034.0000 Family Dryopteridaceae Herter; Revista Sudamer. Bot. 9: 15 (1949), nom. cons.

26 genera, ca. 2100 spp. Phylogeny: Zuo et al. (2024)

Species listed in database: 2174; hybrids: 201

034.7008 Genus Dryopteris Adans.; Fam. Pl. 2: 20 (1763), nom. cons., s. l.

Distribution: Northern temperate zone (over 150 spp. in China and Taiwan), tropical mountains, only 22 New World. Inclusion of all the satellite genera like Acrorumohra, Adenoderris, Peranema, Diacalpe, Dryopsis, Nothoperanema etc. fide Zhang (2012) and others, based on molecular results. Fraser-Jenkins et al. (2018) argue for keeping some genera distinct, like Dryopsis, Stenolepia and Nothoperanema.

Species listed in database: 348; hybrids: 91

= Acrophorus C.Presl; Tent. Pterid. 93 (1836) (fide Zhang (2012))

= Acrorumohra (H.Itô) H.Itô; Nov. Fl. Jap. [Nakai & Honda] 4: 101 (1939) (fide Zhang (2012))

= Anopodium J.Sm.; Cat. Cult. Ferns: 16 (1857)

= Arsenopteris Webb & Berthel.; Hist. Nat. Iles Canaries (Phytogr.) 3: 437 (1847)

= Arthrobotrys Wall.; Numer. List. [Wallich] no. 395 (1829), nomen

= Catapodium J.Sm.; Cat. Cult. Ferns: 16 (1857)

= Ctenitis subgen.Dryopsis Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 280 (1938)

= Desmopodium J.Sm. ex T.Moore; Index Fil. [T. Moore] 308 (1861)

= Diacalpe Blume; Enum. Pl. Javae 2: 241 (1828) (fide Zhang (2012))

= Dichasium (A.Braun) Fée; Mém. Foug. 5 (Gen. Filic.): 302 (1852)

= Diclisodon T.Moore; Index Fil. [T. Moore] 95 (1857)

= Dryopsis Holttum & Edwards; Kew Bull. 41: 179 (1986) (fide Zhang (2012))

= Dryopteris sect.Dryopsis (Holttum & P.J.Edwards) Li Bing Zhang; Phytotaxa 71: 18 (2012)

= Dryopteris subgen.Lophodium (Newman) Tzvelev; Novosti Sist. Vyssh. Rast. 35: 10 (2003)

= Filix Ség.; Pl. Veron. 3: 53 (1754), nom. rej.

= Filix-mas Hill ex Farw.; Amer. Midl. Naturalist 12: 235, 253 (1931)

= Gleichenia Neck.; Elem. Bot. (Necker) 3: 314 (1790)

= Gymnothalamium Zenker ms. ex Kunze; Linnaea 24: 287 (1851)

= Lophodium Newman; Phytologist 4: 371, app. XVI (1851)

= Megalastrum J.Sm.; Hist. Fil. 216 (1875)

= Microdictyon Fée; Mém. Foug. 11 (Hist. Foug. Antil.): 66 (1866)

= Monophlebia T.Moore; Index Fil. [T. Moore] 89 (1857)

= Nematopera Kunze; Bot. Zeitung (Berlin) 797 (1846) (corr. nom.)

= Nephrodium Rich.; Cat. Pl. Jard. Méd. Paris 120 (1801)

= Nothoperanema (Tagawa) Ching; Acta Phytotax. Sin. 11: 25 (1966) (fide Zhang (2012))

= Oligocampia Trevis.; Atti Reale Ist. Veneto Sci. Lett. Arti, ser. 2, 2: 165 (1851)

= Pachyderris J.Sm.ms. ex T.Moore; Index Fil. [T. Moore] 146 (1857)

= Peranema D.Don; Prodr. Fl. Nepal. 12 (1825) (fide Zhang (2012))

= Podeilema R.Br.ms. ex Wall.; Pl. Asiat. Rar. [Wallich] 1: 42 (1830)

= Psidopodium Neck.; Elem. Bot. (Necker) 3: 315 (1790)

= Pteris Gled.; Syst. Pl. [Gleditsch] 289 (1764)

= Pycnopteris T.Moore; Gard. Chron. 1855: 468 (1855)

= Revwattsia D.L.Jones; Fl. Australia 48: 711 (1998)

= Rumohra sect.Acrorumohra H.Itô; J. Jap. Bot. 11: 583 (1935)

= Sphaeropteris R.Br. ex Wall.; Pl. asiat. rar. 1: 41 (1830) (nom. illeg.)

= Syneuron J.Sm.ms. ex Hook.; Hooker´s Icon. Pl. 6: t.950 (1854)

= Stenolepia Alderw.; Bull. Dépt. Agric. Indes Néerl. 27: 45 (1909)

= Dryopteris sect.Stenolepia (Alderw.) L.Y.Kuo & Y.H.Chang; Syst. Bot. 41(3): 603 (2016)

species :

Dryopteris acutodentata Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 432 (1938)

Distribution: China (W-Sichuan, NW-Yunnan); Tibet; Taiwan; Nepal; Bhutan; India (Jammu & Kashmir, Himachal Pradesh, Uttarakhand, Sikkim, Darjeeling, Arunachal Pradesh, West Bengal); Myanmar (Kachin, etc.)

36 CHI TIB 38 TAI 40 BHU IND NEP 41 MYA

Synonyms: hidden

= Dryopteris filix-mas subsp. kingii (C.Hope) C.Chr.; Index Filic. 265 (1905)

= Dryopteris silaensis Ching; Bull. Fan Mem. Inst. Biol. Bot. 11: 65 (1941)

= Nephrodium kingii C.Hope; J. Bombay Nat. Hist. Soc. 12: 621 (1899) (non Dryopteris kingii (Bedd.) C. Chr.)

species :

Dryopteris adnata (Blume) Alderw.; Malayan Ferns 191 (1909) & Suppl. 501 (1917)

Distribution: Sumatra; Borneo (Mt. Kinabalu, Sabah); Sulawesi; Java; Lesser Sunda Isl. (Timor)

42 BOR JAW LSI SUL SUM

Remarks: Subgen. Dryopsis

Synonyms: hidden

= Aspidium adnatum Blume; Enum. Pl. Javae 2: 162 (1828)

= Ctenitis adnata (Blume) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 289 (1938)

= Dryopsis adnata (Blume) Holttum & P.J.Edwards; Kew Bull. 41(1): 190 (1986)

= Polystichum truncatulum Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 16: 60, t. 8 (1914)

species :

Dryopteris aemula (Aiton) Kuntze; Revis. Gen. Pl. 2: 812 (1891)

subspecies :

subsp. aemula

Name: Heuduftender Wurmfarn (DE); Hay-scented Buckler-fern (EN); Ninfa de los bosques (ES)

Distribution: England (N-, W- & S-England, Wales, Scotland, Outer Hebrides, Orkney Isl., Isle of Man); Ireland (Ireland, Northern Ireland); Spain; France; Azores (Santa Maria Isl., Sao Miguel Isl., Terceira, Graciosa, Sao Jorge, Pico, Faial, Flores Isl., Corvo Isl.); Madeira (Madeira Isl.); Canary Isl. (La Gomera)

10 GRB IRE 12 FRA SPA 21 AZO CNY MDR

Remarks: ser. Aemulae

Synonyms: hidden

= Allantodia aemula (Aiton) Desv.; Mem. Soc. Linn. Paris 6(2): 265 (1827)

= Aspidium aemulum (Aiton) Sw.; J. Bot. (Schrader) 1800(2): 42 (1801)

= Aspidium distans var. recurvum Bree; Mag. Nat. Hist. 4: 162 (1831)

= Aspidium recurvum (Newman) Bree; Phytologist 1: 773 (1843)

= Aspidium foenisecii Lowe; J. Bot. (Hooker) 1: 26 (1834), nom. nud.

= Lastrea aemula (Aiton) Brack.; U.S. Expl. Exped., Filic. 16: 200 (1854)

= Lastrea dilatata var. recurva (Bree) H.C.Watson; London Cat. Brit. Pl. [H. C. Watson] 14 (1844)

= Lastrea foenisecii Lowe ex Watson; Phytologist 2: 568 (1846)

= Lastrea recurva Newman; Natur. Alm. 23 (1844)

= Lophodium foenisecii Newman; Phytologist 4: app. 16 (1851)

= Lophodium recurvum (Newman) Newman; Phytologist 4: 371 (1851)

= Nephrodium aemulum (Aiton) Baker; Syn. Fil. [W. J. Hooker & Baker] 279 (1867)

= Nephrodium foenisecii Lowe; Trans. Cambridge Philos. Soc. 4: 7 (1830)

= Nephrodium foenisecii var. tripinnatum (Corb.) Rouy; Fl. France [Rouy & Foucaud] 14: 414 (1913)

= Polypodium aemulum Aiton; Hort. Kew. [W. Aiton] 3: 466 (1789)

= Polystichum aemulum (Aiton) C.Presl; Tent. Pterid. 83 (1836)

= Polystichum aemulum (Aiton) Corb.; Nouv. Fl. Normandie: 676 (1894)

= Polystichum aemulum var. tripinnatum Corb.; Nouv. Fl. Normandie, add. 115 (1895)

= Polystichum foenisecii (Lowe) P.E.Fourn.; Quatre Fl. France: 17 (1934)

= Polystichum recurvum (Newman) Gand.; Fl. Eur. [Gand.] 27: 174 (1891)

= Polystichum spinulosum subsp. aemulum (Aiton) R.C.V.Douin; Bonnier, Fl. Ill. France 12: 95 (1934)

subspecies :

subsp. liliana (Golitsin) Askerov; Dokl. Akad. Nauk Azerbaidzhansk. S.S.R. 1982(9): 59 (1982)

Distribution: Georgia [Caucasus] (Adzharia); Turkey (Anatolia)

33 GEO 34 TUR

Synonyms: hidden

= Dryopteris liliana Golitsin; Repert. Spec. Nov. Regni Veg. 31: 388 (1933)

species :

Dryopteris affinis (Lowe) Fraser-Jenk.; Fern Gaz. 12: 56 (1979)

subspecies :

subsp. affinis

Name: Spreuschuppiger Wurmfarn (DE); Dentabron (ES)

Distribution: England (England, Wales, Scotland, Outer Hebrides, Orkney Isl., Isle of Man); Ireland (Ireland, Northern Ireland); Norway; Netherlands; Belgium; Luxembourg; Germany (Baden-Württemberg, Bayern, Niedersachsen, Rheinland-Pfalz, Saarland, Sachsen, Sachsen-Anhalt); Switzerland; Liechtenstein; Austria; Poland; Slovakia; Hungary; Portugal; Spain; France; Channel Isl. (Alderney, Guernsey, Jersey); Corsica; Sardinia; Italy (incl. Elba Isl.); Sicily; Slovenia; Romania; Greece (Samothraki); European Turkey; Madeira (Madeira Isl.); Canary Isl. (La Gomera, Tenerife); Northern Caucasus; Georgia [Caucasus]; ?Armenia; Azerbaijan; Turkey (Anatolia); New Zealand [I]

10 GRB IRE NOR 11 AUT BGM GER HUN LIE LUX NET POL SLO SWI 12 CIS COR FRA POR SAR SPA 13 GRC ITA ROM SIC SLV TUE 21 CNY MDR 33 AZE GEO NCS 34 TUR 51 nzl

Remarks: affinis agg. The species contains two distinct genetic lines and probably needs splitting in two (Jessen & Bujnoch 2024).

Synonyms: hidden

= Aspidium affine Lowe; J. Bot. (Hooker) 1: 26 (1834), nom. nud.

= Aspidium filix-mas f. subintegrum Döll; Fl. Baden [Döll] 1(1): 27 (1857)

= Dryopteris affinis f. valongensis Res.-Pinto ex Fraser-Jenk.; Anales Jard. Bot. Madrid 41(1): 209 (1984), nom. nov. for D. resendeana M. Pinto 1969

= Dryopteris affinis subsp. cluthensis A.R.Church; Fern Gaz. 21(2): 89 (2019) [fide Fraser-Jenkins et al. 2020]

= Dryopteris affinis var. rubiginosa (Fomin) Askerov; Zametki Sist. Geogr. Rast. 39: 7 (1983)

= Dryopteris affinis var. subintegra (Döll) Askerov; Zametki Sist. Geogr. Rast. 39: 7 (1983)

= Dryopteris borreri subsp. resendeana (M.Pinto) Malag.

= Dryopteris borreri var. atlantica Tavel; 153 (1937), nom. inval.

= Dryopteris borreri var. subintegra (Döll ex Asch. & Graebn.) Tavel; (1937)

= Dryopteris paleacea f. rubiginosa Fomin; Vestn. Tiflissk. Bot. Sada 20: 46 (1911)

= Dryopteris pseudomas (Woll.) Holub & Pouzar; Folia Geobot. Phytotax. 2: 332 (1967)

= Dryopteris resendeana M.Pinto; Portug. Acta Biol. 9: 320, f. 1-2 (1969)

= Lastrea filix-mas var. cristata T.Moore & Houlston; Gard. Mag. Bot. 3: 317 (1851)

= Lastrea filix-mas var. paleacea (T.Moore) Syme & N.E.Br.; Engl. Bot., 3rd ed., vol. XII: 59 (1886)

= Lastrea pseudomas Woll.; Phytologist, n. s. 1: 172 (1856)

= Nephrodium affine Lowe; Trans. Cambridge Philos. Soc. 6: 525 (1838)

= Nephrodium filix-mas var. affine (Lowe) Hook.f.; 465 (1870)

= Nephrodium filix-mas var. affine (Lowe) Rouy; Fl. France [Rouy & Foucaud] 14: 407 (1913)

= Nephrodium paleaceum Lowe; Brit. Ferns [Lowe] 133 (1891)

variant :

var. azorica Fraser-Jenk.; Willdenowia 10(1): 110 (1980)

Distribution: Azores (Sao Miguel Isl., Terceira, Sao Jorge, Pico, Faial, Flores Isl.)

21 AZO

variant :

var. disjuncta (Fomin) Fraser-Jenk.; Willdenowia 10: 110 (1980)

Distribution: Germany; etc.

11 GER

Remarks: the predominant form in Central Europe

Synonyms: hidden

= Dryopteris borreri var. disjuncta (Fomin) Tavel; 52 (1937)

= Dryopteris paleacea f. disjuncta Fomin; Vestn. Tiflissk. Bot. Sada 20: 44 (1911)

subspecies :

subsp. jessenii (Fraser-Jenk.) Fraser-Jenk.; Fern Gaz. 18(1): 5 (2007)

Distribution: Serbia; SC-Romania

13 ROM SRB

Synonyms: hidden

= Dryopteris jessenii (Fraser-Jenk.) Tlalka; Ann. Bot. Fenn. 61: 17-23 (2024)

= Dryopteris affinis var. jessenii Fraser-Jenk.; Ann. Bot. Fenn. 33: 73 (1996)

subspecies :

subsp. kerryensis (Fraser-Jenk.) Fraser-Jenk.; Fern Gaz. 18(1): 5 (2007)

Distribution: Ireland (SW-Ireland)

10 IRE

Synonyms: hidden

= Dryopteris affinis subsp. atropes Fraser-Jenk.; Indian Fern J. 39(1): 127 (2022) (fide Jessen in litt. indistinguishable from kerryensis)

= Dryopteris affinis var. kerryensis Fraser-Jenk.; Ann. Bot. Fenn. 33: 73 (1996)

= Dryopteris atropes (Fraser-Jenk.) Tlalka; Ann. Bot. Fenn. 61: 17-23 (2024)

= Dryopteris kerryensis (Fraser-Jenk.) P.D.Sell; Fl. Gr. Brit. Ireland 1: 671 (2018)

subspecies :

subsp. paleaceolobata (T.Moore) Fraser-Jenk.; Fern Gaz. 18(1): 5 (2007)

Distribution: England (W- & N-England, Wales, SW-Scotland, Isle of Man); Ireland (SW-Ireland); Belgium; ?Germany

10 GRB IRE 11 BGM

Synonyms: hidden

= Dryopteris affinis var. paleaceolobata (T.Moore) Fraser-Jenk.; Indian J. Forest. 43(1): 83 (2020), et Indian J. Forest. 43(1): 83 (2021)

= Dryopteris paleaceolobata (T.Moore) P.D.Sell; Fl. Gr. Brit. Ireland 1: 671 (2018)

= Lastrea filix-mas var. paleaceolobata T.Moore; Handb. Brit. Ferns, ed. 3: 112 (1857)

subspecies :

subsp. punctata (Oberh. & Tavel) Fraser-Jenk.; Fern Gaz. 18(1): 5 (2007)

Name: Punktierter Spreuschuppiger Wurmfarn (DE)

Distribution: Germany (Bayern); Switzerland; Austria; France; N-Italy; Slovenia; Croatia; Turkey (Anatolia)

11 AUT GER SWI 12 FRA 13 CRO ITA SLV 34 TUR

Synonyms: hidden

= Dryopteris affinis var. punctata Oberh. & Tavel ex Fraser-Jenk.; Willdenowia 10(1): 110 (1980)

= Dryopteris affinis var. splendens (Ehrler ex Bech.) Fraser-Jenk.; Willdenowia 10(1): 111 (1980)

= Dryopteris borreri subvar. splendens Ehrler ex Becherer; Ber. Schweiz. Bot. Ges. 60: 469 (1950)

species :

Dryopteris aitoniana Pic.Serm.; Webbia 8: 154, f. 3 (1951), nom. nov.

Distribution: Madeira (Madeira Isl.)

21 MDR

Synonyms: hidden

= Aspidium elongatum (Aiton) Sw.; J. Bot. (Schrader) 1800(2): 38 (1801) (nec Willd.)

= Dryopteris elongata (Aiton) A.Chev.; Rev. Bot. Appl. Agric. Trop. 15: 1060 (1935) (non (Wall. ex Hook.) Kuntze)

= Dryopteris macaronesica Romariz; Revista Fac. Ci. Univ. Lisboa, sér. 2a, C, Ci. Nat. 3: 92, t. 5(1) (1953) (non Dryopteris elongata (Wall. ex Hook.) Kuntze)

= Lastrea elongata (Aiton) C.Presl; Tent. Pterid. 76 (1836)

= Lastrea grandis T.Moore; Index Fil. [T. Moore] 93 (1858)

= Nephrodium boryanum Hook.; Sp. Fil. [W. J. Hooker] 4: 126 (1862) (non Aspidium boryanum Willd. (1810))

= Nephrodium elongatum (Aiton) Hook. & Grev.; Icon. Filic. t. 234 (1831)

= Nephrodium filix-mas var. elongatum (Aiton) Hook.; Sp. Fil. [W. J. Hooker] 4: 117 (1862)

= Polypodium elongatum Aiton; Hort. Kew. [W. Aiton] 3: 465 (1789) (non Dryopteris elongata (Wall. ex Hook.) Kuntze)

= Tectaria elongata (Aiton) Cav.; Anales Ci. Nat. 4: 101 (1801)

species :

Dryopteris alpestris Tagawa; Acta Phytotax. Geobot. 3: 88 (1934)

Distribution: China (Yunnan); Tibet; Taiwan; Nepal; NE-India (Arunachal Pradesh, Sikkim); Myanmar (Kachin, etc.)

36 CHI TIB 38 TAI 40 IND NEP 41 MYA

Conservation Status: IUCN EN for India (Fraser-Jenkins 2012)

Synonyms: hidden

= Dryopteris alpestris Tagawa ex Ching & S.K.Wu; Fl. Xizangica 1: 259 (1983)

= Dryopteris doshunglaensis Ching & S.K.Wu; Enum. Vasc. Pl. Tibet 26 (1981), nom. nud.

species :

Dryopteris alpina Rosenst.; Repert. Spec. Nov. Regni Veg. 12: 173 (1913)

Distribution: Borneo (Sabah: Mt. Kinabalu); Sulawesi; Philippines (Mindanao); Java; New Guinea (Papua New Guinea, Irian Jaya); Bismarck Arch. (New Ireland)

42 BOR JAW NWG PHI SUL 43 BIS

Remarks: Subgen. Nothoperanema, sect. Stenolepia

Synonyms: hidden

= Allantodia scabra Kunze; Bot. Zeitung (Berlin) 192: 192 (1848)

= Alsophila tristis (Blume) Blume; Index Fil. [T. Moore] 58 (1857)

= Aspidium triste Blume; Enum. Pl. Javae 2: 169 (1828)

= Athyrium atropurpureum Copel.; Philipp. J. Sci., C, 12: 59 (1917)

= Athyrium scabrum C.Presl; Abh. Königl. Böhm. Ges. Wiss., ser. 5, 6 [Epim. Bot.]: 67 (1851)

= Ctenitis alpina (Rosenst.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 124 (1947)

= Cyathea tristis (Blume) Domin; Pteridophyta 263 (1929)

= Cystopteris tristis (Blume) Mett.; Ann. Mus. Bot. Lugduno-Batavi 1: 241 (1863)

= Davallia tristis (Blume) Racib.; Pter. Buitenzorg 131 (1898)

= Dryopteris robur Christenh.; Global Fl. 1(4): 40 (2018), nom. nov. for Aspidium triste Blume

= Lastrea tristis (Blume) T.Moore; Index Fil. [T. Moore] 107 (1858)

= Stenolepia tristis (Blume) Alderw.; Bull. Dépt. Agric. Indes Néerl. 27: 46 (1909)

species :

Dryopteris amblyodonta J.P.Roux; Kew Bull. 59(2): 207 (2004)

Distribution: Sierra Leone

22 SIE

species :

Dryopteris amurensis (Milde) Christ; Bull. Acad. Int. Géogr. Bot. 35 (1909), et Mém. 20: 164 (1909)

Distribution: Russian Far East (Amur, Khabarovsk, Primorye, Sakhalin); China (Heilongjiang, Jilin, Liaoning, Nei Mongol); North Korea; ?South Korea; Japan

31 RFE 36 CHI 38 JAP NKO

Synonyms: hidden

= Aspidium spinulosum var. amurense Milde; Fil. Eur. 133 (1867)

= Dryopteris amurensis (Milde) Takeda; Bot. Mag. (Tokyo) 24: 113 (1910)

= Leptorumohra amurensis (Milde) Tzvelev; Novosti Sist. Vyssh. Rast. 26: 6 (1989)

= Nephrodium amurense (Milde) B.Fedtsch.; Trudy Imp. S.-Peterburgsk. Bot. Sada 31: 99 (1912)

species :

Dryopteris anadroma Mitsuta; nom. nud.

Distribution: Japan (Yakushima Isl.)

38 JAP NNS

Conservation Status: VU in Japan

species :

Dryopteris aneitensis (Hook.) C.Chr.; Index Filic. 252 (1905)

Distribution: Vanuatu (Aneityum)

60 VAN

Synonyms: hidden

= Dryopteris layardi (Baker) C.Chr.; Index Filic. 274 (1905)

= Nephrodium layardi Baker; Ann. Bot. (Oxford) 6: 323 (1891)

= Phegopteris aneitensis (Hook.) Kuhn; Verh. K. K. Zool.-Bot. Ges. Wien 19: 577 (1869)

= Polypodium aneitense Hook.; Sp. Fil. [W. J. Hooker] 4: 268 (1862)

species :

Dryopteris angustifrons (Hook.) Kuntze; Revis. Gen. Pl. 2: 812 (1891)

Distribution: China (Yunnan); Myanmar

36 CHI 41 MYA

Remarks: not in India

Synonyms: hidden

= Lastrea angustifrons (Hook.) Bedd.; Ferns Brit. India 2: t. 226 (1867), non (Mett. ex Kunze) T. Moore (1858)

= Nephrodium splendens var. angustifrons Hook.; Sp. Fil. [W. J. Hooker] 4: 126 (1862)

species :

Dryopteris angustipalea Darnaedi, M.Kato & K.Iwats.; J. Jap. Bot. 64: 308 (1989)

Distribution: Moluccas

42 MOL

species :

Dryopteris annamensis (Tagawa) Li Bing Zhang; Taxon 61(6): 1210 (2012), nom. nov.

Distribution: China (Hainan, Yunnan); Tibet; Vietnam

36 CHI TIB 41 VIE

Remarks: subgen. Nothoperanema. Probably to be included in D. aspidioides fide Fraser-Jenkins et al. (2018)

Synonyms: hidden

= Diacalpe annamensis Tagawa; Acta Phytotax. Geobot. 14: 46 (1950)

= Diacalpe caudifolia Ching & S.K.Wu; Fl. Xizangica 1: 194 (1983)

= Peranema annamense (Tagawa) Fraser-Jenk.; Panjab Univ. Res. J., Sci. 59: 18 (2010)

species :

Dryopteris antarctica (Baker) C.Chr.; Index Filic., Suppl. 2 (1906-1912): 29 (1913), et Suppl. III: 81 (1934)

Distribution: D.R.Congo [Zaire]; Rwanda; E-Uganda; Kenya; Tanzania; S-Malawi; E-Zimbabwe; South Africa (W-Cape Prov., E-Cape Prov.); Madagascar; La Réunion; Ile St. Paul; Ile Amsterdam

23 RWA ZAI 25 KEN TAN UGA 26 MLW ZIM 27 SAF 29 MDG REU 91 ASP

Synonyms: hidden

= Aspidium antarcticum (Baker) E.Fourn.; Compt. Rend. Hebd. Séances Acad. Sci. 81: 1140 (1875)

= Dryopteris callolepis C.Chr.; Notizbl. Bot. Gart. Berlin-Dahlem 9: 177 (1924)

= Nephrodium antarcticum Baker; J. Linn. Soc., Bot. 14: 479 (1875)

species :

Dryopteris anthracinisquama Miyam.; J. Jap. Bot. 59(8): 238 (1984)

Distribution: Japan (Kyushu)

38 JAP

Conservation Status: CR in Japan

species :

Dryopteris apiciflora (Wall. ex Mett.) Kuntze; Revis. Gen. Pl. 2: 812 (1891)

Distribution: China (Yunnan); Tibet; Taiwan; Nepal; Bhutan; India (Arunachal Pradesh, Manipur, Nagaland, Sikkim, Uttarakhand, West Bengal); Myanmar (Kachin, etc.)

36 CHI TIB 38 TAI 40 BHU IND NEP 41 MYA

Remarks: Subgen. Dryopsis

Synonyms: hidden

= Aspidium apiciflorum Wall.; Numer. List. [Wallich] no. 345 (1829), nomen

= Aspidium apiciflorum Wall. ex Mett.; Abh. Senckenberg. Naturf. Ges. 2(2) [Pheg. u. Asp.]: 338 (seors. 54) (1858)

= Ctenitis apiciflora (Wall. ex Mett.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 284 (1938)

= Dryopsis apiciflora (Wall. ex Mett.) Holttum & P.J.Edwards; Kew Bull. 41(1): 189 (1986)

= Lastrea apiciflora (Wall. ex Mett.) Bedd.; Ferns Brit. India t. 40 (1865), isonym

= Lastrea apiciflora C.Presl; Tent. Pterid. 76 (1836), nom. nud.

= Lastrea filix-mas subvar. apiciflora (Wall. ex Mett.) Bedd.; Ferns Brit. India 249 (1883)

= Nephrodium apiciflorum (Wall. ex Mett.) Hook.; Sp. Fil. [W. J. Hooker] 4: 112-113, t. 248 (1862)

species :

Dryopteris approximata Sledge; Bull. Brit. Mus. (Nat. Hist.), Bot. 5: 11, f. 1 (1973)

Distribution: S-India (Kerala, Tamil Nadu); Sri Lanka

40 IND SRL

species :

Dryopteris aquilinoides (Desv.) C.Chr.; Index Filic. 252 (1905)

Distribution: New Guinea (Irian Jaya, Papua New Guinea)

42 NWG

Synonyms: hidden

= Aspidium aquilinoides (Desv.) Mett.; Fil. Afr. [Kuhn] 126 (1868)

= Aspidium boryanum Bojer; Hortus Maurit. 391 (1837), non Willd.

= Aspidium grande Fée; Mém. Foug. 5 (Gen. Filic.): 295 (1852)

= Nephrodium aquilinoides Desv.; Mem. Soc. Linn. Paris 6(2): 261 (1827)

= Nephrodium aquilinoides var. angustatum Desv.

species :

Dryopteris arborescens (Baker) Kuntze; Revis. Gen. Pl. 2: 812 (1891)

Distribution: Solomon Isl. (Guadalcanal, Kolombangara, Makira, Vella Lavella); New Caledonia; Vanuatu; American Samoa (Ta´u); [Western] Samoa (Savai´i)

43 SOL 60 ASA NWC SAM VAN

Remarks: the Fijian D. maxima possibly belongs here

Synonyms: hidden

= Aspidium arborescens (Baker) Luerss.; Filic. Graeff. 176 (1871)

= Aspidium ludens (Baker) Christ; Bot. Jahrb. Syst. 28: 352 (1896)

= Aspidium powellii (Baker) Christ; Bot. Jahrb. Syst. 28: 354 (1896)

= Dryopteris ludens (Baker) C.Chr.; Index Filic. 276 (1906)

= Dryopteris powellii (Baker) C.Chr.; Index Filic. 285 (1905)

= Nephrodium arborescens Baker; Syn. Fil. [W. J. Hooker & Baker] 286 (1867)

= Nephrodium ludens Baker; Ann. Bot. (Oxford) 6: 325 (1891)

= Nephrodium powellii Baker; Ann. Bot. (Oxford) 5: 325 (1891)

species :

Dryopteris ardechensis Fraser-Jenk.; Fern Gaz. 12: 184 (1981)

Distribution: ?Spain; France; Corsica; Sardinia; Italy

12 COR FRA SAR 13 ITA

Remarks: tyrrhena agg.

Conservation Status: IUCN R

species :

Dryopteris arguta (Kaulf.) Watt; Canad. Naturalist & Quart. J. Sci., ser. 2, 13: 159 (1867)

Name: Coastal woodfern

Distribution: Canada (Vancouver Isl.); USA (Arizona, California, Oregon, Washington State); Mexico (Baja California Norte)

71 CDN 73 USA 76 USA 79 MEX

Synonyms: hidden

= Aspidium argutum Kaulf.; Enum. Filic. [Kaulfuss] 242 (1824)

= Aspidium rigidum var. argutum (Kaulf.) D.C.Eaton; Rep. U.S. Geogr. Surv., Wheeler 6(Botany): 333 (1878)

= Dryopteris arguta (Kaulf.) Maxon; Amer. Fern J. 11: 3 (1921)

= Dryopteris arguta f. bella Ewan; Amer. Fern J. 34: 112 (1944)

= Dryopteris arguta f. cristata Ewan; Amer. Fern J. 34: 112 (1944)

= Dryopteris arguta f. flocculosa Ewan; Amer. Fern J. 34: 112 (1944)

= Dryopteris arguta f. nudata Ewan; Amer. Fern J. 34: 111 (1944)

= Dryopteris rigida var. arguta (Kaulf.) Underw.; Native Ferns, ed. 4 116 (1893)

= Filix rigida var. arguta (Kaulf.) Farw.; Rep. (Annual) Michigan Acad. Sci. 18: 82 (1916)

= Lastrea arguta (Kaulf.) Brack.; U.S. Expl. Exped., Filic. 16: 196 (1854)

= Nephrodium argutum (Kaulf.) Diels; Nat. Pflanzenfam. [Engler & Prantl] I, 4: 173 (1899)

= Nephrodium rigidum var. americanum Hook.; Brit. Ferns [W. J. Hooker] sub tab. 16 (1861)

= Thelypteris arguta (Kaulf.) Moxley; Bull. South. Calif. Acad. Sci. 19: 57 (1920)

species :

Dryopteris arunachalensis (Fraser-Jenk. & Benniamin) R.Kr.Singh & V.K.Rawat; J. Biodivers. Conservation 8(3): 81 (2024)

Distribution: India (Arunachal Pradesh, ?Meghalaya)

40 IND

Remarks: previously reported as D. paucisora

Synonyms: hidden

= Dryopsis arunachalensis Fraser-Jenk. & Benniamin; Annot. Checkl. Ind. Pterid. 2: 219 (2018)

species :

Dryopteris ascensionis (Hook.) Kuntze; Revis. Gen. Pl. 2: 812 (1891)

Distribution: +Ascension Isl.

