 638 (1909)
species :
Lepisorus loriformis (Wall. ex Mett.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 81 (1933)
Distribution: China (Gansu, Hubei, Shaanxi, Sichuan, Yunnan); Tibet; Nepal; Bhutan; India (N-India, Uttarakhand, Sikkim, Darjeeling, Meghalaya, Arunachal Pradesh); Myanmar (Kachin)
36 CHI TIB 40 BHU IND NEP 41 MYA
Remarks: sect. 16 Sclerophyllon. L. steniste might be a separate species
Synonyms: hidden
= Drynaria loriformis (Wall. ex Mett.) J.Sm.; J. Bot. (Hooker) 4: 61 (1841)
= Lepisorus loriformis f. steniste (C.B.Clarke) Kholia; Ferns Fern Allies Sikkim 2: 126 (2014)
= Lepisorus loriformis var. longifolius Bir & Trikha; Amer. Fern J. 64: 63 (1974)
= Lepisorus loriformis var. steniste (C.B.Clarke) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4(3): 82 (1933)
= Lepisorus steniste (C.B.Clarke) Y.X.Lin; Fl. Reipubl. Popularis Sin. [Fl. China Ed. Committee] 6(2): 69 (2000)
= Lepisorus tenuicauda Ching; Acta Bot. Austro Sin. 1: 22 (1983 (fide Fraser-Jenkins 1997)
= Lepisorus xiphopteris (Baker) W.M.Chu ex Y.X.Lin; Fl. Reipubl. Popularis Sin. [Fl. China Ed. Committee] 6(2): 68 (2000)
= Lepisorus yunnanensis Ching; Bull. Fan Mem. Inst. Biol. Bot. 11: 75 (1941)
= Phymatodes loriformis (Wall. ex Mett.) C.Presl; Tent. Pterid. 196 (1836)
= Pleopeltis linearis var. steniste (C.B.Clarke) Bedd.; Ferns Brit. India 347 (1883)
= Pleopeltis loriformis (Wall. ex Mett.) T.Moore; Index Fil. [T. Moore] 346 (1862)
= Pleopeltis loriformis f. angustifrons (Takeda) C.Chr.; Contr. U. S. Natl. Herb. 26(6): 318 (1931)
= Pleopeltis loriformis var. steniste (C.B.Clarke) Panigrahi & Patnaik; Proc. Nat. Inst. Sci. India B 34: 481 (1964)
= Pleopeltis tenuicauda (Ching) Panigrahi; Pl. Sci. Res. (India), Abstr. 23rd. Ann. Conf. Khurda: 35 (1998)
= Polypodium excavatum var. loriforme (Wall. ex Mett.) C.Chr.; Index Filic. 9: 525 (1906)
= Polypodium lineare f. steniste (C.B.Clarke) Takeda; Notes Roy. Bot. Gard. Edinburgh 8(39): 273 (1915)
= Polypodium lineare subsp. loriforme (Wall. ex Mett.) C.Chr.; Index Filic., Suppl. 2 (1906-1912): 54 (1917)
= Polypodium lineare var. loriforme (Wall. ex Mett.) Takeda; Notes Roy. Bot. Gard. Edinburgh 8: 272 (1915)
= Polypodium lineare var. steniste C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 559 (1880)
= Polypodium loriforme f. angustifrons (Takeda) C.Chr.; Contr. U. S. Natl. Herb. 26: 318 (1931)
= Polypodium loriforme var. steniste (C.B.Clarke) C.Chr.; Contr. U. S. Natl. Herb. 26: 318 (1931)
= Polypodium loriforme Wall.; Numer. List. [Wallich] no. 271 (1829), nomen
= Polypodium loriforme Wall. ex Mett.; Abh. Senckenberg. Naturf. Ges. 2(1) [Polypodium]: 92 (1856)
= Polypodium mengtzeanum Baker; Bull. Misc. Inform. Kew 1906: 14 (1906)
= Polypodium oblongisorum C.Chr.; Index Filic. 549 (1906)
= Polypodium subimmersum Baker; Bull. Misc. Inform. Kew 1895: 55 (1895)
= Polypodium subimmersum f. angustifrons Takeda; Notes Roy. Bot. Gard. Edinburgh 8(39): 276 (1915)
= Polypodium subimmersum f. mengtzeanum (Baker) Takeda; Notes Roy. Bot. Gard. Edinburgh 8(39): 276 (1915)
= Polypodium subintegrum Baker; Bull. Misc. Inform. Kew 1898: 231 (1898) (non Baker (1877))
= Polypodium xiphiopteris Baker; Bull. Misc. Inform. Kew 1906: 13 (1906)
species :
Lepisorus luchunensis Y.X.Lin; Fl. Reipubl. Popularis Sin. [Fl. China Ed. Committee] 6(2): 50, t. 8 (2000)
Distribution: China (Yunnan)
36 CHI
Remarks: sect. 10 Lepisorus. Status not finally confirmed
species :
Lepisorus macrosphaerus (Baker) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 73 (1933)
Distribution: China (Guizhou, Hubei, Sichuan, Yunnan); Tibet; Bhutan
36 CHI TIB 40 BHU
Remarks: sect. 10 Lepisorus. L. asterolepis is included here by Fraser-Jenkins et al. 2015 but kept separate by Zhao et al. 2019
Synonyms: hidden
= Lepisorus macrosphaerus f. maximus (Ching) Y.X.Lin; Fl. Reipubl. Popularis Sin. [Fl. China Ed. Committee] 6(2): 64 (2000)
= Lepisorus macrosphaerus f. minimus (Ching) Y.X.Lin; Fl. Reipubl. Popularis Sin. [Fl. China Ed. Committee] 6(2): 65 (2000)
= Lepisorus macrosphaerus var. maximus Ching; Fl. Xizangica 1: 303 (1983)
= Lepisorus macrosphaerus var. minimus Ching; Fl. Xizangica 1: 304 (1983)
= Pleopeltis macrosphaera (Baker) Panigrahi & Patnaik; Proc. Nat. Acad. Sci. India, B 34: 481 (1964), nom. superfl.
= Pleopeltis macrosphaera (Baker) Pic.Serm.; Webbia 23(1): 189 (1968), nom. superfl.
= Polypodium intramarginale Baker; Christ, Bull. Herb. Boissier, sér. 2, 3: 509 (1903)
= Polypodium macrosphaerum Baker; Bull. Misc. Inform. Kew 1895: 55 (1895)
= Polypodium simplex var. esquirolii Christ; Bull. Acad. Int. Géogr. Bot. 15: 247 (1906)
species :
Lepisorus maculosus (Christ) C.F.Zhao, R.Wei & X.C.Zhang; Cladistics 36(3): 252 (2019)
Distribution: China (Sichuan, Yunnan, etc.); Tibet; Bhutan; India (Arunachal Pradesh, Manipur, Nagaland, Mizoram); Myanmar; Thailand; Laos; Vietnam; peninsular Malaysia; Sumatra
36 CHI TIB 40 BHU IND 41 LAO MYA THA VIE 42 MLY SUM
Remarks: sect. 5 Tricholepidium. Often included in L. normale but kept separate by Zhao et al. 2019. Neocheiropteris venosa could be a separate species but requires a new combination in Lepisorus.
Synonyms: hidden
= Lepisorus chapaensis C.Chr. & Tardieu; Notul. Syst. (Paris) 8: 186 (1939)
= Microsorum subnormale (Nakai) H.Itô; J. Jap. Bot. 11: 97 (1935)
= Neocheiropteris maculosa (Christ) Fraser-Jenk.; Indian J. Forest. 42(4): 314 (2019)
= Neocheiropteris venosa (Ching) Fraser-Jenk.; Indian J. Forest. 42(4): 314 (2019)
= Phymatodes subnormalis Nakai; Bot. Mag. (Tokyo) 43: 3 (1929)
= Pleopeltis subnormalis Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 3, 2: 165 (1920)
= Polypodium maculosum Christ; Bull. Herb. Boissier, sér. 1, 6: 872 (1896)
= Polypodium subnormale (Alderw.) C.Chr.; Index Filic., Suppl. Tert.: 159 (1934)
= Tricholepidium angustifolium Ching; Acta Phytotax. Geobot. 29(1-5): 44 (1978)
= Tricholepidium angustifolium var. falcatolineare Ching; Acta Phytotax. Geobot. 29: 44 (1978)
= Tricholepidium angustifolium var. lanceolatum (Ching & S.K.Wu) Y.X.Lin; Fl. Reipubl. Popularis Sin. [Fl. China Ed. Committee] 6(2): 106 (2000)
= Tricholepidium chapaense (C.Chr. & Tardieu) Ching; Acta Phytotax. Geobot. 29(1-5): 44 (1978)
= Tricholepidium intermedium Ching; Acta Phytotax. Geobot. 29(1-5): 44 (1978)
= Tricholepidium lanceolatum Ching & S.K.Wu; Acta Phytotax. Geobot. 29(1-5): 45 (1978)
= Tricholepidium maculosum (Christ) Ching; Acta Phytotax. Geobot. 29(1-5): 45 (1978)
= Tricholepidium maculosum var. subnormale (Alderw.) Ching; Acta Phytotax. Geobot. 29(1-5): 45 (1978)
= Tricholepidium subnudum Ching & S.K.Wu; Fl. Xizangica 1: 314 (1983)
= Tricholepidium tibeticum Ching & S.K.Wu; Acta Phytotax. Geobot. 29(1-5): 46 (1978)
= Tricholepidium venosum Ching; Acta Phytotax. Geobot. 29(1-5): 45 (1978)
species :
Lepisorus mamas Hovenkamp; Blumea 43: 112 (1998)
Distribution: Sumatra
42 SUM
species :
Lepisorus marginatus Ching; Fl. Tsinling. 2: 233 (1974), without type
Distribution: China (Chongqing, Gansu, Hebei, Henan, Hubei, Shaanxi, Shanxi, Sichuan)
36 CHI
Remarks: sect. 10 Lepisorus
species :
Lepisorus medioximus T.Fujiw., K.Hori & Khine; PhytoKeys 201: 26-27 (2022)
Distribution: Myanmar (Shan)
41 MYA
Remarks: sect. 17 Pseudovittaria
species :
Lepisorus medogensis Ching & Y.X.Lin; Acta Phytotax. Sin. 22(5): 401 (1984)
Distribution: SE-Tibet (Medog)
36 TIB
Remarks: sect. 17 Pseudovittaria
Synonyms: hidden
= Lepisorus zosterifolius Ching & Y.X.Lin; Acta Phytotax. Sin. 22(5): 401 (1984)
species :
Lepisorus megasorus (C.Chr.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 79 (1933)
Distribution: Taiwan
38 TAI
Remarks: sect. 10 Lepisorus
Synonyms: hidden
= Polypodium hypochrysum Hayata; Icon. Pl. Formosan. 6: 314, f. 127 (1915)
= Polypodium megasorum C.Chr.; Index Filic. 544 (1906), based on later homonym P. macrosorum Baker, not Fée (1850)
species :
Lepisorus mehrae Fraser-Jenk.; New Species Syndr. Indian Pteridol. 159 (1997)
Distribution: China (Yunnan, ?Sichuan); Tibet; Nepal; Bhutan; N-India (Himachal Pradesh, Uttarakhand, Sikkim, Darjeeling); Myanmar (Kachin, Chin)
36 CHI TIB 40 BHU IND NEP 41 MYA
Synonyms: hidden
= Lepisorus krameri P.C.Pande & H.C.Pande; Indian Fern J. 11(1-2): 98 (1994 publ. 1995), et Illustr. Fern Fl. Kumaon Himalaya 1: 68 (2003), nom. nud. (fide Fraser-Jenkins 2015)
= Lepisorus paleparaphysus Y.X.Lin; Fl. Reipubl. Popularis Sin. [Fl. China Ed. Committee] 6(2): 75 (2000)
species :
Lepisorus microphyllus (C.Presl) Ebihara & T.Fujiw.; Bull. Nat. Mus. Nat. Sci., Ser. B, 49(3): 101 (2023)
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Hainan, Hubei, Jiangxi, Macao, Yunnan, Zhejiang); South Korea; Japan; Taiwan; NE-India (Arunachal Pradesh); Myanmar (Kachin, etc.); Vietnam
36 CHI 38 JAP KOR TAI 40 IND 41 MYA VIE
Remarks: according to Knapp (Ferns of Taiwan, 2011 ff.) var. obovatum should be included in the typical variety. The Taiwanese plants belong possibly to L. carnosum fide Knapp (Ferns of Taiwan, 2011 ff.)
