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Ver. 24.10
October 9th, 2024

249.0000 Family Urticaceae Juss.; Gen. Pl.: 400 (1789), nom. cons.
ca. 55 genera, ca. 2050 spp. Phylogeny: Wu et al. (2013)
Species listed in database: 2070; hybrids: 6
= Cecropiaceae C.C.Berg; Taxon 27: 39 (1978)
= Parietariaceae Bercht. & J.Presl; Prir. Rostlin: 261 (1820)
249.2143 Genus Gonostegia Turcz.; Bull. Soc. Imp. Nat. Moscou 16: 509 (1846)
Distribution: Tropical Asia, Australia
Species listed in database: 3
species :
Gonostegia parvifolia (Roxb. ex Wight) Miq.; Ann. Mus. Bot. Lugduno-Batavi 4: 303 (1869)
Distribution: Taiwan; Sri Lanka; Philippines (Luzon)
38 TAI 40 SRL 42 PHI
Synonyms: hidden
= Gonostegia matsudae (Yamam.) Yamam. & Masam.; J. Soc. Trop. Agric. 3: 392 (1931)
= Gonostegia neurocarpa (Yamam.) Yamam. & Masam.; J. Soc. Trop. Agric. 3: 392 (1931)
= Memorialis matsudae Yamam.; Icon. Pl. Formosan., Suppl. 1: 23-25, f. 8 (1925)
= Memorialis neurocarpa Yamam.; Icon. Pl. Formosan., Suppl. 1: 25-26, f. 9 (1925)
= Memorialis parvifolia (Wight) Wedd.; Arch. Mus. Par. 8: 432 (1855 publ. 1856)
= Memorialis triandra Wedd.; Prodr. [A. DC.] 16(1): 235(9) (1869)
= Pouzolzia hirta var. parvifolia (Wight) Friis & Wilmot-Dear; Nordic J. Bot. 24(1): 91 (111) (2006)
= Pouzolzia matsudae (Yamam.) S.S.Ying; Coloured Illustr. Fl. Taiwan 3: 479 (1988), as ´matsudai´
= Pouzolzia neurocarpa (Yamam.) S.S.Ying; Coloured Illustr. Fl. Taiwan 3: 481 (1988)
= Pouzolzia parvifolia Wight; Ic. Pl. Ind. Orient. 6: 39, t. 2092, f. 1 (1853)
species :
Gonostegia pentandra (Roxb.) Miq.; Ann. Mus. Bot. Lugduno-Batavi 4: 302 (1868)
Distribution: Afghanistan; China (Guangdong, Guangxi, Yunnan, Hainan); Taiwan; Pakistan (N. W. Frontier Prov., Peshawar, Chitral, Swat, Hazara, Punjab, Rawalpindi); Nepal; India (Andhra Pradesh, Arunachal Pradesh, Bihar, Chhattisgarh, Himachal Pradesh, Jammu & Kashmir, Karnataka, Kerala, Madhya Pradesh, Maharashtra, Northeast, Odisha, Punjab, Sikkim, Tamil Nadu, Telangana, Uttar Pradesh, Uttarakhand, West Bengal); Sri Lanka; Bangladesh; Myanmar (Mandalay, Taninthayi); Vietnam; Philippines (Luzon, Mindanao); Sulawesi; Java; Lesser Sunda Isl.; New Guinea (Irian Jaya, Papua New Guinea)
34 AFG 36 CHI 38 TAI 40 BAN IND NEP PAK SRL 41 MYA VIE 42 JAW LSI NWG PHI SUL
Synonyms: hidden
= Boehmeria melastomoidea Griff.; Notul. 4: 386 (1854)
= Gonostegia alternifolia Turcz.; Bull. Soc. Nat. Mosc. 19: II. 510 (1846)
= Gonostegia aquatica Miq.; Ann. Mus. Bot. Lugduno-Batavi 4: 303 (1869)
= Gonostegia oppositifolia Turcz.; Bull. Soc. Nat. Mosc. 19: II. 510 (1846)
= Gonostegia pentandra var. akoensis (Yamam.) Yamam. & Masam.; J. Soc. Trop. Agric. 3: 392 (1931)
= Gonostegia pentandra var. hypericifolia (Blume) Masam.; J. Soc. Trop. Agric. 3: 114 (1941)
= Gonostegia pentandra var. integrifolia (Hook.f.) Pusalkar & Ingle; Fl. Ind. 24: 90 (2022)