28 ASC

Conservation Status: IUCN Ex/E

Synonyms: hidden

= Aspidium ascensionis (Hook.) Kuhn; Filic. Afr. 126 (1868)

= Lastrea ascensionis (Hook.) J.Sm.; Hist. Fil. 215 (1875)

= Nephrodium ascensionis Hook.; Sp. Fil. [W. J. Hooker] 4: 124, t. 257 (1862)

species :

Dryopteris assamensis (C.Hope) C.Chr. & Ching; Bull. Biol. Dept. Sci. Coll. Sun Yatsen Univ. 6: 4 (1933)

Distribution: China (Guangdong, Hongkong, Guizhou, Guangxi); NE-India (Assam, Arunachal Pradesh, Meghalaya, +West Bengal); Myanmar; Thailand; Vietnam

36 CHI 40 IND 41 MYA THA VIE

Conservation Status: IUCN VU for India (Fraser-Jenkins 2012)

Synonyms: hidden

= Dryopteris filix-mas subsp. assamense (C.Hope) C.Chr.; Index Filic. 5: 266 (1905)

= Lastrea filix-mas var. assamensis (C.Hope) Bedd.; Suppl. Ferns Brit. Ind. 57 (1892)

= Nephrodium assamense C.Hope; J. Bot. 28: 326 (1890)

species :

Dryopteris athamantica (Kunze) Kuntze; Revis. Gen. Pl. 2: 812 (1891)

Distribution: Guinea; Sierra Leone; Liberia; Ivory Coast; Ghana; Nigeria; Cameroon; Congo [Brazzaville]; Central African Republic; D.R.Congo [Zaire]; Rwanda; Burundi; Sudan; Ethiopia; Uganda; Kenya; Tanzania; Angola; Zambia; Malawi; Zimbabwe; Mozambique; South Africa (Limpopo, North West, Gauteng, Mpulamanga, Free State, KwaZulu-Natal, E-Cape Prov.); Lesotho; Eswatini [Swaziland]

22 GHA GUI IVO LBR NGA SIE 23 BUR CAF CMN CON RWA ZAI 24 ETH SUD 25 KEN TAN UGA 26 ANG MLW MOZ ZAM ZIM 27 LES SAF SWZ

Synonyms: hidden

= Aspidium athamanticum Kunze; Linnaea 18: 133 (1844)

= Aspidium filix-panna Lucan.; Allg. Med. Central-Zeitung 25: 619 (1856), nom. prov.

= Lastrea athamantica (Kunze) T.Moore; Hooker´s J. Bot. Kew Gard Misc. 5: 311 (1853)

= Lastrea plantii T.Moore; Hooker´s J. Bot. Kew Gard Misc. 5: 227 (1853)

= Nephrodium athamanticum (Kunze) Hook.; Sp. Fil. [W. J. Hooker] 4: 125, t. 258 (1862)

= Nephrodium eurylepium Peter; Repert. Spec. Nov. Regni Veg. Beih. 40(1): 57 (1929)

species :

Dryopteris atrata (Wall) Ching; Sinensia 3: 326 (1933)

Distribution: China (Anhui, Fujian, Gansu, Guangdong, Guangxi, Guizhou, Hainan, Hubei, Hunan, Jiangsu, Jiangxi, Shaanxi, Shandong, Shanxi, Sichuan, Yunnan, Zhejiang); Tibet; Taiwan; Nepal; Bhutan; India (Arunachal Pradesh, Assam State, Kerala, Manipur, Meghalaya, Mizoram, Nagaland, Sikkim, Tamil Nadu, West Bengal); Myanmar (Kachin, Chin, etc.); Thailand; Vietnam

36 CHI TIB 38 TAI 40 BHU IND NEP 41 MYA THA VIE

Remarks: many Himalayan records of Dryopteris atrata refer to D. stenolepis (gamblei) fide Fraser-Jenkins but true D. atrata occurs at least in China, Nepal and N-India

Synonyms: hidden

= Aspidium atratum Wall. ex Kunze; Linnaea 24: 279 (1851)

= Dryopteris hirtipes subsp. atrata (Wall. ex Kunze) Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.), Bot. 18(5): 333 (1989)

= Dryopteris hirtipes var. atrata (Wall. ex Kunze) C.Chr.; Contr. U. S. Natl. Herb. 26(6): 278 (1931)

= Nephrodium atratum (Wall. ex Kunze) Hand.-Mazz.; Symb. Sin. 6: 22 (1929)

species :

Dryopteris aurantiaca J.P.Roux; Syst. Biodivers. 2(2): 164 (2004 publ. 2005)

Distribution: Annobon Isl.

23 ANN

species :

Dryopteris austroindica Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.), Bot. 18(5): 370 (1989), nom. nov.

Distribution: S-India (Tamil Nadu)

40 IND

Conservation Status: IUCN EN for India (Fraser-Jenkins 2012)

Synonyms: hidden

= Lastrea intermedia Bedd.; Ferns S. India t. 113 (1863) non C. Presl (1836)

species :

Dryopteris bamleriana Rosenst.; Repert. Spec. Nov. Regni Veg. 10: 334 (1912)

Distribution: New Guinea (Irian Jaya, Papua New Guinea)

42 NWG

species :

Dryopteris barbigera (T.Moore ex Hook.) Kuntze; Revis. Gen. Pl. 2: 812 (1891)

Distribution: Kazakhstan; Uzbekistan; Kyrgyzstan; Tajikistan; China (Yunnan, Sichuan, Qinghai); Pakistan (Kurram, Swat, Hazara, Gilgit); Nepal; Bhutan; India (Jammu & Kashmir, Himachal Pradesh, Uttarakhand, Sikkim, Arunachal Pradesh, West Bengal); Myanmar

32 KAZ KGZ TZK UZB 36 CHI 40 BHU IND NEP PAK 41 MYA

Synonyms: hidden

= Aspidium barbigerum (T.Moore ex Hook.) Christ; Farnkr. Erde 260 (1897)

= Dryopteris barbigera var. falconeri (T.Moore ex Hook.) R.R.Stewart; Bull. Torrey Bot. Club 72(4): 407 (1945)

= Dryopteris falconeri (T.Moore ex Hook.) Kuntze; Revis. Gen. Pl. 2: 812 (1891)

= Lastrea barbigera (T.Moore ex Hook.) Bedd.; Ferns Brit. India t. 227 (1867)

= Lastrea falconeri (T.Moore ex Hook.) Bedd.; Ferns Brit. India t. 41 (1865)

= Nephrodium barbigerum T.Moore ex Hook.; Sp. Fil. [W. J. Hooker] 4: 113 (1862)

= Nephrodium falconeri T.Moore ex Hook.; Sp. Fil. [W. J. Hooker] 4: 123, t. 254 (1862)

species :

Dryopteris basisora Christ; Notul. Syst. (Paris) 1: 44 (1909)

Distribution: China (Yunnan, Sichuan); Nepal; Bhutan; N-India (Uttarakhand, Arunachal Pradesh)

36 CHI 40 BHU IND NEP

Remarks: D. lachoongensis is included here by Fraser-Jenkins et al. (2018) but kept separate by Knapp & Hsu (2017)

Conservation Status: IUCN VU for India (Fraser-Jenkins 2012)

Synonyms: hidden

= Aspidium pseudovarium Christ; Mém. Soc. Bot. France 1 [Hist. Acrostich.]: 42 (1905)

= Aspidium varium var. obtusum Christ; Mem. Soc. Bot. France 52(Mem. 1): 43 (1905)

= Dryopteris fructuosa var. integriloba Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 446 (1938) (non Dryopteris integriloba Ching 1933)

= Dryopteris megacarpa Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(4): 3 (1984)

= Dryopteris pseudovaria (Christ) C.Chr.; Index Filic. 287 (1905)

= Dryopteris tieluensis Ching & Y.P.Hsu; Fl. Tsinling. 2: 228 (1974) [fide Fraser-Jenkins (1986)]

species :

Dryopteris bernieri Tardieu; Notul. Syst. (Paris) 16: 161 (1956)

Distribution: La Réunion

29 REU

species :

Dryopteris bhutanica (Fraser-Jenk. & Pariyar) Fraser-Jenk.; Biodivers. & Bioprospecting 28 (2023)

Distribution: China (Yunnan); Nepal; Bhutan; India (Arunachal Pradesh, Meghalaya, Uttarakhand)

36 CHI 40 BHU IND NEP

Synonyms: hidden

= Dryopteris wallichiana subsp. bhutanica Fraser-Jenk.; Ferns and Fern-allies of Nepal 1: 28 (2015)

species :

Dryopteris bissetiana (Baker) C.Chr.; Index Filic. 254 (1905)

Distribution: China (Anhui, Fujian, Guizhou, Henan, Hubei, Hunan, Jiangsu, Jiangxi, Shandong, Shaanxi, Shanxi, Sichuan, Yunnan, Zhejiang); North Korea; South Korea (incl. Cheju Isl.); Japan

36 CHI 38 JAP KOR NKO

Remarks: valid name disputed

Synonyms: hidden

= Aspidium setosum (Thunb. ex Murray) Sw.; J. Bot. (Schrader) 1800 [2]: 39 (1800 publ. 1801)

= Dryopteris setosa (Thunb. ex Murray) Akasawa; Bull. Kochi Women´s Univ., Ser. Nat. Sci. 7: 27 (1959), nom. illeg.

= Dryopteris sinobissetiana Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(4): 8 (1984)

= Dryopteris thunbergii Koidz.; Bot. Mag. (Tokyo) 38: 106 (1924)

= Dryopteris varia subsp. setosa (Thunb. ex Murray) Sugim.; Keys Herb. Pl. Jap. Pterid. 405 (1966)

= Dryopteris varia var. setosa (Thunb. ex Murray) Ohwi; Fl. Jap. [Ohwi] 88 (1957)

= Nephrodium bissetianum Baker; J. Bot. 15: 366 (1877)

= Polypodium setosum Thunb.; Syst. Veg., ed. 14 (J. A. Murray): 938 (1784)

= Polystichum bissetianum (Baker) Nakai; Bot. Mag. (Tokyo) 45: 102 (1931)

= Polystichum setosum (Thunb.) C.Presl; Tent. Pterid. 84 (1836), non (Wall.) Schott

= Polystichum thunbergii Koidz.; Bot. Mag. (Tokyo) 38: 106 (1924)

species :

Dryopteris blanfordii (C.Hope) C.Chr.; Index Filic. 254 (1905)

subspecies :

subsp. blanfordii

Distribution: E-Afghanistan; China (W-Sichuan); Tibet; Pakistan (Kurram, Swat, Hazara); Nepal; India (Jammu & Kashmir, Himachal Pradesh, Uttarakhand)

34 AFG 36 CHI TIB 40 IND NEP PAK

Synonyms: hidden

= Lastrea spinulosa Bedd.; Ferns Brit. India 336 (1870) (non C. Presl (1836))

= Lastrea spinulosa var. remota (C.B.Clarke) Bedd.; Ferns Brit. India 252 (1883)

= Nephrodium blanfordii C.Hope; J. Bombay Nat. Hist. Soc. 12: 624, t. 11 (1899)

= Nephrodium remotum C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 523 (1880)

= Dryopteris mehrae Khullar; (nom. nud., fide Fraser-Jenkins 1997)

subspecies :

subsp. nigrosquamosa (Ching) Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.), Bot. 18(5): 388 (1989)

Distribution: China (Gansu, Sichuan, Yunnan); Tibet

36 CHI TIB

Synonyms: hidden

= Dryopteris blanfordii subsp. gongboensis (Ching) Fraser-Jenk.; Pakistan Syst. 5(1-2): 86 (1991) (fide Fraser-Jenkins 1997)

= Dryopteris gongboensis Ching; Fl. Xizangica 1: 269 (1983)

= Dryopteris gushaingensis Ching; Fl. Xizangica 1: 269 (1983)

= Dryopteris nigrosquamosa Ching; Bull. Fan Mem. Inst. Biol. Bot. 2: 194 (1931)

= Dryopteris sublaeta Ching & Y.P.Hsu; Fl. Tsinling. 2: 227 (1974)

species :

Dryopteris bodinieri (Christ) C.Chr.; Index Filic. 254 (1905)

Distribution: China (Guizhou, Yunnan, Sichuan, Hunan)

36 CHI

Synonyms: hidden

= Aspidium bodinieri Christ; Bull. Acad. Int. Géogr. Bot. 248 (1902)

= Nephrodium cyclodioides Baker; Bull. Misc. Inform. Kew 1900: 10 (1900)

species :

Dryopteris bojeri (Baker) Kuntze; Revis. Gen. Pl. 2: 812 (1891)

Distribution: +Mauritius

29 MAU

Synonyms: hidden

= Nephrodium amplum Bory; Voy. Indes Or. [Bélanger] 2: 62 (1833)

= Nephrodium bojeri Baker; Syn. Fil. [W. J. Hooker & Baker] 280 (1867)

species :

Dryopteris borreri (Newman) Oberh. & Tavel; Verh. Schweiz. Naturf. Ges. 118: 153 (1937)

Name: Borrers Wurmfarn (DE); Borrer´s Male-fern (EN)

Distribution: England (England, Wales, Scotland, Outer Hebrides, Orkney Isl., Isle of Man, Isles of Scilly); Ireland (Ireland, Northern Ireland); Denmark; SW-Norway; Netherlands; Belgium; Luxembourg; Germany (Baden-Württemberg, Bayern, Hessen, Niedersachsen, Nordrhein-Westfalen, Rheinland-Pfalz, Saarland, Sachsen, Sachsen-Anhalt, Thüringen); Switzerland; Austria; Czech Republic; Portugal; Spain; Andorra; France; Channel Isl. (Alderney, Guernsey, Jersey); Corsica; Italy (incl. Elba Isl.); Sicily; Slovenia; Croatia; Bosnia & Hercegovina; Montenegro; Serbia; Kosovo; North Macedonia; Romania; Bulgaria; European Turkey; European Russia (Kaliningrad district); Ukraine; ?Crimea; Morocco; Northern Caucasus (Krasnodar, Daghestan, Severo-Ossetia); Georgia [Caucasus] (incl. Adzharia); Azerbaijan; Turkey (Anatolia); NW-Iran

10 DEN GRB IRE NOR 11 AUT BGM CZE GER LUX NET SWI 12 ADO CIS COR FRA POR SPA 13 BIH BUL CRO ITA KOS MNT NMA ROM SIC SLV SRB TUE 14 RUS UKR 20 MOR 33 AZE GEO NCS 34 IRN TUR

Remarks: affinis agg.

Synonyms: hidden

= Aspidium abbreviatum (Lam. & DC.) Poir.; Encycl. [J. Lamarck et al.], Suppl. 4: 516 (1816)

= Aspidium blackwellianum Ten.; Atti Reale Ist. Incorragg. Sci. Nat. Napoli 6: 133 (1830), nom. prov. inval.

= Aspidium filix-mas subvar. ursinum Walther Zimm.; 53 (1916)

= Aspidium filix-mas var. abbreviatum (Lam. & DC.) Lej.; Lej. & Court., Comp. Fl. Belg. 3: 306 (1836)

= Aspidium filix-mas var. blackwellianum Ten.; Atti Reale Ist. Incoragg. Sci. Nat. Napoli 6: 133 (1832)

= Aspidium filix-mas var. stilluppense Sabr.; Oesterr. Bot. Z. 52: 144 (1902)

= Dryopteris abbreviata (Lam. & DC.) Newman ex Manton; Probl. Cytol. Evol. Pterid.: 48 (1950), nom. illeg., p. p. incl. typo, non D. abbreviata (Schrad.) Kuntze (1891))

= Dryopteris affinis subsp. borreri (Newman) Fraser-Jenk.; Willdenowia 10: 110 (1980)

= Dryopteris affinis subsp. stilluppensis (Sabr.) Fraser-Jenk.; Willdenowia 10: 112 (1980)

= Dryopteris affinis var. cristata (T.Moore & Houlston) Fraser-Jenk.; Indian J. Forest. 43(1): 83 (2020 publ. 2021)

= Dryopteris borreri f. aurea Tavel; 154 (1937), nom. inval.

= Dryopteris borreri var. melanothrix Tavel; 153 (1937), nom. inval.

= Dryopteris borreri var. tenuis Tavel; 154 (1937), nom. inval.

= Dryopteris borreri var. ursina (Walther Zimm.) Tavel; 153 (1937)

= Dryopteris filix-mas nothosubsp. tavelii (Rothm.) Schidlay

= Dryopteris filix-mas f. subintegra (Döll) Stefánsson; 5 (1924)

= Dryopteris filix-mas subsp. abbreviata (DC.) Schidlay; Fl. Slovenska 2: 210 (1966) p. p. incl. typo

= Dryopteris filix-mas subsp. borreri (Newman) Bech. & Tavel; Ber. Schweiz. Bot. Ges. 43: 39 (1934)

= Dryopteris filix-mas var. abbreviata (Lam. & DC.) Newman; Hist. Brit. Ferns, ed. III: 192 (1854)

= Dryopteris filix-mas var. borreri Newman; Hist. Brit. Ferns, ed. III: 189 (1854)

= Dryopteris paleacea (T.Moore) Fomin; Kuzn. et al., Fl. Caucas. Crit. 1(1): 52 (1911) (non (Sw.) C. Chr., nec (D. Don) Hand.-Mazz.)

= Dryopteris paleacea var. borreri (Newman) H.Wolff

= Dryopteris stilluppensis (Sabr.) Holub; Preslia 58(4): 302 (1986)

= Dryopteris tavelii f. concinna (Jones) Rothm.; 93 (1946)

= Dryopteris tavelii f. crispa (Sim) Rothm.; 93 (1946)

= Dryopteris tavelii f. cristata (T.Moore & Houlston) Rothm.; 93 (1946)

= Dryopteris tavelii f. pinderi (T.Moore) Rothm.; 93 (1946)

= Dryopteris ×tavelii Rothm.; Candollea 10: 92 (1946) (fide Fraser-Jenkins (2007))

= Lastrea abbreviata (DC.) Woll.; Phytologist n. s. 1: 172 (1855)

= Lastrea filix-mas subsp. borreri (Newman) Hayw.; Bot. Pocket-Book 180 (1872)

= Lastrea filix-mas var. abbreviata (DC.) T.Moore; 138 (1848)

= Lophodium abbreviatum (Lam. & DC.) Newman; Phytologist 4: app. 21 (1851)

= Nephrodium abbreviatum (Lam. & DC.) Lowe; Fern Grow. 6: 161 (1895)

= Nephrodium borreri Rouy; Fl. France [Rouy & Foucaud] 14: 408 (1913)

= Nephrodium filix-mas proles borreri (Newman) Rouy; Fl. France [Rouy & Foucaud] 14: 408 (1913)

= Nephrodium filix-mas f. paleaceum (Lag. ex Sw.) Fiori; Fl. Italia [Fiori, Béguinot & Paoletti] 1: 8 (1896)

= Nephrodium filix-mas subsp. borreri (Newman) P.E.Fourn.; Fl. Compl. Plaine Franç.: 594 (1928)

= Nephrodium filix-mas var. abbreviatum (DC.) Hook.f.; 465 (1870)

= Nephrodium filix-mas var. borreri (Newman) Hook.f.; 465 (1870)

= Polypodium filix-mas var. abbreviatum (Lam. & DC.) Lej.; Rev. Fl. Spa: 211 (1825)

= Polystichum abbreviatum Lam. & DC.; Fl. Franç. [Lam. & A. DC.], éd. 3, 2: 560 (1805)

= Polystichum filix-mas var. abbreviatum (DC.) Gren.; 631 (1856)

= Polystichum induratum Schur; Enum. Pl. Transsilv. 834 (1866)

variant :

var. robusta Oberh. & Tavel; Verh. Schweiz. Naturf. Ges. 118: 153-154 (1937)

Distribution: England (England, Scotland, Wales); C-Europe; Germany; etc.

10 GRB 11 GER

Remarks: as an apomictic taxon it possibly deserves recognition as species, see Flora of Great Britain.

Synonyms: hidden

= Dryopteris affinis subsp. robusta Oberh. & Tavel ex Fraser-Jenk.; Willdenowia 10: 111 (1980)

= Dryopteris affinis var. robusta (Oberh. & Tavel ex Fraser-Jenk.) Fraser-Jenk. & Salvo; Anales Jard. Bot. Madrid 41(1): 195 (1984)

= Dryopteris mediterranea var. robusta (Fraser-Jenk.) Holub; Preslia 58(4): 301 (1986)

= Dryopteris pseudomas subsp. robusta (Oberh. & Tavel ex Fraser-Jenk.) Holub; Folia Geobot. Phytotax. 19(2): 214 (1984)

= Dryopteris robusta (Oberh. & Tavel ex Fraser-Jenk.) P.D.Sell; Fl. Gr. Brit. Ireland 1: 671 (2018)

species :

Dryopteris cacaina Tagawa; Acta Phytotax. Geobot. 6: 155 (1937)

Distribution: Japan (Yakushima Isl.); Taiwan; Philippines

38 JAP NNS TAI 42 PHI

Remarks: sect. Diclisodon, see Zuo et al. 2022

Synonyms: hidden

= Dryopteris ×yakusilvicola Sa.Kurata; J. Geobot. 15: 85 (1967)

species :

Dryopteris cambrensis (Fraser-Jenk.) Beitel & W.R.Buck; Fiddlehead Forum 15(2): 15 (1988)

subspecies :

subsp. cambrensis

Name: Walisischer Wurmfarn (DE); Narrow Male-fern (EN)

Distribution: England (England, Wales, Scotland, Outer Hebrides, Orkney Isl., Isle of Man); Ireland (Ireland, Northern Ireland); Norway; France; Channel Isl. (Guernsey); Croatia; Turkey (Anatolia)

10 GRB IRE NOR 12 CIS FRA 13 CRO 34 TUR

Remarks: affinis agg.

Synonyms: hidden

= Dryopteris affinis subsp. cambrensis Fraser-Jenk.; Sommerfeltia 6: xi (1987)

= Dryopteris affinis var. paleaceocrispa (T.Moore) Fraser-Jenk.; Indian J. Forest. 43(1): 83 (2021)

= Lastrea filix-mas subvar. paleaceocrispa T.Moore; Handb. Brit. Ferns 1: 196 (1859)

subspecies :

subsp. distans (Viv.) Fraser-Jenk.; Fern Gaz. 18(1): 6 (2007)

Distribution: France; Corsica; Italy

12 COR FRA 13 ITA

Synonyms: hidden

= Aspidium distans Viv.; App. ad Fl. Cors. Prod. 8 (1825)

= Dryopteris affinis var. distans (Viv.) Fraser-Jenk.; Sommerfeltia 6: 11 (1987)

= Nephrodium distans (Viv.) Ces., Pass. & Gibelli; Comp. Fl. ltal. [Cesati] 1: p. ? (1870)

= Nephrodium filix-mas var. distans (Viv.) Fiori; Fl. Italia [Fiori, Béguinot & Paoletti] 4(1): 3 (1907)

= Polystichum distans (Viv.) Trevis.; Atti Soc. Ital. Sci. Nat. 17(2): 227 (1874)

subspecies :

subsp. insubrica (Oberh. & Tavel ex Fraser-Jenk.) Fraser-Jenk.; Fern Gaz. 18(1): 6 (2007)

Name: Insubrischer Wurmfarn (DE)

Distribution: Denmark; Norway; Netherlands; Belgium; Luxembourg; Germany (Baden-Württemberg, Bayern, Hessen, Niedersachsen, Rheinland-Pfalz, Saarland, Sachsen, Sachsen-Anhalt, Thüringen); S-Switzerland (Tessin); Austria; S-Poland; Czech Republic; France; Corsica; Sardinia; Italy (incl. Elba Isl.); Sicily; Slovenia; Croatia; Bulgaria

10 DEN NOR 11 AUT BGM CZE GER LUX NET POL SWI 12 COR FRA SAR 13 BUL CRO ITA SIC SLV

Synonyms: hidden

= Dryopteris affinis var. insubrica Oberh. & Tavel ex Fraser-Jenk.; Ann. Bot. Fenn. 33: 73 (1996)

species :

Dryopteris campyloptera (Kunze) Clarkson; Amer. Fern J. 20: 118 (1930)

Name: Mountain woodfern

Distribution: Canada (Labrador, New Brunswick, Newfoundland, Nova Scotia, Prince Edward Isl., Québec); USA (Alabama, Connecticut, Massachusetts, Maryland, Maine, Michigan, North Carolina, New Hampshire, New York, Pennsylvania, Tennessee, Virginia, Vermont, West Virginia)

72 CDN 75 USA 78 USA

Synonyms: hidden

= Aspidium campylopterum Kunze; Amer. J. Sci. Arts, ser. 2, 6: 83, 84 (1848), nom. prov.

= Aspidium spinulosum var. americanum Fisch. ex Kunze; Amer. J. Sci. Arts, Ser. 2, 6: 84 (1848)

= Aspidium spinulosum var. concordianum (Davenp.) Eastman; Ferns & Fern Allies New England 20 (1904)

= Dryopteris austriaca var. concordiana (Davenp.) C.V.Morton; Amer. Fern J. 40: 222 (1950)

= Dryopteris dilatata subsp. americana (Fisch. ex Kunze) Hultén; Kongl. Svenska Vetensk. Acad. Handl. ser. 4, 7(1): 174 (1958)

= Dryopteris dilatata subsp. campyloptera (Clarkson) C.V.Morton; Rep. Bot. Excurs. Boreal Forest Reg. N. Quebec & Ontario: 83 (1962)

= Dryopteris dilatata var. americana (Fisch. ex Kunze) Benedict; Amer. Fern J. 6: 34 (1916)

= Dryopteris intermedia f. concordiana (Davenp.) Clute; Ferns in Their Haunts 184 (1938)

= Dryopteris intermedia var. concordiana (Davenp.) M.Broun; Index No. Amer. Ferns: 71 (1938)

= Dryopteris spinulosa var. americana (Fisch. ex Kunze) Fernald; Rhodora 17: 48 (1915)

= Dryopteris spinulosa var. concordiana (Davenp.) Eastman; Ferns & Fern Allies New England 20 (1904)

= Filix spinulosa var. americana (Fisch. ex Kunze) Farw.; Rep. (Annual) Michigan Acad. Sci. 18: 82 (1916)

= Filix spinulosa var. concordiana (Davenp.) Farw.; Rep. (Annual) Michigan Acad. Sci. 18: 83 (1916)

= Filix-mas rigida var. americana (Fisch. ex Kunze) Farw.; Amer. Midl. Naturalist 12: 256 (1931)

= Filix-mas spinulosa var. americana (Fisch. ex Kunze) Farw.; Amer. Midl. Naturalist 12: 257 (1931)

= Nephrodium spinulosum f. concordianum (Davenp.) Clute; Fern Bull. 12(3): 88 (1904)

= Nephrodium spinulosum var. concordianum Davenp.; Rhodora 6: 33 (1904)

= Thelypteris dilatata var. americana (Fisch. ex Kunze) House; New York State Mus. Bull. 233-234: 69 (1922)

= Thelypteris spinulosa var. americana (Fisch. ex Kunze) Weath.; Rhodora 21: 178 (1919)

= Thelypteris spinulosa var. concordiana (Davenp.) Weath.; Rhodora 21: 178 (1919)

species :

Dryopteris camusiae Fraser-Jenk.; New Species Syndr. Indian Pteridol. 131 (1997)

Distribution: China (Yunnan); NE-India (Nagaland); ?Myanmar

36 CHI 40 IND

Conservation Status: IUCN CR for India (Fraser-Jenkins 2012)

species :

Dryopteris caperata J.P.Roux; Syst. Biodivers. 2(2): 164 (2004 publ. 2005)

Distribution: São Tomé

23 SAO

species :

Dryopteris caroli-hopei Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.), Bot. 18(5): 422 (1989), nom. nov.

Distribution: China (C-Yunnan); Tibet; Nepal; Bhutan; India (Jammu & Kashmir, Himachal Pradesh, Uttarakhand, Assam State, Arunachal Pradesh, Nagaland, Meghalaya, Manipur, Mizoram, Sikkim, West Bengal); Myanmar (Chin, etc.)

36 CHI TIB 40 BHU IND NEP 41 MYA

Synonyms: hidden

= Aspidium dilatatum var. patuloides Christ; Mém. Soc. Bot. France 1 [Hist. Acrostich.]: 41 (1905)

= Dryopteris pseudomarginata Ching; Sporae Pterid. Sin. 327 (1976), nom. nud.

species :

Dryopteris carpatica S.Jess. & Bujnoch; Fern Gaz. 21(8): 462 (2022)

Name: Carpathian Scaly Male Fern

Distribution: Romania

13 ROM

Remarks: affinis agg.

species :

Dryopteris carthusiana (Vill.) H.P.Fuchs; Bull. Soc. Bot. France 105: 339 (1959)

Name: Dorniger Wurmfarn, Gewöhnlicher Dornfarn (DE), Toothed woodfern, Narrow Buckler-fern (EN)

Distribution: England (England, Wales, Scotland, Outer Hebrides, Orkney Isl., Isle of Man, Isles of Scilly); Ireland (Ireland, Northern Ireland); Denmark; Norway; Sweden; Finland; Netherlands; Belgium; Luxembourg; Germany (Brandenburg, Berlin, Baden-Württemberg, Bayern, Hessen, Hamburg, Mecklenburg-Vorpommern, Niedersachsen, Nordrhein-Westfalen, Rheinland-Pfalz, Schleswig-Holstein, Saarland, Sachsen, Sachsen-Anhalt, Thüringen); Switzerland; Liechtenstein; Austria; Poland; Czech Republic; Slovakia; Hungary; Spain; France; Corsica; Italy; Slovenia; Croatia; Bosnia & Hercegovina; Montenegro; Serbia; Kosovo; North Macedonia; Albania; Romania; Bulgaria; ?European Turkey; Estonia; Latvia; Lithuania; Belarus; C-European Russia; E-European Russia; N-European Russia; W-European Russia; Moldova; Ukraine; Crimea; Siberia (Altai, Buryatia, Chita, Irkutsk, Krasnoyarsk, W-Siberia); Kazakhstan; Northern Caucasus; Georgia [Caucasus]; Armenia; Azerbaijan; Turkey (Anatolia); China (Xinjiang); Canada (Alberta, British Columbia, Labrador, Manitoba, New Brunswick, Newfoundland, Nova Scotia, Northern Territories, Nunavut, Ontario, Prince Edward Isl., Québec, Saskatchewan, Yukon); St. Pierre et Miquelon; USA (Alabama, Arkansas, Connecticut, District of Columbia, Delaware, Georgia, Iowa, Idaho, Illinois, Indiana, Kentucky, Massachusetts, Maryland, Maine, Michigan, Minnesota, Missouri, Montana, North Carolina, North Dakota, Nebraska, New Hampshire, New Jersey, New York, Ohio, Oregon, Pennsylvania, Rhode Island, South Carolina, South Dakota, Tennessee, Virginia, Vermont, Washington State, Wisconsin, West Virginia)

10 DEN FIN GRB IRE NOR SWE 11 AUT BGM CZE GER HUN LIE LUX NET POL SLO SWI 12 COR FRA SPA 13 ALB BIH BUL CRO ITA KOS MNT NMA ROM SLV SRB 14 BLR EST KRY LAT LIT MLD RUS UKR 30 SIB 32 KAZ 33 ARM AZE GEO NCS 34 TUR 36 CHI 70 CDN 71 CDN 72 CDN SPM 73 USA 74 USA 75 USA 78 USA

Synonyms: hidden

= Aspidium callipteris F.H.Wilms; Verh. Rhein. Ges. 9: 577 (1852), nom. illeg., non Fée

= Aspidium groenlandicum Gand.; Bull. Soc. Bot. France 66: 305 (1920)

= Aspidium spinulosum (O.F.Müll.) Sw.; J. Bot. (Schrader) 1800(2): 38 (1801), nom. illeg.