Conservation Status: IUCN VU for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Drymoglossum carnosum var. obovatum Harr.; J. Linn. Soc., Bot. 16: 33 (1877)
= Drymoglossum microphyllum (C.Presl) C.Chr.; Index Filic. 246 (1905)
= Drymoglossum nobukoanum Makino; J. Jap. Bot. 7: 8 (1931)
= Drymoglossum obovatum (Harr.) Christ; J. Bot. (Morot) 19: 73 (1905)
= Drymoglossum obovatum var. lutchuense Nakai; Bot. Mag. (Tokyo) 40(475): 396 (1926)
= Drymoglossum spatulatum C.Presl; Tent. Pterid. 227 (1836)
= Lemmaphyllum carnosum subsp. microphyllum (C.Presl) Fraser-Jenk. & Kandel; Indian J. Forest. 42(4): 310 (2019)
= Lemmaphyllum carnosum var. microphyllum (C.Presl) X.P.Wei; J. Syst. Evol. 51(4): 493 (2013)
= Lemmaphyllum microphyllum C.Presl; Abh. Königl. Böhm. Ges. Wiss., ser. 5, 6 [Epim. Bot.]: 263 (1851)
= Lemmaphyllum microphyllum var. lutchuense (Nakai) C.Chr.; Dansk Bot. Ark. 6(3): 47, pl. 5, f. 4 (1929)
= Lemmaphyllum microphyllum var. nobukoanum (Makino) Nakaike; Enum. Pterid. Jap., Filic. 332 (1975)
= Lemmaphyllum microphyllum var. obovatum (Harr.) C.Chr.; Dansk Bot. Ark. 6(3): 47 (1929)
= Lemmaphyllum microphyllum var. spathulatum (C.Presl) Nakaike; J. Nippon Fernist Club 3(Suppl. 2): 104 (2004)
= Lemmaphyllum minimum S.H.Fu; Acta Phytotax. Sin. 5: 75, f. 1-4 (1956)
= Lemmaphyllum nobukoanum (Makino) Ching; Icon. Filic. Sin. ad., t. 80 (1934)
= Lemmaphyllum spathulatum C.Presl; Abh. Königl. Böhm. Ges. Wiss., ser. 5, 6: 158 (1851)
= Lemmaphyllum squamosum C.Chr.; Dansk Bot. Ark. 6(3): 48 (1929)
= Taenitis microphylla (C.Presl) Mett.; Fil. Afr. [Kuhn] 58 (1868)
species :
Lepisorus mikawanus Sa.Kurata; Sci. Rep. Yokosuka City Mus. 11: 38 (1965)
Distribution: Japan
38 JAP
Remarks: hexaploid species in the thunbergianus complex
Conservation Status: IUCN R
species :
Lepisorus minor (W.M.Chu) C.F.Zhao, R.Wei & X.C.Zhang; Cladistics 36(3): 251 (2019)
Distribution: China (Yunnan)
36 CHI
Remarks: sect. 3 Neolepisorus. Sometimes included in L. ensatus
Synonyms: hidden
= Neolepisorus minor W.M.Chu; Acta Bot. Yunnan. 1(2): 95 (1979) (fide Fraser-Jenkins 2008)
species :
Lepisorus mishmica Benniamin & Fraser-Jenk.; Indian J. Forest. 42(4): 310 (2019)
Distribution: India (Arunachal Pradesh)
40 IND
Remarks: sect. 17 Pseudovittaria
species :
Lepisorus miyoshianus (Makino) Fraser-Jenk. & Subh.Chandra; Taxon. Revis. Indian Subcontinental Pteridophytes 37 (2008)
Distribution: China (Guangdong, Guizhou, Hubei, Hunan, Shaanxi, Sichuan, Yunnan, Zhejiang); Tibet; Japan; Taiwan; India (Arunachal Pradesh)
36 CHI TIB 38 JAP TAI 40 IND
Remarks: sect. 14 Drymotaenium
Conservation Status: IUCN CR for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Drymotaenium miyoshianum (Makino) Makino; Bot. Mag. (Tokyo) 15: 109 (1901)
= Drymotaenium nakaii Hayata; Bull. Soc. Bot. France 58: 565, t. 19 (1911)
= Monogramma robusta (Christ) C.Chr.; Index Filic. 431 (1906)
= Pleurogramme robusta Christ; Bull. Herb. Boissier, sér. 1, 6: 867 (1898)
= Taenitis miyoshiana Makino; Bot. Mag. (Tokyo) 11: 26 (1898)
species :
Lepisorus monilisorus (Hayata) Tagawa; Acta Phytotax. Geobot. 11: 303 (1942)
Distribution: Taiwan
38 TAI
Remarks: sect. 15 Obtusi
Synonyms: hidden
= Polypodium lineare var. monilisorum Hayata; Icon. Pl. Formosan. 4: 248, f. 173 (1914)
species :
Lepisorus morrisonensis (Hayata) H.Itô; J. Jap. Bot. 11: 92 (1935)
Distribution: China (Sichuan, Yunnan); Tibet; Taiwan; Nepal; Bhutan; India (Uttarakhand, Sikkim, ?Darjeeling, Meghalaya, Himachal Pradesh, Jammu & Kashmir, Arunachal Pradesh); Myanmar; N-Thailand
36 CHI TIB 38 TAI 40 BHU IND NEP 41 MYA THA
Remarks: sect. 8 Pachyphyllon
Synonyms: hidden
= Lepisorus mediocris Ching & Khullar; (1988) (nom. nud., fide Fraser-Jenkins (1997))
= Lepisorus reichsteinii Khullar; (1988) (nom. nud., fide Fraser-Jenkins (1997))
= Polypodium morrisonense Hayata; Bot. Mag. (Tokyo) 23: 77 (1909)
species :
Lepisorus mucronatus (Fée) Li Wang; Bot. J. Linn. Soc. 162(1): 35 (2010)
Name: Needle fern, Tailed fern
Distribution: China (Yunnan); Tibet; Taiwan; Sri Lanka; Myanmar (Kachin, etc.); Thailand; Laos; Cambodia; Vietnam; peninsular Malaysia (widespread); Sumatra; Borneo (Mt. Kinabalu, Sabah, Sarawak); Sulawesi; Palawan; Philippines; Java; Lesser Sunda Isl. (Bali, Lombok, Timor, etc.); Moluccas (Seram, etc.); New Guinea (Irian Jaya, Papua New Guinea); Bismarck Arch. (New Ireland, New Hanover); Solomon Isl. (Bougainville, Guadalcanal, Kolombangara, Malaita, New Georgia, Rendova, Rennell, Santa Ysabel, Vella Lavella, Makira); Australia (E-Queensland, NE-New South Wales); New Caledonia; Santa Cruz Isl.; Vanuatu; Tonga (Kao, Tafahi); Fiji (Viti Levu, Vanua Levu, Kadavu, Gau, Koro, Ovalau, Taveuni); [Western] Samoa (Savai´i); Cook Isl. (Rarotonga); Marquesas Isl. (Nuku Hiva, Ua Huka, Ua Pou, Hiva Oa, Tahuata, Fatu Hiva); Austral Isl. (Tubuai, Raivavae); Palau (Babeldaob, Koror, Malakal); Northern Marianas (Rota Isl.); Guam
36 CHI TIB 38 TAI 40 SRL 41 CBD LAO MYA THA VIE 42 BOR JAW LSI MLY MOL NWG PHI PLW SUL SUM 43 BIS SOL 50 AUS 60 FIJ NWC SAM SCZ TON VAN 61 AUI COO MRQ 62 GUM MRN PLU
Remarks: sect. 13 Belvisia
Synonyms: hidden
= Acrostichum spicatum var. schneideri F.M.Bailey; Compr. Cat. Queensland Pl. 649, 849, f. 613 (1913)
= Belvisia callifolia (Christ) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 192 (1947)
= Belvisia formosana (M.Ogata) Ching; Acta Phytotax. Sin. 10: 301 (1965)
= Belvisia melanesica Brownlie; Nova Hedwigia Beih. 55 (Pterid. Fl. Fiji): 369 (1977) (probably a valid species)
= Belvisia minor (Copel.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 192 (1947)
= Belvisia mucronata (Fée) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 192 (1947)
= Belvisia vaupelii (C.Chr.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 192 (1947)
= Hymenolepis callifolia Christ; Ann. Jard. Bot. Buitenzorg, ser. 2, 5: 128 (1905)
= Hymenolepis formosana M.Ogata; J. Jap. Bot. 11: 28, t. 1(A) (1935)
= Hymenolepis minor Copel.; Occas. Pap. Bernice P. Bishop Mus. 14: 69, t. 19 (1938)
= Hymenolepis mucronata f. australiensis C.Chr.; Dansk Bot. Ark. 6(3): 64 (1929)
= Hymenolepis mucronata f. graminifolia (Rosenst.) C.Chr.; Dansk Bot. Ark. 6(3): 64 (1929)
= Hymenolepis mucronata f. helocharidioides C.Chr.; Dansk Bot. Ark. 6(3): 64 (1929)
= Hymenolepis mucronata f. latior C.Chr.; Dansk Bot. Ark. 6(3): 65 (1929)
= Hymenolepis mucronata Fée; Mém. Foug. 5 (Gen. Filic.): 82 (1851-52)
= Hymenolepis mucronata var. nigropunctata C.Chr.; Dansk Bot. Ark. 6(3): 64 (1929)
= Hymenolepis ophioglossoides Kunze; Farnkräuter 99, t. 47, f. 1 (1842) (non Kaulf.)