= Gonostegia pentandra var. ramosissima (Wight) Pusalkar & Ingle; Fl. Ind. 24: 91 (2022)
= Gonostegia pentandra var. stocksii (Wight) Pusalkar & Ingle; Fl. Ind. 24: 91 (2022)
= Hyrtanandra javanica Miq.; Pl. Jungh. 25 (1851)
= Hyrtanandra pentandra (Roxb.) Miq.; Pl. Jungh. 25 (1851)
= Hyrtanandra ternata Miq. ex Bedd.; Trans. Linn. Soc. 25: 224 (1865)
= Memorialis ciliaris Buch.-Ham. ex Wall.; Numer. List no. 4598 (1831)
= Memorialis pentandra (Roxb.) Wedd.; Prodr. [A. DC.] 16: I. 2352 (1869)
= Memorialis pentandra var. akoensis Yamam.; Icon. Pl. Formosan., Suppl. 1: 26-27 (1925)
= Memorialis pentandra var. hypericifolia (Blume) Wedd.; Prodr. 16(1): 2353 (1869)
= Memorialis ternata Wedd.; Arch. Mus. Par. 8: 422 (1855 publ. 1856)
= Phenax aquatica Wedd.; Prodr. [A. DC.] 16(1): 235(2) (1869)
= Phenax pentandrus (Roxb.) Wedd.; Prodr. [A. DC.] 16(1): 235(2) (1869), nom. illeg.
= Phenax stocksii Wight ex Wedd.; Prodr. [A. DC.] 16(1): 492 (1869), nomen
= Phenax ternata Benn. ex Wedd.; Prodr. [A. DC.] 16(1): 492 (1869), nomen
= Pouzolzia acuta Wight; Icon. Pl. Ind. Orient. 6: 39 (1853)
= Pouzolzia dalzellii Wight; Ic. Pl. Ind. Orient. 6: 41 (1853)
= Pouzolzia hypericifolia Blume; Mus. Bot. 2: 242 (1857)
= Pouzolzia longifolia Wight; Icon. Pl. Ind. Orient. 6: 40 (1853)
= Pouzolzia pentandra (Roxb.) Benn.; Pl. Jáv. Rar. 64 (1838)
= Pouzolzia pentandra var. akoensis (Yamam.) S.S.Ying; Coloured Illustr. Fl. Taiwan 3: 477 (1988)
= Pouzolzia pentandra var. dalzellii M.R.Almeida; Fl. Maharashtra 4B: 392 (2003), nom. inval.
= Pouzolzia pentandra var. hypericifolia (Blume) S.S.Ying; Coloured Illustr. Fl. Taiwan (I) 3: 476 (1988)
= Pouzolzia pentandra var. integrifolia Hook.f.; Fl. Brit. India 5: 583 (1888)
= Pouzolzia pentandra var. ramosissima (Wight) Hook.f.; Fl. Brit. India 5: 583 (1888)
= Pouzolzia pentandra var. stocksii (Wight) Hook.f.; Fl. Brit. India 5: 583 (1888)
= Pouzolzia ramosissima Wight; Ic. Pl. Ind. Orient. 6: 41 (1853)
= Pouzolzia stocksii Wight; Ic. Pl. Ind. Orient. 6: 41 (1853)
= Pouzolzia ternata Benn.; Pl. Jáv. Rar. 66 (1838)
= Urtica hippurioides Griff.; Itin. Notes, 362 (1848)
= Urtica pentandra Roxb.; Hort. Beng. 67 (1814), nomen
= Urtica ternata B.Heyne ex Wall.; Numer. List no. 4599 (1831), nomen
variant :
var. gracilis (Miq.) Pusalkar; Fl. Pl. India Annot. Checkl. 2: 523 (2020)
Distribution: S-India (Karnataka, Kerala, Tamil Nadu)
40 IND
Synonyms: hidden
= Gonostegia gracilis (Miq.) Miq.; Ann. Mus. Bot. Lugduno-Batavi 4: 303 (1869)
= Gonostegia meeboldii (W.W.Sm. & Ramaswami) Pusalkar & Ingle; Fl. Ind. 24: 89 (2022)
= Hyrtanandra gracilis Miq.; Ann. Mus. Bot. Lugduno-Batavi 4: 303 (1869)
= Pouzolzia aspera Wight; Ic. Pl. Ind. Orient. 6: 40, t. 2095, f. 18 (1853)
= Pouzolzia meeboldii W.W.Sm. & Ramaswami; Rec. Bot. Surv. India 6: 40 (1914)