= Aspidium spinulosum f. anadenium B.L.Rob.; Rhodora 9: 84 (1907)

= Aspidium spinulosum var. elevatum A.Braun ex Döll; Rhein. Fl. 18 (1843)

= Aspidium spinulosum var. exaltatum Lasch; Verh. Bot. Vereins Prov. Brandenburg 2: 79 (1860)

= Aspidium spinulosum var. subcoriaceum Rupr.; Distr. Crypt. Vasc. Ross. 37 (1845)

= Aspidium spinulosum var. tanacetifolium (Hoffm.) Rupr.; Distr. Crypt. Vasc. Ross. 37 (1845)

= Dryopteris aristata Charit.; Dopoln. Fl. i Rastitel´n. Yuga Zapadnoi Sibiri 89 (2020) (tentative synonym)

= Dryopteris austriaca subsp. spinulosa (O.F.Müll.) Schinz & Thell.; Vierteljahrsschr. Naturf. Ges. Zürich 60: 340 (1915)

= Dryopteris austriaca var. spinulosa (O.F.Müll.) Fiori; Filic. Equiset. Lycopod. 115 (1943)

= Dryopteris lanceolatocristata (Hoffm.) Alston; Watsonia 4: 41 (1957)

= Dryopteris spinulosa (O.F.Müll.) Kuntze; Revis. Gen. Pl. 2: 810 (1891), isonym

= Dryopteris spinulosa var. dumetorum Underw.; Native Ferns, ed. 4: 117 (1893)

= Dryopteris spinulosa var. elevata (A.Braun ex Döll) Vollm.; Fl. Bayern: p. ? (1914)

= Dryopteris spinulosa var. exaltata (Lasch) Vollm.; Fl. Bayern: p. ? (1914)

= Dryopteris spinulosa var. glandulosa Gilbert; Fern Bull. 8: 11 (1900)

= Dryopteris spinulosa var. patuloides Christ; Mém. Soc. Bot. France 1 [Hist. Acrostich.]: 41 (1905)

= Filix spinulosa (O.F.Müll.) Farw.; Rep. (Annual) Michigan Acad. Sci. 18: 81-83 (1916)

= Filix spinulosa var. intermedia Farw.; Rep. (Annual) Michigan Acad. Sci. 18: 83 (1916)

= Filix-mas spinulosa (O.F.Müll.) Farw.; Amer. Midl. Naturalist 12: 234-251 (1931)

= Lastrea dilatata var. linearis Bab.; 386 (1843)

= Lastrea spinosa (Schrank) Newman; Natur. Alm. 21 (1844)

= Lastrea spinulosa (O.F.Müll.) C.Presl; Tent. Pterid. 76 (1836)

= Lastrea spinulosa var. dilatata G.Lawson; Fern Fl. Canada 240 (1889)

= Lastrea spinulosa var. elevata (A.Braun ex Döll) Syme & N.E.Br.; Engl. Bot., 3rd ed., vol. XII: 78 (1886)

= Lastrea spinulosa var. exaltata (Lasch) Syme & N.E.Br.; Engl. Bot., 3rd ed., vol. XII: 78 (1886)

= Lophodium spinosum (Weiss) Newman; Phytologist 4: 371 (1851)

= Nephrodium dilatatum proles tanacetifolium (Hoffm.) Rouy; Fl. France [Rouy & Foucaud] 14: 413 (1913)

= Nephrodium dilatatum subsp. tanacetifolium (Hoffm.) P.E.Fourn.; Fl. Compl. Plaine Franç.: 595 (1928)

= Nephrodium jordanii Rouy; Fl. France [Rouy & Foucaud] 14: 411 (1913)

= Nephrodium spinulosum (O.F.Müll.) Strempel; Fil. Berol. Syn.: 30 (1822) (nom. illeg.)

= Nephrodium spinulosum f. anadenium (B.L.Rob.) Clute; Fern Bull. 17: 20 (1909)

= Nephrodium spinulosum var. bipinnatum Hook.; Brit. Ferns [W. J. Hooker] tab. 18 (1861)

= Nephrodium spinulosum var. elevatum (A.Br.) Rouy; Fl. France [Rouy & Foucaud] 14: 411 (1913)

= Nephrodium spinulosum var. exaltatum (Lasch) Rouy; Fl. France [Rouy & Foucaud] 14: 411 (1913)

= Nephrodium spinulosum var. humile Rouy; Fl. France [Rouy & Foucaud] 14: 411 (1913)

= Nephrodium spinulosum var. swarzii F.W.Schultz; 566 (1845), nom. inval.

= Polypodium carthusianum Vill.; Hist. Pl. Dauphiné [Villars] 1: 292 (1786)

= Polypodium filix-femina var. spinosa F.G.Weiss; Pl. Crypt. Fl. Gött. 316 (1770)

= Polypodium lanceolatocristatum Hoffm.; Bot. Mag. (Römer & Usteri) 3(9): 9 (1790)

= Polypodium spinosum Schrank; Baier. Fl. 2: 424 (1789)

= Polypodium tanacetifolium Hoffm.; Deutschl. Fl., Theil 2 (Hoffm.) 2: 8 (1795)

= Polypodium ×spinulosum O.F.Müll.; Fl. Dan. [Oeder] 12: 7 (1777) (non Burm.f.)

= Polystichum angustatum Schumach.; Enum. Pl. [Schumacher] 2: 22 (1803)

= Polystichum carthusianum (Vill.) Sanio; Verh. Bot. Vereins Prov. Brandenburg 25: 84 (1884)

= Polystichum coniifolium Schumach.; Enum. Pl. [Schumacher] 2: 22 (1803)

= Polystichum muelleri Schumach.; Enum. Pl. [Schumacher] 2: 20 (1803), as ´mülleri´

= Polystichum pseudocristatum Schur; Verh. Mitth. Siebenbürg. Vereins Naturwiss. Hermannstadt 3: 94 (1852)

= Polystichum spinosum (F.G.Weiss) Roth; Tent. Fl. Germ. 3(1.1): 91 (1799)

= Polystichum spinulosum (O.F.Müll.) Lam. & DC.; Fl. Franç. [Lam. & A. DC.], éd. 3, 2: 561 (1805)

= Polystichum spinulosum var. tanacetifolium P.E.Fourn.; Quatre Fl. France: 17 (1934)

= Thelypteris spinulosa (O.F.Müll.) Nieuwl.; Midl. Naturalist (Notre Dame) 1: 226 (1910)

variant :

var. enigmatica Harmaja; Memoranda Soc. Fauna Fl. Fenn. 88: 23 (2012)

Distribution: Finland

10 FIN

species :

Dryopteris caucasica (A.Braun) Fraser-Jenk. & Corley; Brit. Fern Gaz. 10: 222 (1972)

Distribution: European Turkey; Moldova; Crimea; Northern Caucasus; Georgia [Caucasus]; Armenia; ?Azerbaijan; Turkey (Anatolia); NW-Iran

13 TUE 14 KRY MLD 33 ARM GEO NCS 34 IRN TUR

Remarks: filix-mas agg.

Synonyms: hidden

= Aspidium affine Fisch. & C.A.Mey.; Bull. Soc. Imp. Naturalistes Moscou 11: 240 (1838), nom. illeg., non Blume (1828)

= Aspidium caucasicum A.Braun; Flora 24: 707 (1841)

= Dryopteris affinis (Newman) Kinahan; Phytologist 5: 197 (1854 publ. 1856), nom. illeg.

= Dryopteris filix-mas subsp. caucasica (A.Braun) Askerov; Zametki Sist. Geogr. Rast. 39: 6 (1983)

= Dryopteris filix-mas var. affinis (Fisch. & C.A.Mey.) Askerov; Zametki Sist. Geogr. Rast. 39: 7 (1983)

= Dryopteris filix-mas var. affinis (Fisch. & C.A.Mey.) B.Fedtsch.; 9 (1927), nom. illeg.

= Dryopteris filix-mas var. affinis Newman; Hist. Brit. Ferns, ed. III: 187 (1854), nom. nov. for Aspidium affine Fisch. & C. A. Mey.

= Dryopteris filix-mas var. athyriformis Fomin; Vestn. Tiflissk. Bot. Sada 20: 35 (1911)

= Lastrea filix-mas var. incisa T.Moore; Phytologist 3: 137 (1850)

= Polystichum affine Ledeb.; Fl. Ross. (Ledeb.) 4(14): 515 (1853), nom. illeg., non Wall. ex C. Presl

species :

Dryopteris caudipinna Nakai; Bot. Mag. (Tokyo) 45: 96 (1931)

Distribution: Japan

38 JAP

Synonyms: hidden

= Dryopteris erythrosora var. caudipinna (Nakai) H.Itô; (1939)

species :

Dryopteris celsa (W.Palmer) Knowlt., T.S.Palmer & Pollard ex Small; Proc. Biol. Soc. Wash. 13: 202 (1900)

Name: Log fern

Distribution: USA (Alabama, Arkansas, District of Columbia, Delaware, Georgia, Illinois, Indiana, Kentucky, Louisiana, Maryland, Michigan, Missouri, North Carolina, New Jersey, New York, Pennsylvania, South Carolina, Tennessee, Virginia, West Virginia)

74 USA 75 USA 78 USA

Synonyms: hidden

= Aspidium goldieanum f. celsum (W.Palmer) Clute; Ferns in Their Haunts 313 (1901)

= Aspidium goldieanum var. celsum (W.Palmer) B.L.Rob.; Rhodora 10: 30 (1908)

= Dryopteris clintoniana f. atropalustris (Small) Wherry; Amer. Fern J. 40: 120 (1950)

= Dryopteris clintoniana f. celsa (W.Palmer) Wherry; Amer. Fern J. 40: 119 (1950)

= Dryopteris clintoniana var. atropalustris (Small) C.F.Reed; Bull. Nat. Hist. Soc. Maryland 13: 51 (1943)

= Dryopteris goldieana f. celsa (W.Palmer) Clute; Ferns in Their Haunts 172, 385 (1938)

= Dryopteris goldieana subsp. celsa W.Palmer; Proc. Biol. Soc. Wash. 13: 65 (1899)

= Dryopteris wherryi Crane; Amer. Fern J. 50: 271, t. 23(6) (1960)

= Dryopteris ×atropalustris Small; Ferns S. E. States [Small] 274 cum tab. (1938)

= Filix goldieana var. celsa (W.Palmer) Farw.; Rep. (Annual) Michigan Acad. Sci. 18: 82 (1917)

= Nephrodium goldieanum var. celsum (W.Palmer) Waters; Ferns (Waters) 235 (1903)

species :

Dryopteris chaerophyllifolia (Zippel) C.Chr.; Index Filic. 257 (1905)

Distribution: Sumatra; Borneo (Mt. Kinabalu, Sabah); Java; Vanuatu

42 BOR JAW SUM 60 VAN

Synonyms: hidden

= Aspidium chaerophyllifolium Zippel; Mett., Ann. Mus. Bot. Lugduno-Batavi 1: 228 (1864)

= Phegopteris heterolepia Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 16: 23 (1914) [synonymy fide Backer 1939]

species :

Dryopteris championii (Benth.) C.Chr. ex Ching; Sinensia 8: 327 (1933)

Distribution: China (Fujian, Guangdong, Guangxi, Guizhou, Henan, Hubei, Hunan, Jiangsu, Jiangxi, Macao, Shandong, Sichuan, Yunnan, Zhejiang); South Korea (incl. Cheju Isl.); Japan; Taiwan

36 CHI 38 JAP KOR TAI

Synonyms: hidden

= Aspidium championii Benth.; Fl. Hongkong [Bentham] 456 (1861)

= Aspidium erythrosorum var. amoyense Christ; Warb., Monsunia 1: 80 (1900)

= Dryopteris bullatipaleacea Ching; Bull. Bot. Res., Harbin 2(2): 65-66, pl. 2, f. 1 (1982)

= Dryopteris changii Ching; Wuyi Sci. J. 1: 7 (1981)

= Dryopteris conferta Ching & P.S.Chiu; Bot. Res. Acad. Sin. 1987(2): 17 (1987) (nom. illeg.)

= Dryopteris erythrosora subsp. chiliolepis Hand.-Mazz.; Anz. Akad. Wiss. Wien, Math.-Naturwiss. Kl. no. 7 (1922)

= Dryopteris erythrosora var. cavaleriei Rosenst.; Repert. Spec. Nov. Regni Veg. 13(355-358): 131-132 (1914)

= Dryopteris grandiosa Ching & P.S.Chiu; Bot. Res. Acad. Sin. 1987(2): 8 (1987)

= Dryopteris huanglungensis Ching; Bot. Res. Acad. Sin. 1987(2): 10 (1987)

= Dryopteris infrahirtella Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 3(4): 22 (1983)

= Dryopteris jinfoshanensis Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 6(1): 179 (1986), nom. nov.

= Dryopteris laoshanensis J.X.Li & S.T.Ma; Bull. Bot. Res., Harbin 3(4): 139 (1983)

= Dryopteris lepigera C.Chr.; Index Filic. 274 (1905) (non (Baker) Kuntze)

= Dryopteris linganensis Ching & C.F.Zhang; Bull. Bot. Res., Harbin 3(3): 23 (1983)

= Dryopteris nanchuanensis Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(3): 19 (1984) (non 3(4) (1983))

= Dryopteris neofuscipes Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(3): 22 (1984)

= Dryopteris paleifera Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(4): 2 (1984)

= Dryopteris pseudoerythrosora Kodama; Matsum., Icon. Pl. Koisikav. 1: 165, t. 83 (1913)

= Dryopteris qinyuanensis Ching & P.S.Chiu; Bot. Res. Acad. Sin. 1987(2): 7 (1987)

= Dryopteris wangii Ching; Bot. Res. Acad. Sin. 1987(2): 13 (1987)

= Dryopteris wuyishanensis Ching; Fl. Fujianica 1: 600 (1982)

= Dryopteris yandongensis Ching & C.F.Zhang; Bull. Bot. Res., Harbin 3(3): 13 (1983)

= Dryopteris zunyiensis Ching; Bot. Res. Acad. Sin. 1987(2): 18 (1987)

species :

Dryopteris chinensis (Baker) Koidz.; Fl. Symb. Orient.-Asiat. 39 (1930)

Distribution: Russian Far East (Primorye); China (Anhui, Henan, Jiangsu, Jiangxi, Liaoning, Shandong, Zhejiang); North Korea; South Korea (incl. Cheju Isl.); Japan

31 RFE 36 CHI 38 JAP KOR NKO

Synonyms: hidden

= Aspidium forbesii Hance; J. Bot. 13: 198 (1875)

= Nephrodium chinense Baker; Syn. Fil. [W. J. Hooker & Baker] 278 (1867)

species :

Dryopteris christenseniae (Ching) Li Bing Zhang; Taxon 61(6): 1209 (2012), nom. nov.

Distribution: China (Yunnan)

36 CHI

Remarks: subgen. Nothoperanema

Synonyms: hidden

= Diacalpe christenseniae Ching; Bull. Fan Mem. Inst. Biol. Bot., ser. 2, 1: 291 (1949), as ´christensenae´

species :

Dryopteris chrysocoma (Christ) C.Chr.; Index Filic. 257 (1905)

Distribution: China (Guizhou, Sichuan, Yunnan); Tibet; Pakistan (Hazara); Nepal; Bhutan; India (Jammu & Kashmir, Himachal Pradesh, Arunachal Pradesh, Meghalaya, Mizoram, Sikkim, Uttarakhand, West Bengal); Myanmar; Vietnam; Philippines

36 CHI TIB 40 BHU IND NEP PAK 41 MYA VIE 42 PHI

Synonyms: hidden

= Aspidium chrysocoma (Christ) Christ; Bull. Acad. Int. Géogr., sér. 3, 11: 253 (1902)

= Aspidium filix-mas var. chrysocoma Christ; Bull. Herb. Boissier, sér. 1, 6: 966 (1898)

= Dryopteris alpicola Ching & Z.R.Wang; Acta Phytotax. Sin. 23(5): 349 (1985), nom. nov. for D. chrysocoma var. alpina Ching 1937

= Dryopteris chrysocoma var. alpina Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 438 (1937), not D. alpina Rosenst. (1913) (fide Fraser-Jenkins et al. 2015)

= Dryopteris chrysocoma var. major Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 438 (1938)

= Dryopteris chrysocoma var. squamosa (C.Chr.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 438 (1938)

= Dryopteris cochleata var. squamosa C.Chr.; Acta Horti Gothob. 1: 59 (1924)

= Dryopteris daliensis Z.R.Wang ex Fraser-Jenk.; New Species Syndr. Indian Pteridol. 119 (1997) (pro hybr.)

= Dryopteris gracilofrons C.Chr.; Index Filic. 268 (1905) (nom. nud., fide Fraser-Jenkins (1997))

= Dryopteris macrocarpa Stewart; Bull. Torrey Bot. Club 72: 406 (1945) (fide Fraser-Jenkins 1997)

= Dryopteris parachrysocoma Ching & Z.R.Wang; Acta Phytotax. Sin. 23(5): 344 (1985) (fide Fraser-Jenkins 1997)

= Dryopteris pseudochrysocoma Ching; Acta Phytotax. Sin. 23(5): 349 (1985), nom. nud.

= Dryopteris ×daliensis Z.R.Wang; Acta Phytotax. Sin. 23(5): 346 (1985)

= Lastrea filix-mas var. elongata Bedd.; Ferns Brit. India 250 (1883) (non Dryopteris elongata (Wall. ex Hook.) Kuntze, fide Fraser-Jenkins (1997))

= Nephrodium chrysocoma (Christ) Hand.-Mazz.; Symb. Sin. 6: 21-25 (1929)

species :

Dryopteris cicatricata J.P.Roux; Syst. Biodivers. 2(2): 167 (2004 publ. 2005)

Distribution: São Tomé

23 SAO

species :

Dryopteris cinnamomea (Cav.) C.Chr.; Amer. Fern J. 1: 95 (1911)

Name: Cinnamon woodfern

Distribution: USA (Arizona, SW-Texas); Mexico (Aguascalientes, Chiapas, Chihuahua, Coahuila, Colima, Ciudad de Mexico, Durango, Guanajuato, Guerrero, Hidalgo, Jalisco, Mexico State, Michoacan, Morelos, Nayarit, Oaxaca, Puebla, Queretaro, San Luis Potosi, Sinaloa, Sonora, Tamaulipas, Tlaxcala, Veracruz, Zacatecas)

76 USA 77 USA 79 MEX

Synonyms: hidden

= Aspidium agatholepis Fée; Mém. Foug. 8 (Icon. Esp. Nouv.): 106 (1857), as ´agatolepis´

= Aspidium athyrioides M.Martens & Galeotti; Nouv. Mém. Acad. Roy. Sci. Bruxelles 15 (Mém. Lycopod.): 67, t. 18 (1842)

= Aspidium flaccidum E.Fourn.; Bull. Soc. Bot. France 27: 328 (1880) (non Blume (1828))

= Aspidium indecorum (Liebm.) E.Fourn.; Mexic. Pl. 1: 97 (1872)

= Aspidium inquinans (T.Moore) Fée; Mém. Foug. 8 (Icon. Esp. Nouv.): 106 (1867)

= Aspidium mexicanum (C.Presl) Kunze; Linnaea 13: 147 (1839)

= Aspidium mexicanum var. acutilobum Kunze; Linnaea 13: 147 (1839)

= Aspidium mexicanum var. obtusilobum Kunze; Linnaea 13: 147 (1839)

= Aspidium roseum E.Fourn.; Mexic. Pl. 1: 97 (1872)

= Aspidium sphaerocarpon (Fée) Christ; Bull. Herb. Boissier, sér. 2, 6: 726 (1905)

= Athyrium sphaerocarpon Fée; Mém. Foug. 5 (Gen. Filic.): 186 (1852)

= Dryopteris athyrioides (M.Martens & Galeotti) Kuntze; Revis. Gen. Pl. 2: 811 (1891)

= Dryopteris fournieri (Baker) C.Chr.; Index Filic. 266 (1905)

= Dryopteris glandulifera (Liebm.) C.Chr.; Index Filic. 267 (1905)

= Dryopteris indecora (Liebm.) C.Chr.; Index Filic. 272 (1905)

= Dryopteris indecora var. obtusa Hieron.; Hedwigia 46: 346 (1907)

= Dryopteris mexicana (C.Presl) C.Chr.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 7, 10(2): 68 (1913)

= Dryopteris mexicana (C.Presl) Kuntze; Revis. Gen. Pl. 2: 811 (1891)

= Dryopteris patula var. mexicana (C.Presl) Bonap.; Notes Ptéridol. 2: 130 (1915)

= Dryopteris rosea (E.Fourn.) Mickel & Beitel; Mem. New York Bot. Gard. 46 [Pterid. Fl. Oaxaca]: 165 (1988)

= Lastrea agatholepis (Fée) T.Moore; Index Fil. [T. Moore] 85 (1858)

= Lastrea athyrioides (M.Martens & Galeotti) Liebm.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 5, 1: 274 (1849)

= Lastrea indecora Liebm.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 5, 1: 272 (seors. 120) (1849)

= Lastrea inquinans T.Moore; Index Fil. [T. Moore] 94 (1858)

= Lastrea mexicana (C.Presl) C.Presl; Tent. Pterid. 76 (1836)

= Lophodium glanduliferum (Liebm.) Newman; Phytologist 4: 371 (1851)

= Nephrodium agatholepis (Fée) Kiaersk.; Cat. Fil. hb. hort. Haun. 62 (1874)

= Nephrodium fournieri Baker; Ann. Bot. (Oxford) 6: 317 (1891) (non p. 328)

= Nephrodium glanduliferum (Liebm.) Kiaersk.; Cat. Fil. hb. hort. Haun. 63 (1874)

= Nephrodium indecorum (Liebm.) Kiaersk.; Cat. Fil. hb. hort. Haun. 63 (1874)

= Nephrodium mexicanum C.Presl; Reliq. Haenk. 1: 38 (1825), nom. illeg., non auct.

= Nephrodium sphaerocarpon Hook.; Sp. Fil. [W. J. Hooker] 4: 139 (1862)

= Nephrodium sphaerocarpon var. glandulosum Hook.; Sp. Fil. [W. J. Hooker] 4: 139 (1862)

= Phegopteris glanduligera (Liebm.) E.Fourn.; Mexic. Pl. 1: 90 (1872) (err.)

= Polypodium glanduliferum Liebm.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 5, 1: 206 (1849)

= Polystichum cystopteroides Nees; Linnaea 19: 685 (1847)

= Tectaria cinnamomea Cav.; Descr. Pl. (Cavanilles) 252 (1801)

species :

Dryopteris clarkei (Baker) Kuntze; Revis. Gen. Pl. 2: 812 (1891)

Distribution: China (Sichuan, Guizhou, Yunnan, Guangxi); Tibet; Nepal; Bhutan; India (Arunachal Pradesh, West Bengal, Sikkim); Myanmar (Kachin, etc.)

36 CHI TIB 40 BHU IND NEP 41 MYA

Remarks: Subgen. Dryopsis

Synonyms: hidden

= Ctenitis clarkei (Baker) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 287 (1938)

= Dryopsis clarkei (Baker) Holttum & P.J.Edwards; Kew Bull. 41(1): 181 (1986)

= Dryopteris fibrillosa Hand.-Mazz.; Anz. Akad. Wiss. Wien, Math.-Naturwiss. Kl. no. 7: 2 (1922) (non Dryopteris fibrillosa (Baker) C. Chr.)

= Lastrea filix-mas subvar. clarkei (Baker) Bedd.; Ferns Brit. India 250 (1883)

= Lastrea filix-mas var. clarkei (Baker) Bedd.; Suppl. Ferns Brit. Ind. 17 (1876)

= Nephrodium clarkei Baker; Syn. Fil. [W. J. Hooker & Baker] 497 (1874)

species :

Dryopteris clintoniana (D.C.Eaton) Dowell; Proc. Staten Island Assoc. Arts 1: 64 (1906)

Name: Clinton´s woodfern

Distribution: Canada (New Brunswick, Ontario, Québec); USA (Connecticut, District of Columbia, Delaware, Illinois, Indiana, Massachusetts, Maine, Michigan, New Hampshire, New Jersey, New York, Ohio, Pennsylvania, Rhode Island, Vermont, Wisconsin)

72 CDN 74 USA 75 USA 78 USA

Synonyms: hidden

= Aspidium cristatum var. clintonianum D.C.Eaton; Manual (Gray), ed. 5: 665 (1867)

= Dryopteris cristata var. clintoniana (D.C.Eaton) Underw.; Native Ferns, ed. 4 115 (1893)

= Dryopteris ×poyseri Wherry; Amer. Fern J. 50: 90 (1960)

= Filix cristata var. clintoniana (D.C.Eaton) Farw.; Rep. (Annual) Michigan Acad. Sci. 18: 81 (1916)

= Filix-mas cristata var. clintoniana (D.C.Eaton) Farw.; Amer. Midl. Naturalist 12: 253 (1931)

= Nephrodium cristatum var. clintonianum (D.C.Eaton) Gilbert; Working List N. Amer. Pterid.: 17 (1901)

= Thelypteris clintoniana (D.C.Eaton) House; New York State Mus. Bull. 233-234: 69 (1920)

= Thelypteris cristata var. clintoniana (D.C.Eaton) Weath.; Rhodora 21: 177 (1919)

species :

Dryopteris cochleata (D.Don) C.Chr.; Index Filic. 258 (1905)

Distribution: China (Yunnan, Guizhou, Sichuan); Pakistan (Swat); Nepal; Bhutan; India (Jammu & Kashmir, Andhra Pradesh, Arunachal Pradesh, Assam State, Bihar, Chhattisgarh, Himachal Pradesh, Jharkhand, Karnataka, Kerala, Madhya Pradesh, Maharashtra, Manipur, Meghalaya, Mizoram, Nagaland, Sikkim, Tamil Nadu, Tripura, Uttarakhand, Uttar Pradesh, West Bengal); Bangladesh; Myanmar (Chin, etc.); Thailand; Vietnam; peninsular Malaysia; Philippines; Java; Lesser Sunda Isl. (Bali)

36 CHI 40 BAN BHU IND NEP PAK 41 MYA THA VIE 42 JAW LSI MLY PHI

Synonyms: hidden

= Arthrobotrys avara Wall.; Numer. List. [Wallich] no. 1034 (1829), nomen

= Arthrobotrys macrocarpus Wall.; Numer. List. [Wallich] no. 395 (1829), nomen

= Aspidium cochleatum (D.Don) Christ; Bull. Herb. Boissier, sér. 1, 6: 967 (1898), isonym

= Aspidium cochleatum (D.Don) Spreng.; Syst. Veg., ed. 16 [Sprengel] 4(1): 106 (1827)

= Aspidium erythrosorum var. souliei Christ; Mém. Soc. Bot. France 1 [Hist. Acrostich.]: 40 (1905)

= Aspidium filix-mas var. cochleatum (D.Don) Christ; Farnkr. Erde 257 (1897)

= Dryopteris filix-mas var. cochleata (D.Don) Alderw.; Malayan Ferns 193 (1909)

= Dryopteris heleopteroides Christ; Philipp. J. Sci. 2: 212 (1907)

= Lastrea cochleata (D.Don) T.Moore; Index Fil. [T. Moore] 88 (1857)

= Lastrea filix-mas var. cochleata (D.Don) Bedd.; Ferns Brit. India 250 (1883)

= Lastrea macrocarpa (Wall.) C.Presl; Tent. Pterid. 77 (1836)

= Nephrodium cochleatum D.Don; Prodr. Fl. Nepal. 6 (1825)

= Nephrodium filix-mas var. cochleatum (D.Don) Hook.; Sp. Fil. [W. J. Hooker] 4: 117 (1862)

species :

Dryopteris cognata (C.Presl) Kuntze; Revis. Gen. Pl. 2: 812 (1891)

Name: Large Kidney Fern (EN)

Distribution: St. Helena

28 STH

Conservation Status: IUCN E

Synonyms: hidden

= Aspidium cognatum (C.Presl) Fée; Mém. Foug. 5 (Gen. Filic.): 292 (1852), nomen, ex Mett., Pheg. u. Asp. 61, no. 145 (1858)

= Dryopteris fuliginosa C.Chr.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 8, 6: 120, f. 29 (1920)

= Lastrea cognata C.Presl; Abh. Königl. Böhm. Ges. Wiss., ser. 5, 6 [Epim. Bot.]: 40 (1851)

= Nephrodium cognatum (C.Presl) Hook.; Sp. Fil. [W. J. Hooker] 4: 123, t. 256 (1862)

species :

Dryopteris commixta Tagawa; Acta Phytotax. Geobot. 2(3): 190 (1933)

Distribution: China (Fujian, Guangxi, Jiangxi, Sichuan, Zhejiang); Japan

36 CHI 38 JAP

Synonyms: hidden

= Dryopteris shangqianensis Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(3): 20 (1984)

= Dryopteris sinodickinsii Ching ex K.H.Shing & J.F.Cheng; Jiangxi Sci. 8(3): 46 (1990)

species :

Dryopteris comorensis (Tardieu) Fraser-Jenk.; Thaiszia 4(1): 34 (1994)

Distribution: Madagascar; Comores (Grande Comore)

29 COM MDG

Synonyms: hidden

= Dryopteris inaequalis var. comorensis Tardieu; Notul. Syst. (Paris) 15(2): 162 (1956)

species :

Dryopteris conjugata Ching; Bull. Fan Mem. Inst. Biol. Bot. 11: 63 (1941)

Distribution: China (Yunnan); Nepal; Bhutan; India (Himachal Pradesh, Uttarakhand, Arunachal Pradesh, Meghalaya); Myanmar

36 CHI 40 BHU IND NEP 41 MYA

Synonyms: hidden

= Dryopteris subconjugata S.G.Lu; Guihaia 11(3): 224 (1991)

species :

Dryopteris conversa Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 20: 13 (1915)

Distribution: Sumatra

42 SUM

species :

Dryopteris corleyi Fraser-Jenk.; Bol. Soc. Brot., ser. 2, 55: 248 (1982)

Distribution: Spain (N-Spain: Asturias)

12 SPA

Conservation Status: IUCN I

species :

Dryopteris costalisora Tagawa; Acta Phytotax. Geobot. 3: 88 (1934)

Distribution: Taiwan; E-Nepal; India (West Bengal, ?Arunachal Pradesh, Sikkim)

38 TAI 40 IND NEP

Remarks: bona sp. fide Fraser-Jenkins (1997) and Knapp (Ferns of Taiwan)

Conservation Status: IUCN EN for India (Fraser-Jenkins 2012)

Synonyms: hidden

= Dryopteris chrysocoma var. costalisora (Tagawa) W.C.Shieh; J. Sci. Engin. (Natl. Chung-Hsing Univ.) 2: 306 (1975)

species :

Dryopteris crassirhizoma Nakai; Cat. Sem. Hort. Bot. Univ. Imp. Tokyo 32 (1920)

Distribution: Russian Far East (Amur, Khabarovsk, Kuril Isl., Primorye, Sakhalin); China (Hebei, Heilongjiang, Jilin, Liaoning); North Korea; South Korea (incl. Cheju Isl.); Japan

31 RFE 36 CHI 38 JAP KOR NKO

Synonyms: hidden

= Aspidium filix-mas var. maackii Milde; Fil. Eur. 122 (1867)

= Dryopteris buschiana Fomin; Fl. Sibir. Orient. Extremi 5: 52, fig. (1930)

= Dryopteris buschiana var. maackii (Milde) Fomin; Fl. Sibir. Orient. Extremi 5: 54 (1930)

= Dryopteris crassirhizoma var. maackii (Milde) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 426 (1938)

species :

Dryopteris crinalis (Hook. & Arn.) C.Chr.; Index Filic. 269 (1905)

Distribution: Hawaii (Kauai, Oahu, Molokai, Maui, Hawaii Isl.)