= Hymenolepis spicata f. longipaleacea Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 7: 19 (1912)
= Hymenolepis spicata f. schneideri (F.M.Bailey) Alderw.; Malayan Ferns Fern Allies Suppl., 432 (1917)
= Hymenolepis spicata var. bakhuizenii Alderw.; Malayan Ferns Fern Allies Suppl., 432, corr. 59 (1917)
= Hymenolepis spicata var. graminifolia Rosenst.; Repert. Spec. Nov. Regni Veg. 12: 530 (1913)
= Hymenolepis spicata var. novoguineensis Rosenst.; Hedwigia 56: 353 (1915)
= Hymenolepis spicata var. squamulifera Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 7: 19 (1912)
= Hymenolepis vaupelii Hieron. ex C.Chr.; Dansk Bot. Ark. 6(3): 65 (1929)
= Macroplethus callifolius (Christ) Tagawa; Acta Phytotax. Geobot. 11: 234 (1942)
= Macroplethus mucronatus (Fée) Tagawa; Acta Phytotax. Geobot. 11: 234 (1942)
= Macroplethus vaupelii (Hieron. ex C.Chr.) Tagawa; Acta Phytotax. Geobot. 11: 235 (1942)
variant :
var. durus (Copel.) Rouhan; Phytotaxa 497(1): 54 (2021)
Distribution: Austral Isl. (Rapa Iti)
61 AUI
Synonyms: hidden
= Belvisia dura (Copel.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 192 (1947)
= Belvisia mucronata var. dura (Copel.) Hovenkamp & Franken; Blumea 37(2): 523 (1993)
= Hymenolepis dura Copel.; Occas. Pap. Bernice P. Bishop Mus. 14: 69, t. 20 (1938)
species :
Lepisorus nigripes T.Fujiw. & Seriz.; J. Pl. Res. 131(6): 956 (2018)
Distribution: Japan (Hokkaido, Honshu, Shikoku, Kyushu); Ryukyu Isl. (Yaku Isl.)
38 JAP NNS
Remarks: an allotetraploid species derived from L. thunbergianus and L. angustus
species :
Lepisorus normalis (D.Don) C.F.Zhao, R.Wei & X.C.Zhang; Cladistics 36(3): 252 (2019)
Distribution: China (Guangxi, Yunnan); Tibet; Japan; Nepal; Bhutan; NE-India (Arunachal Pradesh, Sikkim, Darjeeling); Myanmar (Kachin, etc.); Thailand; Laos; ?Cambodia; Vietnam; peninsular Malaysia (Pahang); Sumatra
36 CHI TIB 38 JAP 40 BHU IND NEP 41 LAO MYA THA VIE 42 MLY SUM
Remarks: sect. 5 Tricholepidium
Synonyms: hidden
= Colysis normalis (D.Don) J.Sm.; Hist. Fil. 100 (1875)
= Drynaria longifrons (Wall. ex Hook. & Grev.) J.Sm.; J. Bot. (Hooker) 3: 397 (1841)
= Drynaria normalis (D.Don) J.Sm.; J. Bot. (Hooker) 4: 61 (1841)
= Microsorum normale (D.Don) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 295-311 (1933)
= Neocheiropteris normalis (D.Don) Tagawa; J. Jap. Bot. 27: 217 (1952)
= Neolepisorus normalis (D.Don) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 13 (1940)
= Phymatodes normalis (D.Don) C.Presl; Tent. Pterid. 196 (1836)
= Pleopeltis longifrons (Wall. ex Hook. & Grev.) T.Moore; Index Fil. [T. Moore] 346 (1862)
= Pleopeltis normalis (D.Don) T.Moore; Index Fil. [T. Moore] 347 (1862)
= Polypodium longifrons Wall.; Numer. List. [Wallich] no. 274 (1829), nomen
= Polypodium longifrons Wall. ex Hook. & Grev.; Icon. Filic. 1: t. 65 (1828)
= Polypodium normale D.Don; Prodr. Fl. Nepal. 1 (1825)
= Tricholepidium mutense Ching & S.K.Wu; Acta Phytotax. Geobot. 29(1-5): 43 (1978)
= Tricholepidium normale (D.Don) Ching; Acta Phytotax. Geobot. 29(1-5): 43 (1978)
= Tricholepidium pteropodum Ching; Acta Phytotax. Geobot. 29(1-5): 43 (1978)
species :
Lepisorus novoguineensis (Rosenst.) Li Wang; Bot. J. Linn. Soc. 162(1): 35 (2010)
Distribution: Sulawesi; New Guinea (Irian Jaya, Papua New Guinea); Bismarck Arch. (New Ireland)
42 NWG SUL 43 BIS
Remarks: sect. 13 Belvisia
Synonyms: hidden
= Belvisia novoguineensis (Rosenst.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 192 (1947)
= Hymenolepis novoguineensis (Rosenst.) C.Chr.; Index Filic., Suppl. Tert.: 113 (1934)
= Lemmaphyllum novoguineense (Rosenst.) C.Chr.; Dansk Bot. Ark. 6(3): 51 (1929)
= Macroplethus novoguineensis (Rosenst.) Tagawa; Acta Phytotax. Geobot. 11: 234 (1942)
= Paltonium novoguineense Rosenst.; Nova Guinea 8: 729 (1912)
species :
Lepisorus nudus (Hook.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 83 (1933)
Distribution: China (Yunnan); Tibet; Japan; Pakistan (Hazara); Nepal; Bhutan; India (throughout, incl. Jammu & Kashmir, Uttarakhand, Himachal Pradesh, Arunachal Pradesh, Darjeeling, Sikkim); Sri Lanka; Bangladesh; Myanmar (Kachin, Chin, etc.); Thailand; Vietnam; Sumatra
36 CHI TIB 38 JAP 40 BAN BHU IND NEP PAK SRL 41 MYA THA VIE 42 SUM
Remarks: sect. 10 Lepisorus
Synonyms: hidden
= Drynaria nuda (Hook.) Fée; Mém. Foug. 5 (Gen. Filic.): 270 (1852)
= Drynaria sesquipedalis (Wall.) J.Sm.; Compan. Bot. Mag. 13 (1846) (fide Fraser-Jenkins 2008)
= Drynaria zeylanica Fée; Mém. Foug. 5 (Gen. Filic.): 272 (1852)
= Lepisorus amaurolepidus var. longifolius Bir & Trikha; Amer. Fern J. 64: 63, f. 53-57 (1974) (fide Fraser-Jenkins 2008)
= Lepisorus birii Khullar; Khullar et al. 1991 (nom. nud., synonymy fide Fraser-Jenkins (1997))
= Lepisorus gyirongensis Ching & S.K.Wu; Fl. Xizangica 1: 304 (1983) (fide Fraser-Jenkins 1997)
= Lepisorus intermedius Ching & Khullar; Khullar, Illustr. Fern Fl. W. Himalaya 1: 93 (1994) (fide Fraser-Jenkins 1997)
= Lepisorus leiopteris (Kunze) Bir & Trikha; Amer. Fern J. 64: 54, f. 21-26 (1974) (fide Fraser-Jenkins 2008)
= Lepisorus parvus Khullar; (1988) (nom. nud., synonymy fide Fraser-Jenkins (1997))
= Lepisorus pseudolinearis Ching & Khullar; (1988) (nom. nud., fide Fraser-Jenkins (1997))
= Lepisorus sequipedalis (Wall. ex Mett.) Fraser-Jenk.; Pakistan Syst. 5: 91 (1992), nom. superfl.
= Lepisorus tenuipes Ching & Khullar; Indian Fern J. 1(1-2): 91 (1984 publ. 1985) (fide Fraser-Jenkins 1997)
= Niphobolus sesquipedalis (Wall. ex Mett.) Keyserl.; Polyp. Herb. Bunge 39 (1873) (fide Fraser-Jenkins 2008)
= Phymatodes nuda (Hook.) J.Sm.; Bot. Voy. Herald [Seemann] 425 (1857)
= Pleopeltis leiopteris (Kunze) S.Singh & Panigrahi; J. Hill Res. 3: 18 (1990) (fide Fraser-Jenkins 2008)
= Pleopeltis linearis var. leiopteris (Kunze) C.Chr.; Index Fil. [T. Moore] 540 (1906)
= Pleopeltis nuda Hook.; Exot. Fl. 1: t. 63 (1823)
= Pleopeltis tenuipes (Ching & Khullar) Panigrahi; Pl. Sci. Res. (India) 20: 22 (1999)
= Polypodium leiopteris Kunze; Linnaea 23: 279 319 (1859) (fide Fraser-Jenkins 2008)
= Polypodium lineare var. wightianum C.Chr.; Index Fil. [T. Moore] 540 (1906)
= Polypodium nudiusculum Kunze; Linnaea 24: 253 (1851)
= Polypodium nudum (Hook.) Kunze; Linnaea 23: 281 (1850), nom. illeg.
= Polypodium sesquipedale f. leiopteris (Kunze) Makino; Phan. Pter. Jap. Icon. 1: t. 8 (1899)
= Polypodium sesquipedale Wall.; Numer. List. [Wallich] no. 275 (1829), nomen
= Polypodium sesquipedale Wall. ex Mett.; Abh. Senckenberg. Naturf. Ges. 2(1) [Polypodium]: 91 (1856)
= Polypodium wightianum Thwaites; Enum. Pl. Zeyl.: 394 (1864), nom. superfl.
= Polypodium wightianum Wall.; Numer. List. [Wallich] no. 2222 (1829), nomen
species :
Lepisorus obscurevenulosus (Hayata) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 76 (1933)
Distribution: China (Anhui, Chongqing, Fujian, Guangdong, Guangxi, Guizhou, Hunan, Jiangxi, Sichuan, Yunnan, Zhejiang); Taiwan; Vietnam
36 CHI 38 TAI 41 VIE
Remarks: sect. 17 Pseudovittaria. If synonymy of P. caudiceps is confirmed, this name would have priority
Synonyms: hidden
= Goniophlebium caudiceps T.Moore; Gard. Chron., ser. 2, 25: 234 (1886)
= Polypodium caudiceps (T.Moore) G.Nicholson; Ill. Dict. Gard. 4: 592 (1888)
= Polypodium feei var. caudiceps (T.Moore) Baker; Ann. Bot. (Oxford) 5: 469 (1891)
= Polypodium obscurevenulosum Hayata; Icon. Pl. Formosan. 6: 322, f. 134 (1915)
= Polypodium suprapunctatum Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 76 (1933)
species :
Lepisorus oligolepidus (Baker) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 80 (1933)
Distribution: China (Anhui, Chongqing, Fujian, Guangdong, ?Guangxi, Guizhou, ?Henan, Hunan, Jiangxi, Shaanxi?, Sichuan, Yunnan, Zhejiang); ?Tibet; Japan; NE-India (Arunachal Pradesh, Manipur, Meghalaya, Tripura); Myanmar
36 CHI 38 JAP 40 IND 41 MYA
Remarks: sect. 17 Pseudovittaria. Not in India
Synonyms: hidden
= Lepisorus ellipticus Ching; Bull. Bot. Res., Harbin 2(2): 74 (1982)
= Pleopeltis oligolepida (Baker) Á.Löve & D.Löve; Taxon 26(2-3): 325 (1977)
= Polypodium lineare var. oligolepidum (Baker) Christ; Bull. Acad. Int. Géogr. Bot. sér. 3, 11(153-154): 208 (1902), et Mém. Soc. Bot. France 1 [Hist. Acrostich.]: 15 (1905)
= Polypodium oligolepidum Baker; Gard. Chron., ser. 2, 14: 594 (1880)
= Polypodium trabeculatum Copel.; Philipp. J. Sci. 3: 283 (1908)
species :
Lepisorus onoei (Franch. & Sav.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 87 (1933)
Distribution: South Korea (incl. Cheju Isl.); Japan; Ryukyu Isl.