= Pouzolzia pentandra var. gracilis (Miq.) Friis & Wilmot-Dear; Nordic J. Bot. 24(1): 102 (2006)
variant :
var. wightii (Benn.) Pull. & Karupp.; Fl. E. Ghats 5: 320 (2019)
Distribution: Soqotra [I]; India (Andhra Pradesh, Karnataka, Kerala, Maharashtra, Odisha, Telangana, Tamil Nadu); Sri Lanka; Vietnam
24 soc 40 IND SRL 41 VIE
Synonyms: hidden
= Gonostegia ambigua (Wight) Miq.; Ann. Mus. Bot. Lugduno-Batavi 4: 303 (1869)
= Gonostegia bracteata (Benn.) Miq.; Ann. Mus. Bot. Lugduno-Batavi 4: 303 (1869)
= Gonostegia caudata (Benn.) Miq.; Ann. Mus. Bot. Lugduno-Batavi 4: 303 (1869)
= Gonostegia concinna (Wight) Miq.; Ann. Mus. Bot. Lugduno-Batavi 4: 303 (1869)
= Gonostegia lawsoniana (C.E.C.Fisch.) Pusalkar & Ingle; Fl. Ind. 24: 88 (2022)
= Gonostegia nilghirensis (Wight) Miq.; Ann. Mus. Bot. Lugduno-Batavi 4: 303 (1869)
= Gonostegia pentandra subsp. wightii (Benn.) Pusalkar; Fl. Pl. India Annot. Checkl. 2: 523 (2020)
= Gonostegia pentandra var. wightii (Benn.) Pusalkar; Fl. Pl. India Annot. Checkl. 2: 523 (2020), isonym
= Gonostegia scabra (Wight) Miq.; Ann. Mus. Bot. Lugduno-Batavi 4: 303 (1869)
= Gonostegia wightii (Benn.) Pusalkar & Ingle; Fl. Ind. 24: 91 (2022)
= Gonostegia wightii var. neilgherrensis (Wight) Pusalkar & Ingle; Fl. Ind. 24: 93 (2022)
= Gonostegia wightii var. ovata (Wight) Pusalkar & Ingle; Fl. Ind. 24: 93 (2022)
= Gonostegia wightii var. wallichiana (Wight) Pusalkar & Ingle; Fl. Ind. 24: 93 (2022)
= Hyrtanandra caudata (Benn.) Miq. ex Bedd.; Trans. Linn. Soc. London 25: 224 (1865)
= Hyrtanandra scabra (Wight) Miq. ex Bedd.; Trans. Linn. Soc. 25: 224 (1865)
= Hyrtanandra walkeriana (Wight) Thwaites; Enum. Pl. Zeyl. 261 (1859)
= Memorialis ambigua (Wight) Wedd.; Arch. Mus. Hist. Nat. 9: 430 (1857)
= Memorialis aquatica Moon ex Wedd.; Arch. Mus. Par. 8: 418 (1855 publ. 1856)
= Memorialis bracteata (Benn.) Wedd.; Arch. Mus. Hist. Nat. 9: 421 (1857)
= Memorialis caudata (Benn.) Wall. ex Wedd.; Arch. Mus. Hist. Nat. 9: 425 (1857)
= Memorialis caudata var. velutina Wedd.; Arch. Mus. Hist. Nat. 9: 569 (1957)
= Memorialis concinna (Wight) Wedd.; Prodr. [A. DC.] 16: I. 235(4) (1869)
= Memorialis concinna var. velutina (Wedd.) Wedd.; Prodr. [A. DC.] 16(1): 235(5) (1869)
= Memorialis neilgherrensis (Wight) Wedd.; Arch. Mus. Hist. Nat. 9: 420 (1857)
= Memorialis neilgherrensis var. ternata Wedd.; Arch. Mus. Hist. Nat. 9: 420 (1857)
= Memorialis neilgherrensis var. trialata (Wight) Wedd.; Arch. Mus. Hist. Nat. 9: 420 (1857)
= Memorialis neilgherrensis var. wallichiana (Wight) Wedd.; Arch. Mus. Hist. Nat. 9: 420 (1857)
= Memorialis scabra (Wight) Wedd.; Arch. Mus. Par. 8: 423 (1855 publ. 1856)
= Memorialis ternata var. concinna (Wight) Wedd.; Arch. Mus. Hist. Nat. 9: 422 (1857)
= Parietaria bracteata Wight ex Wall.; Numer. List no. 4600 (1831)