63 HAW

Conservation Status: IUCN R

Synonyms: hidden

= Polypodium crinale Hook. & Arn.; Bot. Beechey Voy. 105 (1832)

= Phegopteris crinalis (Hook. & Arn.) Mann; Proc. Amer. Acad. Arts 7: 218 (1868)

variant :

var. podosora (W.H.Wagner) D.D.Palmer; Amer. Fern J. 92(2): 100 (2002)

Distribution: Hawaii (Kauai)

63 HAW

Synonyms: hidden

= Dryopteris podosora W.H.Wagner & Flynn; Contr. Univ. Michigan Herb. 20: 252 (1995)

species :

Dryopteris crispifolia Rasbach, Reichst. & G.Vida; Bot. J. Linn. Soc. 74: 270 (1977)

Distribution: Azores (Sao Miguel Isl., Pico, Faial)

21 AZO

species :

Dryopteris cristata (L.) A.Gray; Manual (Gray) 631 (1848)

Name: Kamm-Wurmfarn, Kammfarn (DE), Crested Buckler-fern (EN)

Distribution: England (E-England, +NE-England); Denmark; Norway; Sweden; Finland; Netherlands; Belgium; Germany (Brandenburg, Berlin, Baden-Württemberg, Bayern, Hessen, Hamburg, Mecklenburg-Vorpommern, Niedersachsen, Nordrhein-Westfalen, Rheinland-Pfalz, Schleswig-Holstein, Saarland, Sachsen, Sachsen-Anhalt, Thüringen); Switzerland; Liechtenstein; Austria; Poland; Czech Republic; Slovakia; Hungary; France; Italy; Slovenia; Croatia; Serbia; Kosovo; Romania; Estonia; Latvia; Lithuania; Belarus; C-European Russia; E-European Russia; N-European Russia; W-European Russia; Ukraine; Siberia (Altai, Krasnoyarsk, W-Siberia); Kazakhstan; Canada (Alberta, British Columbia, Manitoba, New Brunswick, Newfoundland, Nova Scotia, Ontario, Prince Edward Isl., Québec, Saskatchewan); St. Pierre et Miquelon; USA (Alabama, Connecticut, District of Columbia, Delaware, Georgia, Iowa, Idaho, Illinois, Indiana, Massachusetts, Maryland, Maine, Michigan, Minnesota, Missouri, Montana, North Carolina, North Dakota, Nebraska, New Hampshire, New Jersey, New York, Ohio, Pennsylvania, Rhode Island, Tennessee, Virginia, Vermont, Washington State, Wisconsin, West Virginia)

10 DEN FIN GRB NOR SWE 11 AUT BGM CZE GER HUN LIE NET POL SLO SWI 12 FRA 13 CRO ITA KOS ROM SLV SRB 14 BLR EST LAT LIT RUS UKR 30 SIB 32 KAZ 71 CDN 72 CDN SPM 73 USA 74 USA 75 USA 78 USA

Synonyms: hidden

= Acrostichum callipteris (Ehrh.) Ehrh.; Beitr. Naturk. [Ehrhart] 3: 77 (1788)

= Aspidium cristatum (L.) Sw.; J. Bot. (Schrader) 1800(2): 37 (1801)

= Aspidium cristatum var. crenatum Christ; 145 (1900)

= Aspidium lancastriense Spreng.; Anleit. Kenntn. Gew. 3: 134 (1804)

= Dryopteris cristata var. lancastriensis (Spreng.) Tidestr.; Elys. Marian., Ferns 1: 44, pl. 6 (1906)

= Dryopteris cristata var. mariana Tidestr.; Elys. Marian., Ferns 1: 44, pl. 6 (1906)

= Filix cristata (L.) Farw.; Rep. (Annual) Michigan Acad. Sci. 18: 81-83 (1916)

= Filix-mas cristata (L.) Farw.; Amer. Midl. Naturalist 12: 234-251 (1931)

= Lastrea callipteris (Ehrh.) Newman; Hist. Brit. Ferns, ed. II: 12 (1844)

= Lastrea cristata (L.) C.Presl; Tent. Pterid. 77 (1836)

= Lastrea lancastriensis (Spreng.) Houlston; Gard. Mag. Bot. 3: 318 (1851)

= Lastrea lancastriensis (Spreng.) J.Sm.; Cat. Ferns Kew 6 (1856), isonym

= Lophodium callipteris (Ehrh.) Newman; Phytologist 4: 421 (1851)

= Nephrodium cristatum (L.) Michx.; Fl. Bor.-Amer. (Michaux) 2: 269 (1803)

= Nephrodium cristatum f. lancastriense (Spreng.) Gilbert; List N. Amer. Pterid.: 17 (1901)

= Nephrodium cristatum f. silvaticum Poyser; Fern Bull. 16: 13 (1908)

= Nephrodium lancastriense (Spreng.) Desv.; Mém. Soc. Linn. Paris 6(3): 260 (1827)

= Polypodium callipteris Ehrh.; Hannover. Mag. 8-9: 127, 138 (1784)

= Polypodium cristatum L.; Sp. Pl. 2: 1090 (1753)

= Polystichum callipteris (Ehrh.) Bernh.; J. Bot. (Schrader) 1799(1): 307 (1799)

= Polystichum cristatum (L.) Roth; Tent. Fl. Germ. 3: 84 (1799)

= Tectaria cristata (L.) Lag., Garcia & Clem.; Anales Ci. Nat. 5: 147 (1802)

= Thelypteris cristata (L.) Nieuwl.; Midl. Naturalist (Notre Dame) 1: 226 (1910)

species :

Dryopteris cycadina (Franch. & Sav.) C.Chr.; Index Filic. 260 (1905)

Distribution: China (Fujian, Guangxi, Guizhou, Hubei, Hunan, Jiangxi, Sichuan, Yunnan, Zhejiang); South Korea; Japan; Taiwan

36 CHI 38 JAP KOR TAI

Synonyms: hidden

= Aspidium cycadinum Franch. & Sav.; Enum. Pl. Jap. 2: 236, 630 (1877)

= Dryopteris cycadina var. melanolepis Nakai; Bot. Mag. (Tokyo) 45: 99 (-100) (1931)

= Dryopteris fengyangshanensis Ching & C.F.Zhang; Bull. Bot. Res., Harbin 3(3): 33 (1983)

= Dryopteris longirostrata Ching ex K.H.Shing & J.F.Cheng; Jiangxi Sci. 8(3): 47 (1990)

= Dryopteris pseudatrata Ching; Fl. Fujianica 1: 601 (1982)

= Dryopteris rigidiuscula Ching ex K.H.Shing & J.F.Cheng; Jiangxi Sci. 8(3): 47 (1990)

species :

Dryopteris cyclopeltidiformis C.Chr.; Dansk Bot. Ark. 9(3): 60, t. 6(3-4) (1937)

Distribution: China (Hainan); Vietnam

36 CHI 41 VIE

species :

Dryopteris damingshanensis Li Bing Zhang & H.M.Liu; Novon 23(1): 119 (2014)

Distribution: China (Guangxi)

36 CHI

species :

Dryopteris daozhenensis P.S.Wang & X.Y.Wang; Pterid. Fl. Guizhou 310 (2001)

Distribution: China (Guizhou)

36 CHI

Remarks: provisionally accepted name, status uncertain

species :

Dryopteris decipiens (Hook.) Kuntze; Revis. Gen. Pl. 2: 812 (1891)

Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Jiangxi, Hunan, Sichuan, Zhejiang); Japan; Ryukyu Isl.; Taiwan

36 CHI 38 JAP NNS TAI

Remarks: relationship to D. integripinnula unclear, both are frequently combined or treated as subspecies

Synonyms: hidden

= Aspidium decipiens (Hook.) Luerss.; Bot. Jahrb. Syst. 4: 360 (1883)

= Dryopteris cordipinna Ching & K.H.Shing; Fl. Fujianica 1: 599 (1982)

= Dryopteris decipiens var. subbipinnata Ching; Wuyi Sci. J. 1: 8 (1981)

= Dryopteris metafuscipes Ching & C.F.Zhang; Bull. Bot. Res., Harbin 3(3): 19 (1983)

= Dryopteris mimetica Ching & C.F.Zhang; Bull. Bot. Res., Harbin 3(3): 21 (1983)

= Dryopteris retrorsopaleacea Ching & C.F.Zhang; Bull. Bot. Res., Harbin 3(3): 20 (1983)

= Nephrodium decipiens Hook.; Sp. Fil. [W. J. Hooker] 4: 86, t. 243 (1862)

variant :

var. diplazioides (Christ) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 476 (1938)

Distribution: China (Anhui, Fujian, Guizhou, Jiangsu, Jiangxi, Sichuan, Zhejiang); South Korea; Japan; Taiwan

36 CHI 38 JAP KOR TAI

Remarks: sometimes treated as full species, but included in nominate variety by Knapp (Ferns of Taiwan)

Synonyms: hidden

= Dryopteris fuscipes var. diplazioides (Christ) Ching; Fl. Jiangsuensis 1: 66 (1977), nom. nud. ex L. K. Ling, Q. Q. Zhang & Y. R. Hwang (1991)

= Dryopteris integripinnula Ching; Wuyi Sci. J. 1: 7 (1981)

= Polystichum diplazioides Christ; Bull. Acad. Int. Géogr., sér. 3, 11: 260 (1902)

species :

Dryopteris dehuaensis Ching & K.H.Shing; Fl. Fujianica 1: 601 (1982)

Distribution: China (Fujian, Jiangxi, Zhejiang)

36 CHI

Synonyms: hidden

= Dryopteris gushanica Ching & K.H.Shing; Fl. Fujianica 1: 601 (1982)

= Dryopteris neosordidipes Ching ex K.H.Shing & J.F.Cheng; Jiangxi Sci. 8(3): 48 (1990)

species :

Dryopteris deparioides (T.Moore) Kuntze; Revis. Gen. Pl. 2: 812 (1891)

subspecies :

subsp. deparioides

Distribution: +S-India (+Tamil Nadu); Sri Lanka

40 IND SRL

Remarks: sect. Diclisodon

Conservation Status: IUCN CR or EX for India (Fraser-Jenkins 2012)

Synonyms: hidden

= Aspidium deparioides (T.Moore) Hook.; Fil. Exot.: t. 3 (1858)

= Diclisodon deparioides T.Moore; Index Fil. [T. Moore] 96 (1857)

= Lastrea deparioides (T.Moore) Bedd.; Ferns S. India 104 (1865)

= Nephrodium deparioides (T.Moore) Hook.; Fil. Exot.: t. 53, t. 60 (1858), nom. illeg.

subspecies :

subsp. ambigua (Sledge) Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.), Bot. 18(5): 447 (1989)

Distribution: Sri Lanka

40 SRL

Synonyms: hidden

= Dryopteris ambigua Sledge; Bull. Brit. Mus. (Nat. Hist.), Bot. 5: 23 (1973)

subspecies :

subsp. concinna [Bedd.] C.Chr.; Index Filic., Suppl. Tert.: 84 (1934)

Distribution: Sri Lanka

40 SRL

Synonyms: hidden

= Aspidium concinnum Thwaites; Enum. Pl. Zeyl. [Thwaites]: 392 (1864) (non Link (1833))

= Dryopteris emigrans Copel.; Univ. Calif. Publ. Bot. 12: 392 (1931)

= Dryopteris thwaitesii (Baker) Kuntze; Revis. Gen. Pl. 2: 813 (1891)

= Lastrea concinna Bedd.; Ferns Brit. India t. 207 (1864) (non L. concinna (Willd.) T. Moore)

= Lastrea thwaitesii (Baker) Bedd.; Suppl. Ferns S. Ind. 18 (1876)

= Nephrodium thwaitesii Baker; Syn. Fil. [W. J. Hooker & Baker] 277 (1867)

species :

Dryopteris devriesei Rosenst.; Meded. Rijks-Herb. no. 31: 1 (1917)

Distribution: Java

42 JAW

Remarks: provisionally accepted name

species :

Dryopteris diacalpioides (Ching) Li Bing Zhang; Taxon 61(6): 1206 (2012)

Distribution: China (S-Yunnan)

36 CHI

Synonyms: hidden

= Nothoperanema diacalpioides Ching; Acta Phytotax. Sin. 11: 28 (1966)

species :

Dryopteris dickinsii (Franch. & Sav.) C.Chr.; Index Filic. 262 (1905)

Distribution: China (Anhui, Fujian, Guangxi, Guizhou, Hubei, Jiangxi, Sichuan, Yunnan, Zhejiang); South Korea (Cheju Isl.); Japan; Taiwan; Pakistan (Azad Kashmir); NW-India (Jammu & Kashmir)

36 CHI 38 JAP KOR TAI 40 IND PAK

Conservation Status: IUCN CR or EX for India (Fraser-Jenkins 2012)

Synonyms: hidden

= Aspidium dickinsii Franch. & Sav.; Enum. Pl. Jap. 2: 236 (1877)

= Aspidium thibeticum Franch.; Nouv. Arch. Mus. Hist. Nat., sér. 2, 10: 118 (1887)

= Dryopteris basiaurita Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 405 (1938)

= Dryopteris hirtipes var. japonica Nakai; Bot. Mag. (Tokyo) 45: 100 (1931)

= Dryopteris okushirensis Miyabe & Kudô; Trans. Sapporo Nat. Hist. Soc. 7: 23 (1918)

= Dryopteris thibetica (Franch.) C.Chr.; Index Filic. 298 (1905) (fide Fraser-Jenkins et al. 2018)

= Dryopteris yungtzeensis Ching; Bull. Fan Mem. Inst. Biol. Bot. 11: 60 (1941) (fide Fraser-Jenkins et al. 2018)

= Lastrea thibetica (Franch.) Bedd.; Suppl. Ferns Brit. Ind. 52 (1892)

= Nephrodium dickinsii (Franch. & Sav.) Baker; Hooker´s Icon. Pl. t. 1651 (1886)

= Nephrodium tibeticum (Franch.) Baker; Ann. Bot. (Oxford) 6: 318 (1891)

= Thelypteris thibetica (Franch.) B.K.Nayar & Kaur; Companion Ferns Brit. India 57 (1974)

species :

Dryopteris dicksonioides (Mett.) Copel.; Bull. Bernice P. Bishop Mus. 93: 35 (1932)

Distribution: Society Isl. (Tahiti, Raiatea, Moorea, Huahine)

61 SCI

Synonyms: hidden

= Phegopteris dicksonioides Mett.; Kuhn, Linnaea 36: 118 (1869)

= Thelypteris dicksonioides (Mett.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 251 (1941)

species :

Dryopteris diffracta (Baker) C.Chr.; Index Filic. 262 (1905)

Distribution: China (Guangxi, Guizhou, Hainan, Yunnan); Tibet; Taiwan; NE-India (Arunachal Pradesh); Myanmar (Kachin, etc.); N-Thailand; Vietnam

36 CHI TIB 38 TAI 40 IND 41 MYA THA VIE

Conservation Status: IUCN CR for India (Fraser-Jenkins 2012)

Synonyms: hidden

= Acrorumohra diffracta (Baker) H.Itô; Nov. Fl. Jap. [Nakai & Honda] 4: 101 (1939)

= Arachniodes diffracta (Baker) Ching; Acta Bot. Sin. 10: 257 (1962)

= Aspidium diffractum (Baker) Christ; Bull. Herb. Boissier, sér. 1, 7: 17 (1899)

= Dryopteris reflexipinna Hayata; Icon. Pl. Formosan. 4: 174, f. 113 (1914)

= Nephrodium diffractum Baker; Bull. Misc. Inform. Kew 1898: 230 (1898)

= Polystichum diffractum (Baker) Masam.; Short Fl. Formosa 26 (1936)

= Rumohra diffracta (Baker) Ching; Sinensia 6: 69, t. 18 (1934)

species :

Dryopteris dilatata (Hoffm.) A.Gray; Manual (Gray) 631 (1848)

subspecies :

subsp. dilatata

Name: Breitblättriger Wurmfarn, Breitblättriger Dornfarn (DE); Broad Buckler-fern (EN)

Distribution: Faeroe Isl.; England (England, Wales, Scotland, Outer Hebrides, Orkney Isl., Shetland Isl., Isle of Man, St. Kilda Isl., Iles of Scilly); Ireland (Ireland, Northern Ireland); Denmark; Norway; Sweden; Finland; Netherlands; Belgium; Luxembourg; Germany (Brandenburg, Berlin, Baden-Württemberg, Bayern, Hessen, Hamburg, Mecklenburg-Vorpommern, Niedersachsen, Nordrhein-Westfalen, Rheinland-Pfalz, Schleswig-Holstein, Saarland, Sachsen, Sachsen-Anhalt, Thüringen); Switzerland; Liechtenstein; Austria; Poland; Czech Republic; Slovakia; Hungary; Portugal; Spain; France; Channel Isl. (Alderney, Guernsey, Jersey, Sark); Corsica; Italy; Slovenia; Croatia; Bosnia & Hercegovina; Montenegro; Serbia; Kosovo; North Macedonia; Romania; Bulgaria; Greece (mountains in N-Greece); Estonia; Latvia; Lithuania; Belarus; C-European Russia; N-European Russia; W-European Russia; Moldova; Ukraine; Azores (Pico); Siberia (Altai); Northern Caucasus; Georgia [Caucasus]; Armenia; Azerbaijan; Turkey (Anatolia); Mongolia; Japan; New Zealand [I]; Argentina [I] (Tierra del Fuego [I]); Falkland Isl. [I]

10 DEN FIN FOR GRB IRE NOR SWE 11 AUT BGM CZE GER HUN LIE LUX NET POL SLO SWI 12 CIS COR FRA POR SPA 13 BIH BUL CRO GRC ITA KOS MNT NMA ROM SLV SRB 14 BLR EST LAT LIT MLD RUS UKR 21 AZO 30 SIB 33 ARM AZE GEO NCS 34 TUR 37 MON 38 JAP 51 nzl 85 arg 91 fal

Remarks: carthusiana agg.

Synonyms: hidden

= Aspidium dilatatum (Hoffm.) Sm.; Fl. Brit. 3: 1125 (1804)

= Aspidium dilatatum var. deltoideum (Milde) Borbás; Math. Term. Közlem. 14: 449 (1877)

= Aspidium dilatatum var. majus Gray; Nat. Arr. Brit. Pl. 2: 8 (1821 publ. 1822)

= Aspidium dilatatum var. minus Gray; Nat. Arr. Brit. Pl. 2: 8 (1821 publ. 1822)

= Aspidium dilatatum var. oblongum (Milde) Borbás; Math. Term. Közlem. 14: 449 (1877)

= Aspidium distans var. chanteriae (T.Moore) Milde; Fil. Eur. 139 (1867)

= Aspidium distans var. oblongum Milde; Sporenpfl. 57 (1865)

= Aspidium drepanum Schkuhr; 24. Kl. Linn. Pfl.-Syst. 1: t. 47 (1806)

= Aspidium dumetorum Sm.; Engl. Fl. 4: 281 (1828)

= Aspidium spinulosum f. deltoideum (Milde) Zapal.; Conspic. Fl. Gallic. Crit. 1: 79 (1905)

= Aspidium spinulosum f. oblongum (Milde) Zapal.; Conspic. Fl. Gallic. Crit. 1: 79 (1905)

= Aspidium spinulosum f. recurvatum (Lasch) Asch.; 926 (1864)

= Aspidium spinulosum subsp. dilatatum (Hoffm.) Celak.; Prodr. Fl. Böhmen 10 (1867)

= Aspidium spinulosum subsp. recurvatum Lasch; 80 (1861)

= Aspidium spinulosum var. deltoideum Milde; Sporenpfl. 57 (1865)

= Aspidium spinulosum var. dilatatum (Hoffm.) Hampe; 63 (1836)

= Aspidium spinulosum var. dilatatum (Hoffm.) Link; Fil. Spec. 106 (1841), isonym

= Aspidium spinulosum var. muticum A.Braun ex Döll; Rhein. Fl. 18 (1843)

= Aspidium spinulosum var. oblongum Milde; Fil. Eur. 137 (1867)

= Aspidium tanacetifolium (Lam. & DC.) Opiz; Kratos 2: 10 (1820)

= Dryopteris alexeenkoana Fomin; Kuzn. et al., Fl. Caucas. Crit. 1(1): 67 (1911), et Vestn. Tiflissk. Bot. Sada 20: 46 (1911)

= Dryopteris aristata (Vill.) Druce; List of Brit. Plants 87 (1908)

= Dryopteris austriaca (Jacq.) Woyn.; Vierteljahrsschr. Naturf. Ges. Zürich 60: 339 (1915), nom. illeg.

= Dryopteris austriaca f. deltoidea (Milde) Tavel; 73 (1918)

= Dryopteris austriaca f. oblonga (Milde) Tavel; 74 (1918)

= Dryopteris austriaca subsp. dilatata (Hoffm.) Schinz & Thell.; Vierteljahrsschr. Naturf. Ges. Zürich 60: 340 (1915), nom. illeg.

= Dryopteris austriaca var. dilatata Hoffm.; (1943)

= Dryopteris carthusiana subsp. dilatata (Hoffm.) O.Bolòs, Vigo, Masalles & Ninot; Fl. Man. Paísos Catalans: 1213 (1990)

= Dryopteris dilatata f. deltoideum (Milde) Diklic & V.Nikolic; Fl. SR Srbije 10: 10 (1986)

= Dryopteris dilatata f. oblongum (Milde) Diklic & V.Nikolic; Fl. SR Srbije 10: 10 (1986)

= Dryopteris dilatata subsp. alexeenkoana (Fomin) Askerov; Zametki Sist. Geogr. Rast. 39: 6 (1983)

= Dryopteris dilatata subsp. remotiloba (Fomin) Askerov; Zametki Sist. Geogr. Rast. 39: 6 (1983)

= Dryopteris dilatata var. remotiloba Fomin

= Dryopteris spinulosa subsp. austriaca Bergdolt; 29 (1936)

= Dryopteris spinulosa subsp. dilatata (Hoffm.) C.Chr.; Index Filic. 293 (1905)

= Dryopteris spinulosa subsp. dilatata (Hoffm.) Piper & Beattie; Fl. N.W. Coast [Piper & Beattie]: 7 (1915), isonym

= Dryopteris spinulosa var. dilatata (Hoffm.) Underw.; Native Ferns, ed. 4: 116 (1893)

= Filix spinulosa var. dilatata (Hoffm.) Farw.; Rep. (Annual) Michigan Acad. Sci. 6: 209 (1904)

= Lastrea aristata (Vill.) Britton & Rendle; List Brit. Pl. 39 (1907)

= Lastrea collina Newman; Hist. Brit. Ferns, ed. II: 224 (1844)

= Lastrea dilatata (Hoffm.) C.Presl; Tent. Pterid. 77 (1836)

= Lastrea dilatata var. chanteriae T.Moore; Ferns Gr. Brit. (Moore): 24 (1855)

= Lastrea glandulosa (Newman) Newman; Phytologist 4: 258 (1851)

= Lastrea lepidota T.Moore; Handb. Brit. Ferns, ed. 3: 130 (1857)

= Lastrea maculata Deakin; Fl. Brit. 4: 110 (1848)

= Lastrea multiflora (Roth) Newman; Hist. Brit. Ferns, ed. II: 215 (1844)

= Lophodium collinum (Newman) Newman; Phytologist 4: app. 18 (1851)

= Lophodium glandulosum Newman; Phytologist 4: app. 18 (1851)

= Lophodium multiflorum (Roth) Newman; Phytologist 4: 371 (1851)

= Nephrodium austriacum (Jacq.) Fritsch; Mitt. Naturwiss. Vereines Steiermark 46 (1908)

= Nephrodium dilatatum (Hoffm.) Desv.; Mem. Soc. Linn. Paris 6(2): 261 (1827)

= Nephrodium dilatatum var. chanteriae (T.Moore) Rouy; Fl. France [Rouy & Foucaud] 14: 412 (1913)

= Nephrodium dilatatum var. dumetorum (Sm.) Rouy; Fl. France [Rouy & Foucaud] 14: 412 (1913)

= Nephrodium dilatatum var. oblongum (Milde) Rouy; Fl. France [Rouy & Foucaud] 14: 412 (1913)

= Nephrodium spinulosum subsp. dilatatum (Hoffm.) Arcang.; Comp. Fl. Ital. [Arcangeli] 811 (1882), isonym

= Nephrodium spinulosum subsp. dilatatum (Hoffm.) Hook.f.; Student. Fl. Brit. Isl.: 466 (1870)

= Nephrodium spinulosum var. dilatatum (Hoffm.) F.W.Schultz; 566 (1845)

= Nephrodium spinulosum var. dilatatum (Hoffm.) Hook.; Brit. Ferns: t. 19 (1861)

= Nephrodium spinulosum var. lepidotum (T.Moore) Baker; Syn. Fil. [W. J. Hooker & Baker] 276 (1867)

= Nephrodium tanacetifolium (Lam. & DC.) Rouy; Fl. France [Rouy & Foucaud] 14: 413 (1913)

= Polypodium aristatum Vill.; Hist. Pl. Dauphiné [Villars] 3: 844 (1789) (non G. Forst. (1786))

= Polypodium cristatum Hoffm.; Deutschl. Fl. 2: 8 (1796), nom. illeg.

= Polypodium dilatatum Hoffm.; Deutschl. Fl. 2: 7 (1796)

= Polypodium filix-femina var. cristatum Weiss; 317 (1770)

= Polypodium lanceolata-cristatum Hoffm.; Bot. Mag. (Römer & Usteri) 9: 9 (1790)

= Polypodium multiflorum Raeusch.; Nomencl. Bot. [Raeuschel], ed. 3: 311 (1797)

= Polypodium multiflorum Roth; Catal. Bot. 1: 135 (1797)

= Polypodium spinulosum var. dilatatum (Hoffm.) Wahlenb.; Fl. Suecica 2: 673 (1826)

= Polystichum dilatatum (Hoffm.) Schumach.; Enum. Pl. [Schumacher] 2: 24 (1803)

= Polystichum multiflorum (Roth) Roth; Tent. Fl. Germ. 3: 87 (1799)

= Polystichum spinulosum subsp. dilatatum (Hoffm.) Corb.; Nouv. Fl. Normandie: 676 (1894), isonym

= Polystichum spinulosum subsp. dilatatum (Hoffm.) F.Aresch.; 552 (1881)

= Polystichum spinulosum subsp. dilatatum (Hoffm.) P.E.Fourn.; Quatre Fl. France: 17 (1934), isonym

= Polystichum spinulosum var. dilatatum (Hoffm.) W.D.J.Koch; Syn. Fl. Germ. Helv., ed. 2, 3: 979 (1845)

= Polystichum spinulosum var. muticum (A.Braun ex Döll) Schur; 168 (1851)

= Polystichum tanacetifolium Lam. & DC.; Fl. Franç. [Lam. & A. DC.], éd. 3, 2: 562 (1805)

= Thelypteris dilatata (Hoffm.) House; New York State Mus. Bull. 233-234: 69 (1920)

= Thelypteris spinulosa var. dilatata (Hoffm.) H.St.John; Prelim. List Pl. Kaniksu Natl. Forest 1: 1 (1925)

subspecies :

subsp. lusitanica Fraser-Jenk., Andreis & Vasco Silva; Indian J. Forest. 43(1): 82 (2020), et Indian J. Forest. 43(1): 82 (2021)

Distribution: Portugal; Spain; France

12 FRA POR SPA

species :

Dryopteris dracomontana Schelpe & N.C.Anthony; Contr. Bolus Herb. 10: 147 (1982)

Distribution: South Africa (KwaZulu-Natal, E-Cape Prov., northeastern W-Cape Prov., Free State); Lesotho

27 LES SAF

Synonyms: hidden

= Aspidium inaequale var. montanum Kunze; Linnaea 10: 549 (1836)

= Dryopteris pentheri var. montana (Kunze) Alston; Bol. Soc. Brot., ser. 2, 30: 14 (1956)

species :

Dryopteris drakei C.Chr.; Index Filic. 262 (1906)

Distribution: Society Isl. (Tahiti)

61 SCI

Synonyms: hidden

= Polypodium vescoi Drake; Fl. Polynésie Franç. [Drake] 307 (1893) (non Nephrodium vescoi Drake (1893))

species :

Dryopteris dulongensis (S.K.Wu & X.Cheng) Zhang; Phytotaxa 71: 17-27 (2012)

Distribution: China (Yunnan)

36 CHI

Remarks: Possibly a hybrid D. apiciflora x heterolaena. Subgen. Dryopsis

Synonyms: hidden

= Dryopsis ×dulongensis (S.K.Wu & X.Cheng) S.Y.Dong; Acta Bot. Yunnan. 23: 187 (2001)

= Ctenitis dulongensis S.K.Wu & X.Cheng; Acta Phytotax. Sin. 23: 402 (1985)

species :

Dryopteris edwardsii Fraser-Jenk.; New Species Syndr. Indian Pteridol. 139 (1997)

Distribution: Tibet; Taiwan; Pakistan; Nepal; India (Uttarakhand, Himachal Pradesh, Jammu & Kashmir)

36 TIB 38 TAI 40 IND NEP PAK

Remarks: not mentioned in Flora of China but recorded for Taiwan by Knapp (Ferns of Taiwan, 2011 ff.). Records for Bhutan were a misidentification of D. yigongensis

species :

Dryopteris enneaphylla (Baker) C.Chr.; Index Filic. 263 (1905)

Distribution: China (Zhejiang, Hubei); Taiwan

36 CHI 38 TAI

Synonyms: hidden

= Nephrodium enneaphyllum Baker; J. Bot. 25: 170 (1887)

species :

Dryopteris erythrosora (D.C.Eaton) Kuntze; Revis. Gen. Pl. 2: 812 (1891)

Name: Autumn fern

Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hubei, Hunan, Jiangsu, Jiangxi, Sichuan, Yunnan, Zhejiang); South Korea (incl. Cheju Isl.); Japan; Ryukyu Isl.

36 CHI 38 JAP KOR NNS

Synonyms: hidden

= Aspidium erythrosorum D.C.Eaton; Narr. Exped. China Japan [Perry] 2: 330 (1856)

= Aspidium filix-mas var. erythrosorum (D.C.Eaton) Christ; Farnkr. Erde 257 (1897)

= Aspidium prolificum Maxim. ex Franch. & Sav.; Enum. Pl. Jap. 2: 239, 632 (1877)

= Dryopteris bulligera Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 483 (1938)

= Dryopteris distantipinna Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(3): 27, f. 27 (1984)

= Dryopteris erythrosora (D.C.Eaton) C.Chr.; Index Filic. 260 (1905)

= Dryopteris erythrosora f. viridisora (Nakai ex H.Itô) H.Itô; Nov. Fl. Jap. [Nakai & Honda] 4: 41 (1939)

= Dryopteris erythrosora var. dilatata (Koidz.) Sugim.; (1962)

= Dryopteris erythrosora var. viridisora Nakai ex H.Itô; Bot. Mag. (Tokyo) 50: 69 (1936)

= Dryopteris linyingensis Ching & C.F.Zhang; Bull. Bot. Res., Harbin 3(3): 30 (1983)

= Dryopteris oblongipinnula Ching & P.S.Chiu; Bot. Res. Acad. Sin. 1987(2): 5 (1987)

= Dryopteris oppositipinna Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(3): 25 (1984), nom. illeg., non (Hayata) Hayata (1911)

= Dryopteris paraerythrosora Ching & C.F.Zhang; Bull. Bot. Res., Harbin 3: 27 (1983)

= Dryopteris pseudoerythrosora Ching & C.F.Zhang; Bull. Bot. Res., Harbin 3(3): 27 (1983), nom. illeg., non Kodama 1913

= Dryopteris remotipinnula Ching & C.F.Zhang; Bull. Bot. Res., Harbin 3(3): 15 (1983), non Bonap. 1925

= Dryopteris sinoerythrosora Ching & K.H.Shing; Fl. Fujianica 1: 600 (1982)

= Dryopteris squamistipes Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(4): 7 (1984)

= Lastrea erythrosora (D.C.Eaton) T.Moore; Index Fil. [T. Moore] 91 (1858)

= Lastrea prolifica (Maxim.) T.Moore; Gard. Chron., ser. 2, 18: 744 (1882)

= Nephrodium erythrosorum (D.C.Eaton) Hook.; Sp. Fil. [W. J. Hooker] 4: 120, t. 253 (1862)

= Nephrodium filix-mas var. erythrosorum (D.C.Eaton) Christ ex Matsum.; Index Pl. Jap. 1: 319 (1904)

= Nephrodium prolificum (Maxim.) Diels; Nat. Pflanzenfam. [Engler & Prantl] I, 4: 173 (1899), isonym

= Nephrodium prolificum (Maxim.) G.Nicholson; Ill. Dict. Gard. 4: 573 (1888)

species :

Dryopteris erythrovaria K.Hori & N.Murak.; Acta Phytotax. Geobot. 69(2): 86 (2018)

Distribution: China (Jiangsu, Zhejiang, Anhui); South Korea; Japan (Honshu, Kyushu)

36 CHI 38 JAP KOR

species :

Dryopteris esterhuyseniae Schelpe & N.C.Anthony; Contr. Bolus Herb. 10: 148 (1982)

Distribution: South Africa (W-KwaZulu-Natal: Drakensberg)

27 SAF

Remarks: included by Roux (2009) in D. dracomontana but kept separate by Ferns of South Africa

species :

Dryopteris expansa (C.Presl) Fraser-Jenk. & Jermy; Fern Gaz. 11(5): 338 (1977)

Name: Feingliedriger Wurmfarn (DE), Spreading woodfern, Northern Buckler-fern (EN)

Distribution: Iceland; England (N-England, N-Wales, Scotland, Outer Hebrides, Orkney Isl., Shetland Isl., St. Kilda Isl.); Norway; Sweden; Germany (Brandenburg, Berlin, Baden-Württemberg, Bayern, Hessen, Mecklenburg-Vorpommern, Niedersachsen, Nordrhein-Westfalen, Sachsen, Sachsen-Anhalt, Thüringen); Switzerland; Liechtenstein; Czech Republic; Slovakia; Hungary; Portugal; Spain; Andorra; France; Corsica; Italy; Slovenia; Croatia; Montenegro; Serbia; Kosovo; Bulgaria; Greece (mountains in N-Greece); Estonia; Latvia; Lithuania; Belarus; C-European Russia; N-European Russia; Moldova; Ukraine; Siberia (Altai, Buryatia, Chita, Irkutsk, Krasnoyarsk, Tuva, W-Siberia, Yakutia); Russian Far East (Amur, Kamchatka, Khabarovsk, Kuril Isl., Magadan, Primorye, Sakhalin); Northern Caucasus; Georgia [Caucasus]; Turkey (Anatolia); China (Hebei, Heilongjiang, Jilin, Liaoning); Mongolia; North Korea; South Korea; Japan; Taiwan; Alaska; Greenland; Canada (Alberta, British Columbia, Labrador, Newfoundland, Northern Territories, Ontario, Québec, Yukon); USA (California, Colorado, Idaho, Michigan, Minnesota, Montana, Oregon, Washington State, Wisconsin, Wyoming)

10 GRB ICE NOR SWE 11 CZE GER HUN LIE SLO SWI 12 ADO COR FRA POR SPA 13 BUL CRO GRC ITA KOS MNT SLV SRB 14 BLR EST LAT LIT MLD RUS UKR 30 SIB 31 RFE 33 GEO NCS 34 TUR 36 CHI 37 MON 38 JAP KOR NKO TAI 70 ASK CDN GNL 71 CDN 72 CDN 73 USA 74 USA 75 USA 76 USA

Remarks: carthusiana agg.