38 JAP KOR NNS
Remarks: sect. 15 Obtusi
Synonyms: hidden
= Pleopeltis onoei (Franch. & Sav.) Okuyama ex Ohwi; Bull. Natl. Sci. Mus., Tokyo, 3: 99 (1956)
= Polypodium lineare var. onoei (Franch. & Sav.) Makino; Bot. Mag. (Tokyo) 11: 282 (1897)
= Polypodium onoei Franch. & Sav.; Enum. Pl. Jap. 2: 246, 642 (1877)
species :
Lepisorus oosphaerus (C.Chr.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 73 (1933)
Distribution: N-Thailand
41 THA
Remarks: not in India
Conservation Status: VU in Thailand
Synonyms: hidden
= Pleopeltis oosphaera (C.Chr.) Panigrahi & Patnaik; Proc. Nat. Acad. Sci. India, B, 34: 482 (1964)
= Polypodium oosphaerum C.Chr.; Contr. U. S. Natl. Herb. 26: 334, t. 29 (1931)
species :
Lepisorus ovatus (C.Presl) R.Kr.Singh & V.K.Rawat; J. Biodivers. Conservation 8(3): 81 (2024)
Distribution: China (Anhui, Chongqing, Guangdong, Guangxi, Guizhou, Hubei, Hunan, Jiangsu, Jiangxi, Sichuan, Yunnan, Zhejiang); Japan; Nepal; Bhutan; India (Meghalaya, Sikkim, Arunachal Pradesh, Darjeeling); Bangladesh; Myanmar; Vietnam
36 CHI 38 JAP 40 BAN BHU IND NEP 41 MYA VIE
Remarks: sect. 3 Neolepisorus. There is confusion between the synonyms of L. ovatus and L. fortun(e)i which need to be sorted out better.
Synonyms: hidden
= Colysis ovata (C.Presl) J.Sm.; Hist. Fil. 100 (1875)
= Drynaria ovata (C.Presl) Fée; Mém. Foug. 5 (Gen. Filic.): 270 (1852)
= Lepisorus ovatus (C.Presl) C.F.Zhao, R.Wei & X.C.Zhang; Cladistics 36(3): 251 (2019), nom. inval. (without basionym ref.)
= Microsorum ensatum var. phyllomanes (Christ) Tagawa; Acta Phytotax. Geobot. 8(3): 172 (1939)
= Microsorum ovatum (C.Presl) N.C.Nair & Bennet; Bull. Bot. Surv. India 11: 432 (1972)
= Microsorum phyllomanes (Christ) Koidz.; Acta Phytotax. Geobot. 5: 50 (1936), nom. superfl.
= Neocheiropteris ovata (C.Presl) Fraser-Jenk.; New Species Syndr. Indian Pteridol. 180 (1997), isonym
= Neocheiropteris ovata (C.Presl) B.K.Nayar & S.Kaur; Companion Ferns Brit. India 86 (1974)
= Neocheiropteris ovata f. deltoidea (Ching) Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 68 (2008)
= Neocheiropteris ovata f. doryopteris (Christ) Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 68 (2008)
= Neocheiropteris ovatus (Wall. ex Hook. & Grev.) B.K.Nayar & Kaur; Companion Ferns Brit. India 86 (1974) (comb. inval.)
= Neocheiropteris phyllomanes (Christ) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 110 (1933), nom. superfl.
= Neocheiropteris phyllomanes f. deltoidea (Hand.-Mazz.) Ching
= Neocheiropteris phyllomanes var. doryopteris (Christ) Ching
= Neolepisorus basicordatus P.S.Wang; Acta Bot. Yunnan. 9(4): 399 (1987)
= Neolepisorus crenatus S.F.Wu; Keys Vasc. Pl. Wuling Mts. 573 (1995)
= Neolepisorus cuneatus S.F.Wu; Keys Vasc. Pl. Wuling Mts. 577 (1995)
= Neolepisorus dengii Ching & P.S.Wang; Acta Phytotax. Sin. 21(3): 272 (1983)
= Neolepisorus dengii f. hastatus Ching & P.S.Wang; Acta Phytotax. Sin. 21(3): 273 (1983)
= Neolepisorus emeiensis Ching & K.H.Shing; Acta Phytotax. Sin. 21(3): 271 (1983)
= Neolepisorus emeiensis f. dissectus Ching & K.H.Shing; Acta Phytotax. Sin. 21(3): 271 (1983)
= Neolepisorus lancifolius Ching & K.H.Shing; Acta Phytotax. Sin. 21(3): 271 (1983)
= Neolepisorus ovatus (C.Presl) Ching; Acta Phytotax. Sin. 9: 99 (1964)
= Neolepisorus ovatus f. deltoideus Ching; Acta Phytotax. Sin. 9: 99 (1964)
= Neolepisorus ovatus f. doryopteris (Christ) Ching; Acta Phytotax. Sin. 9: 99 (1964)
= Neolepisorus ovatus f. gracilis Ching & K.H.Shing; Acta Phytotax. Sin. 21: 269 (1983)
= Neolepisorus ovatus f. monstrosus Ching & K.H.Shing; Acta Phytotax. Sin. 21(3): 269 (1983)
= Neolepisorus ovatus f. truncatus (Ching & P.S.Wang) L.Shi & X.C.Zhang; Fl. China 2-3: 807 (2013)
= Neolepisorus phyllomanes (Christ) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 14 (1940)
= Neolepisorus phyllomanes var. doryopteris (Christ) Ching
= Neolepisorus sinensis Ching; Acta Phytotax. Sin. 21(3): 274 (1983)
= Neolepisorus truncatus Ching & P.S.Wang; Acta Phytotax. Sin. 21(3): 270 (1983)
= Neolepisorus truncatus f. laciniatus Ching & K.H.Shing; Acta Phytotax. Sin. 21(3): 270 (1983)
= Neolepisorus tsaii Ching & K.H.Shing; Acta Phytotax. Sin. 21(3): 273 (1983)
= Phymatodes ovata C.Presl; Tent. Pterid. 197 (1836)
= Pleopeltis ovata (C.Presl) T.Moore; Index Fil. [T. Moore] lxxviii (1857)
= Pleopeltis ovata (Wall. ex Hook. & Grev.) Bedd.; Ferns Brit. India 354 (1883), based on Polypodium ovatum Wall. ex Hook. & Grev., Icon. Filic. 1: t. 41: 1827, not Thunb. (1768)
= Polypodium hemitomum var. deltoideum (Baker) Hand.-Mazz.; Symb. Sin. 6: 44 (1929), based on Polypodium deltoideum Baker, J. Bot. 26: 230 (1880), not Sw. (1788), nor Liebmann (1849)
= Polypodium ovatum Wall. ex Hook. & Grev.; Icon. Filic. t. 41 (1827) (non Burm.f. (1768))
= Polypodium phyllomanes Christ; Bull. Acad. Int. Géogr. Bot. 1902: 210 c. figg. (1902)
= Polypodium phyllomanes var. doryopteris Christ; Bull. Acad. Int. Géogr. Bot. 11(153-154): 214 (1902)
= Polypodium phyllomanes var. ovatum (Wall. ex Hook. & Grev.) Christ; Bull. Acad. Int. Géogr., sér. 3, 11: 210 (1902), nom. inval.
species :
Lepisorus palmatopedatus (Baker) C.F.Zhao, R.Wei & X.C.Zhang; Cladistics 36(3): 253 (2019)
Distribution: China (Guizhou, Sichuan, Yunnan)
36 CHI
Remarks: sect. 6 Neocheiropteris
Conservation Status: IUCN V
Synonyms: hidden
= Cheiropteris henryi Christ; Bull. Herb. Boissier, sér. 1, 6: 876 (1898), non Polypodium henryi (Baker) Christ 1898
= Cheiropteris palmatopedata (Baker) Christ; Bull. Herb. Boissier, sér. 1, 7: 21, t. 1 (1899)
= Microsorum palmatopedatum (Baker) Noot.; Blumea 42(2): 343 (1997)
= Neocheiropteris palmatopedata (Baker) Christ; Mém. Soc. Bot. France 1 [Hist. Acrostich.]: 21 (1905)
= Polypodium palmatopedatum Baker; Bull. Misc. Inform. Kew 1898: 232 (1898)
species :
Lepisorus perrierianus (C.Chr.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 51 (1933)
Distribution: Madagascar
29 MDG
Remarks: sect. 8 Pachyphyllon
Synonyms: hidden
= Pleopeltis perrieriana (C.Chr.) Tardieu; Humbert, Fl. Madagasc. Fam. 5, 2: 112 (1960)
= Polypodium perrierianum C.Chr.; Notes Ptéridol. [Bonaparte] 16: 193 (1925)
species :
Lepisorus platyrhynchos (Kunze) Li Wang; Bot. J. Linn. Soc. 162(1): 35 (2010)
Distribution: Philippines (Luzon, Mindoro, Agusan del Norte, Bukidnon, Davao); New Guinea (Irian Jaya, Yapen Isl.)