= Phenax concinna (Wight) Wedd.; Prodr. [A. DC.] 16(1): 235(2) (1869)
= Phenax neilgherrensis (Wight) Wedd.; Prodr. [A. DC.] 16(1): 235(2) (1869)
= Pouzolzia ambigua Wight; Ic. Pl. Ind. Orient. 6: 39 (1853)
= Pouzolzia bracteata (Wight ex Wall.) Benn.; Pl. Jáv. Rar. 67 (1838)
= Pouzolzia caudata (Wall.) Benn.; Pl. Jáv. Rar. 66 (1838)
= Pouzolzia concinna Wight; Ic. Pl. Ind. Orient. 6: 40 (1853)
= Pouzolzia courtallensis Wight; Ic. Pl. Ind. Orient. 6: 38, t. 2093, f. 10 (1853)
= Pouzolzia glabra Wight; Ic. Pl. Ind. Orient. 6: 41 (1853)
= Pouzolzia neilgherrensis Wight; Ic. Pl. Ind. Orient. 6: 42 (1853)
= Pouzolzia oblongifolia Wight; Icon. Pl. Ind. Orient. 6: 42 (1853)
= Pouzolzia ovata Wight; Icon. Pl. Ind. Orient. 6: 41 (1853)
= Pouzolzia pentandra subsp. wightii (Benn.) Friis & Wilmot-Dear; Nordic J. Bot. 24(1): 100 (108) (2006)
= Pouzolzia pentandra var. walkeriana (Wight) Hook.f.; Fl. Brit. India 5: 583 (1888)
= Pouzolzia scabra Wight; Ic. Pl. Ind. Orient. 6: 41 (1853)
= Pouzolzia trialata Wight; Icon. Pl. Ind. Orient. 6: 42 (1853)
= Pouzolzia walkeriana Wight; Ic. Pl. Ind. Orient. 6: 41, t. 2095 f. 16 (1853)
= Pouzolzia wightii Benn.; Pl. Jáv. Rar. 66 (1838)
= Pouzolzia wightii var. caudata (Benn.) C.E.C.Fisch.; Fl. Madras: 1384 (1928)
= Pouzolzia wightii var. laevifolia Hook.f.; Fl. Brit. India 5: 584 (1888)
= Pouzolzia wightii var. lawsoniana C.E.C.Fisch.; Fl. Madras 3: 1384 (1928)
= Pouzolzia wightii var. nilghirensis (Wight) Hook.f.; Fl. Brit. India 5: 584 (1888)
= Pouzolzia wightii var. wallichiana (Wight) Hook.f.; Fl. Brit. India 5: 584 (1888)
= Urtica aquatica Moon; Cat. Pl. Ceyl. 62 (1824) (nom. nud.)
= Urtica bracteata Wight ex Hook.f.; Fl. Brit. Ind. 5: 585 (1888)
= Urtica caudata Wall.; Numer. List no. 4600 (1831)
= Urtica foliolosa Wall. ex Wedd.; Arch. Mus. Par. 9: 103 (1856)
species :
Gonostegia triandra (Blume) Miq.; Ann. Mus. Bot. Lugduno-Batavi 4: 303 (1869)
Distribution: Madagascar; China (Yunnan, Sichuan, Guangxi, Hainan, Guangdong, Fujian, Jiangxi, Zhejiang, Anhui, Jiangsu, Shaanxi, Henan); SE-Tibet; Japan (Honshu, Kyushu); Ryukyu Isl.; Taiwan; Pakistan; Nepal; Bhutan; India (Andhra Pradesh, Arunachal Pradesh, Bihar, Chhattisgarh, Himachal Pradesh, Jammu & Kashmir, Jharkhand, Karnataka, Kerala, Madhya Pradesh, Maharashtra, Northeast, Odisha, Punjab, Sikkim, Tamil Nadu, Telangana, Uttar Pradesh, Uttarakhand, West Bengal); Sri Lanka; Nicobar Isl. (Central Nicobars); Myanmar (Mandalay); Thailand; Laos; Vietnam; peninsular Malaysia (Pahang); Philippines (Luzon, Mindanao); Java; Lesser Sunda Isl. (Bali, etc.); Moluccas; New Guinea (Irian Jaya, Papua New Guinea); Solomon Isl.; Australia (Queensland: Lynd R. and Mackay areas)
29 MDG 36 CHI TIB 38 JAP NNS TAI 40 BHU IND NEP PAK SRL 41 LAO MYA NCB THA VIE 42 JAW LSI MLY MOL NWG PHI 43 SOL 50 AUS
Synonyms: hidden
= Driessenia sinensis H.Lév.