Synonyms: hidden

= Dryopteris assimilis S.Walker; Amer. J. Bot. 48: 607 (1961)

= Dryopteris austriaca subsp. assimilis (S.Walker) O.Bolòs & Vigo; Fl. Paisos Catalans: 170 (1984)

= Dryopteris carthusiana subsp. assimilis (S.Walker) O.Bolòs, Vigo, Masalles & Ninot; Fl. Man. Paisos Catalans: 1213 (1990)

= Dryopteris expansa subsp. assimilis (S.Walker) Tzvelev; Sosud. Rast. Sovet. Dal´nego Vostoka 5: 54 (1991)

= Dryopteris extremiorientalis V.N.Vassil.; nom. inval.

= Dryopteris manshurica Ching; Fl. Pl. Herb. Chin. Bor.-Or. 1: 69 (1958)

= Dryopteris minimisora Nakai; Bull. Natl. Sci. Mus., Tokyo, 27: 19 (1949)

= Dryopteris morrisonensis (Hayata) Hayata; J. Coll. Sci. Imp. Univ. Tokyo 30(1): 450 (1911), Ic. Fl. Taiwan 5: 281 f. 108 (1915)

= Dryopteris siranensis Nakai; Bull. Natl. Sci. Mus., Tokyo, 33: 3 (1953)

= Dryopteris spinulosa subsp. assimilis (S.Walker) Schidlay; Fl. Slovenska 2: 217 (1966)

= Dryopteris spinulosa var. morrisonensis Hayata; J. Coll. Sci. Imp. Univ. Tokyo 30: 450 (1911) [fide Fraser-Jenkins]

= Lastrea expansa (C.Presl) C.Presl; Tent. Pterid. 76 (1836)

= Nephrodium expansum C.Presl; Reliq. Haenk. 1: 38 (1825)

variant :

var. alpina (T.Moore) Viane; Pl. Syst. Evol. 153(1-2): 90 (1986)

Distribution: England (Scotland)

10 GRB

Synonyms: hidden

= Lastrea dilatata var. alpina T.Moore; Ferns Brit. & Ireland Nature-Printed, sub t. 22 (1856) (non Dryopteris alpina Rosenst.)

variant :

var. willeana (Lid) Elven; Nordic J. Bot. 16(1): 4 (1996)

Distribution: Scandinavia

Synonyms: hidden

= Dryopteris austriaca var. willeana Lid

species :

Dryopteris fadenii Pic.Serm.; Webbia 37(2): 333 (1984)

Distribution: Ethiopia; Uganda; Rwanda; Kenya; Tanzania

23 RWA 24 ETH 25 KEN TAN UGA

species :

Dryopteris fangii Ching ex Fraser-Jenk. & Z.R.Wang; Acta Phytotax. Sin. 23(5): 349 (1985), nom. nov.

Distribution: China (Sichuan, etc.); Tibet; Bhutan; NE-India (Arunachal Pradesh)

36 CHI TIB 40 BHU IND

Remarks: included in D. chrysocoma by Flora of China but Fraser-Jenkins et al. (2018) keep it separate

Synonyms: hidden

= Dryopteris chrysocoma var. gracilis Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 437 (1938) (non (T. Moore ex Bedd.) Ching)

species :

Dryopteris fatuhivensis E.D.Br.; Bull. Bernice P. Bishop Mus. 89 [Fl. S. E. Polynesia II]: 27, t. 8 (1931) (fide Fraser-Jenkins et al. 2018)

Distribution: Cook Isl. (Rarotonga); Marquesas Isl. (Nuku Hiva, Fatu Hiva, Ua Huka, Ua Pou, Hiva Oa)

61 COO MRQ

Remarks: belongs possibly to D. hirtipes

Synonyms: hidden

= Thelypteris fatuhivensis (E.D.Br.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 251 (1941)

species :

Dryopteris filipaleata J.P.Roux; Bothalia 34(1): 28 (2004)

Distribution: Kenya; Tanzania

25 KEN TAN

species :

Dryopteris filix-mas (L.) Schott; Gen. Fil. (Vindob.) t. 9 (1834)

subspecies :

subsp. filix-mas

Name: Gewöhnlicher Wurmfarn, Männlicher Wurmfarn (DE), Male Fern (EN)

Distribution: Iceland; England (England, Wales, Scotland, Outer Hebrides, Orkney Isl., Shetland Isl., Isle of Man); Ireland (Ireland, Northern Ireland); Denmark; Norway; Sweden; Finland; Netherlands; Belgium; Luxembourg; Germany (Brandenburg, Berlin, Baden-Württemberg, Bayern, Hessen, Hamburg, Mecklenburg-Vorpommern, Niedersachsen, Nordrhein-Westfalen, Rheinland-Pfalz, Schleswig-Holstein, Saarland, Sachsen, Sachsen-Anhalt, Thüringen); Switzerland; Liechtenstein; Austria; Poland; Czech Republic; Slovakia; Hungary; Portugal; Spain; Andorra; France; Channel Isl. (Alderney, Guernsey, Jersey); Corsica; ?Sardinia; Italy; Sicily; Malta; San Marino; Slovenia; Croatia; Bosnia & Hercegovina; Montenegro; Serbia; Kosovo; North Macedonia; Albania; Romania; Bulgaria; European Turkey; Greece (widespread on mainland, Samothraki); Estonia; Latvia; Lithuania; Belarus; C-European Russia; E-European Russia; N-European Russia; W-European Russia; Moldova; Ukraine; Crimea; Morocco; Algeria; Tunisia; Siberia (Altai, Buryatia, Chita, Irkutsk, Tuva, W-Siberia, Yakutia); Kazakhstan; Uzbekistan; Kyrgyzstan; Tajikistan; Northern Caucasus; Georgia [Caucasus]; Armenia; Azerbaijan; Turkey (Anatolia); Iran; E-Afghanistan; China (Xinjiang); Pakistan (Dir, Chitral, Swat, Hazara, Baltistan); NW-India (Jammu & Kashmir: Ladakh); New Zealand [I]; Chatham Isl. [I]; S-Greenland; Canada (Alberta, British Columbia, Saskatchewan); USA (Arizona, California, Colorado, Idaho, Montana, New Mexico, Nevada, Oklahoma, Oregon, South Dakota, Texas, Utah, Washington State, Wyoming); Mexico (Coahuila); Argentina [I] (Neuquen [I]); Chile [I] (Los Lagos [I])

10 DEN FIN GRB ICE IRE NOR SWE 11 AUT BGM CZE GER HUN LIE LUX NET POL SLO SWI 12 ADO CIS COR FRA POR SPA 13 ALB BIH BUL CRO GRC ITA KOS MLT MNT NMA ROM SIC SLV SMA SRB TUE 14 BLR EST KRY LAT LIT MLD RUS UKR 20 ALG MOR TUN 30 SIB 32 KAZ KGZ TZK UZB 33 ARM AZE GEO NCS 34 AFG IRN TUR 36 CHI 40 IND PAK 51 ctm nzl 70 GNL 71 CDN 73 USA 74 USA 76 USA 77 USA 79 MEX 85 arg cle

Remarks: filix-mas agg.

Synonyms: hidden

= Aspidium depastum Schkuhr; Deutschl. Krypt. Gew. 1: 50 (1809)

= Aspidium erosum Schkuhr; Deutschl. Krypt. Gew. 1: 46 (1809)

= Aspidium expansum D.Dietr.; Fl. Jenensis: s. p. (1826)

= Aspidium filix-mas (L.) Sw.; J. Bot. (Schrader) 1800(2) 38 (1801)

= Aspidium filix-mas f. crenatum Milde; Sporenpfl. 51 (1865)

= Aspidium filix-mas f. erosum (Döll) Döll; Fl. Baden [Döll] 1(1): 27 (1855)

= Aspidium filix-mas f. erosum (Döll) Zapal.; Conspic. Fl. Gallic. Crit. 1: 78 (1905), isonym

= Aspidium filix-mas f. heleopteris (Borkh.) Milde; Sporenpfl. 52 (1865)

= Aspidium filix-mas f. incisum Döll; Fl. Baden [Döll] 1(1): 27 (1855)

= Aspidium filix-mas var. crenatum Milde; Nov. Actorum Acad. Caes. Leop.-Carol. German. Nat. Cur. 26: 508 (1858)

= Aspidium filix-mas var. erosum (Schkuhr) Pursh; Fl. Amer. Sept. 2: 663 (1813)

= Aspidium filix-mas var. erosum Döll; Rhein. Fl. 16 (1843)

= Aspidium filix-mas var. heleopteris (Borkh.) Christ; Mém. Soc. Bot. France 1 [Hist. Acrostich.]: 38 (1905)

= Aspidium filix-mas var. heleopteris (Borkh.) Milde; 121 (1867)

= Aspidium mildeanum Göpp.; Denkschr. 50jähr. Best. Vaterl. Ges. Schles. 193 (1853)

= Aspidium nemorale Gray; Nat. Arr. Brit. Pl. 2: 7 (1821 publ. 1822)

= Aspidium opizii Wierzb.; Flora 23: 379 (1840)

= Aspidium umbilicatum (Poir.) Desv.; Mag. Neuesten Entdeck. Gesammten Naturk. Ges. Naturf. Freunde Berlin 5: 320 (1811)

= Aspidium veselskii Hassl. ex Domin; Vestn. Ceské Spolecn. Nauk, Tr. Mat.-Prír. 1941(21): 4 (1942)

= Dryopteris filix-mas f. crenata (Milde) Diklic & V.Nikolic; Fl. SR Srbije 10: 10 (1986)

= Dryopteris filix-mas var. recurva (Francis) Rothm.; 97 (1946)

= Dryopteris patagonica Diem; Darwiniana 12: 70, f. 2 (1960)

= Dryopteris ×bohemica Domin; Vestn. Ceské Spolecn. Nauk, Tr. Mat.-Prír. 1941(21): 2-3 (1942)

= Dryopteris ×digenea Domin; Preslia 13-15: 8 (1936)

= Filix filix-mas (L.) Farw.; Rep. (Annual) Michigan Acad. Sci. 6: 201 (1904)

= Filix pinnata Gilib.; Exerc. Phyt. 2: 557 (1792), opus utique oppr.

= Filix-mas filix-mas (L.) Farw.; Amer. Midl. Naturalist 12: 234-251 (1931)

= Filix-mas vulgaris Hill; Brit. Herb. (Hill) 527, t. 74 (1756)

= Lastrea erosa (Schkuhr) Deakin; Fl. Brit. 4: 101 (1848)

= Lastrea filix-mas (L.) C.Presl; Tent. Pterid. 76 (1836)

= Lastrea officinalis Bubani; Fl. Pyren. (Bubani) 4: 433 (1901)

= Lastrea propinqua Woll. ex Lowe; Our nat. Ferns 1: 234, 280 (1865)

= Lophodium erosum (Schkuhr) Newman; Phytologist 4: app. 21 (1851)

= Lophodium filix-mas (L.) Newman; Phytologist 4: app. 20 (1851)

= Nephrodium crenatum Stokes; Bot. Mat. Med. 4: 606 (1812), based on Polystichum filix-mas (L.) Roth, nom. superfl.

= Nephrodium filix-mas (L.) Rich.; Cat. Pl. Jard. Méd. Paris 129 (1801)

= Nephrodium filix-mas (L.) Strempel; Fil. Berol. Syn.: 30 (1822)

= Nephrodium filix-mas var. crenatum (Milde) Murr; 8 (1923), isonym

= Nephrodium filix-mas var. crenatum (Milde) Rouy; Fl. France [Rouy & Foucaud] 14: 407 (1913)

= Nephrodium filix-mas var. deorsilobatum (T.Moore) Murr; 8 (1923)

= Nephrodium filix-mas var. erosum (Schkuhr) Murr; 8 (1923)

= Nephrodium filix-mas var. subintegrum (Döll) Murr; 8 (1923)

= Nephrodium propinquum (Woll. ex Lowe) Lowe; Brit. Ferns [Lowe] 133 (1891), nom. illeg.

= Nephrodium rigidoforme Rouy; Fl. France [Rouy & Foucaud] 14: 408 (1913)

= Nephrodium umbilicatum (Poir.) Desv.; Mem. Soc. Linn. Paris 6(2): 260 (1827)

= Polypodium filix-mas L.; Sp. Pl. 2: 1090 (1753)

= Polypodium heleopteris Borkh.; Arch. Bot. (Leipzig) 1(3): 19 (1798)

= Polypodium mas L.; Mant. Pl. 2: 506 (1771)

= Polypodium nemorale Salisb.; Prodr. Stirp. Chap. Allerton 403 (1796), nom. nov. for P. filix-mas L., nom. superfl.

= Polypodium paleaceum Borkh.; Roem. Arch. 1(3): 20 (1798)

= Polypodium umbilicatum Poir.; Encycl. [J. Lamarck et al.] 5: 528 (1804)

= Polystichum filix-mas (L.) Roth; Tent. Fl. Germ. 3: 82 (1799)

= Polystichum filix-mas var. erosum (Döll) Sendtn.; 907 (1854)

= Polystichum obtusum Dulac; Fl. Hautes-Pyrenées 33 (1867)

= Polystichum paleaceum (Borkh.) O.Schwarz; Mitt. Thüring. Bot. Ges. 1: 85 (1949)

= Polystichum patentissimum Trevis.; Atti Soc. Ital. Sci. Nat. 17(2): 227 (1874)

= Polystichum polysorum Tod.; Giorn. Sci. Nat. Econ. Palermo 1: 240 (1866)

= Tectaria filix-mas (L.) Cav.; Descr. Pl. (Cavanilles) 251 (1801)

= Thelypteris filix-mas (L.) Nieuwl.; Midl. Naturalist (Notre Dame) 1: 226 (1910)

variant :

var. pseudorigida Christ; Farnkr. Schweiz 1: 134 (1900)

Distribution: Switzerland; Northern Caucasus

11 SWI 33 NCS

Synonyms: hidden

= Dryopteris filix-mas subsp. pseudorigida (Christ) Askerov; Zametki Sist. Geogr. Rast. 39: 6 (1983)

subspecies :

subsp. brittonii Fraser-Jenk. & Widén; Advances Forest. Res. India 29: 139-160 (2006)

Distribution: Canada (New Brunswick, Newfoundland, Nova Scotia, Ontario, Québec); St. Pierre et Miquelon; USA (Maine, Michigan, New Hampshire, New York, Pennsylvania, Rhode Island, Vermont, Wisconsin)

72 CDN SPM 74 USA 75 USA

Synonyms: hidden

= Thelypteris filix-mas f. incisa (T.Moore) Dole; Fl. Vermont, ed. 3: 11 (1937)

= Thelypteris filix-mas f. producta (T.Moore) Dole; Fl. Vermont, ed. 3: 11 (1937)

species :

Dryopteris flaccisquama A.Rojas; Revista Biol. Trop. 49(2): 436 (2001)

Distribution: Costa Rica; Panama; Hispaniola (Dominican Republic, Haiti); Colombia (Cundinamarca); Bolivia (Cochabamba, La Paz)

80 COS PAN 81 HIS 82 CLM 83 BOL

species :

Dryopteris flemingii (Fraser-Jenk.) Fraser-Jenk.; New Species Syndr. Indian Pteridol. 125 (1997), pro hybr.

Distribution: China (Yunnan); Tibet; C-Nepal; Bhutan; India (Himachal Pradesh, Uttarakhand)

36 CHI TIB 40 BHU IND NEP

Conservation Status: IUCN VU for India (Fraser-Jenkins 2012)

Synonyms: hidden

= Dryopteris ×flemingii Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.), Bot. 18(5): 458 (1989)

species :

Dryopteris formosana (Christ) C.Chr.; Index Filic. 266 (1905)

Distribution: China (Sichuan, Zhejiang); South Korea (Cheju Isl.); Japan; Ryukyu Isl. (Tokunoshima); Taiwan; Philippines

36 CHI 38 JAP KOR NNS TAI 42 PHI

Synonyms: hidden

= Aspidium formosanum Christ; Bull. Herb. Boissier, sér. 2, 4: 615 (1904)

= Dryopteris constantissima (Hayata) Hayata; Icon. Pl. Formosan. 4: 191 (1914)

= Dryopteris kodamae Hayata; Icon. Pl. Formosan. 4: 156 (1914)

= Dryopteris liui Ching; Bull. Bot. Res., Harbin 4(4): 12 (1984)

= Dryopteris phaeolepis Hayata; Icon. Pl. Formosan. 4: 169, t. 109 (1914)

= Polystichum constantissimum Hayata; Icon. Pl. Formosan. 4: 191, f. 129 (1914)

species :

Dryopteris fragrans (L.) Schott; Gen. Fil. (Vindob.) ad t. 9 (1834)

Name: Duftender Wurmfarn, Fragrant woodfern

Distribution: N-Finland; N-European Russia; Siberia (Altai, Buryatia, Chita, Irkutsk, Krasnoyarsk, Tuva, W-Siberia, Yakutia); Russian Far East (Amur, Kamchatka, Khabarovsk, Magadan, Primorye, Sakhalin); Kazakhstan; China (Hebei, Heilongjiang, Jilin, Liaoning, Nei Mongol, Xinjiang); Mongolia; North Korea; South Korea (incl. Cheju Isl.); Japan; Alaska; USA (Minnesota, New York)

10 FIN 14 RUS 30 SIB 31 RFE 32 KAZ 36 CHI 37 MON 38 JAP KOR NKO 70 ASK 75 USA

Synonyms: hidden

= Aspidium fragrans (L.) Sw.; J. Bot. (Schrader) 1800(2): 35 (1801)

= Dryopteris aquilonaris Maxon; Bull. Torrey Bot. Club 27: 638 (1900)

= Dryopteris fragrans f. aquilonaris (Maxon) M.Broun; Index No. Amer. Ferns: 68 (1938)

= Dryopteris fragrans var. hookeriana (Fernald) A.Prince ex Weath.; Amer. Fern J. 26: 62 (1936)

= Dryopteris fragrans var. remotiuscula (Kom.) Kom.; Fl. URSS 1: 38 (1934)

= Filix aquilonaris (L.) Farw.; Rep. (Annual) Michigan Acad. Sci. 18: 81-83 (1916)

= Filix fragrans (L.) Farw.; Rep. (Annual) Michigan Acad. Sci. 18: 81-83 (1916)

= Filix-mas fragrans (L.) Farw.; Amer. Midl. Naturalist 12: 234-251 (1931)

= Filix-mas fragrans var. hookeriana (Fernald) Farw.; Amer. Midl. Naturalist 12: 254 (1931)

= Lastrea fragrans (L.) C.Presl; Tent. Pterid. 76 (1836)

= Nephrodium fragrans (L.) Desv.; Mem. Soc. Linn. Paris 6: 260 (1827), isonym

= Nephrodium fragrans (L.) Richards.; Narr. Journey Polar Sea [Franklin] 754 (1823)

= Nephrodium fragrans f. aquilonare (Maxon) Clute; Fern Bull. 15: 21 (1907)

= Nephrodium fragrans var. aquilonare (Maxon) Gilbert; List N. Amer. Pterid.: 17 (1901)

= Nephrodium fragrans var. remotiusculum Kom.; Repert. Spec. Nov. Regni Veg. 9: 94 (1910)

= Polypodium fragrans L.; Sp. Pl. 2: 1089 (1753)

= Polystichum fragrans (L.) Ledeb.; Fl. Ross. (Ledeb.) 4(14): 514 (1853)

= Polystichum fragrans (L.) Roth; Catal. Bot. 2: 147 (1800)

= Thelypteris fragrans (L.) Nieuwl.; Midl. Naturalist (Notre Dame) 1: 226 (1910)

= Thelypteris fragrans var. hookeriana Fernald; Rhodora 25: 3 (1923)

= Woodsia xanthosporangia Ching; Bull. Fan Mem. Inst. Biol. Bot. 1: 101, c. t (1929)

species :

Dryopteris fragrantiformis Tzvelev; Novosti Sist. Vyssh. Rast. 26: 5 (1989)

Distribution: Russian Far East (Kuril Isl.)

31 RFE

species :

Dryopteris fructuosa (Christ) C.Chr.; Index Filic. 267 (1905)

Distribution: China (Hubei, Sichuan, Yunnan); Tibet; Taiwan; Nepal; Bhutan; India (Arunachal Pradesh, Nagaland, Sikkim); Myanmar

36 CHI TIB 38 TAI 40 BHU IND NEP 41 MYA

Remarks: D. lachoongensis is kept separate by Flora of China and probably a valid species fide Knapp (in litt. 2016)

Synonyms: hidden

= Aspidium fructuosum (Christ) Christ; Mém. Soc. Bot. France 1 [Hist. Acrostich.]: 38 (1905)

= Aspidium varium var. fructuosum Christ; Mém. Soc. Bot. France 1 [Hist. Acrostich.]: 38 (1905)

= Dryopteris adenorachis C.Chr.; Index Filic., Suppl. 2 (1906-1912): 13 (1917), based on D. cavaleriei H. Lévl.

= Dryopteris apicifixa Ching, Boufford & K.H.Shing; J. Arnold Arbor. 64: 27 (1983)

= Dryopteris cavaleriei H.Lév.; Fl. Kouy-Tcheou 490 (1915), nom. illeg., non C. Chr. (1905)

= Dryopteris hypophlebia Hayata; Icon. Pl. Formosan. 4: 154, f. 96 (1914)

= Dryopteris pseudosabaei Hayata; Icon. Pl. Formosan. 5: 283, f. 110 (1915)

= Nephrodium fructuosum (Christ) Hand.-Mazz.; Symb. Sin. 6: 21-25 (1929)

species :

Dryopteris fuscipes C.Chr.; Index Filic., Suppl. 2 (1906-1912): 14 (1917)

Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hubei, Hunan, Jiangsu, Jiangxi, Sichuan, Yunnan, Zhejiang); South Korea; Japan; Taiwan (MaZu Isl.); Vietnam; New Guinea (Papua New Guinea)

36 CHI 38 JAP KOR TAI 41 VIE 42 NWG

Synonyms: hidden

= Aspidium erythrosorum var. obtusum Christ; Mém. Soc. Bot. France 1 [Hist. Acrostich.]: 40 (1905)

= Dryopteris bipinnata C.Chr.; H. Lév., Cat. Pl. Yun-Nan 102 (1916) (non Copel. (1914))

= Dryopteris confertipinna Ching & K.H.Shing; Fl. Fujianica 1: 600 (1982)

= Dryopteris erythrosora var. obtusa (Makino) Makino; Bot. Mag. (Tokyo) 23(270): 144 (1909)

= Dryopteris fuscipes f. major Ching ex K.H.Shing & J.F.Cheng; Jiangxi Sci. 8(3): 49 (1990)

= Dryopteris makinoi f. latipinnula H.Itô; Bot. Mag. (Tokyo) 50(590): 71 (1936)

= Dryopteris makinoi Koidz.; Acta Phytotax. Geobot. 1: 26 (1932), nom. nov.

= Dryopteris medialisora Ching & P.S.Chiu; Bot. Res. Acad. Sin. 1987(2): 6 (1987)

= Dryopteris medioxima Koidz.; Acta Phytotax. Geobot. 1: 32 (1932)

= Dryopteris multijugata Ching & C.F.Zhang; Bull. Bot. Res., Harbin 3(3): 12 (1983)

= Dryopteris neofuscipes Ching & P.S.Chiu; Bot. Res. Acad. Sin. 1987(2): 4 (1987), non Ching & Z. Y. Liu (1984)

= Dryopteris obtusissima Makino; J. Jap. Bot. 2: 47 (1918), nom. nov.

= Dryopteris parafuscipes Ching & Z.Y.Liu; (1984), not Ching & P. S. Chiu (1987)

= Dryopteris persimilis Ching & C.F.Zhang; Bull. Bot. Res., Harbin 3(3): 16 (1983), nom. illeg., non (Baker) C. Chr.

= Dryopteris stenochlamys Ching ex K.H.Shing & J.F.Cheng; Jiangxi Sci. 8(3): 49 (1990)

= Nephrodium erythrosorum var. obtusum Makino; Bot. Mag. (Tokyo) 13: 79 (1899)

= Nephrodium filix-mas f. obtusum (Makino) Makino ex Matsum.; Index Pl. Jap. 1: 319 (1904)

= Nephrodium fuscipes (C.Chr.) Hand.-Mazz.; Symb. Sin. 6: 21-25 (1929)

species :

Dryopteris fuscoatra (Hillebr.) W.J.Rob.; Bull. Torrey Bot. Club 39: 592, t. 42 (1912)

Distribution: Hawaii (Kauai, Oahu, Lanai, Molokai, Maui, Hawaii Isl.)

63 HAW

Synonyms: hidden

= Aspidium filix-mas var. fuscoatrum Hillebr.; Fl. Hawaiian Isl. [Hillebrand] 575 (1888)

variant :

var. lamoureuxii Fraser-Jenk.; Thaiszia 4(1): 22 (1994)

Distribution: Hawaii (Kauai, Maui, Hawaii Isl.)

63 HAW

species :

Dryopteris futura A.R.Sm.; Proc. Calif. Acad. Sci. ser, 4, 40: 216 (1975)

Distribution: Mexico (Chiapas, Guerrero, Hidalgo, Michoacan, Morelos, Oaxaca); Guatemala; Honduras

79 MEX 80 GUA HON

species :

Dryopteris gamblei (C.Hope) C.Chr.; Index Filic. 267 (1906)

Distribution: China; Tibet; Japan; Taiwan; Nepal; Bhutan; India (Arunachal Pradesh, Manipur, Meghalaya, Nagaland, Uttarakhand, Sikkim, West Bengal); Myanmar

36 CHI TIB 38 JAP TAI 40 BHU IND NEP 41 MYA

Remarks: not a synonym of D. stenolepis fide Fraser-Jenkins et al. (2015, 2018)

Synonyms: hidden

= Dryopteris darjeelingensis Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.), Bot. 18(5): 335 (1989)

= Nephrodium gamblei C.Hope; J. Bombay Nat. Hist. Soc. 12: 533, t. 7 (1899)

species :

Dryopteris gaoligongensis Z.Y.Zuo, J.Mei Lu & D.Z.Li; Pl. Diversity 44(2): 185 (2021)

Distribution: China (Yunnan)

36 CHI

Remarks: sect. Diclisodon, see Zuo et al. 2022

species :

Dryopteris gemmifera S.Y.Dong; Bot. Stud. (Taipei) 51(4): 537 (2010)

Distribution: China (Hainan)

36 CHI

species :

Dryopteris glabra (Brack.) Kuntze; Revis. Gen. Pl. 2: 812 (1891)

Distribution: Hawaii (Kauai, Oahu, Molokai, Lanai, Maui, Hawaii Isl.)

63 HAW

Remarks: ser. Aemulae

Synonyms: hidden

= Aspidium glabrum (Brack.) Mett.; Abh. Senckenberg. Naturf. Ges. 2(2) [Pheg. u. Asp.]: 343 [Pheg. u. Asp. 59, t. 140] (1858)

= Lastrea glabra Brack.; U.S. Expl. Exped., Filic. 16: 200 (1854)

= Nephrodium glabrum (Brack.) Baker; Syn. Fil. [W. J. Hooker & Baker] 278 (1867)

= Thelypteris glabra (Brack.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 251 (1941)

variant :

var. alboviridis (W.H.Wagner) D.D.Palmer; Amer. Fern J. 92(2): 100 (2002)

Distribution: Hawaii (Kauai)

63 HAW

Synonyms: hidden

= Dryopteris alboviridis W.H.Wagner; Contr. Univ. Michigan Herb. 22: 177 (1999)

variant :

var. flynnii D.D.Palmer; Amer. Fern J. 92(2): 101 (2002)

Distribution: Hawaii (Kauai)

63 HAW

variant :

var. hobdyana (W.H.Wagner) D.D.Palmer; Amer. Fern J. 92(2): 101 (2002)

Distribution: Hawaii (Maui)

63 HAW

Synonyms: hidden

= Dryopteris hobdyana W.H.Wagner; Contr. Univ. Michigan Herb. 22: 177 (1999)

variant :

var. nuda (Underw.) Fraser-Jenk.; Thaiszia 4(1): 38 (1994)

Distribution: Hawaii (Kauai, Oahu, Molokai, Maui, Hawaii Isl.)

63 HAW

Synonyms: hidden

= Dryopteris nuda Underw. ex A.Heller; Minnesota Bot. Stud. 1: 780 (1897)

variant :

var. pusilla (Hillebr.) Fraser-Jenk.; Thaiszia 4(1): 35 (1994)

Name: Na Pali-Kona Woodfern

Distribution: Hawaii (Kauai)

63 HAW

Synonyms: hidden

= Aspidium glabrum var. pusillum Hillebr.; Fl. Hawaiian Isl. [Hillebrand] 577 (1888)

= Dryopteris parvula W.J.Rob.; Bull. Torrey Bot. Club 39: 593, t. 44 (1912)

= Dryopteris tenebrosa W.H.Wagner; Contr. Univ. Michigan Herb. 19: 68 (1993)

variant :

var. soripes (Hillebr.) Herat ex Fraser-Jenk.; Thaiszia 4(1): 36 (1994)

Distribution: Hawaii (Kauai, Oahu, Molokai, Maui, Hawaii Isl.)