42 NWG PHI
Remarks: sect. 13 Belvisia
Synonyms: hidden
= Acrostichum platyrhynchos (Kunze) Hook.; Sp. Fil. [W. J. Hooker] 6: 280 (1864)
= Belvisia glauca (Copel.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 192 (1947)
= Belvisia platyrhynchos (Kunze) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 192 (1947)
= Gymnopteris platyrhynchos J.Sm.; J. Bot. (Hooker) 3: 403 (1841), nomen
= Hymenolepis glauca (Copel.) C.Chr.; Dansk Bot. Ark. 6(3): 54-70 (1929)
= Hymenolepis platyrhynchos Kunze; Farnkräuter 1: 101 (1842)
= Hymenolepis platyrhynchos var. glauca Copel.; Leafl. Philipp. Bot. 3: 847 (1910)
= Macroplethus glaucus (Copel.) Tagawa; Acta Phytotax. Geobot. 11: 234 (1942)
= Macroplethus platyrhynchos (Kunze) C.Presl; Abh. Königl. Böhm. Ges. Wiss., ser. 5, 6 [Epim. Bot.]: 142 (1851)
= Taenitis macrorhynchos Mett.; Fil. Hort. Bot. Lips. 28 (1856)
= Taenitis platyrhyncha (Kunze) Mett.; Ann. Mus. Bot. Lugduno-Batavi 4: 173 (1869)
species :
Lepisorus pseudonudus (Ching) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 83 (1933)
Distribution: China (Yunnan, Sichuan, Gansu); Tibet
36 CHI TIB
Remarks: sect. 16 Sclerophyllon
Synonyms: hidden
= Lepisorus bilouensis Ching & Y.X.Lin ex K.H.Shing; Vasc. Pl. Hengduan Mount. 31(6): 573 (1993), without latin descr. or type, et Acta Phytotax. Sin. 1: 166 (1993)
= Pleopeltis pseudonuda (Ching) Panigrahi & Patnaik; J. Indian Bot. Soc. 43: 317 (1964)
species :
Lepisorus pseudoussuriensis Tagawa; Acta Phytotax. Geobot. 5: 110 (1936)
Distribution: Taiwan
38 TAI
Remarks: sect. 12 Ussurienses
Synonyms: hidden
= Lepisorus angustifrons Tagawa; Acta Phytotax. Geobot. 6: 110 (1936)
species :
Lepisorus pyriformis (Ching) C.F.Zhao, R.Wei & X.C.Zhang; Cladistics 36(3): 252 (2019)
Distribution: China (C-China, S-China); Japan (Honshu, Yakushima Isl.)
36 CHI 38 JAP NNS
Remarks: sect. 4 Lemmaphyllum. Often included in L. rostratus but kept separate by Zhao et al. 2019
Synonyms: hidden
= Lemmaphyllum pyriforme (Ching) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 99 (1933)
= Lepidogrammitis pyriformis (Ching) Ching; Sunyatsenia 5: 258 (1940)
= Polypodium pyriforme Ching; Bull. Fan Mem. Inst. Biol. Bot. 2: 212, t. 29 (1931)
species :
Lepisorus rostratus (Bedd.) C.F.Zhao, R.Wei & X.C.Zhang; Cladistics 36(3): 252 (2019)
Distribution: China (Gansu, Guangdong, Guangxi, Guizhou, Hainan, Hubei, Sichuan, Yunnan, Zhejiang); S-Japan (Yakushima Isl.); Taiwan; Nepal; Bhutan; NE-India (Arunachal Pradesh, Sikkim, Darjeeling, NE-India); Myanmar (Kachin, etc.); Thailand; Laos; Cambodia; Vietnam; N-Sumatra
36 CHI 38 JAP NNS TAI 40 BHU IND NEP 41 CBD LAO MYA THA VIE 42 SUM
Remarks: sect. 4 Lemmaphyllum
Synonyms: hidden
= Lemmaphyllum carnosum var. rostratum (Bedd.) Hovenkamp; Fl. Males. Ser. 2, Pterid. 3: 76 (1998)
= Lemmaphyllum rostratum (Bedd.) Tagawa; Fl. E. Himal.: 493 (1966)
= Lemmaphyllum subrostratum (C.Chr.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 97 (1933)
= Lepidogrammitis rostrata (Bedd.) Ching; Acta Phytotax. Sin. 9: 372 (1964)
= Lepidogrammitis sikkimensis S.R.Ghosh; Pterid. Fl. E. India 508 (-510, fig. 154) (2004)
= Lepidogrammitis subrostrata (C.Chr.) Ching; Sunyatsenia 6: 258 (1940)
= Lepisorus rostratus (Hook.) B.K.Nayar & Kaur; Companion Ferns Brit. India 84 (1974)
= Lepisorus subrostratus (C.Chr.) C.Chr. & Tardieu; Notul. Syst. (Paris) 8: 187 (1939)
= Phymatopsis rostrata (Bedd.) J.Sm.; Hist. Fil. 104 (1875)
= Pleopeltis rostrata Bedd.; Ferns Brit. India t. 159 (1866)
= Pleopeltis subrostrata Lacaita; J. Linn. Soc., Bot. 48: 490 (1916)
= Polypodium rostratum Hook.; Hooker´s Icon. Pl. 6: t. 953 (1854), nom. illeg.
= Polypodium subrostratum C.Chr.; Index Filic. 567 (1906)
species :
Lepisorus rufofuscus T.Fujiw.; Acta Phytotax. Geobot. 73(3): 178 (2022)
Distribution: Japan (Honshu)
38 JAP
species :
Lepisorus schraderi (Mett.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 51 (1933)
Distribution: Rwanda; Ethiopia; Uganda; Kenya; Tanzania; Zambia; Malawi; E-Zimbabwe; Mozambique; South Africa (Limpopo, Gauteng, Mpulamanga, Free State, KwaZulu-Natal, W-Cape Prov., E-Cape Prov.); Lesotho; Eswatini [Swaziland]; Madagascar; Mauritius; La Réunion
23 RWA 24 ETH 25 KEN TAN UGA 26 MLW MOZ ZAM ZIM 27 LES SAF SWZ 29 MAU MDG REU
Remarks: sect. 8 Pachyphyllon. In South Africa L. schraderi and L. excavatus are well separated, but in East Africa the differences disappear partially.
Synonyms: hidden
= Lepisorus gueinzii (Mett.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 51 (1933), as ´gueintzii´
= Niphobolus schraderi (Mett.) Keyserl.; Polyp. Herb. Bunge 39 (1873)
= Phymatodes elongata (Schrad.) Pappe & Rawson; Syn. Filic. Afr. Austr. 41 (1858)
= Pleopeltis schraderi (Mett.) Tardieu; Humbert, Fl. Madagasc. Fam. 5, 2: 110 (1960)
= Polypodium elongatum Schrad.; Gött. Gel. Anz. 918 (1818) (non Mett. (1857), nec Aiton (1789))
= Polypodium gueintzii Mett.; Abh. Senckenberg. Naturf. Ges. 2(1) [Polypodium]: 91 (1856)
= Polypodium lineare var. gueinzii (Mett.) C.Chr.; Index Filic. 540 (1906), as ´gueintzii´
= Polypodium lineare var. schraderi (Mett.) Sim; Ferns S. Afr. [Sim], ed. 2: 276, t. 140 (1915)
= Polypodium schraderi Mett.; Abh. Senckenberg. Naturf. Ges. 2(2) [Pheg. u. Asp.]: 98, no. 179, t. 2, f. 11 (1857)
species :
Lepisorus scolopendrium (Buch.-Ham. ex Ching) Mehra & Bir; Res. Bull. Panjab Univ. Sci. 15: 168 (1964)
Distribution: China (Guizhou, Hainan, Sichuan, Yunnan); Tibet; N-Pakistan; Nepal; Bhutan; India (Himachal Pradesh, Uttarakhand, Sikkim, Darjeeling Hills, Jammu & Kashmir, Arunachal Pradesh); Myanmar (Chin, etc.); Thailand; Laos
36 CHI TIB 40 BHU IND NEP PAK 41 LAO MYA THA
Remarks: sect. 8 Pachyphyllon
Synonyms: hidden
= Lepisorus excavatus f. himalayensis (Bir & Trikha) Benl; Acta Bot. Barcinon. 33: 18 (1982)
= Lepisorus excavatus var. himalayensis Bir & Trikha; Amer. Fern J. 64: 58, f. 33, 37 (1974)
= Lepisorus excavatus var. mortonianus Bir & Trikha; Amer. Fern J. 64: 56, f. 25, 32 (1974)
= Lepisorus excavatus var. scolopendrium Buch.-Ham. ex Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 69 (1933) (validation of originally invalid epithet scolopendrium)
= Lepisorus himalayensis (Bir & Trikha) Khullar & Pangtey; Khullar et al., Ferns of Nainital: 165 (1991) (fide Fraser-Jenkins 2008)
= Lepisorus mortonianus (Bir & Trikha) Khullar & Pangtey; Khullar et al., Ferns of Nainital: 169 (1991) (fide Fraser-Jenkins 2008)
= Lepisorus scolopendrium (Buch.-Ham. ex D.Don) Tagawa; H. Hara, Fl. E. Himal. 494 (1966), nom. superfl.
= Lepisorus scolopendrium var. himalayensis (Bir & Trikha) Pangtey & Punetha; Illustr. Fern Fl. W. Himalaya 2: 399 (1987)
= Lepisorus virescens Ching & S.K.Wu; Fl. Xizangica 1: 306 (1983)
= Lepisorus ×kumaonensis Kholia, Bhakuni, R.Punetha & Bankotia; NeBIO 3(3): 31 (2012)
= Pleopeltis excavata var. himalayensis (Bir & Trikha) Benl; Acta Bot. Barcinon. 33: 18 (1982)
= Pleopeltis excavata var. mortoniana (Bir & Trikha) Benl; Acta Bot. Barcinon. 33: 18 (1982)
= Pleopeltis himalayensis (Bir & Trikha) Á.Löve & D.Löve; Taxon 26: 324 (1977), nom. illeg.
= Pleopeltis mortonianus (Bir & Trikha) Á.Löve & D.Löve; Taxon 26: 324 (1977)
= Pleopeltis scolopendrium (Buch.-Ham. ex Ching) Alston & Bonner; Candollea 15: 207 (1956)
= Polypodium scolopendrium Buch.-Ham. ex D.Don; Prodr. Fl. Nepal. 1 (1825), non P. scolopendria Burm.f. (1768)
species :
Lepisorus sinensis (Christ) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 63 (1933)
Distribution: China (SE-Yunnan); ?Bhutan; Myanmar; N-Thailand; Vietnam
36 CHI 41 MYA THA VIE
Remarks: sect. 17 Pseudovittaria
Synonyms: hidden
= Lemmaphyllum sinense (Christ) C.Chr.; Dansk Bot. Ark. 6(3): 51 (1929)
= Lepisorus vittarioides Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 64 (1933)
= Neurodium sinense Christ; Bull. Herb. Boissier, sér. 1, 6: 880 (1898)
= Paltonium sinense (Christ) C.Chr.; Index Filic. 477 (1906)
= Pleopeltis sinensis (Christ) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 184 (1947)
= Polypodium neurodioides C.Chr.; Contr. U. S. Natl. Herb. 26: 318 (1931)
= Polypodium neurodioides var. vittarioides (Ching) C.Chr.; Index Filic., Suppl. Tert.: 118, 154 (1934)
species :
Lepisorus sordidus (C.Chr.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 78 (1933)
Distribution: China (Yunnan, Sichuan); Tibet; India (Arunachal Pradesh, Nagaland, Manipur); Myanmar
36 CHI TIB 40 IND 41 MYA
Remarks: sect. 17 Pseudovittaria
Conservation Status: IUCN VU for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Lepisorus sordidus f. rostratus Ching; Bull. Fan Mem. Inst. Biol. Bot. 4(3): 79 (1933)
= Pleopeltis sordida (C.Chr.) Panigrahi & Patnaik; Curr. Sci. 34: 127 (1965)
= Polypodium sordidum C.Chr.; Contr. U. S. Natl. Herb. 26: 320 (1931)
species :
Lepisorus soulieanus (Christ) Ching & S.K.Wu; Acta Bot. Yunnan. 5(1): 11 (1983), et Fl. Xizangica 1: 307 (1983)
Distribution: China; etc.