= Gonostegia bennettiana (Wight) Pusalkar & Ingle; Fl. Ind. 24: 82 (2022)
= Gonostegia bennettiana var. acuta (Wight) Pusalkar & Ingle; Fl. Ind. 24: 84 (2022)
= Gonostegia bennettiana var. longifolia (Wight) Pusalkar & Ingle; Fl. Ind. 24: 84 (2022)
= Gonostegia bennettiana var. mysorensis (Wight) Pusalkar & Ingle; Fl. Ind. 24: 84 (2022)
= Gonostegia bennettiana var. tomentosa (Wight); Fl. Ind. 24: 84 (2022)
= Gonostegia gardneri (Wight) Pusalkar & Ingle; Fl. Ind. 24: 94 (2022)
= Gonostegia hirta (Hassk.) Miq.; Ann. Mus. Bot. Lugduno-Batavum 4: 303. 1868-69
= Gonostegia hirta var. tenggerensis Hochr.; Candollea 2: 347 (1925)
= Gonostegia integrifolia (Dalzell) Pusalkar & Ingle; Fl. Ind. 24: 88 (2022), isonym
= Gonostegia integrifolia (Dalzell) Miq.; Ann. Mus. Bot. Lugduno-Batavi 4: 303 (1869)
= Gonostegia quinquenervis (Benn.) Miq.; Ann. Mus. Bot. Lugduno-Batavi 4: 303 (1869)
= Hyrtanandra hirta (Blume) Miq.; Pl. Jungh. 26 (1851)
= Hyrtanandra hirta var. bennettiana (Wight) Thwaites; Enum. Pl. Zeyl.: 261 (1864)
= Hyrtanandra hirta var. gardneriana (Wight) Thwaites; (1864)
= Hyrtanandra lythroides (F.Muell.) F.Muell.; Fragm. 6: 253 (1868)
= Hyrtanandra pouzolzia F.Muell.; Fragm. 5: 194 (1866)
= Hyrtanandra triandra (Blume) Miq.; Fl. Ned. Ind. 1(2): 262 (1859)
= Memorialis dalzellii Wedd.; Arch. Mus. Par. 8: 431 (1855 publ. 1856), nom. illeg.
= Memorialis hirta (Blume) Wedd.; Prodr. 16(1): 2356 (1869)
= Memorialis hispida Buch.-Ham. ex Wall.; Numer. List no. 4601 (1831), nomen
= Memorialis hispida var. bennettiana (Wight) Wedd.; Arch. Mus. Hist. Nat. 9: 427 (1857)
= Memorialis hispida var. gardneriana (Wight) Wedd.; Arch. Mus. Hist. Nat. 9: 427 (1857)
= Memorialis integrifolia (Dalzell) Wedd.; Prodr. [A. DC.] 16: I. 235s (1869)
= Memorialis integrifolia var. pubescens Griseb.; Index Seminum (GOET, Gottingensis) 1878: 8 (1878)
= Memorialis lythroides F.Muell.; Fragm. 5: 194 (1866)
= Memorialis mysorensis (Wight) Wedd.; Arch. Mus. Hist. Nat. 9: 428 (1857)
= Memorialis quinquenervis Buch.-Ham. ex Wall.; Numer. List no. 4601 (1831), nomen
= Phenax tomentosa Wight ex Wedd.; Prodr. [A. DC.] 16(1): 492 (1869), nomen
= Pouzolzia bennettiana var. acuta Fischer; Gamble, Fl. Pres. Madras 1385 (1928)
= Pouzolzia bennettiana var. gardneriana (Wight) Hook.f.; Fl. Brit. India 5: 586 (1888)
= Pouzolzia bennettiana var. macrophylla Hook.f.; Fl. Brit. India 5: 586 (1888)
= Pouzolzia bennettiana var. mysorensis (Wight) Hook.f.; Fl. Brit. India 5: 586 (1888)
= Pouzolzia bennettiana var. ovalifolia (Wight) Hook.f.; Fl. Brit. India 5: 586 (1888)
= Pouzolzia bennettiana var. quadrialata (Wight) Hook.f.; Fl. Brit. India 5: 586 (1888)
= Pouzolzia bennettiana var. tomentosa (Wight) Hook.f.; Fl. Brit. India 5: 586 (1888)
= Pouzolzia bennettiana Wight; Ic. Pl. Ind. Orient. 6: 40 (1853)
= Pouzolzia gardneri Wight; Icon. Pl. Ind. Orient. 6: t. 2092, f. 5 (1853), orth. var.