63 HAW

Synonyms: hidden

= Aspidium glabrum var. soripes Hillebr.

species :

Dryopteris glandulosopaleata J.P.Roux; Kew Bull. 59(2): 211 (2004)

Distribution: Nigeria; Cameroon

22 NGA 23 CMN

species :

Dryopteris goeringiana (Kunze) Koidz.; Bot. Mag. (Tokyo) 48: 382 (1929)

Distribution: Russian Far East (Primorye); China (Gansu, Hebei, Henan, Liaoning, Nei Mongol, Shaanxi, Shanxi, Sichuan, Xinjiang); North Korea; South Korea

31 RFE 36 CHI 38 KOR NKO

Remarks: see also D. laeta which is frequently included here

Synonyms: hidden

= Aspidium filix-mas var. giraldii Christ; Baroni & Christ, Nuovo Giorn. Bot. Ital., n. s. 4(1): 94-95 (1897)

= Aspidium goeringianum Kunze; Bot. Zeitung (Berlin) 557 (1848)

= Asplenium goeringianum (Kunze) Mett.; Abh. Senckenberg. Naturf. Ges. 3(1) [Aspl.]: 198, no. 243, t. 6, f. 11-12 (1859)

= Athyrium goeringianum (Kunze) T.Moore; Index Fil. [T. Moore] 185 (1860)

= Dryopteris giraldii (Christ) C.Chr.; J. Wash. Acad. Sci. 17: 500 (1927)

= Dryopteris laeta var. giraldii (C.Chr.) Ching; Fl. Tsinling. 2: 170 (1974)

= Dryopteris subramosa Christ; Notul. Syst. (Paris) 1: 42 (1909)

= Lastrea goeringiana (Kunze) T.Moore; Index Fil. [T. Moore] 93 (1858)

species :

Dryopteris goldieana (Hook. ex Goldie) A.Gray; Manual (Gray): 631 (1848)

Name: Goldie´s woodfern

Distribution: Canada (New Brunswick, Ontario, Québec, Nova Scotia); USA (Alabama, Arkansas, Connecticut, District of Columbia, Delaware, Georgia, Iowa, Illinois, Indiana, Kentucky, Massachusetts, Maryland, Maine, Michigan, Minnesota, Missouri, North Carolina, New Hampshire, New Jersey, New York, Ohio, Pennsylvania, Rhode Island, South Carolina, Tennessee, Virginia, Vermont, Wisconsin, West Virginia)

72 CDN 74 USA 75 USA 78 USA

Synonyms: hidden

= Aspidium americanum Torrey; Cat. Pl. New York 80 (1818), nomen

= Aspidium filix-mas var. goldieanum (Hook. ex Goldie) Christ; Farnkr. Erde: 258 (1897)

= Aspidium goldieanum Hook. ex Goldie; Edinburgh New Philos. J. 6: 333 (1822)

= Dryopteris goldieana var. prolifera Gilbert; Fern Bull. 8: 9 (1900)

= Filix goldieana (Hook. ex Goldie) Farw.; Rep. (Annual) Michigan Acad. Sci. 18: 81 (1917)

= Filix-mas goldieana (Hook. ex Goldie) Farw.; Amer. Midl. Naturalist 12: 254 (1931)

= Lastrea goldieana (Hook. ex Goldie) C.Presl; Tent. Pterid. 76 (1836)

= Nephrodium goldieanum (Hook. ex Goldie) Hook. & Grev.; Icon. Filic. [W. J. Hooker & Greville] 1: t. 102 (1829)

= Polystichum goldieanum (Hook. ex Goldie) Keyserl.; Polyp. Herb. Bunge 44 (1873)

= Thelypteris goldieana (Hook. ex Goldie) Nieuwl.; Amer. Midl. Naturalist 1: 226 (1910)

species :

Dryopteris gonggaensis H.S.Kung, Li Bing Zhang & X.S.Guo; Acta Bot. Yunnan. 17(2): 159-160, f. 1 (1995)

Distribution: China (Sichuan)

36 CHI

Remarks: included by Fraser-Jenkins (1997) in D. blanfordii ssp. nigrosquamosa, but kept separate in Flora of China

species :

Dryopteris gorgonea J.P.Roux; Bothalia 34(1): 33 (2004)

Distribution: Cape Verde Isl. (Fogo)

21 CVI

species :

Dryopteris grandifrons Li Bing Zhang; Taxon 61(6): 1207 (2012), nom. nov.

Distribution: China (NW-Yunnan)

36 CHI

Synonyms: hidden

= Nothoperanema giganteum Ching; Acta Phytotax. Sin. 11: 28 (1966)

species :

Dryopteris guanchica Gibby & Jermy; J. Linn. Soc., Bot. 74: 256 (1977)

Name: Guanchen-Wurmfarn (DE); Helecho penco dentado (ES)

Distribution: Portugal; Spain; Canary Isl. (Tenerife, La Gomera, Hierro)

12 POR SPA 21 CNY

Remarks: carthusiana agg.

species :

Dryopteris guangxiensis S.G.Lu; Guihaia 19(1): 27 (1999)

Distribution: China (Guangxi)

36 CHI

Synonyms: hidden

= Dryopteris parafuscipes Ching & P.S.Chiu; Bot. Res. Acad. Sin. 1987(2): 4 (1987), nom. illeg., non Ching & Z. Y. Liu (1984)

species :

Dryopteris gymnophylla (Baker) C.Chr.; Index Filic. 269 (1905)

Distribution: China (Anhui, Guizhou, Hebei, Henan, Hubei, Jiangsu, Jiangxi, Shandong, Zhejiang); North Korea; South Korea; Japan; N-Thailand

36 CHI 38 JAP KOR NKO 41 THA

Synonyms: hidden

= Dryopteris koraiensis Tagawa; Acta Phytotax. Geobot. 6: 254 (1936)

= Dryopteris sakuraii (Rosenst.) Tagawa; J. Jap. Bot. 13: 185 (1937)

= Dryopteris subtripinnata var. sakuraii Rosenst.; Repert. Spec. Nov. Regni Veg. 13: 132 (1914)

= Nephrodium gymnophyllum Baker; J. Bot. 25: 170 (1887)

species :

Dryopteris gymnosora (Makino) C.Chr.; Index Filic. 269 (1905)

Distribution: China (Anhui, Fujian, Guizhou, Hubei, Hunan, Jiangxi, Sichuan, Yunnan, Zhejiang); Japan

36 CHI 38 JAP

Synonyms: hidden

= Nephrodium gymnosorum Makino; Bot. Mag. (Tokyo) 13: 64 (1899)

variant :

var. angustata H.Itô; (1938)

Distribution: Japan (Yakushima Isl.)

38 JAP NNS

Conservation Status: CR in Japan

species :

Dryopteris habaensis Ching; Sunyatsenia 6: 2 (1941)

Distribution: China (NW-Yunnan)

36 CHI

species :

Dryopteris hadanoi Sa.Kurata; J. Geobot. 14: 83 (1966)

Distribution: Japan

38 JAP

Conservation Status: IUCN R. NT in Japan

species :

Dryopteris handeliana C.Chr.; Dansk Bot. Ark. 9(3): 62, t. 6(5-6) (1937)

Distribution: China (Guizhou, Hubei, Hunan, Sichuan, Yunnan, Zhejiang); Japan

36 CHI 38 JAP

Synonyms: hidden

= Dryopteris tasiroi Tagawa; Acta Phytotax. Geobot. 9: 236 (1940)

species :

Dryopteris hangchowensis Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 414 (1938)

Distribution: China (NE-Zhejiang); South Korea

36 CHI 38 KOR

species :

Dryopteris hasseltii (Blume) C.Chr.; Index Filic. 269 (1905)

Distribution: China (Yunnan, Hainan); S-Japan (Yakushima Isl.); Ryukyu Isl. (Tokara Isl.); Taiwan; India (Assam, Arunachal Pradesh, Meghalaya, Mizoram); Thailand; Vietnam; peninsular Malaysia (Malacca, Perak, Pahang, Johor); Sumatra; Borneo (Mt. Kinabalu, Sabah); Sulawesi; Philippines; Java; ?Lesser Sunda Isl.; Moluccas; New Guinea (Irian Jaya, Papua New Guinea); Solomon Isl. (Guadalcanal); Australia (NE-Queensland); Vanuatu; Fiji (Viti Levu, Vanua Levu, Ovalau, Taveuni); Fed. States of Micronesia (Pohnpei)

36 CHI 38 JAP NNS TAI 40 IND 41 THA VIE 42 BOR JAW MLY MOL NWG PHI SUL SUM 43 SOL 50 AUS 60 FIJ VAN 62 MIC

Conservation Status: IUCN EN for India (Fraser-Jenkins 2012)

Synonyms: hidden

= Acrorumohra hasseltii (Blume) Ching; Acta Phytotax. Sin. 9: 385 (1964)

= Arachniodes hasseltii (Blume) Ching; Acta Bot. Sin. 10: 258 (1962)

= Athyrium hasseltii (Blume) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 6 (1940)

= Byrsopteris hasseltii (Blume) C.V.Morton; Amer. Fern J. 50: 154 (1960)

= Dryopteris adiantoides T.Suzuki; J. Jap. Bot. 11: 644 (1935)

= Dryopteris gillespiei Copel.; Bull. Bernice P. Bishop Mus. 69: 10 (1929)

= Dryopteris hatusimae H.Itô; Bot. Mag. (Tokyo) 67: 216, f. 2 (1954)

= Dryopteris laserpitiifolia (Scort. ex Bedd.) C.Chr.; Index Filic. 273 (1905), nom. illeg., non (Mett.) Kunze

= Dryopteris obovata (Baker) C.Chr.; Index Filic. 280 (1905)

= Dryopteris odontophora Copel.; Philipp. J. Sci. 60: 106 (1936)

= Dryopteris psilosora Tagawa; Acta Phytotax. Geobot. 2(3): 191 (1933)

= Dryopteris sparsa var. raapii Alderw.; Malayan Ferns 197 (1909)

= Lastrea anisoptera (Kunze) T.Moore; Index Fil. [T. Moore] 85 (1858)

= Lastrea obovata (Baker) Bedd.; Suppl. Ferns Brit. Ind. 60 (1892)

= Nephrodium obovatum Baker; J. Bot. 28: 265 (1890)

= Phegopteris hasseltii (Blume) Mett.; Abh. Senckenberg. Naturf. Ges. 2(2): [Pheg. u. Asp. 13, no. 14] (1858)

= Phegopteris laserpitiifolia (Scort. ex Bedd.) Bedd.; Suppl. Ferns Brit. Ind. 84 (1892)

= Polypodium anisopterum Kunze; Bot. Zeitung (Berlin) 118 (1848)

= Polypodium hasseltii Blume; Fl. Javae, Fil. 195 (1851)

= Polypodium laserpitiifolium Scort. ex Bedd.; J. Bot. 25: 324 (1887)

= Polystichopsis hasseltii (Blume) Holttum; Rev. Fl. Malaya 2: 487, f. 288 (1955)

= Rumohra hasseltii (Blume) Ching; Sinensia 6: 61 (1934)

species :

Dryopteris hawaiiensis (Hillebr.) W.J.Rob.; Bull. Torrey Bot. Club 39: 594 (1912)

Distribution: Hawaii (Kauai, Oahu, Molokai, Maui, Hawaii Isl.)

63 HAW

Remarks: ser. Aemulae

Synonyms: hidden

= Aspidium hawaiiense Hillebr.; Fl. Hawaiian Isl. [Hillebrand] 575 (1888)

species :

Dryopteris hendersonii (Bedd.) C.Chr.; Index Filic. 270 (1905)

Distribution: China (Yunnan, Guizhou); Japan; Taiwan; Nepal; Bhutan; India (Arunachal Pradesh, Manipur, Meghalaya, Nagaland, Sikkim, West Bengal); Myanmar; Thailand; Vietnam; Borneo (Mt. Kinabalu, Sabah); Philippines (Luzon)

36 CHI 38 JAP TAI 40 BHU IND NEP 41 MYA THA VIE 42 BOR PHI

Remarks: subgen. Nothoperanema

Synonyms: hidden

= Aspidium spectabile Wall.; Numer. List. [Wallich] no. 372 (1829), nom. inval., non Blume

= Ctenitis hendersonii (Bedd.) H.Itô; Nov. Fl. Jap. [Nakai & Honda] 4: 89 (1939)

= Ctenitis leptorhachis (Hayata) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 280 (1938)

= Ctenitis mearnsii Copel.; Philipp. J. Sci. 81: 24 (1952)

= Dryopteris leptorhachis Hayata; Icon. Pl. Formosan. 4: 162, f. 102 (1914)

= Lastrea hendersonii Bedd.; Ferns Brit. India Suppl., t. 377 (1876)

= Lastrea spectabilis (C.B.Clarke) Bedd.; Ferns Brit. India 257 (1883), non (Blume) J. Sm. 1841

= Lastrea wallichii T.Moore; Index Fil. [T. Moore] 104 (1858)

= Nephrodium spectabile C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 526 (1880), nom. illeg.

= Nothoperanema hendersonii (Bedd.) Ching; Acta Phytotax. Sin. 11: 28 (1966)

= Nothoperanema hendersonii (Bedd.) Nakaike; Enum. Pterid. Jap., Filic. 239 (1975), isonym

species :

Dryopteris herbacea Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 20: 13 (1915)

Distribution: Java

42 JAW

Remarks: provisionally accepted name

species :

Dryopteris heterolaena C.Chr.; Acta Horti Gothob. 1: 62, t. 17 (1924)

Distribution: China (Guangdong, Guangxi, Guizhou, Hunan, Sichuan, Yunnan, Zhejiang); Tibet; NE-India (Arunachal Pradesh)

36 CHI TIB 40 IND

Remarks: Subgen. Dryopsis

Conservation Status: IUCN CR for India (as C. fengiana, Fraser-Jenkins 2012)

Synonyms: hidden

= Ctenitis chingii Z.Y.Liu & J.I.Chang; Bull. Bot. Res., Harbin 4(4): 14, f. 42 (1984)

= Ctenitis fengiana Ching; Bull. Fan Mem. Inst. Biol. Bot. 11: 69 (1941)

= Ctenitis heterolaena (C.Chr.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 293 (1938)

= Ctenitis kwangsiensis Ching & P.S.Chiu; Acta Phytotax. Sin. 19(1): 120 (1981)

= Ctenitis omeiensis Ching & Chu H.Wang; Acta Phytotax. Sin. 19(1): 121 (1981)

= Ctenitis tibetica Ching & S.K.Wu; Fl. Xizangica 1: 274 (1983)

= Ctenitis zhejiangensis Ching & C.F.Zhang; Bull. Bot. Res., Harbin 3(3): 34-35, f. 25 (1983)

= Dryopsis fengiana (Ching) Holttum & P.J.Edwards; Kew Bull. 41(1): 183 (1986)

= Dryopsis heterolaena (C.Chr.) Holttum & P.J.Edwards; Kew Bull. 41(1): 185 (1986)

= Dryopsis kwangsiensis (Ching & Chu H.Wang) Holttum & P.J.Edwards; Kew Bull. 41(1): 186 (1986)

= Dryopteris nobilis var. fengiana Ching; Bull. Fan Mem. Inst. Biol. Bot. 11(2): 67 (1941)

species :

Dryopteris hezhangensis P.S.Wang; Pterid. Fl. Guizhou 315 (-317; fig. 81.1-2) (2001)

Distribution: China (Guizhou)

36 CHI

Remarks: provisionally accepted name, not mentioned in Fl. of China

species :

Dryopteris himachalensis Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.), Bot. 18(5): 367 (1989)

Distribution: China (NW-Yunnan, SW-Sichuan); India (Himachal Pradesh)

36 CHI 40 IND

Conservation Status: IUCN CR for India (Fraser-Jenkins 2012)

Synonyms: hidden

= Dryopteris pectinatopinnata Ching; Acta Phytotax. Sin. 23(5): 349-350 (1985), nom. nud.

species :

Dryopteris hirtipes (Blume) Kuntze; Revis. Gen. Pl. 2: 813 (1891)

Distribution: S-India (Kerala, Tamil Nadu); Sri Lanka; Myanmar; N-Thailand; Vietnam; peninsular Malaysia (Cameron Highlands, Pahang); Sumatra; Borneo (Sabah); Sulawesi; Philippines; Java; Lesser Sunda Isl. (Bali); New Guinea (Irian Jaya, Papua New Guinea); Vanuatu; Fiji (Viti Levu); [Western] Samoa (Savai´i, ´Upolu)

40 IND SRL 41 MYA THA VIE 42 BOR JAW LSI MLY NWG PHI SUL SUM 60 FIJ SAM VAN

Synonyms: hidden

= Aspidium aduncum Wall.; Numer. List. [Wallich] no. 384 (1829), nomen

= Aspidium hirtipes Blume; Enum. Pl. Javae 2: 148 (1828)

= Lastrea hirtipes (Blume) T.Moore; Index Fil. [T. Moore] 85 (1858)

species :

Dryopteris hondoensis Koidz.; Acta Phytotax. Geobot. 1: 31 (1932)

Distribution: China (Sichuan, Zhejiang); Japan

36 CHI 38 JAP

Synonyms: hidden

= Dryopteris scabripes Ching; Bot. Res. Acad. Sin. 1987(2): 9 (1987)

species :

Dryopteris huberi (Christ) C.Chr.; Index Filic., Suppl. 2 (1906-1912): 34 (1913)

Distribution: Brazil (Goiás, Mato Grosso, Pará)

84 BRA

Remarks: a species close to D. futura, recognized by Sessa et al. 2012

Synonyms: hidden

= Aspidium huberi Christ; Hedwigia 46: 192 (1906)

species :

Dryopteris huongii C.W.Chen; Syst. Bot. 48(2): 170 (2023)

Distribution: Vietnam

41 VIE

species :

Dryopteris hypolepioides Rosenst.; Repert. Spec. Nov. Regni Veg. 12: 175 (1913)

Distribution: New Guinea (Irian Jaya, Papua New Guinea)

42 NWG

Synonyms: hidden

= Phegopteris hypolepioides (Rosenst.) Alderw.; Malayan Ferns Fern Allies, Suppl. 309 (1917)

= Ctenitis hypolepioides (Rosenst.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 124 (1947)

species :

Dryopteris immixta Ching; Fl. Tsinling. 2: 225 (1974)

Distribution: China (Anhui, Fujian, Gansu, Guizhou, Hubei, Hunan, Jiangsu, Jiangxi, Shaanxi, Sichuan, Yunnan, Zhejiang); South Korea (incl. Cheju Isl.); Japan

36 CHI 38 JAP KOR

Remarks: D. pudouensis is frequently kept separate in China

Synonyms: hidden

= Dryopteris bissetiana var. hikonensis (H.Itô) H.Itô; Bot. Mag. (Tokyo) 50: 36 (1936)

= Dryopteris consimilis Ching & P.C.Chiu; Bot. Res. Acad. Sin. 1987(2): 25 (1987), nom. illeg.

= Dryopteris fuyangensis Ching & P.S.Chiu; Bot. Res. Acad. Sin. 1987(2): 26 (1987)

= Dryopteris hikonensis (H.Itô) Nakaike; New Fl. Japan, Pterid. 841 (1992)

= Dryopteris hikonensis monstr. cristata (T.Ueno) Nakaike; New Fl. Japan, Pterid. 841 (1992)

= Dryopteris lungjingensis Ching & P.S.Chiu; Bot. Res. Acad. Sin. 1987(2): 27 (1987)

= Dryopteris pacifica (Nakai) Tagawa; Col. Ill. Jap. Pterid. 211 (1959), nom. illeg., non Christ (1912)

= Dryopteris paravaria Ching & P.C.Chiu; Bot. Res. Acad. Sin. 1987(2): 22 (1987)

= Dryopteris pudouensis Ching; Bull. Bot. Res., Harbin 3(3): 11 (1983)

= Dryopteris quadrifida Ching ex K.H.Shing & J.F.Cheng; Jiangxi Sci. 8(3), 49 (1990)

= Dryopteris shanghaiensis Ching & P.C.Chiu; Bot. Res. Acad. Sin. 1987(2): 24 (1987)

= Dryopteris tieanzuensis Ching & P.C.Chiu; Bot. Res. Acad. Sin. 1987(2): 24 (1987)

= Dryopteris varia subsp. hikonensis (H.Itô) Sugim.; Keys Herb. Pl. Jap. Pterid. 281, 404 (1966)

= Dryopteris varia var. hikonensis (H.Itô) Sa.Kurata; Enum. Jap. Pterid. 304 (1961)

= Dryopteris yushanensis Ching & P.S.Chiu; Bot. Res. Acad. Sin. 1987(2): 28 (1987)

= Polystichum hololepis var. hikonense H.Itô; J. Jap. Bot. 10: 451 (1961)

= Polystichum pacificum Nakai; Bot. Mag. (Tokyo) 39: 119 (1925)

species :

Dryopteris inaequalis (Schltdl.) Kuntze; Revis. Gen. Pl. 2: 813 (1891)

Distribution: South Africa (KwaZulu-Natal, W-Cape Prov., E-Cape Prov.); Lesotho; Eswatini [Swaziland]

27 LES SAF SWZ

Synonyms: hidden

= Aspidium amplum Kuhn; Filic. Afr. 125 (1868)

= Aspidium inaequale Schltdl.; Adumbr. Pl. 23, t. 12 (1825)

= Lastrea inaequalis (Schltdl.) C.Presl; Tent. Pterid. 77 (1836)

= Nephrodium inaequale (Schltdl.) Hook.; Sp. Fil. [W. J. Hooker] 4: 125 (1862) (non Schrad. (1824))

= Polystichum inaequale (Schltdl.) Keyserl.; Polyp. Herb. Bunge 44 (1873)

species :

Dryopteris indonesiana Darnaedi, M.Kato & K.Iwats.; J. Jap. Bot. 64: 304 (1989)

Distribution: Sumatra; Java; Moluccas

42 JAW MOL SUM

species :

Dryopteris indusiata (Makino) Makino & Yamam.; Suppl. Ic. Pl. Formos. V: 2 (1932)

Distribution: China (Fujian, Guangxi, Guizhou, Hunan, Jiangxi, Sichuan, Yunnan, Zhejiang); Japan

36 CHI 38 JAP

Synonyms: hidden

= Dryopteris gymnosora var. indusiata (Makino) Makino ex Bonap.

= Dryopteris labordei var. indusiata (Makino) S.Sherizawa; J. Jap. Bot. 52: 16 (1977)

= Dryopteris subfuscipes Ching ex K.H.Shing & J.F.Cheng; Jiangxi Sci. 8(3): 49 (1990)

= Nephrodium gymnosorum var. indusiatum Makino; Bot. Mag. (Tokyo) 13: 65 (1899)

species :

Dryopteris insularis Kodama; Matsum., Icon. Pl. Koisikav. 2: 49, t. 109 (1913)

subspecies :

subsp. insularis

Distribution: Bonin Isl.

38 OGA

Conservation Status: VU in Japan. IUCN R

Synonyms: hidden

= Dryopteris varia var. insularis (Kodama) H.Ohba; Sci. Rep. Tohoku Imp. Univ., Ser. 4, Biol. 36: 113 (1971)

subspecies :

subsp. chichishimensis (Nakai ex H.Itô) H.Itô; Nov. Fl. Jap. [Nakai & Honda] 4: 57 (1939)

Distribution: Japan (Izu Isl.); Bonin Isl. (Chichijima)

38 JAP OGA

Synonyms: hidden

= Dryopteris chichishimensis Nakai ex H.Itô; Bot. Mag. (Tokyo) 40: 435, f. 6) 7(14) (1935)

species :

Dryopteris integriloba C.Chr.; Bull. Biol. Dept. Sci. Coll. Sun Yatsen Univ. 6: 5 (1933)

Distribution: China (Guangdong, Hainan, Guangxi, Yunnan); Japan; Taiwan; Thailand; N-Vietnam; peninsular Malaysia (Negeri Sembilan)

36 CHI 38 JAP TAI 41 THA VIE 42 MLY

Synonyms: hidden

= Dryopteris subintegriloba Seriz.; J. Jap. Bot. 46: 17, f.1a (1971)

species :

Dryopteris intermedia (Muhl. ex Willd.) A.Gray; Manual (Gray) 630 (1848)

subspecies :

subsp. intermedia

Name: Fancy fern, Intermediate woodfern (EN)

Distribution: Canada (New Brunswick, Newfoundland, Nova Scotia, Ontario, Prince Edward Isl., Québec); St. Pierre et Miquelon; USA (Alabama, Connecticut, District of Columbia, Delaware, Georgia, Iowa, Illinois, Indiana, Kentucky, Massachusetts, Maryland, Maine, Michigan, Minnesota, Missouri, North Carolina, New Hampshire, New Jersey, New York, Ohio, Pennsylvania, Rhode Island, South Carolina, Tennessee, Virginia, Vermont, Wisconsin, West Virginia)

72 CDN SPM 74 USA 75 USA 78 USA

Synonyms: hidden

= Aspidium americanum Davenp.; Amer. Naturalist 12: 714 (1878)

= Aspidium grayanum Regel; Index Sem. (LE, Petropolitanus) (1855)

= Aspidium intermedium Muhl. ex Willd.; Sp. Pl., ed. 4 [Willdenow] 5(1): 262 (1810)

= Dryopteris austriaca var. intermedia (Muhl. ex Willd.) C.V.Morton; Amer. Fern J. 40: 222 (1950)

= Dryopteris intermedia f. furcata C.F.Reed; Castanea 16: 70, fig. 5 (1951)

= Dryopteris intermedia f. sparsifrondosa C.F.Reed; Castanea 16: 71, fig. 6 (1951)

= Dryopteris spinulosa var. intermedia (Muhl. ex Willd.) Underw.; Native Ferns, ed. 4 116 (1893)

= Filix spinulosa var. intermedia (Muhl. ex Willd.) Farw.; Rep. (Annual) Michigan Acad. Sci. 18: 83 (1916)

= Filix-mas spinulosa var. intermedia (Muhl. ex Willd.) Farw.; Amer. Midl. Naturalist 12: 257 (1931)

= Lastrea intermedia (Muhl. ex Willd.) C.Presl; Tent. Pterid. 77 (1836)

= Nephrodium intermedium (Muhl. ex Willd.) C.Presl; Reliq. Haenk. 1: 38 (1826)

= Nephrodium spinulosum f. intermedium (Muhl. ex Willd.) Gilbert; Fern Bull. 13: 37 (1905)

= Nephrodium spinulosum var. intermedium (Muhl. ex Willd.) Davenp.; Working List N. Amer. Pterid. 17 (1901)

= Thelypteris intermedia (Muhl. ex Willd.) House; New York State Mus. Bull. 233-234: 69 (1920)

= Thelypteris spinulosa var. intermedia (Muhl. ex Willd.) Nieuwl.; Amer. Midl. Naturalist 2: 278 (1912)

subspecies :

subsp. azorica (Christ) Jermy; Webbia 38: 408 (1984)

Distribution: Azores (Santa Maria Isl., Sao Miguel Isl., Terceira, Sao Jorge, Pico, Faial, Flores Isl., Corvo Isl.)

21 AZO

Synonyms: hidden

= Aspidium dilatatum var. azoricum Christ; Bull. Acad. Int. Géogr. Bot. 17 (212): 157 (1907)

= Dryopteris azorica (Christ) Alston; Bol. Soc. Brot., ser. 2, 30: 14 (1956)

subspecies :

subsp. maderensis (Milde ex Alston) Fraser-Jenk.; Arquipelago, Sér. Ci. Nat. 2: 85 (1981)

Distribution: Madeira (Madeira Isl.)

21 MDR

Synonyms: hidden

= Aspidium spinulosum f. maderensis Milde; Fil. Eur. 140 (1867)

= Dryopteris maderensis Milde ex Alston; Bol. Soc. Brot., ser. 2, 30: 14 (1956)

species :

Dryopteris inuyamensis H.Itô; J. Jap. Bot. 9: 54 (1933)

Distribution: Japan

38 JAP

species :

Dryopteris iranica Fraser-Jenk.; Fern Gaz. 18(1): 9 (2007)

Distribution: Turkey (Anatolia); Iran

34 IRN TUR

Remarks: affinis agg.

Synonyms: hidden

= Dryopteris affinis subsp. coriacea Fraser-Jenk.; Willdenowia 10(1): 112 (1980)

= Dryopteris affinis var. gigantea Fraser-Jenk.; Willdenowia 10: 113 (1980)

= Dryopteris coriacea (Fraser-Jenk.) Tzvelev; Novosti Sist. Vyssh. Rast. 35: 17 (2003), nom. illeg., non (Brause) Ching

= Dryopteris wallichiana subsp. coriacea (Fraser-Jenk.) Fraser-Jenk.; Thaiszia 4(1): 23 (1994)

species :

Dryopteris jinpingensis Z.Y.Zuo, Jin Mei Lu & D.Z.Li; PhytoKeys 239: 197 (2024)

Distribution: China (Yunnan)

36 CHI

Remarks: sect. Diclisodon

species :

Dryopteris jishouensis G.X.Chen & D.G.Zhang; Life Sci. Res. 17(5): 412, figs. 1-2 (2013)

Distribution: China (Hunan)

36 CHI

species :

Dryopteris jiucaipingensis P.S.Wang, Q.Luo & Li Bing Zhang; Novon 22(2): 183 (2012)

Distribution: China (NW-Guizhou)

36 CHI

species :

Dryopteris juxtaposita Christ; Bull. Acad. Int. Géogr. Bot. 138 (1907)

Distribution: China (Yunnan, Sichuan, Guizhou, Gansu); Tibet; Nepal; Bhutan; India (Himachal Pradesh, Arunachal Pradesh, Karnataka, Kerala, Manipur, Meghalaya, Nagaland, Sikkim, Tamil Nadu, Uttarakhand, West Bengal, Jammu & Kashmir); Myanmar; Vietnam

36 CHI TIB 40 BHU IND NEP 41 MYA VIE

Synonyms: hidden

= Aspidium filix-mas var. normale (C.B.Clarke) Christ; Mém. Soc. Bot. France 1 [Hist. Acrostich.]: 38 (1905)

= Nephrodium filix-mas var. normale C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 519, t. 68, f. 2 (1880)

species :

Dryopteris karwinskyana (Mett.) Kuntze; Revis. Gen. Pl. 2: 813 (1891)

Distribution: +Easter Isl.; Mexico (Chiapas, Ciudad de Mexico, Durango, Guerrero, Jalisco, Mexico State, Michoacan, Morelos, Nayarit, Oaxaca, Sinaloa); Guatemala; El Salvador; Honduras; Nicaragua; Costa Rica

61 EAS 79 MEX 80 COS ELS GUA HON NIC

Synonyms: hidden

= Aspidium karwinskyanum Mett.; Abh. Senckenberg. Naturf. Ges. 2(2) [Pheg. u. Asp.]: 343 (1858)

= Dryopteris espinosae Hicken; Revista Chilena Hist. Nat. 17: 93, t. 9 (1913)

= Lastrea karwinskiana (Mett.) T.Moore; Index Fil. [T. Moore] 94 (1858)

= Nephrodium karwinskyanum (Mett.) Baker; Syn. Fil. [W. J. Hooker & Baker] 279 (1867)

= Nephrodium karwinskyanum f. grande Donn.Sm.; Enum. Pl. Guatem. 4: 172 (1895)

= Thelypteris espinosae (Hicken) R.R.Rodriguez; Bol. Soc. Biol. Concepcion 45: 153 (1972)

species :

Dryopteris kashmiriana Fraser-Jenk. & Widén; Ferns and Fern-allies of Nepal 1: 26 (2015)

Distribution: India (Jammu & Kashmir, Uttarakhand)

40 IND

species :

Dryopteris katangaensis J.P.Roux; Bot. J. Linn. Soc. 142(2): 237 (2003)

Distribution: D.R.Congo [Zaire]; Zambia

23 ZAI 26 ZAM

species :

Dryopteris kawakamii Hayata; J. Coll. Sci. Imp. Univ. Tokyo 30: 416 (1911)

Distribution: China (Chongqing, Fujian, Guangdong, Guangxi, Guizhou, Hunan, Jiangxi, Sichuan, Yunnan, Zhejiang); Tibet; Taiwan; Myanmar

36 CHI TIB 38 TAI 41 MYA

Remarks: Subgen. Dryopsis. D. mariformis is possibly a separate species

Synonyms: hidden

= Aspidium polylepis Christ; (1905) (non Franch. & Sav. 1879)

= Ctenitis jiulungshanensis P.C.Chiu & G.H.Yao; Bull. Bot. Res., Harbin 2(2): 66 (1982)

= Ctenitis kawakamii (Hayata) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 286 (1938)

= Ctenitis mariformis (Rosenst.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 286 (1938)

= Ctenitis sacholepis (Hayata) H.Itô; Nov. Fl. Jap. [Nakai & Honda] 4: 80 (1939)

= Ctenitis truncata Ching & Chu H.Wang; Acta Phytotax. Sin. 19(1): 123 (1981)

= Dryopsis kawakamii (Hayata) Holttum & P.J.Edwards; Kew Bull. 41(1): 186 (1986)

= Dryopsis mariformis (Rosenst.) Holttum & P.J.Edwards; Kew Bull. 41(1): 188 (1986)

= Dryopsis truncata (Ching & H.S.Kung) Holttum & P.J.Edwards; Kew Bull. 41(1): 188 (1986)

= Dryopteris mariformis Rosenst.; Repert. Spec. Nov. Regni Veg. 13: 131 (1914)

= Dryopteris sacholepis Hayata; Icon. Pl. Formosan. 6: 285, f. 111 (1916)

species :

Dryopteris khullarii Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.), Bot. 18(5): 362 (1989)

Distribution: Nepal; NW-India (Himachal Pradesh, Uttarakhand)

40 IND NEP

Conservation Status: IUCN EN for India (Fraser-Jenkins 2012)

species :

Dryopteris kilemensis (Kuhn) Kuntze; Revis. Gen. Pl. 2: 813 (1891) (kilmensis)

Distribution: Cameroon; D.R.Congo [Zaire]; Rwanda; Burundi; Sudan; Ethiopia; Uganda; Kenya; Tanzania; Zambia; Malawi; Zimbabwe; Mozambique; Madagascar

23 BUR CMN RWA ZAI 24 ETH SUD 25 KEN TAN UGA 26 MLW MOZ ZAM ZIM 29 MDG

Synonyms: hidden

= Aspidium kilemense Kuhn; Reisen Ost-Afr. [Decken] 3(3): Bot. 46 (1879)

= Aspidium lastii (Baker) Hieron.; Pflanzenw. Ost-Afrikas [Engler] C: 85 (1895)

= Dryopteris cordipinnula C.Chr.; Cat. Pl. Madag., Pterid. 27 (1932) (nomen), Dansk Bot. Ark. 7: 55, t. 11, f. 10-12 (1932)

= Dryopteris lastii (Baker) C.Chr.; Index Filic. 274 (1905)

= Dryopteris platylepis Rosenst.; Repert. Spec. Nov. Regni Veg. 4: 4, 5 (1907)

= Dryopteris remotipinnula Bonap.; Notes Ptéridol. [Bonaparte] 16: 178 (1925)

= Nephrodium kilemense (Kuhn) Baker; Syn. Fil. [W. J. Hooker & Baker], ed. 2: 498 (1874)

= Nephrodium lastii Baker; Ann. Bot. (Oxford) 6: 324 (1891)

species :

Dryopteris kinkiensis Koidz.; Tagawa, Acta Phytotax. Geobot. 2(3): 200 (1933)

Distribution: China (Fujian, Guangdong, Zhejiang, Jiangxi, Sichuan); South Korea (incl. Cheju Isl.); Japan; Myanmar (Kachin, etc.)