36 CHI
Remarks: sect. 9 Hymenophyton. Separate from L. clathratus fide Zhao et al. 2019. Distribution uncertain because this was for a long time included in L. clathratus.
Synonyms: hidden
= Polypodium soulieanum Christ; Mém. Soc. Bot. France 1 [Hist. Acrostich.]: 15 (1905)
= Platygyria soulieana (Christ) X.C.Zhang & Q.R.Liu; Acta Phytotax. Sin. 41(5): 407 (2003)
species :
Lepisorus spicatus (L.f.) Li Wang; Bot. J. Linn. Soc. 162(1): 35 (2010)
Distribution: Guinea; Liberia; Ivory Coast; Cameroon; Bioko Isl. [Fernando Poo]; São Tomé; Gabon; Congo [Brazzaville]; D.R.Congo [Zaire]; Tanzania; Malawi; Zimbabwe; Mozambique; South Africa; Madagascar; Comores; La Réunion; Seychelles; Sri Lanka; Myanmar (Kachin, etc.); SW-Thailand; Laos; Vietnam; peninsular Malaysia (Kedah, Langkawi Isl., Perak, Negeri Sembilan, Kelantan, Pahang); Sumatra; Borneo (Mt. Kinabalu, Sabah, Kalimantan); Sulawesi; Palawan; Philippines; Java; Lesser Sunda Isl. (Bali, Lombok, etc.); Moluccas; New Guinea (Irian Jaya, Papua New Guinea); Solomon Isl. (Guadalcanal); New Caledonia; Vanuatu; Fiji; Society Isl. (Tahiti, Moorea); Austral Isl. (Tubuai)
22 GUI IVO LBR 23 BIO CMN CON GAB SAO ZAI 25 TAN 26 MLW MOZ ZIM 27 SAF 29 COM MDG REU SEY 40 SRL 41 LAO MYA THA VIE 42 BOR JAW LSI MLY MOL NWG PHI PLW SUL SUM 43 SOL 60 FIJ NWC VAN 61 AUI SCI
Remarks: sect. 13 Belvisia
Synonyms: hidden
= Acrostichum spicatum L.f.; Suppl. Pl. 444 (1782)
= Belvisia revoluta (Blume) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 192 (1947)
= Belvisia spicata (L.f.) Mirb.; Hist. Nat. Veg. [Lamarck & Mirbel] 3: 473 (1802), et ex Copel., Gen. Fil. 192 (1947)
= Gymnopteris ophioglossoides (Kaulf.) Paxton; Pocket Bot. Dict., ed. 2, Suppl.: 33 (1849)
= Gymnopteris spicata (L.f.) C.Presl; Tent. Pterid. 244, t. 11, f. 7 (1836)
= Hyalolepis ophioglossoides (Kaulf.) Kunze; Linnaea 28: 258 (1850)
= Hyalolepis revoluta (Blume) Kunze; Linnaea 23: 258 (1850)
= Hyalolepis revoluta var. planiuscula Kunze; (1850)
= Hymenolepis ophioglossoides Kaulf.; Enum. Filic. [Kaulfuss] 146, t. 1, f. 9 (1824)
= Hymenolepis revoluta Blume; Enum. Pl. Javae 2: 201 (1828)
= Hymenolepis revoluta var. planiuscula (Mett.) Hieron. ex C.Chr.; Hedwigia 57: 58 (1916)
= Hymenolepis rigidissima Christ; Bull. Herb. Boissier, sér. 2, 6: 990 (1906)
= Hymenolepis spicata (L.f.) C.Presl; Abh. Königl. Böhm. Ges. Wiss., ser. 5, 6 [Epim. Bot.]: 159 (1851)
= Hymenolepis spicata f. minima Alderw.; (1917)
= Hymenolepis spicata var. costulata Alderw.; (1912)
= Hymenolepis spicata var. occultivenia C.Chr.; Dansk Bot. Ark. 6(3): 57 (1929)
= Hymenolepis spicata var. tenella Alderw.; (1908)
= Hymenolepis spicata var. usambarensis C.Chr.; Dansk Bot. Ark. 6(3): 57 (1929)
= Lomaria spicata (L.f.) Willd.; Sp. Pl., ed. 4 [Willdenow] 5(1): 289 (1810)
= Macroplethus revolutus (Blume) Tagawa; Acta Phytotax. Geobot. 11: 234 (1942)
= Macroplethus spicatus (L.f.) Tagawa; Acta Phytotax. Geobot. 11: 235 (1942)
= Oetosis ophioglossoides (Kaulf.) Kuntze; Revis. Gen. Pl. 2: 817 (1891)
= Onoclea spicata (L.f.) Sw.; J. Bot. (Schrader) 1801(2): 299 (1803)
= Schizaea spicata (L.f.) Sm.; Mém. Acad. Roy. Sci. (Turin) 5: 43 (1793)
= Taenitis ophioglossoides (Kaulf.) Mett.; Fil. Hort. Bot. Lips. 28 (1856)
= Taenitis revoluta (Blume) Mett.; Fil. Hort. Bot. Lips. 28 (1856)
= Taenitis revoluta var. planiuscula Mett.; Fil. Hort. Bot. Lips. 28 (1856)
= Taenitis spicata (L.f.) Mett.; Ann. Mus. Bot. Lugduno-Batavi 4: 173 (1869)
species :
Lepisorus squamatus (A.R.Sm. & X.C.Zhang) C.F.Zhao, R.Wei & X.C.Zhang; Cladistics 36(3): 252 (2019)
Distribution: China (Guangxi); N-Vietnam
36 CHI 41 VIE
Remarks: sect. 4 Lemmaphyllum
Synonyms: hidden
= Caobangia squamata A.R.Sm. & X.C.Zhang; Novon 12(4): 549 (2002)
= Lemmaphyllum squamatum (A.R.Sm. & X.C.Zhang) Li Wang; Bot. J. Linn. Soc. 162(1): 36 (2010)
species :
Lepisorus subhemionitideus (Christ) C.F.Zhao, R.Wei & X.C.Zhang; Cladistics 36(3): 253 (2019)
Distribution: China (Guangxi, Hunan, Sichuan, Yunnan); Tibet; Nepal; Bhutan; N-India; Myanmar; N-Vietnam; S-Vietnam
36 CHI TIB 40 BHU IND NEP 41 MYA VIE
Remarks: sect. 7 Lepidomicrosorium. Not in Taiwan
Synonyms: hidden
= Lepidomicrosorium caudifrons Ching & W.M.Chu; Bot. Res. Acad. Sin. 1983(1): 10 (1983)
= Lepidomicrosorium crenatum Ching & K.H.Shing; Bot. Res. Acad. Sin. 1983(1): 11 (1983)
= Lepidomicrosorium emeiense Ching & K.H.Shing; Bot. Res. Acad. Sin. 1983(1): 10 (1983)
= Lepidomicrosorium hongchunpingense Ching & K.H.Shing; Bot. Res. Acad. Sin. 1983(1): 9 (1983)
= Lepidomicrosorium hongchunpingense var. laceratum Ching & K.H.Shing; Bot. Res. Acad. Sin. 1983(1): 9 (1983)
= Lepidomicrosorium hunanense Ching & K.H.Shing; Bot. Res. Acad. Sin. 1983(1): 6 (1983)
= Lepidomicrosorium laojunense Ching & K.H.Shing; Bot. Res. Acad. Sin. 1983(1): 10 (1983)
= Lepidomicrosorium lineare Ching & P.S.Chiu; Bot. Res. Acad. Sin. 1983(1): 5 (1983)
= Lepidomicrosorium longshengense Ching & K.H.Shing; Bot. Res. Acad. Sin. 1983(1): 9 (1983)
= Lepidomicrosorium subhemionitideum (Christ) P.S.Wang; P. S. Wang & X. Y. Wang Pterid. Fl. Guizhou 382 (2001)
= Lepidomicrosorium undulatum Ching & P.S.Chiu; Bot. Res. Acad. Sin. 1983(1): 11 (1983)
= Leptochilus subhemionitideus (Christ) Bosman; Leiden Bot. Ser. 14: 117 (1991)
= Microsorum hymenodes var. marginale (Takeda) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4(10): 301 (1933)
= Microsorum jinfoshanense Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 3(4): 12 (1983)
= Microsorum rubripes Ching & W.M.Chu; Bull. Bot. Res., Harbin 3(4): 11 (1983)
= Neocheiropteris sapaensis V.N.Tu; Bot. Zhurn. (Moscow & Leningrad) 65(4): 584 (1980)
= Neocheiropteris subhemionitidea (Christ) Fraser-Jenk.; New Species Syndr. Indian Pteridol. 179 (1997)
= Polypodium hymenodes var. marginale Takeda; Notes Roy. Bot. Gard. Edinburgh 8(39): 288 (1915)
= Polypodium subhemionitideum Christ; Bull. Herb. Boissier, sér. 1, 7: 5 (1809)
= Polypodium superficiale var. attenuatum Rosenst.; Repert. Spec. Nov. Regni Veg. 13(355-358): 134 (1914)
= Polypodium superficiale var. semilineare C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 558 (1880)
= Tricholepidium subhemionitideum (Christ) Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 69 (2008)
species :
Lepisorus sublinearis (Baker ex Takeda) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 78 (1933)
Distribution: China (Yunnan); Tibet; Nepal; Bhutan; N-India (Arunachal Pradesh, Sikkim, Darjeeling, Uttarakhand, NE-India); Myanmar (Kachin, Chin, etc.); N-Thailand; Laos; Vietnam
36 CHI TIB 40 BHU IND NEP 41 LAO MYA THA VIE
Remarks: sect. 16 Sclerophyllon
Synonyms: hidden
= Lepisorus cespitosus Y.X.Lin; Fl. Reipubl. Popularis Sin. [Fl. China Ed. Committee] 6(2): 77 (2000)
= Lepisorus lancifolius Ching; Bull. Fan Mem. Inst. Biol. Bot. 11: 75 (1941)
= Pleopeltis sublinearis (Baker ex Takeda) Tagawa & K.Iwats.; Acta Phytotax. Geobot. 22: 100 (1967)
= Polypodium sublineare Baker ex Takeda; Notes Roy. Bot. Gard. Edinburgh 8: 276 (1915)
species :
Lepisorus suboligolepidus Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 77 (1933)
Distribution: China (Hubei, Sichuan, Yunnan); Taiwan; NE-India (Meghalaya, Manipur, Tripura, Arunachal Pradesh); Myanmar; NE-Thailand
36 CHI 38 TAI 40 IND 41 MYA THA
Remarks: sect. 17 Pseudovittaria. Sometimes included in L. oligolepidus
Synonyms: hidden