= Pouzolzia gardneriana Wight; Ic. Pl. Ind. Orient. 6: 39 (1853)
= Pouzolzia heterocarpa Wight; Ic. Pl. Ind. Orient. 6: 40 (1853)
= Pouzolzia hirta (Blume) Hassk.; Cat. Hort. Bot. Bogor. 80 (1844) non Sw.
= Pouzolzia hirta var. lythroides (F.Muell.) Domin; Biblioth. Bot. 89: 23 (1921)
= Pouzolzia hirta var. quinquenervis (Benn.) Domin; Biblioth. Bot. 89: 23 (1921)
= Pouzolzia hispida Benn.; Pl. Jáv. Rar. 66 (1838)
= Pouzolzia huegeliana Blume; Mus. Bot. Lugduno-Batavi 2: 241(1857)
= Pouzolzia huegeliana var. ovalifolia (Wight) M.R.Almeida; Fl. Maharashtra 4B: 391 (2003)
= Pouzolzia integrifolia Dalzell; Hook. Kew J. 3: 134 (1851)
= Pouzolzia lucida Hassk.; Cat. Hort. Bot. Bogor. Alt.: 80 (1844)
= Pouzolzia lythroides F.Muell.; Fragm. 5: 194 (1866)
= Pouzolzia mysorensis Wight; Ic. Pl. Ind. Orient. 6: 39 (1853)
= Pouzolzia obscura Blume; Mus. Bot. Lugduno-Batavi 2: 241(1857)
= Pouzolzia ovalifolia Wight; Ic. Pl. Ind. Orient. 6: 39 (1853)
= Pouzolzia quadrialata Wight; Ic. Pl. Ind. Orient. 6: 37, t. 2094, f. 12 (1853)
= Pouzolzia quinquenervis Benn.; Pl. Jáv. Rar. 66 (1838)
= Pouzolzia sponiifolia Schltdl. ex Blume; Mus. Bot. Lugduno-Batavi 2: 244(1857)
= Pouzolzia tomentosa Wight; Ic. Pl. Ind. Orient. 6: 40 (1853)
= Pouzolzia triandra (Blume) Blume; Mus. Bot. 2: 241 (1857)
= Pouzolzia wightii var. longifolia Hook.f.; Fl. Brit. India 5: 584 (1888)
= Urtica hirta Blume; Bijdr. Fl. Ned. Ind. 495 (1825)
= Urtica quinquenervis Wall.; Numer. List no. 4601 (1831), nomen
= Urtica triandra Blume; Bijdr. Fl. Ned. Ind.: 496 (1826)
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A complete, synonymic checklist of the Higher Plants of the World

Version 24.10 (October 9th, 2024)

Covering 355,419 accepted species,
8,711 hybrids, 47,863 accepted infraspecific taxa and 1,000,150 synonyms,
in total 1,412,143* names

[*about 25,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus" are deliberately omitted]

With complete, detailed distribution data (countries and regions) and literature references

Arranged in linear phylogenetic sequence reflecting true relationships

Fully searchable and continously updated

With 38,209 pictures of 15,787 species

Assembled through 40 years independently from other databases

In cooperation with Catalogue of Life