36 CHI 38 JAP KOR 41 MYA

Synonyms: hidden

= Dryopteris championii var. tenuifrons (H.Itô) H.Itô; Nov. Fl. Jap. [Nakai & Honda] 4: 48 (1939)

= Dryopteris hunanica Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 179 (1940) (fide Fraser-Jenkins 1986)

= Dryopteris neoassamensis Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 480 (1938)

= Dryopteris pseudoerythrosora var. tenuifrons H.Itô

= Dryopteris zhenangensis Ching & P.S.Chiu; Bot. Res. Acad. Sin. 1987(2): 11 (1987)

species :

Dryopteris kinokuniensis Sa.Kurata; J. Geobot. 18: 6 (1970)

Distribution: Japan

38 JAP

species :

Dryopteris knoblochii A.R.Sm.; Mem. New York Bot. Gard. 88 [Pterid. Mexico]: 276 (2004)

Distribution: USA (Arizona); Mexico (Chihuahua, Durango, Sinaloa, Sonora)

76 USA 79 MEX

species :

Dryopteris koidzumiana Tagawa; Acta Phytotax. Geobot. 2(3): 190 (1933)

Distribution: Japan

38 JAP

Synonyms: hidden

= Dryopteris erythrosora var. koidzumiana (Tagawa) H.Hara; (1939)

species :

Dryopteris komarovii Kossinsky; Bot. Mater. Gerb. Glavn. Bot. Sada R.S.F.S.R.2: 1 (1921)

Distribution: Turkmenistan; Uzbekistan; Kyrgyzstan; Tajikistan; China (Gansu, Qinghai, Shaanxi, Sichuan, Yunnan); Tibet; Taiwan; Pakistan (Kurram, Hazara); Nepal; Bhutan; India (Arunachal Pradesh, Jammu & Kashmir, Himachal Pradesh, Sikkim, Uttarakhand); Myanmar

32 KGZ TKM TZK UZB 36 CHI TIB 38 TAI 40 BHU IND NEP PAK 41 MYA

Synonyms: hidden

= Dryopteris barbigera subsp. komarovii (Kossinsky) Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.), Bot. 18(5): 384 (1989)

= Dryopteris subbarbigera Ching; Fl. Xizangica 1: 260 (1983)

species :

Dryopteris kwanzanensis Tagawa; Acta Phytotax. Geobot. 7: 186 (1938)

Distribution: Taiwan

38 TAI

Synonyms: hidden

= Dryopteris amoena Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 454 (1938) (fide Fraser-Jenkins 1986)

species :

Dryopteris labordei (Christ) C.Chr.; Index Filic. 273 (1905)

Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Hubei, Hunan, Jiangxi, Sichuan, Yunnan, Zhejiang); Japan

36 CHI 38 JAP

Remarks: not in Taiwan

Synonyms: hidden

= Aspidium labordei Christ; Mém. Soc. Bot. France 1 [Hist. Acrostich.]: 40 (1905)

species :

Dryopteris lacera (Thunb.) Kuntze; Revis. Gen. Pl. 2: 813 (1891)

Distribution: China (Heilongjiang, Hubei, Hunan, Jiangxi, Sichuan, Zhejiang); North Korea; South Korea; Japan; Taiwan

36 CHI 38 JAP KOR NKO TAI

Synonyms: hidden

= Aspidium filix-mas var. lacerum (Thunb.) Christ; Farnkr. Erde 257 (1897)

= Aspidium lacerum (Thunb.) Sw.; J. Bot. (Schrader) 1800(2): 39 (1801)

= Aspidium lacerum var. bipinnatum Franch. & Sav.; Enum. Pl. Jap. 2: 238 (1876)

= Aspidium lacerum var. obtusum Christ; Mém. Soc. Bot. France 1 [Hist. Acrostich.]: 39 (1905)

= Dryopteris filix-mas var. lacerum (Thunb.) Christ; (1897)

= Dryopteris lacera var. chinensis Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 439 (1938)

= Lastrea lacera (Thunb.) D.C.Eaton; Proc. Amer. Acad. Arts 4: 110 (1858)

= Nephrodium lacerum (Thunb.) Baker; Syn. Fil. [W. J. Hooker & Baker] 273 (1867)

= Polypodium lacerum Thunb.; Fl. Jap. [Thunberg] 337 (1784) et Syst. Veg., ed. 14 (J. A. Murray): 938 (1784)

= Polystichum lacerum C.Presl; Abh. Königl. Böhm. Ges. Wiss., ser. 5, 6 [Epim. Bot.]: 56 (1851)

species :

Dryopteris lachoongensis (Bedd.) B.K.Nayar & Kaur; Companion Ferns Brit. India 61 (1974)

Distribution: China (Yunnan); Tibet; Taiwan; Nepal; ?Bhutan; India

36 CHI TIB 38 TAI 40 IND NEP

Remarks: valid species separate from D. fructuosa - where it was included by Fraser-Jenkins - fide Knapp & Hsu (2017)

Synonyms: hidden

= Lastrea filix-mas var. lachoongensis Bedd.; Ferns Brit. India Suppl. 58 (1892) (fide Fraser-Jenkins 1997)

= Dryopteris pseudodontoloma (Christ) Ching; Fl. Xizangica 1: 265 (1983)

= Dryopteris venosa Ching & S.K.Wu; Fl. Xizangica 1: 264 (1983)

species :

Dryopteris lacunosa S.Jess., Zenner, Chr.Stark & Bujnoch; Kochia 5: 21 (2011)

Name: Lückiger Wurmfarn, Lückiger Schuppen-Wurmfarn (DE)

Distribution: England (England, Scotland); Ireland (Ireland); Germany (Baden-Württemberg, Bayern, Niedersachsen, Nordrhein-Westfalen, Rheinland-Pfalz, Saarland); Switzerland; Austria; France; Italy

10 GRB IRE 11 AUT GER SWI 12 FRA 13 ITA

Remarks: affinis agg.

species :

Dryopteris laeta (Kom.) C.Chr.; Index Filic. 267 (1905)

Distribution: Russian Far East (Khabarovsk, Primorye); China (Gansu, Hebei, Henan, Qinghai, Liaoning, Nei Mongol); Japan

31 RFE 36 CHI 38 JAP

Remarks: often included in D. goeringiana

Synonyms: hidden

= Aspidium oxyodon Franch.; Nouv. Arch. Mus. Hist. Nat., sér. 2, 7: 163 (1883) (non Dryopteris oxyodon (Baker) C. Chr.)

= Athyrium nakaii Tagawa; Acta Phytotax. Geobot. 2(3): 195 (1933), nom. nov.

= Athyrium triangulare Nakai; Bot. Mag. (Tokyo) 41: 91 (1931) (non Alderw. (1915))

= Dryopteris filix-mas subsp. oxyodon (Franch.) C.Chr.; Index Filic. 5: 266 (1905)

= Dryopteris laeta f. oblongifrons (Kitag.) Kitag.; J. Jap. Bot. 41: 368 (1966)

= Dryopteris laeta f. triangularis (Nakai) Kitag.; Neo-Lineam. Fl. Manshur.: 35 (1979)

= Dryopteris laeta var. alpina Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 466 (1938)

= Dryopteris laeta var. oblongifrons Kitag.; Bot. Mag. (Tokyo) 48(566): 91, f. 8 (1934)

= Dryopteris oxyodon (Franch.) Kitag.; Rep. First Sci. Exped. Manchoukuo 4(2): 58 (1935)

= Dryopteris oxyodon var. oblongifrons (Kitag.) Kitag.; Rep. First Sci. Exped. Manchoukuo 4(2): 59 (1935)

= Dryopteris wladiwostokensis B.Fedtsch.; Trudy Imp. S.-Peterburgsk. Bot. Sada 31: 4 (1912)

= Nephrodium laetum Kom.; Fl. Mansh. 1: 124 (1901)

= Nephrodium wladiwostokense (B.Fedtsch.) B.Fedtsch.; Trudy Imp. S.-Peterburgsk. Bot. Sada 31: 99 (1912)

species :

Dryopteris lepidopoda Hayata; Icon. Pl. Formosan. 4: 161 (1914)

Distribution: China (N-Yunnan, W-Sichuan); Tibet; Taiwan; Pakistan (Hazara); Nepal; Bhutan; India (Jammu & Kashmir, Himachal Pradesh, Arunachal Pradesh, Assam State, ?Manipur, Meghalaya, Nagaland, Sikkim, Uttarakhand, West Bengal); Myanmar (Chin)

36 CHI TIB 38 TAI 40 BHU IND NEP PAK 41 MYA

Synonyms: hidden

= Christella khasiana (C.B.Clarke) H.Lév.; Fl. Kouy-Tcheou 474-476 (1915)

= Dryopteris filix-mas subsp. khasiana (C.B.Clarke) C.Chr.; Index Filic. 5: 265 (1905)

= Dryopteris junlianensis H.S.Kung; Acta Bot. Yunnan. 4(4): 340 (1982)

= Dryopteris latibasis Ching; Bull. Fan Mem. Inst. Biol. Bot. 11: 61 (1941) (fide Fraser-Jenkins et al. 2018)

= Dryopteris lepidopoda var. phaeocoma Ching & S.K.Wu; Fl. Xizangica 1: 258 (1983)

= Dryopteris longistipes Ching; Bull. Fan Mem. Inst. Biol. Bot. 11: 59 (1941)

= Dryopteris neolepidopoda Ching & S.K.Wu; Fl. Xizangica 1: 258 (1983)

= Dryopteris nigra Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 430 (1938), nom. superfl.

= Dryopteris paleacea var. khasiana (C.B.Clarke) C.Chr.; (1931) (non D. khasiana C. Chr. 1905)

= Dryopteris taiwanicola Tagawa; Acta Phytotax. Geobot. 8: 230 (1939)

= Nephrodium filix-mas var. khasianum C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 519 (1880)

= Nephrodium parallelogrammum var. khasianum (C.B.Clarke) C.Hope; J. Bombay Nat. Hist. Soc. 14: 729 (1903)

species :

Dryopteris lepidorachis C.Chr.; Index Filic. 5: 274 (1905), nom. nov.

Distribution: China (Anhui, Fujian, Jiangsu, Jiangxi, Zhejiang)

36 CHI

Synonyms: hidden

= Dryopteris basitripinnatifida Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(4): 6 (1984)

= Dryopteris championii var. rheosora (Baker) K.H.Shing; Fl. Jiangxi 1: 282 (1993)

= Dryopteris lepidocaulon Ching & P.S.Chiu; Bot. Res. Acad. Sin. 1987(2): 19 (1987), non Kuntze (1891)

= Nephrodium rheosorum (Baker) Hand.-Mazz.; Symb. Sin. 6: 24 (1929), based on Polypodium rheosorum Baker (1891), nom. nud.

= Polypodium rheosorum Baker; Ann. Bot. (Oxford) 5: 457 (1891) (non Baker (1884))

species :

Dryopteris lepinei (Kuhn) Kuntze; Revis. Gen. Pl. 2: 813 (1891)

Distribution: Society Isl. (Tahiti)

61 SCI

Synonyms: hidden

= Aspidium lepinei Kuhn; Linnaea 36: 118 (1869)

= Nephrodium lepinei (Kuhn) Baker; Syn. Fil. [W. J. Hooker & Baker], ed. 2: 501 (1874)

species :

Dryopteris leucorhachis (Cheeseman) C.Chr.; Index Filic. 274 (1905)

Distribution: Cook Isl. (Rarotonga)

61 COO

Synonyms: hidden

= Acrophorus leucorhachis (Cheeseman) Brownlie; Brownlie & Philipson, Pacific Sci. 25: 507 (1971)

= Nephrodium leucorhachis Cheeseman; Trans. Linn. Soc. London, Bot., ser. 2, 6: 309 (1903)

species :

Dryopteris lewalleana Pic.Serm.; Bull. Jard. Bot. Natl. Belg. 55(1-2): 158 (1985)

Distribution: D.R.Congo [Zaire]; Rwanda; Burundi; Ethiopia; Uganda; Kenya; Tanzania; Zambia; Malawi; Zimbabwe; Mozambique; South Africa (Limpopo, Mpulamanga, KwaZulu-Natal, E-Cape Prov., W-Cape Prov.); Lesotho; Eswatini [Swaziland]

23 BUR RWA ZAI 24 ETH 25 KEN TAN UGA 26 MLW MOZ ZAM ZIM 27 LES SAF SWZ

Synonyms: hidden

= Aspidium pentagonum (T.Moore) Kuhn; Filic. Afr. 139 (1868)

= Dryopteris inaequalis var. atropaleacea Schelpe; Bol. Soc. Brot., ser. 2, 41: 213 (1967)

= Lastrea pentagona T.Moore; Hooker´s J. Bot. Kew Gard Misc. 5: 227 (1853)

species :

Dryopteris liangkwangensis Ching; Icon. Filic. Sin. 4: t. 180 (1937) (liankwangensis)

Distribution: China (Guangxi, Guangdong, SE-Yunnan); Vietnam

36 CHI 41 VIE

species :

Dryopteris liboensis P.S.Wang, X.Y.Wang & Li Bing Zhang; Novon 22(2): 256 (2012)

Distribution: China (S-Guizhou)

36 CHI

species :

Dryopteris lijianxiui Xiao J.Li; Bangladesh J. Bot. 50(3): 747 (2021)

Distribution: China (Shandong)

36 CHI

species :

Dryopteris loxoscaphoides (Baker) C.Chr.; Index Filic. 276 (1905)

Distribution: New Guinea (Papua New Guinea)

42 NWG

Remarks: the genus and family of this species is disputed. It might belong to Davallia.

Synonyms: hidden

= Acrophorus loxoscaphoides (Baker) Alston; J. Bot. 77: 288 (1939)

= Davallia forbesii A.Gepp; J. Bot. 61(Suppl.): 59 (1923)

= Leucostegia loxoscaphoides (Baker) C.Chr.; Index Filic., Suppl. Tert.: 120 (1934)

= Phegopteris loxoscaphoides (Baker) Alderw.; Malayan Ferns 493 (1909)

= Polypodium loxoscaphoides Baker; J. Bot. 28: 107 (1890)

species :

Dryopteris ludoviciana (Kunze) Small; Ferns S. E. States [Small] 281 (1938)

Name: Southern woodfern

Distribution: USA (Alabama, Arkansas, Florida, Georgia, Kentucky, Louisiana, Mississippi, North Carolina, South Carolina, Texas)

77 USA 78 USA

Synonyms: hidden

= Aspidium cristatum var. floridanum (Hook.) Christ; Farnkr. Erde: 258 (1897)

= Aspidium floridanum (Hook.) D.C.Eaton; Fl. South. U.S. [Chapman] 595 (1860)

= Aspidium ludovicianum Kunze; Amer. J. Sci. Arts, ser. 2, 6: 84 (1848)

= Dryopteris floridana (Hook.) Kuntze; Revis. Gen. Pl. 2: 812 (1891)

= Filix floridana (Hook.) Farw.; Rep. (Annual) Michigan Acad. Sci. 18: 81-83 (1916)

= Filix-mas cristata var. floridana (Hook.) Farw.; Amer. Midl. Naturalist 12: 254 (1931)

= Lastrea floridana (Hook.) K.Koch; Wochenschr. Vereines Beförd. Gartenbaues Königl. Preuss. Staaten 12: 143 (1869)

= Lastrea floridanum (Hook.) J.Sm.; Ferns Brit. Foreign, ed. 2: 306 (1877), isonym

= Lastrea petiolata Riddell; New Orleans Med. Surg. J. 9: 617 (1853)

= Nephrodium floridanum Hook.; Fil. Exot.: t. 99 (1859)

species :

Dryopteris lunanensis (Christ) C.Chr.; Index Filic. 276 (1905)

Distribution: China (Gansu, Guizhou, Sichuan, Yunnan); Bhutan; India (Arunachal Pradesh)

36 CHI 40 BHU IND

Synonyms: hidden

= Aspidium lunanense Christ; Bull. Herb. Boissier, sér. 1, 6: 966 (1898)

= Dryopteris lungnanensis Ching ex K.H.Shing & J.F.Cheng; Jiangxi Sci. 8(3): 48 (1990), orth. var.

= Dryopteris paralunanensis W.M.Chu ex S.G.Lu; Guihaia 11: 225 (1991)

= Dryopteris semipinnata Ching; Fl. Tsinling. 2: 226 (1974)

species :

Dryopteris lustrata (Hieron.) C.Chr.; Index Filic. 276 (1905)

Distribution: Colombia (Nariño)

82 CLM

Synonyms: hidden

= Nephrodium lustratum Hieron.; Bot. Jahrb. Syst. 34(4): 443 (1904)

species :

Dryopteris macrochlamys (Fée) Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.), Bot. 18(5): 453 (1989)

Distribution: Sri Lanka

40 SRL

Remarks: D. undulata is sometimes treated as separate species

Synonyms: hidden

= Acrorumohra obtusissima (Mett.) Ching; Acta Phytotax. Sin. 9: 385 (1964) (fide Fraser-Jenkins 1997)

= Acrorumohra undulata (Bedd.) Ching; Acta Phytotax. Sin. 9: 385 (1964)

= Arachniodes obtusissima (Mett.) Ching; Acta Bot. Sin. 10: 259 (1962)

= Arachniodes zeylanica (Ching) Ching; Acta Bot. Sin. 10: 260 (1962) (fide Fraser-Jenkins 1997)

= Aspidium macrochlamys Fée; Mém. Foug. 5 (Gen. Filic.): 296 (1852)

= Aspidium obtusissimum Mett.; Kuhn, Linnaea 36: 119 (1869)

= Aspidium undulatum (Bedd.) Thwaites; Enum. Pl. Zeyl. [Thwaites]: 444 (1864) (non Afzel. ex Sw. (1801))

= Dryopteris deparioides subsp. gracillima (Ching) Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.), Bot. 18(5): 449 (1989) (fide Fraser-Jenkins 1997)

= Dryopteris gracillima Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 452 (1938) (fide Fraser-Jenkins 1997)

= Dryopteris gracillima var. prolongata Sledge; Bull. Brit. Mus. (Nat. Hist.), Bot. 5(1): 19 (1973)

= Dryopteris gracillima var. triangularis Sledge; Bull. Brit. Mus. (Nat. Hist.), Bot. 5(1): 19 (1973)

= Dryopteris obtusissima (Mett.) Christ; J. Bot. (Morot) 21: 231 (1908)

= Dryopteris simulans Kuntze; Revis. Gen. Pl. 2: 813 (1891)

= Dryopteris sri-lankensis Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.), Bot. 18(5): 451 (1989), nom. nov. superfl.

= Dryopteris undulata (Bedd.) Kuntze; Revis. Gen. Pl. 2: 814 (1891)

= Lastrea deltoidea Bedd.; Ferns S. India t. 248 (1863) (non (Sw.) T. Moore)

= Lastrea filix-mas var. minor Bedd.; Ferns Brit. India 254 (1883)

= Lastrea undulata Bedd.; Ferns S. India t. 271 (1863)

= Nephrodium sparsum var. minus Thwaites ex Baker; Syn. Fil. [W. J. Hooker & Baker], ed. 2: 498 (1874)

= Nephrodium undulatum (Bedd.) Baker; Syn. Fil. [W. J. Hooker & Baker] 276 (1867)

= Rumohra obtusissima (Mett.) Ching; Sinensia 6: 62 (1934)

= Rumohra zeylanica Ching; Sinensia 5: 70 (1934) (fide Fraser-Jenkins 1997)

species :

Dryopteris macrolepidota Copel.; Bull. Bernice P. Bishop Mus. 98: 8, t. 7 B (1932)

Distribution: Society Isl. (Tahiti)

61 SCI

species :

Dryopteris macropholis Lorence & W.L.Wagner; PhytoKeys 4: 15 (2011)

Distribution: Marquesas Isl. (Ua Huka, Nuku Hiva, Ua Pou, Hiva Oa, Tahuata)

61 MRQ

species :

Dryopteris mangindranensis Tardieu; Notul. Syst. (Paris) 15: 161 (1956)

Distribution: Madagascar

29 MDG

species :

Dryopteris manipurensis (Bedd.) C.Chr.; Index Filic. 276 (1905)

Distribution: Nepal; Bhutan; NE-India (Manipur, Arunachal Pradesh, Meghalaya); Philippines; New Guinea (Papua New Guinea)

40 BHU IND NEP 42 NWG PHI

Remarks: subgen. Nothoperanema, sect. Shiehia

Conservation Status: IUCN EN for India (Fraser-Jenkins 2012)

Synonyms: hidden

= Ctenitis copelandii (Christ) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 124 (1947)

= Ctenitis manipurensis (Bedd.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 297 (1938)

= Dryopsis manipurensis (Bedd.) Holttum & P.J.Edwards; Kew Bull. 41(1): 200 (1986)

= Dryopteris copelandii Christ; Philipp. J. Sci. 2: 216 (1907)

= Phegopteris copelandii (Christ) Alderw.; Malayan Ferns 492 (1909)

= Phegopteris manipurensis (Bedd.) Bedd.; Suppl. Ferns Brit. Ind. 83 (1892)

= Polypodium manipurense Bedd.; J. Bot. 26: 235 (1888)

species :

Dryopteris manniana (Hook.) C.Chr.; Index Filic. 276 (1905)

Distribution: Guinea; Sierra Leone; Liberia; Ivory Coast; Nigeria; Cameroon; Bioko Isl. [Fernando Poo]; São Tomé; Equatorial Guinea (Rio Muni); D.R.Congo [Zaire]; Rwanda; Burundi; Uganda; Kenya; Tanzania; Malawi; Zimbabwe; Mozambique; Madagascar

22 GUI IVO LBR NGA SIE 23 BIO BUR CMN EQG RWA SAO ZAI 25 KEN TAN UGA 26 MLW MOZ ZIM 29 MDG

Synonyms: hidden

= Dryopteris raynalii Tardieu; Adansonia, sér. 2, 5: 200, t. 2, f. 1-4 (1965)

= Phegopteris manniana (Hook.) Kuhn; Filic. Afr. 123 (1868)

= Polypodium mannianum Hook.; Sp. Fil. [W. J. Hooker] 4: 253 (1862)

species :

Dryopteris marginalis (L.) A.Gray; Manual (Gray) ed. 1: 632 (1848)

Name: Marginal woodfern

Distribution: Greenland; Canada (British Columbia, New Brunswick, Newfoundland, Nova Scotia, Ontario, Québec); USA (Alabama, Arkansas, Connecticut, District of Columbia, Delaware, Georgia, Iowa, Illinois, Indiana, Kansas, Kentucky, Massachusetts, Maryland, Maine, Michigan, Minnesota, Missouri, Mississippi, North Carolina, New Hampshire, New Jersey, New York, Ohio, Oklahoma, Pennsylvania, Rhode Island, South Carolina, Tennessee, Virginia, Vermont, Wisconsin, West Virginia)

70 GNL 71 CDN 72 CDN 74 USA 75 USA 78 USA

Synonyms: hidden

= Aspidium filix-mas var. marginale (L.) Christ; Farnkr. Erde 257 (1897)

= Aspidium marginale (L.) Sw.; J. Bot. (Schrader) 1800(2): 35 (1801)

= Aspidium marginale f. davenportii Eastman; Ferns & Fern Allies New England 27 (1904)

= Aspidium marginale var. elegans J.Rob.; Bull. Essex Inst. 7(3): 51 (1875)

= Aspidium marginatum (L.) Schkuhr; Deutschl. Krypt. Gew. 1: 195, t. 45b (1809), emend.

= Dryopteris marginalis f. churchiae (Gilbert) Leeds; Amer. Fern J. 23: 12 (1933)

= Dryopteris marginalis f. davenportii (Eastman) A.Prince ex Weath.; Amer. Fern J. 26: 62 (1936)

= Dryopteris marginalis f. elegans (J.Rob.) F.W.Gray; Amer. Fern J. 14: 9 (1924)

= Dryopteris marginalis f. traillae (G.Lawson) Gruber; Amer. Fern J. 30: 89 (1940)

= Dryopteris marginalis f. tripinnatifida (Clute) Weath.; Amer. Fern J. 26: 61 (1936)

= Dryopteris marginalis var. elegans (J.Rob.) Carhart; Amer. Fern J. 6: 52 (1916)

= Filix marginalis (L.) Farw.; Rep. (Annual) Michigan Acad. Sci. 18: 81-83 (1916)

= Filix marginalis var. bipinnatifida Farw.; Rep. (Annual) Michigan Acad. Sci. 18: 82 (1916)

= Filix-mas marginalis (L.) Farw.; Amer. Midl. Naturalist 12: 234-251 (1931)

= Lastrea marginalis (L.) C.Presl; Tent. Pterid. 77 (1836)

= Lastrea marginalis var. traillae G.Lawson; Edinburgh New Philos. J. 19: 275 (1864)

= Nephrodium marginale (L.) Michx.; Fl. Bor.-Amer. (Michaux) 2: 267 (1803)

= Nephrodium marginale f. bipinnatifidum Clute; Fern Bull. 19: 50 (1911)

= Nephrodium marginale f. davenportii (Eastman) Floyd; Rhodora 4: 245 (1902)

= Nephrodium marginale f. tripinnatifidum Clute; Fern Bull. 15: 21 (1907)

= Nephrodium marginale var. distans Waters; Ferns (Waters) 238 (1903)

= Nephrodium marginale var. elegans (J.Rob.) Gilbert; Working List N. Amer. Pterid.: 17 (1901)

= Nephrodium marginale var. traillae (G.Lawson) Gilbert; List N. Amer. Pterid.: 17 (1901)

= Nephrodium marginale var. umbilicatum Desv.; Mém. Soc. Linn. Paris 6(3): 260 (1827)

= Polypodium marginale L.; Sp. Pl. 2: 1091 (1753)

= Polypodium vulgare var. churchiae Gilbert; Fern Bull. 14: 39 (1906)

= Polystichum marginale (L.) Keyserl.; Polyp. Herb. Bunge 44 (1873)

= Thelypteris marginalis (L.) Nieuwl.; Midl. Naturalist (Notre Dame) 1: 226 (1910)

= Thelypteris marginalis f. churchiae (Gilbert) Dole; Fl. Vermont, ed. 3: 11 (1937)

= Thelypteris marginalis f. davenportii (Eastman) L.B.Sm.; Rhodora 30: 17 (1928)

= Thelypteris marginalis f. elegans (J.Rob.) Weath.; Proc. Boston Soc. Nat. Hist. 36: 197 (1922)

species :

Dryopteris marginata (Wall. ex Christ) Christ; Philipp. J. Sci. 2: 212 (1907)

Distribution: China (Guangxi, Guizhou, Jiangxi, Sichuan, Yunnan); Taiwan; Pakistan; Nepal; Bhutan; India (Arunachal Pradesh, Assam State, Manipur, Meghalaya, Mizoram, Nagaland, Sikkim, West Bengal); Myanmar; Vietnam; peninsular Malaysia

36 CHI 38 TAI 40 BHU IND NEP PAK 41 MYA VIE 42 MLY

Synonyms: hidden

= Aspidium filix-mas var. marginatum (Wall. ex C.B.Clarke) Christ; Bull. Herb. Boissier, sér. 1, 6: 967 (1898)

= Aspidium marginatum (Wall. ex C.B.Clarke) Wall. ex Christ; Mém. Soc. Bot. France 1 [Hist. Acrostich.]: 39 (1905), nom. illeg.

= Aspidium marginatum Wall.; Numer. List. [Wallich] no. 391 (1829), nomen

= Dryopteris blinii H.Lév.; Fl. Kouy-Tcheou 490 (1915)

= Dryopteris chiui Ching; Bot. Res. Acad. Sin. 1987(2): 32 (1987)

= Dryopteris grandissima Tagawa; Acta Phytotax. Geobot. 3: 89 (1934)

= Dryopteris leveillei Nakai; Bull. Natl. Sci. Mus., Tokyo, 31: 17 (1952), non Christ (1909)

= Dryopteris metcalfei Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 463 (1938), nom. illeg.

= Lastrea marginata (Wall. ex C.B.Clarke) T.Moore; Index Fil. [T. Moore] 96 (1858)

= Nephrodium filix-mas var. marginatum Wall. ex C.B.Clarke; Trans. Linn. Soc. II, Bot., ser. 2, 1: 521, t. 71 (1880)

= Nephrodium marginatum (Wall. ex C.B.Clarke) C.Hope; J. Bombay Nat. Hist. Soc. 14: 740, t. 33 (1903)

= Polystichum blinii (H.Lév.) H.Lév. & C.Chr.; H. Lév., Cat. Pl. Yun-Nan 110 (1916)

species :

Dryopteris mauiensis C.Chr.; Index Filic. 277 (1905)

Distribution: Hawaii (Kauai, Oahu, Molokai, Lanai, Maui, Hawaii Isl.)