= Polypodium suboligolepidum (Ching) C.Chr.; Index Filic., Suppl. Tert.: 159 (1934)
= Pleopeltis suboligolepida (Ching) Tagawa & K.Iwats.; Acta Phytotax. Geobot. 26: 172 (1975)
species :
Lepisorus subsessilis Ching & Y.X.Lin; Fl. Reipubl. Popularis Sin. [Fl. China Ed. Committee] 6(2): 77 (2000)
Distribution: China (Guangdong, Guangxi)
36 CHI
Remarks: sect. 10 Lepisorus
species :
Lepisorus superficialis (Blume) C.F.Zhao, R.Wei & X.C.Zhang; Cladistics 36(3): 253 (2019)
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hubei, Hunan, Jiangxi, Sichuan, Yunnan, Zhejiang); Tibet; South Korea (incl. Cheju Isl.); Japan; Bonin Isl. (Hahajima); Taiwan; Bhutan; NE-India; Myanmar (Kachin, etc.); Thailand; ?Laos; Vietnam; peninsular Malaysia (Pahang); Sumatra; Sulawesi; Java; Moluccas
36 CHI TIB 38 JAP KOR OGA TAI 40 BHU IND 41 MYA THA VIE 42 JAW MLY MOL SUL SUM
Remarks: sect. 7 Lepidomicrosorium. L. ningpoense is probably a separate species fide Kuo et al. (2019)
Synonyms: hidden
= Colysis superficialis (Blume) J.Sm.; Hist. Fil. 101 (1875)
= Lepidomicrosorium angustifolium Ching & K.H.Shing; Bot. Res. Acad. Sin. 1983(1): 8 (1983)
= Lepidomicrosorium nanchuanense Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(4): 27 (1984)
= Lepidomicrosorium ningpoense (Baker) L.Y.Kuo; Taiwania 64(4): 378 (2019)
= Lepidomicrosorium sichuanense Ching & K.H.Shing; Bot. Res. Acad. Sin. 1983(1): 8 (1983)
= Lepidomicrosorium superficiale (Blume) Li Wang; Bot. J. Linn. Soc. 162(1): 36 (2010)
= Microsorum brachylepis (Baker) Nakaike; Ill. Pterid. Japan 2: 492 (1981)
= Microsorum buergerianum f. laciniatum Ching; Bull. Fan Mem. Inst. Biol. Bot. 4(10): 303 (1933)
= Microsorum masaskei (Nakai) H.Itô; J. Jap. Bot. 11: 96 (1935)
= Microsorum ovalifolium Ching & S.K.Wu; Fl. Xizangica 1: 327 (1983) (fide Fraser-Jenkins 2008)
= Microsorum simulans Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(4): 26 (1984)
= Microsorum superficiale (Blume) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 295-311 (1933)
= Microsorum takhtajanii V.N.Tu; Bot. Zhurn. (Moscow & Leningrad) 64(12): 1769 (1979)
= Microsorum tibeticum (Ching & S.K.Wu) Ching & S.K.Wu; Fl. Xizangica 1: 328 (1983) (fide Fraser-Jenkins 2008)
= Neocheiropteris ningpoensis (Baker) Bosman; Leiden Bot. Ser. 14: 119 (1991)
= Neocheiropteris superficialis (Blume) Bosman; Leiden Bot. Ser. 14: 121 (1991)
= Phymatodes masaskei Nakai; Bot. Mag. (Tokyo) 43: 4 (1929)
= Pleopeltis superficialis (Blume) Bedd.; Ferns Brit. India t. 75 (1865)
= Pleopeltis superficialis var. latifrons Bedd.; J. Bot. 31: 226 (1893)
= Polypodium brachylepis Baker; Gard. Chron., ser. 2, 14: 494 (1880)
= Polypodium buergerianum var. ningpoense (Baker) Takeda; Notes Roy. Bot. Gard. Edinburgh 8(39): 291 (1915)
= Polypodium masaskei (Nakai) Ogata; Icon. Fil. Jap. 8, t. 138 (1930)
= Polypodium nigrocinctum Christ; Bull. Herb. Boissier, sér. 1, 6: 874 (1896)
= Polypodium ningpoense Baker; Ann. Bot. (Oxford) 5: 474 (1891)
= Polypodium superficiale Blume; Enum. Pl. Javae 2: 123 (1828)
= Polypodium superficiale var. anguinum Christ; Mém. Soc. Bot. France 1 [Hist. Acrostich.]: 16 (1905)
= Tricholepidium superficiale (Blume) Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 69 (2008)
= Tricholepidium superficiale (Blume) Fraser-Jenk., Kandel & Pariyar; Ferns and Fern-allies of Nepal 1: 43 (2015), isonym (isonym)
species :
Lepisorus thunbergianus (Kaulf.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 88 (1933), nom. cons. prop.
Distribution: Russian Far East (Kuril Isl.); China (Anhui, Fujian, Gansu, Guizhou, Hebei, Hubei, Hunan, Jiangsu, Jiangxi, Macao, Shaanxi, Sichuan, Yunnan, Zhejiang); Tibet; South Korea (Cheju Isl.); Japan; Ryukyu Isl.; Taiwan; N-Pakistan; Nepal; Bhutan; N-India (Arunachal Pradesh, Sikkim, Uttarakhand, Himachal Pradesh, Jammu & Kashmir); Myanmar; Laos; Vietnam; Philippines (N-Luzon); Hawaii (Kauai, Oahu, Molokai, Lanai, Maui, Hawaii Isl.)
31 RFE 36 CHI TIB 38 JAP KOR NNS TAI 40 BHU IND NEP PAK 41 LAO MYA VIE 42 PHI 63 HAW
Remarks: sect. 17 Pseudovittaria
Synonyms: hidden
= Drynaria elongata (Kaulf.) Brack.; U.S. Expl. Exped., Filic. 16: 42 (1854)
= Drynaria subspathulata Hook.; Hooker´s J. Bot. Kew Gard Misc. 9: 356 (1857)
= Lepisorus atropunctatus (Keyserl.) Ching; (1933)
= Lepisorus calcifer Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(4): 25 (1984)
= Lepisorus chingii Khullar; (1988) (nom. nud., fide Fraser-Jenkins (1997))
= Lepisorus elongatus Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 89 (1933), nom. rej. prop.
= Lepisorus linearifolius Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 3(4): 5 (1983)
= Lepisorus linearis (Thunb.) Ching; (nom. illeg.)
= Lepisorus mussooriensis Khullar; (1988) (nom. nud., fide Fraser-Jenkins (1997))
= Lepisorus myriosorus Ching; Wuyi Sci. J. 1: 11 (1981)
= Lepisorus pygmaeus Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(4): 24 (1984)
= Lepisorus simulans Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(4): 23 (1984), et Phytotaxa 205(4): 299 (2015)
= Lepisorus stewartii Ching; Acta Bot. Austro Sin. 1: 23 (1983) (fide Fraser-Jenkins 1997)
= Lepisorus thunbergianus var. subspathulatus (Hook.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4(3): 89 (1933)
= Lepisorus thunbergianus var. tsutaei Akasawa & E.Yokote; Bull. Kochi Women´s Univ., Ser. Nat. Sci. 27: 4 (1979)
= Microsorum lineare (Thunb.) Copel.; Univ. Calif. Publ. Bot. 16: 111-116 (1929)
= Niphobolus atropunctatus Keyserl.; Polyp. Herb. Bunge 39 (1873)
= Niphobolus linearis (Thunb.) Keyserl.; Polyp. Herb. Bunge 39 (1873)
= Phymatodes elongata (Kaulf.) C.Presl; Tent. Pterid. 196 (1836)
= Pleopeltis elongata Kaulf.; Enum. Filic. [Kaulfuss] 246 (1824), nom. rej. prop.
= Pleopeltis linearis (Thunb.) T.Moore; Index Fil. [T. Moore] 346 (1862)
= Pleopeltis linearis Bedd.; Ferns Brit. India 346 (1883), nom. illeg.
= Pleopeltis subspathulata (Hook.) T.Moore; Index Fil. [T. Moore] 348 (1862)
= Pleopeltis thunbergiana Kaulf.; Wesen Farrenkr. 113 (1827), based on Polypodium lineare Thunb., Syst. Veg., ed. 14 (J. A. Murray): 934 (1784), not N. F. Burman (1768), nom. cons. prop.
= Pleopeltis wightiana Bedd.; Ferns S. India t. 180 (1863), nom. superfl.
= Pleopeltis zeylanica T.Moore; Index Fil. [T. Moore] 349 (1862)
= Polypodium coraiense Christ; Repert. Spec. Nov. Regni Veg. 6: 285 (1908)
= Polypodium elongatum Goldm.; Nov. Actorum Acad. Caes. Leop.-Carol. Nat. Cur. 19(Suppl. 1): 454 (1843), nom. illeg.
= Polypodium gladiatum Wall.; Numer. List. [Wallich] no. 279 (1829), nomen
= Polypodium lineare Thunb.; Fl. Jap. [Thunberg] 335 (1784) (non Burm.f. (1768) nec Houtt. (1783)), et Syst. Veg., ed. 14 (J. A. Murray): 934 (1784)
= Polypodium lineare var. coraiense Christ; Repert. Spec. Nov. Regni Veg. 5: 10 (1908) (tentative synonym)
= Polypodium lineare var. elongatum Takeda; Notes Roy. Bot. Gard. Edinburgh 8: 273 (1915) (tentative synonym)
= Polypodium lineare var. glaucosorum Christ; Bull. Acad. Int. Géogr. Bot. sér. 3, 11(153-154): 209 (1902) (tentative synonym)
= Polypodium lineare var. subspathulatum (Hook.) Takeda; Notes Roy. Bot. Gard. Edinburgh 8(39): 270-271 (1915)
= Polypodium lineare var. thunbergianum (Kaulf.) Takeda; Notes Roy. Bot. Gard. Edinburgh 8(39): 268 (1915)
= Polypodium loriforme Hook.; Gard. Ferns: t. 14 (1862), non Wall. ex Mett.
= Polypodium thunbergianum (Kaulf.) C.Chr.; Index Filic., Suppl. Tert.: 160 (1934)
species :
Lepisorus tibeticus Ching & S.K.Wu; Fl. Xizangica 1: 311 (1983)
Distribution: China (Sichuan, Yunnan); Tibet
36 CHI TIB
Remarks: sect. 17 Pseudovittaria. Included in L. thunbergianus by Fraser-Jenkins et al. 2015 but kept separate on genetic basis by Zhao et al. 2019
Synonyms: hidden
= Pleopeltis tibetica (Ching & S.K.Wu) Panigrahi; Pl. Sci. Res. (India), Abstr. 23rd Ann. Conf. Khurda: 36 (1998)
species :
Lepisorus tosaensis (Makino) H.Itô; J. Jap. Bot. 11: 93 (1935)
Distribution: China (Anhui, Chongqing, Guangdong, Guangxi, Guizhou, Hunan, Jiangsu, Jiangxi, Sichuan, Xinjiang, Yunnan, Zhejiang); Tibet; Japan; Taiwan; Vietnam
36 CHI TIB 38 JAP TAI 41 VIE
Remarks: sect. 17 Pseudovittaria
Synonyms: hidden
= Lepisorus infraplanicostalus (Hayata) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4(3): 92 (1933)
= Lepisorus paohuashanensis Ching; Rec. Fl. Jiangsu (Fl. Kiang Su Prov.) 1: 467 (1977)
= Lepisorus sinicus Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 3(4): 2 (1983)
= Lepisorus tosaensis f. bifidus Akasawa; Bull. Kochi Women´s Univ., Ser. Nat. Sci. 27: 4 (-5) (1979)
= Pleopeltis tosaensis (Makino) Ohwi; Bull. Natl. Sci. Mus., Tokyo, 3: 99 (1956)
= Polypodium hoozanense Hayata; Icon. Pl. Formosan. 8: 152, f. 81-82 (1919)
= Polypodium infra-planicostale Hayata; Icon. Pl. Formosan. 6: 31 5, f. 128 (1915)
= Polypodium lineare var. caudatum Makino; Bot. Mag. (Tokyo) 17: 78 (1903)
= Polypodium morii Hayata; Icon. Pl. Formosan. 8: 153, f. 83-84 (1919)
= Polypodium tosaense Makino; Bot. Mag. (Tokyo) 27: 127 (1913)
species :
Lepisorus tricholepis K.H.Shing & Y.X.Lin; Fl. Reipubl. Popularis Sin. [Fl. China Ed. Committee] 6(2): 70 (2000)
Distribution: Tibet; Bhutan
36 TIB 40 BHU
Remarks: sect. 9 Hymenophyton
species :
Lepisorus triglossus (Baker) C.F.Zhao, R.Wei & X.C.Zhang; Cladistics 36(3): 253 (2019)
Distribution: China (Yunnan)
36 CHI
Remarks: sect. 3 Neolepisorus. Possibly a hybrid L. palmatopedatus x macrosphaerus
Conservation Status: very rare and possibly extinct
Synonyms: hidden
= Neocheiropteris triglossa (Baker) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 108-112 (1933)
= Neolepisorus triglossus (Baker) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 14 (1940)
= Polypodium triglossum Baker; Bull. Misc. Inform. Kew 1898: 232 (1898)
= Selliguea triphylla Christ; Bull. Herb. Boissier, sér. 1, 6: 878 (1898)
species :
Lepisorus uchiyamae (Makino) H.Itô; J. Jap. Bot. 11: 93 (1935)
Distribution: Japan (Honshu, Shikoku, Kyushu); Ryukyu Isl.
38 JAP NNS
Remarks: sect. 12 Ussurienses
Synonyms: hidden
= Pleopeltis uchiyamae (Makino) Ohwi; Bull. Natl. Sci. Mus., Tokyo, 3: 99 (1956)
= Polypodium uchiyamae Makino; Bot. Mag. (Tokyo) 20: 30 (1906)
species :
Lepisorus ussuriensis (Regel & Mack.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 91 (1933)
Distribution: Russian Far East (Kuril Isl., Primorye, Sakhalin); China (Anhui, Hebei, Heilongjiang, Henan, Jilin, Liaoning, Shandong); North Korea; South Korea (incl. Cheju Isl.)
31 RFE 36 CHI 38 KOR NKO
Remarks: sect. 12 Ussurienses. Not in India
Synonyms: hidden
= Pleopeltis ussuriensis Regel & Maack; Mém. Acad. Imp. Sci. St. Pétersb. 7: 44-175 (1861)
= Polypodium lineare var. ussuriense (Regel & Maack) C.Chr.; Index Filic. 9: 540 (1906)
= Polypodium lineare var. ussuriense (Regel & Maack) Christ; Index Filic. 540 (1906)
= Polypodium schraderi Milde; Fil. Eur. 16 (1867), nom. illeg.
= Polypodium ussuriense (Regel & Maack) Regel; Trudy Imp. S.-Peterburgsk. Bot. Sada 7: 663 (1881)
species :
Lepisorus validinervis (Kunze) Li Wang; Bot. J. Linn. Soc. 162(1): 36 (2010)
Distribution: peninsular Malaysia (Pahang); Sumatra; Borneo (Sabah); Sulawesi; Philippines; Java; Lesser Sunda Isl. (Lombok); Moluccas (Seram); New Guinea (Irian Jaya, Papua New Guinea); Bismarck Arch. (New Ireland); Vanuatu
42 BOR JAW LSI MLY MOL NWG PHI SUL SUM 43 BIS 60 VAN
Remarks: sect. 13 Belvisia
Synonyms: hidden
= Acrostichum brachystachys (Hook.) Racib.; Pteridoph. Buitenzorg 51 (1898)
= Belvisia longissima Holttum; Blumea 14: 328 (1967)
= Belvisia squamata (Hieron. ex C.Chr.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 192 (1947)
= Belvisia squamata var. borneensis (C.Chr.) Parris; Plants Mt. Kinabalu [J. H. Beaman et al.] 98 (1992)
= Belvisia validinervis (Kunze) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 192 (1947)
= Belvisia validinervis var. longissima (Holttum) Hovenkamp & Franken; Blumea 37(2): 527 (1993)
= Hymenolepis brachystachys (Hook.) J.Sm.; Ferns Brit. Foreign 92 (1866)
= Hymenolepis brachystachys var. mirabilis Alderw.; (1918)
= Hymenolepis spicata var. brachystachys Hook.; Gard. Ferns, t. 3 (1862)
= Hymenolepis squamata Hieron.; C. Chr., Dansk Bot. Ark. 6(3): 54-70 (1929)
= Hymenolepis squamata var. borneensis (C.Chr.) Parris; Parris et al., Plants Mt. Kinabalu, 1. Ferns & Fern Allies: 98 (1992)
= Hymenolepis validinervis Kunze; Bot. Zeitung (Berlin) 6: 132 (1848)
= Macroplethus squamatus (Hieron. ex C.Chr.) Tagawa; Acta Phytotax. Geobot. 11: 235 (1942)
= Macroplethus validinervis (Kunze) Tagawa; Acta Phytotax. Geobot. 11: 235 (1942)
= Taenitis validinervis (Kunze) Mett.; Ann. Mus. Bot. Lugduno-Batavi 4: 173, t. 7 (1869)
species :
Lepisorus waltonii (Ching) S.L.Yu; Acta Phytotax. Sin. 35: 343 (1997)
Distribution: Tibet
36 TIB
Remarks: sect. 9 Hymenophyton. Separate from L. clathratus fide Zhao et al. 2019
Synonyms: hidden
= Lepisorus clathratus f. waltonii (Ching) Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 40 (2008)
= Neocheiropteris waltonii Ching; Hooker´s Icon. Pl. t. 3158 (1932)
= Phymatopteris waltonii (Ching) Fraser-Jenk.; New Species Syndr. Indian Pteridol. 196 (1997)
= Platygyria waltonii (Ching) Ching & S.K.Wu; Acta Bot. Yunnan. 2(1): 71 (1980)
= Platygyria ×inaequibasis Ching & S.K.Wu; Acta Bot. Yunnan. 2(1): 72 (1980)
= Polypodium clathratum var. lobatum Takeda; Notes Roy. Bot. Gard. Edinburgh 8(39): 282 (1915)
= xEllipisorus inaequibasis (Ching & S.K.Wu) Li Bing Zhang & Liang Zhang; Phytotaxa 464(4): 299 (2020)
= xLepinema inaequibasis (Ching & S.K.Wu) Li Bing Zhang & Liang Zhang; Phytotaxa 455(4): 265 (2020)
species :
Lepisorus yamaokae Seriz.; Shidekobushi 3(1): 57 (2015)
Distribution: Japan (Honshu, Shikoku, Kyushu)
38 JAP
species :
Lepisorus youxingii R.H.Jiang & X.C.Zhang; Phytotaxa 437(2): 102 (2020)
Distribution: China (Guangxi)
36 CHI
species :
Lepisorus zippelii (Blume) C.F.Zhao, R.Wei & X.C.Zhang; Cladistics 36(3): 251 (2019)
Distribution: China (Guangdong, Guangxi, Guizhou, Hainan, Yunnan); Bhutan; India (N-India, Himachal Pradesh, Uttarakhand, Sikkim, Darjeeling, Meghalaya); Myanmar (Kachin, etc.); Thailand; Laos; Cambodia; Vietnam; peninsular Malaysia (Pahang, Perak, Selangor, Negeri Sembilan); Sumatra; Borneo (Mt. Kinabalu, Sabah); Sulawesi; Philippines (Luzon); Java; Lesser Sunda Isl. (Lombok); New Guinea (Papua New Guinea)
36 CHI 40 BHU IND 41 CBD LAO MYA THA VIE 42 BOR JAW LSI MLY NWG PHI SUL SUM
Remarks: sect. 3 Neolepisorus
Synonyms: hidden
= Bathmium subfalcatum (J.Sm.) Fée; Mém. Foug. 5 (Gen. Filic.): 287 (1852)
= Colysis zippelii (Blume) J.Sm.; Hist. Fil. 101 (1875)
= Drynaria subfalcata J.Sm.; J. Bot. (Hooker) 3: 397 (1841)
= Microsorum luzonicum (Copel.) Tagawa; Acta Phytotax. Geobot. 16: 51 (1955)
= Microsorum zippelii (Blume) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 308 (1933)
= Neocheiropteris zippelii (Blume) Bosman; Leiden Bot. Ser. 14: 123 (1991)
= Neolepisorus zippelii (Blume) Li Wang; Bot. J. Linn. Soc. 162(1): 36 (2010)
= Pleopeltis heterocarpa Bedd.; Ferns Brit. India t. 319 (1869), nom. illeg.
= Pleopeltis luzonica (Copel.) Alderw.; Bull. Dépt. Agric. Indes Néerl. 27: 7 (1909)
= Pleopeltis zippelii (Blume) T.Moore; Index Fil. [T. Moore] 308 (1862)
= Polypodium heterocarpum var. zippelii (Blume) Baker; Syn. Fil. [W. J. Hooker & Baker] 360 (1868)
= Polypodium luzonicum Copel.; Philipp. J. Sci. 1, Suppl. II: 162, t. 23 (1906), not C. Presl (1825)
= Polypodium oxyphyllum Kunze; Bot. Zeitung (Berlin) 116 (1848)
= Polypodium zippelii Blume; Enum. Pl. Javae 2: 172, t. 80 (1828)
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A complete, synonymic checklist of the Higher Plants of the World

Version 24.11 (November 5th, 2024)

Covering 355,576 accepted species,
8,715 hybrids, 47,863 accepted infraspecific taxa and 1,000,581 synonyms,
in total 1,412,785* names

[*about 25,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus" are deliberately omitted]

With complete, detailed distribution data (countries and regions) and literature references

Arranged in linear phylogenetic sequence reflecting true relationships

Fully searchable and continously updated

With 38,209 pictures of 15,787 species

Assembled through 40 years independently from other databases

In cooperation with Catalogue of Life