63 HAW

Synonyms: hidden

= Goniopteris clarkei (Baker) Ching; Sunyatsenia 5: 239 (1940)

= Polypodium clarkei Baker; Ann. Bot. (Oxford) 5: 457 (1891) (non Thelypteris clarkei (Bedd.) C. F. Reed)

= Thelypteris mauiensis (C.Chr.) C.F.Reed; Phytologia 17: 291 (1968)

= Thelypteris pacifica C.F.Reed; Phytologia 17: 300 (1968), nom. nov.

species :

Dryopteris maxima (Baker) C.Chr.; Index Filic. 277 (1905)

Distribution: Fiji (Viti Levu, Ovalau, Taveuni)

60 FIJ

Remarks: possibly to D. arborescens

Synonyms: hidden

= Nephrodium maximum Baker; J. Bot. 24: 182 (1886)

= Arachniodes maxima (Baker) Brownlie; Nova Hedwigia Beih. 55 (Pterid. Fl. Fiji): 280 (1977)

species :

Dryopteris maximowicziana (Miq.) C.Chr.; Acta Horti Gothob. 1(2): 63 (1924)

Distribution: China (Chongqing, Fujian, Guangxi, Guizhou, Hunan, Jiangxi, Sichuan, Zhejiang); South Korea; Japan; Taiwan

36 CHI 38 JAP KOR TAI

Remarks: subgen. Nothoperanema, sect. Shiehia

Synonyms: hidden

= Aspidium matsumurae Makino; Bot. Mag. (Tokyo) 6(60): 46 (1892)

= Aspidium maximowiczianum Miq.; Ann. Mus. Bot. Lugduno-Batavi 8: 178 (1867)

= Ctenitis aureovestita (Rosenst.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: t. 295 (1938)

= Ctenitis matsumurae (Makino) Koidz.; Acta Phytotax. Geobot. 8: 189 (1939)

= Ctenitis maximowicziana (Miq.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 294 (1938)

= Ctenitis whankanshanensis Ching & Chu H.Wang; Acta Phytotax. Sin. 19(1): 123 (1981)

= Dryopsis maximowicziana (Miq.) Holttum & P.J.Edwards; Kew Bull. 41(1): 197 (1986)

= Dryopteris aureovestita Rosenst.; Hedwigia 56: 343 (1915)

= Dryopteris matsumurae (Makino) C.Chr.; Index Filic. 277 (1905)

= Dryopteris maximowicziana (Miq.) Koidz.; Fl. Symb. Orient.-Asiat. 21 (1931) (non Dryopteris maximowiczii)

= Nephrodium matsumurae Makino; Bot. Mag. (Tokyo) 13: 48 (1899)

species :

Dryopteris maximowiczii (Baker) Kuntze; Revis. Gen. Pl. 2: 813 (1891)

Distribution: South Korea (incl. Cheju Isl.); Japan

38 JAP KOR

Remarks: sect. Diclisodon

Synonyms: hidden

= Arachniodes maximowiczii (Baker) Ohwi; J. Jap. Bot. 37: 76 (1962)

= Aspidium callopsis Franch. & Sav.; Enum. Pl. Jap. 2: 240, 634 (1877)

= Aspidium commutatum Franch. & Sav.; Enum. Pl. Jap. 2: 240 (1876)

= Byrsopteris maximowiczii (Baker) C.V.Morton; Amer. Fern J. 50: 154 (1960)

= Dryopteris callipteris C.Chr.; Bull. Acad. Int. Géogr. Bot. 1909, Mem. 20: 153 (1909)

= Dryopteris callopsis (Franch. & Sav.) C.Chr.; Index Filic. 256 (1905)

= Lastrea maximowiczii (Baker) T.Moore; Gard. Chron., ser. 2, 15: 626 (1881)

= Nephrodium callopsis (Franch. & Sav.) Palib.; Trudy Imp. S.-Peterburgsk. Bot. Sada 19: 140 (1901)

= Nephrodium maximowiczii Baker; Syn. Fil. [W. J. Hooker & Baker], ed. 2: 499 (1874)

= Polystichopsis maximowiczii (Baker) Tagawa; J. Jap. Bot. 33: 94 (1958)

= Rumohra callipteris (C.Chr.) C.Chr. ex Tardieu & C.Chr.; Notul. Syst. (Paris) 7: 84 (1938)

= Rumohra maximowiczii (Baker) Ching; Sinensia 5: 62, t. 13 (1934)

species :

Dryopteris maxonii Underw. & C.Chr.; Amer. Fern J. 1: 96 (1911)

Distribution: Mexico (Colima, Guanajuato, Guerrero, Jalisco, Mexico State, Michoacan, Morelos, Nayarit, Sinaloa); Guatemala; El Salvador; Costa Rica

79 MEX 80 COS ELS GUA

Synonyms: hidden

= Dryopteris patula var. moreliae Christ; Notul. Syst. (Paris) 1: 234 (1910)

species :

Dryopteris meghalaica Fraser-Jenk. & Gibby; Acta Bot. Fenn. 164: 6 (1999)

Distribution: NE-India (Manipur, Meghalaya, Nagaland); Myanmar (Kachin)

40 IND 41 MYA

Conservation Status: IUCN NT for India (Fraser-Jenkins 2012)

species :

Dryopteris melanocarpa Hayata; Icon. Pl. Formosan. 4: 163, f. 104 (1914)

Distribution: South Korea; Japan; Taiwan

38 JAP KOR TAI

Remarks: sect. Diclisodon, see Zuo et al. 2022

species :

Dryopteris mengshanensis J.X.Li & Xiao J.Li; Bangladesh J. Bot. 53(3): 712 (2024)

Distribution: China (Shandong)

36 CHI

species :

Dryopteris microlepis (Bak) C.Chr.; Index Filic., Suppl. 2 (1906-1912): 36 (1913)

Distribution: China (Guizhou, Yunnan); India (Arunachal Pradesh); Myanmar

36 CHI 40 IND 41 MYA

Conservation Status: IUCN CR for India (Fraser-Jenkins 2012)

Synonyms: hidden

= Dryopteris nakanensis Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 413 (1938)

= Nephrodium microlepis Baker; Bull. Misc. Inform. Kew 1908: 10 (1908)

species :

Dryopteris mindshelkensis Pavlov; Vestn. Akad. Nauk Kazakhsk. S.S.R. 8: 129, f. 31 (1954)

Name: Rigid buckler-fern (EN)

Distribution: England (NW-England, Wales); Spain; France; Italy; Croatia; Montenegro; Serbia; Kosovo; North Macedonia; Albania; Greece (mountains in NW-Greece, Peloponnisos); Algeria; Kazakhstan; ?Northern Caucasus; Georgia [Caucasus]; ?Armenia; ?Azerbaijan; Turkey (Anatolia); N-Iran; E-Afghanistan

10 GRB 12 FRA SPA 13 ALB CRO GRC ITA KOS MNT NMA SRB 20 ALG 32 KAZ 33 GEO 34 AFG IRN TUR

Remarks: affinis agg.

Synonyms: hidden

= Dryopteris submontana (Fraser-Jenk. & Jermy) Fraser-Jenk.; Candollea 32(2): 311 (1977)

= Dryopteris villarii subsp. mindshelkensis (Pavlov) Fraser-Jenk.; Candollea 32(2): 311 (1977)

= Dryopteris villarii subsp. submontana Fraser-Jenk. & Jermy; Fern Gaz. 11: 339 (1977)

species :

Dryopteris monticola (Makino) C.Chr.; Index Filic. 278 (1905)

Distribution: Russian Far East (Kuril Isl., Sakhalin); China (Liaoning); North Korea; South Korea (incl. Cheju Isl.); Japan

31 RFE 36 CHI 38 JAP KOR NKO

Synonyms: hidden

= Aspidium goldieanum Christ; Bull. Herb. Boissier, sér. 2, 2: 828 (1902) (non Hook. (1822))

= Aspidium monticola (Makino) Christ; Mém. Soc. Bot. France 1 [Hist. Acrostich.]: 39 (1905)

= Dryopteris goldieana subsp. monticola (Makino) Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.), Bot. 14(3): 192 (1986)

= Dryopteris submonticola Nakai; Bot. Mag. (Tokyo) 46: 98 (1931)

= Nephrodium erythrosorum var. manshuricum Kom.; Trudy Imp. S.-Peterburgsk. Bot. Sada 20: 120 (1901)

= Nephrodium monticola Makino; Bot. Mag. (Tokyo) 13: 80 (1899)

species :

Dryopteris montigena Ching; Bull. Fan Mem. Inst. Biol. Bot. 11: 64 (1941)

Distribution: China (Yunnan)

36 CHI

Synonyms: hidden

= Dryopteris likiangensis Ching; Sunyatsenia 6: 1 (1941) (non Dryopteris lichiangensis (Wright) C. Chr. 1913)

species :

Dryopteris munchii A.R.Sm.; Proc. Calif. Acad. Sci. ser. 4, 40: 218 (1975)

Distribution: Mexico (Chiapas, Hidalgo, Puebla)

79 MEX

species :

Dryopteris namegatae (Sa.Kurata) Sa.Kurata; J. Geobot. 17: 89 (1969)

Distribution: China (Gansu, Hunan, Jiangxi, Sichuan, Yunnan, Zhejiang); Japan; Taiwan; N-India (Arunachal Pradesh)

36 CHI 38 JAP TAI 40 IND

Conservation Status: IUCN VU for India (Fraser-Jenkins 2012)

Synonyms: hidden

= Dryopteris dickinsii var. namegatae Sa.Kurata; J. Geobot. 7: 115 (1958)

= Dryopteris infrapuberula Ching, Boufford & K.H.Shing; J. Arnold Arbor. 64(1): 28 (1983)

species :

Dryopteris napoleonis (Bory) Kuntze; Revis. Gen. Pl. 2: 813 (1891)

Distribution: St. Helena

28 STH

Conservation Status: IUCN R

Synonyms: hidden

= Aspidium napoleonis (Bory) Fée; Mém. Foug. 5 (Gen. Filic.): 292 (1852)

= Aspidium petersenii Kunze ex Kuhn; (1868)

= Lastrea napoleonis (Bory) C.Presl; Tent. Pterid. 77 (1836)

= Nephrodium napoleonis Bory; Voy. Indes Or. [Bélanger] 2: 61, t. 6 (1833)

= Polystichum napoleonis (Bory) Keyserl.; Polyp. Herb. Bunge 44 (1873)

= Polystichum riparium C.Presl; Tent. Pterid. 8 (1836)

species :

Dryopteris nidus (Baker) Zhang; Phytotaxa 71: 17-27 (2012)

Distribution: China (Guizhou, Sichuan, Yunnan); Tibet; E-Nepal; Bhutan; India (Arunachal Pradesh, Sikkim, West Bengal); Myanmar

36 CHI TIB 40 BHU IND NEP 41 MYA

Remarks: Subgen. Dryopsis. The lumping in Flora of China might be too much in this case, and some species might be valid (Fraser-Jenkins et al. 2018)

Synonyms: hidden

= Aspidium filix-mas var. nidus (Baker) Christ; Mém. Soc. Bot. France 1 [Hist. Acrostich.]: 38 (1905)

= Ctenitis contigua Ching; Acta Phytotax. Sin. 19(1): 119 (1981)

= Ctenitis crassirachis Ching; Acta Phytotax. Sin. 19(1): 119 (1981)

= Ctenitis crenata Ching; Acta Phytotax. Sin. 19(1): 120 (1981)

= Ctenitis dentisora Ching; Bull. Fan Mem. Inst. Biol. Bot. 11: 67 (1941)

= Ctenitis nidus (Baker) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 287 (1938)

= Ctenitis silaensis Ching; Bull. Fan Mem. Inst. Biol. Bot. 11: 68 (1941)

= Ctenitis submariformis Ching & Chu H.Wang; Acta Phytotax. Sin. 19(1): 122 (1981)

= Ctenitis wantsingshanica Ching & K.H.Shing; Acta Phytotax. Sin. 19: 123 (1981)

= Ctenitis zayuensis Ching & S.K.Wu; Fl. Xizangica 1: 273 (1983)

= Dryopsis contigua (Ching) Holttum & P.J.Edwards; Kew Bull. 41(1): 193 (1986)

= Dryopsis crassirachis (Ching) Holttum & P.J.Edwards; Kew Bull. 41(1): 195 (1986)

= Dryopsis crenata (Ching) Holttum & P.J.Edwards; Kew Bull. 41(1): 193 (1986)

= Dryopsis dentisora (Ching) Holttum & P.J.Edwards; Kew Bull. 41(1): 195 (1986)

= Dryopsis nidus (Baker) Holttum & P.J.Edwards; Kew Bull. 41(1): 192 (1986)

= Dryopsis silaensis (Ching) Holttum & P.J.Edwards; Kew Bull. 41(1): 197 (1986)

= Dryopsis submariformis (Ching & Chu H.Wang) Holttum & P.J.Edwards; Kew Bull. 41(1): 194 (1986)

= Dryopsis wantsingshanica (Ching & K.H.Shing) Holttum & P.J.Edwards; Kew Bull. 41(1): 194 (1986)

= Dryopteris crassirachis (Ching) Li Bing Zhang; Phytotaxa 71: 21 (2012)

= Dryopteris filix-mas subsp. nidus (Baker) C.Chr.; Index Filic. 5: 265 (1905)

= Dryopteris leiboensis Li Bing Zhang; Phytotaxa 71: 22 (2012), nom. nov.

= Dryopteris nokoensis Tagawa; Acta Phytotax. Geobot. 3: 91 (1934)

= Dryopteris nushanensis Li Bing Zhang; Phytotaxa 71: 24 (2012)

= Dryopteris submariformis (Ching & Chu H.Wang) Li Bing Zhang; Phytotaxa 71: 25 (2012)

= Dryopteris wantsingshanica (Ching & K.H.Shing) Li Bing Zhang; Phytotaxa 71: 26 (2012)

= Lastrea filix-mas subvar. nidus (Baker) Bedd.; Ferns Brit. India 250 (1883)

= Lastrea filix-mas var. nidus (Baker) Bedd.; Ferns Brit. India Suppl. 171, t. 372 (1876)

= Lastrea filix-mas var. parallelogramma subvar.nidus Bedd.; Ferns Brit. India t. 249, t. 129 (1883)

= Nephrodium apiciflorum var. nidus (Baker) C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 518 (1880)

= Nephrodium filix-mas var. nidus Baker; Syn. Fil. [W. J. Hooker & Baker], ed. 2: 498 (1874)

= Nephrodium nidus (Baker) C.B.Clarke; J. Linn. Soc., Bot. 16: 156 (1876)

species :

Dryopteris nigropaleacea (Fraser-Jenk.) Fraser-Jenk.; Bol. Soc. Brot. 55: 238 (1982)

Distribution: ?Tibet; Pakistan (Kurram, Peshawar, Waziristan, Swat, Chitral, Pakistani Punjab, Hazara, Chitral, Dir, Swat, Murree, Azad Kashmir, Gilgit); W-Nepal; India (Jammu & Kashmir, Himachal Pradesh, Uttarakhand)

40 IND NEP PAK

Remarks: reports of D. odontoloma for the S-Himalaya belong here fide Fraser-Jenkins 2008

Synonyms: hidden

= Dryopteris juxtaposita subsp. nigropaleacea (Fraser-Jenk.) Khullar; J. Indian Bot. Soc. 88(3-4): 207 (2009), et in Indian Fern J. 26(1-2): 91 (2010), without basionym reference

= Dryopteris pallida subsp. nigropaleacea Fraser-Jenk.; Candollea 32(2): 316 (1977)

species :

Dryopteris nipponensis Koidz.; Mayebara, Fl. Austro-Higoensis 5 (1931)

Distribution: Japan (Hokkaido, Kyushu, Shikoku)

38 JAP

Remarks: often treated under the name D. ambigens

Synonyms: hidden

= Aspidium cystolepidotum Miq.; Ann. Mus. Bot. Lugduno-Batavi 3: 177 (1867)

= Dryopteris ambigens (Nakai) Koidz.; Fl. Symb. Orient.-Asiat. 39 (1930)

= Dryopteris cystolepidota (Miq.) C.Chr.; Index Filic. 260 (1905)

= Dryopteris erythrosora var. ambigens Nakai; Bot. Mag. (Tokyo) 39: 119 (1925)

= Dryopteris taquetii Christ; Repert. Spec. Nov. Regni Veg. 5: 284 (1908)

= Polystichum cystolepidotum (Miq.) Franch. ex H.Lév.; Bull. Acad. Int. Géogr. Bot. 20: 9 (1910)

species :

Dryopteris nobilis Ching; Bull. Fan Mem. Inst. Biol. Bot. 11: 65 (1941), nom. illeg., non J. Douglas 1866

Distribution: China (NW-Yunnan); ?Tibet; Bhutan; N-India (West Bengal, Sikkim, Arunachal Pradesh, Assam State, Meghalaya)

36 CHI 40 BHU IND

Conservation Status: IUCN VU for India (Fraser-Jenkins 2012)

species :

Dryopteris nodosa (C.Presl) Li Bing Zhang; Taxon 61(6): 1208 (2012)

Distribution: Japan (Yakushima Isl.); Myanmar; N-Thailand; Vietnam; Sumatra; Borneo (Mt. Kinabalu, Sabah, Sarawak); Sulawesi; Philippines; Lesser Sunda Isl. (Bali, Flores); New Guinea (Irian Jaya, Papua New Guinea); Bismarck Arch. (New Ireland); Solomon Isl. (Kolombangara, Rendova, Guadalcanal); Vanuatu; Fiji (Viti Levu)

38 JAP NNS 41 MYA THA VIE 42 BOR LSI NWG PHI SUL SUM 43 BIS SOL 60 FIJ VAN

Remarks: subgen. Nothoperanema. Separated from D. paleolata by molecular data (Zhang et al. 2012). Cited for Taiwan by Flora of China but excluded by Knapp (Ferns of Taiwan, 2011 ff.)

Synonyms: hidden

= Acrophorus blumei Ching ex C.Chr. & Holttum; Gard. Bull. Straits Settlem. 7: 226 (1934), nom. nov.

= Acrophorus nodosus (Blume) C.Presl; Tent. Pterid. 94, t. 3, f. 2 (1836)

= Acrophorus stipellatus (Wall.) T.Moore; Gard. Chron. 1864: 135 (1864)

= Aspidium nodosum Blume; Enum. Pl. Javae 171 (1828), non Willd.

= Cystopteris gigantea C.Presl; Tent. Pterid. 93 (1836)

= Cystopteris nodosa (C.Presl) Mett.; Ann. Mus. Bot. Lugduno-Batavi 1: 241 (1863)

= Davallia ligulata Wall.; Numer. List. [Wallich] no. 254 (1829), nomen

= Davallia nodosa (C.Presl) Hook.; Sp. Fil. [W. J. Hooker] 1: 157 (1846)

= Leucostegia ligulata J.Sm.; London J. Bot. 1: 426 (1842)

= Leucostegia nodosa (C.Presl) Bedd.; Ferns Brit. India Suppl. 4 (1870)

= Peranema nodosum (C.Presl) Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 317 (2008)

species :

Dryopteris nubigena Maxon & C.V.Morton; Proc. Biol. Soc. Wash. 50: 179 (1937)

Distribution: Mexico (Chiapas, Guerrero, Mexico State, Oaxaca); Guatemala; El Salvador; Honduras; Costa Rica; Panama

79 MEX 80 COS ELS GUA HON PAN

species :

Dryopteris nyingchiensis Ching; Fl. Xizangica 1: 262 (1983)

Distribution: Tibet; Taiwan; etc.

36 TIB 38 TAI

Remarks: kept separate from D. sublacera by Taiwan checklist (2019)

species :

Dryopteris obtusiloba (Baker ex Hook. & Baker) Kuntze; Revis. Gen. Pl. 2: 813 (1891)

Distribution: Sri Lanka

40 SRL

Remarks: subgen. Nothoperanema, sect. Shiehia

Synonyms: hidden

= Aspidium obtusilobum (Baker ex Hook. & Baker) Prantl; Verh. K. K. Zool.-Bot. Ges. Wien 81: 119 (1882)

= Ctenitis ferruginea var. obtusiloba (Baker) Sledge; Kew Bull. 27: 409 (1972)

= Ctenitis obtusiloba (Baker ex Hook. & Baker) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 296 (1938)

= Dryopsis obtusiloba (Baker ex Hook. & Baker) Holttum & P.J.Edwards; Kew Bull. 41(1): 202 (1986)

= Dryopteris peranemiformis subsp. obtusiloba (Baker ex Hook. & Baker) C.Chr.; Index Filic. 284 (1905)

= Dryopteris zeylanica Alderw.; Malayan Ferns 203 (1909) (nom. nov. superfl.)

= Lastrea obtusiloba (Baker ex Hook. & Baker) Bedd.; Ferns Brit. India t. 296 (1868)

= Nephrodium obtusilobum Baker; Syn. Fil. [W. J. Hooker & Baker] 284 (1867)

species :

Dryopteris obtusipinnula (Fraser-Jenk.) Fraser-Jenk.; Biodivers. & Bioprospecting 29 (2023)

Distribution: Nepal; Bhutan; India (Arunachal Pradesh, Meghalaya, Mizoram, Sikkim, Tamil Nadu); Sri Lanka; Myanmar; ?Java

40 BHU IND NEP SRL 41 MYA

Remarks: sect. Diclisodon

Synonyms: hidden

= Dryopteris sparsa subsp. obtusipinnula Fraser-Jenk.; Indian J. Forest. 45(1): 13 (2022)

species :

Dryopteris occidentalis J.P.Roux; Kew Bull. 59(2): 214 (2004)

Distribution: Guinea; Ivory Coast; Nigeria

22 GUI IVO NGA

species :

Dryopteris odontoloma (T.Moore) C.Chr.; Acta Horti Gothob. 1: 59 (1924)

Distribution: S-India (Tamil Nadu)

40 IND

Conservation Status: IUCN NT for India (Fraser-Jenkins 2012)

Synonyms: hidden

= Lastrea filix-mas var. odontoloma (T.Moore) Bedd.; Ferns Brit. India Suppl. 55 (1892)

= Lastrea odontoloma (T.Moore) Bedd.; Ferns S. India 39, pl. 114 (1864)

= Lastrea odontoloma T.Moore; Index Fil. [T. Moore] 90 (1858)

= Nephrodium odontoloma (T.Moore) C.Hope; J. Bombay Nat. Hist. Soc. 14: 736 (1903)

species :

Dryopteris oligodonta (Desv.) Pic.Serm.; Webbia 8: 150, f. 2 (1951)

Name: Kanaren-Wurmfarn (DE); Helecho macho (ES)

Distribution: Canary Isl. (Gran Canaria, Tenerife, La Gomera, Hierro, La Palma Isl.); Cape Verde Isl.

21 CNY CVI

Conservation Status: IUCN R (Capverdes)

Synonyms: hidden

= Allantodia oligantha (Desv.) Desv.; Mem. Soc. Linn. Paris 6(2): 265 (1827)

= Aspidium aquilinoides Bolle; Giorn. Sci. Nat. Econ. Palermo 1: 221 (1866)

= Aspidium canariense A.Braun; Flora 24: 708 (1841)

= Aspidium elongatum Willd.; Sp. Pl., ed. 4 [Willdenow] 5(1): 269 (1810) (non Sw. (1801) nec All.)

= Aspidium oliganthum Desv.; Mag. Neuesten Entdeck. Gesammten Naturk. Ges. Naturf. Freunde Berlin 5: 321 (1811)

= Aspidium oligodonton Desv.; Mag. Neuesten Entdeck. Gesammten Naturk. Ges. Naturf. Freunde Berlin 5: 322 (1811)

= Dryopteris elongata var. simplex A.Chev.; Rev. Bot. Appl. Agric. Trop. 15(170/171): 328 (seors 1060) (1935)

= Dryopteris oligantha (Desv.) C.Chr.; Index Filic., Suppl. Tert.: 93 (1934)

= Lastrea canariensis (A.Braun) T.Moore; Index Fil. [T. Moore] 87 (1858)

= Nephrodium canariense (A.Braun) Kiaersk.; Cat. Fil. hb. hort. Haun. 62 (1874)

= Nephrodium oligodontum (Desv.) Desv.; Mem. Soc. Linn. Paris 6(2): 261 (1827)

= Polystichum canariense (A.Braun) Keyserl.; Polyp. Herb. Bunge 45 (1873)

species :

Dryopteris oreades Fomin; Vestn. Tiflissk. Bot. Sada 18: 20 (1911)

Name: Geröll-Dornfarn (DE)

Distribution: Faeroe Isl.; England (N-England, Wales, Scotland, Outer Hebrides, Orkney Isl.); Ireland (NW-Ireland); Norway; Germany (Nordrhein-Westfalen); Portugal; Spain; Andorra; France; Corsica; Sardinia; Italy; Northern Caucasus; Georgia [Caucasus]; Armenia; Azerbaijan; Turkey (Anatolia)

10 FOR GRB IRE NOR 11 GER 12 ADO COR FRA POR SAR SPA 13 ITA 33 ARM AZE GEO NCS 34 TUR

Remarks: filix-mas agg.

Synonyms: hidden

= Aspidium pumilum (T.Moore) E.J.Lowe; Ferns 6: t. 15 (1857)

= Dryopteris abbreviata (DC.) Newman ex Manton; (1950) (p. p. excl. typo, non (Schrad.) Kuntze (1891), nec Polystichum abbreviatum DC. (1805))

= Dryopteris filix-mas subsp. oreades (Fomin) O.Bolòs & Vigo; Fl. Paisos Catalans: 171 (1984)

= Lastrea filix-mas var. pumila T.Moore

= Lastrea pumila (T.Moore) J.Sm.; Hist. Fil. 214 (1875)

= Nephrodium filix-mas proles abbreviatum Rouy; Fl. France [Rouy & Foucaud] 14: 407 (1913)

= Nephrodium filix-mas subsp. abbreviatum P.E.Fourn.; Fl. Compl. Plaine Franç.: 594 (1928)

= Polystichum filix-mas var. abbreviatum P.E.Fourn.; Quatre Fl. France: 16 (1934)

species :

Dryopteris paleolata (Pic.Serm.) Li Bing Zhang; Taxon 61(6): 1208 (2012)

Distribution: China (Fujian, Guangdong, Guangxi, Guizhou, Hainan, Jiangxi, Sichuan, Yunnan); Tibet; Taiwan; Nepal; Bhutan; India (Arunachal Pradesh, Manipur, Meghalaya, Mizoram, Nagaland, Sikkim, Uttarakhand, West Bengal); Myanmar (Kachin, etc.); Vietnam; Philippines

36 CHI TIB 38 TAI 40 BHU IND NEP 41 MYA VIE 42 PHI

Remarks: subgen. Nothoperanema. The name Acrophorus stipellatus (Wall.) T. Moore, based on Davallia stipellata Wall., a nom. nud., is not validly published. Synonymy fide Fraser-Jenkins et al. 2018

Synonyms: hidden

= Acrophorus diacalpioides Ching & S.H.Wu; S. H. Wu, Acta Phytotax. Sin. 21(4): 381 (1983)

= Acrophorus dissectus Ching & S.H.Wu; S. H. Wu, Acta Phytotax. Sin. 21(4): 382 (1983) (fide Fraser-Jenkins et al. 2018)

= Acrophorus emeiensis Ching; S. H. Wu, Acta Phytotax. Sin. 21(4): 380 (1983) (fide Fraser-Jenkins et al. 2018)

= Acrophorus exstipellatus Ching & S.H.Wu; S. H. Wu, Acta Phytotax. Sin. 21(4): 383 (1983) (fide Fraser-Jenkins et al. 2018)

= Acrophorus paleolatus Pic.Serm.; Webbia 31(1): 252 (1977) (paleolulatus auct., errore)

= Davallia stipellata Wall.; Numer. List. [Wallich] no. 260 (1829), nomen

= Dryopteris acrophorus Li Bing Zhang; Taxon 61(6): 1208 (2012), nom. nov. for Acrophorus diacalpoides Ching & S. H. Wu (fide Fraser-Jenkins et al. 2018)

= Dryopteris emeiensis (Ching) Li Bing Zhang; Taxon 61(6): 1208 (2012) (fide Fraser-Jenkins et al. 2018), non Dryopteris omeiensis (Baker) C. Chr. 1905

= Dryopteris exstipellata (Ching & S.H.Wu) Li Bing Zhang; Taxon 61(6): 1209 (2012) (fide Fraser-Jenkins et al. 2018)

= Dryopteris wusugongii Li Bing Zhang; Taxon 61(6): 1208 (2012), nom. nov. for Acrophorus dissectus Ching & S. H. Wu (fide Fraser-Jenkins et al. 2018)

= Peranema paleolatum (Pic.Serm.) Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 317 (2008)

species :

Dryopteris pallida (Bory) Maire & Petitm.; Bull. Séances Soc. Sci. Nancy, ser. 3, 9: 480 (1908)

subspecies :

subsp. pallida

Name: Fahler Wurmfarn (DE); Pale Male-fern (EN)

Distribution: Baleares; Sardinia; Italy; Sicily; Pantelleria; Croatia; Bosnia & Hercegovina; Montenegro; North Macedonia; Albania; Greece (widespread, incl. Kiklades, Samothraki); Crete (widespread); Algeria; Tunisia; East Aegaean Isl. (Lesvos, Chios, Ikaria, Samos, Dodekanese, Simi, Karpathos, Rhodos, Kastellorhizo Isl.); Turkey (Anatolia); Iran; Cyprus; Lebanon; Syria; Israel; Palestine

12 BAL SAR 13 ALB BIH CRO GRC ITA KRI MNT NMA PTL SIC 20 ALG TUN 34 CYP EAI IRN ISR LEB PAL SYR TUR

Synonyms: hidden

= Aspidium affine Reichb.; Fl. Dalmat. 1: 39 (1842), nom. illeg.

= Aspidium pallidum (Bory) Link; Fil. Spec. 107 (1841) (non Blume (1828))

= Aspidium rigidum var. australe Ten.; Atti Reale Ist. Incorragg. Sci. Nat. Napoli 6: 144, t. 2, f. 4 B (1830)

= Dryopteris pallida (Bory) Fomin; Vestn. Tiflissk. Bot. Sada 20, 3: 2 (1910)

= Dryopteris villarii subsp. pallida (Bory) Heywood; Feddes Repert. Spec. Nov. Regni Veg. 69: 44 (1964)

= Lastrea nivalis T.Moore; Index Fil. [T. Moore] 98 (1858)

= Nephrodium aristatum C.Presl; Fl. Sicul. [Presl] 1: 46 (1826)

= Nephrodium pallidum Bory; Exp. Sci. Morée, Bot. 3(2): 287 (1832)

= Nephrodium rigidum subsp. pallidum (Bory) Arcang.; Comp. Fl. Ital. [Arcangeli] 811 (1882)

= Nephrodium villarii subsp. pallidum (Bory) Hayek; Repert. Spec. Nov. Regni Veg. Beih. 30(1): 27 (1924)

= Polystichum pallidum (Bory) Tod.; Giorn. Sci. Nat. Econ. Palermo 1: 240 (1866) non Gardner 1842

= Polystichum rigidum var. australe (Ten.) P.E.Fourn.; Quatre Fl. France: 17 (1934)

subspecies :

subsp. balearica (Litard.) Fraser-Jenk.; Candollea 32: 314 (1977)

Distribution: Baleares

12 BAL

Conservation Status: IUCN R

Synonyms: hidden

= Dryopteris balearica (Litard.) E.Nardi; Webbia 30: 28 (1976)

= Dryopteris rigida var. balearica Litard.; Bull. Acad. Int. Géogr. Bot. 21: 23 (1911)

= Dryopteris villarii subsp. balearica (Litard.) O.Bolòs & Vigo; Fl. Paisos Catalans: 172 (1984)

subspecies :

subsp. libanotica (Rosenst.) E.Nardi; Webbia 32: 97 (1900)

Distribution: Turkey (Anatolia); Cyprus; Lebanon; Syria; Israel; ?Jordania

34 CYP ISR LEB SYR TUR

Synonyms: hidden

= Aspidium libanoticum Rosenst.; Mém. Herb. Boissier 9: 1 (1900)

= Dryopteris libanotica (Rosenst.) C.Chr.; Index Filic. 275 (1905)

= Dryopteris pallida subsp. libanotica (Rosenst.) Fraser-Jenk.; Candollea 32(2): 315 (1977)

= Nephrodium libanoticum (Rosenst.) Bornm.; Beih. Bot. Centralbl. 36(11): 279 (1914)

subspecies :

subsp. raddeana (Fomin) Fraser-Jenk.; Candollea 32(2): 315 (1977)

Distribution: Azerbaijan (Talysh)

33 AZE

Synonyms: hidden

= Dryopteris pallida subsp. raddeana (Fomin) E.Nardi; Webbia 32(1): 97 (1977), isonym

= Dryopteris raddeana (Fomin) Fomin; Kuzn. et al., Fl. Caucas. Crit. 1(1): 57 (1911), et Vestn. Tiflissk. Bot. Sada 20: 33 (1911)

= Dryopteris raddeana var. talyschensis Askerov & A.E.Bobrov; Bot. Zhurn. (Moscow & Leningrad) 57(10): 1297 (1972)

= Dryopteris talyschensis (Askerov & A.E.Bobrov) Askerov & Aktcay; Turczaninowia 19(1): 81 (2017)

= Nephrodium raddeanum Fomin; Vestn. Tiflissk. Bot. Sada 12: 8, t. 2 (1908)

species :

Dryopteris panda (C.B.Clarke) C.Chr.; Index Filic., Suppl. Tert.: 94 (1934)

Distribution: China (Gansu, Guizhou, Sichuan, Yunnan); Tibet; Pakistan (Azad Kashmir); Nepal; Bhutan; India (Himachal Pradesh, Uttarakhand, Sikkim, West Bengal, Jammu & Kashmir)

36 CHI TIB 40 BHU IND NEP PAK

Synonyms: hidden

= Aspidium filix-mas var. pandum (C.B.Clarke) Christ; Mém. Soc. Bot. France 1 [Hist. Acrostich.]: 38 (1905)

= Dryopteris bonatiana (Brause) Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.), Bot. 18(5): 363 (1989) (fide Fraser-Jenkins 1997)

= Dryopteris filix-mas subsp. panda (C.B.Clarke) C.Chr.; Index Filic. 5: 265 (1905) ex Ching (1938)

= Dryopteris yui Ching; Bull. Fan Mem. Inst. Biol. Bot. 11: 60 (1941) (fide Fraser-Jenkins 1997)

= Lastrea filix-mas var. panda (C.B.Clarke) Bedd.; Ferns Brit. India 251 (1883)

= Nephrodium filix-mas var. pandum C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 519, t. 68, f. 1 (1880)

= Nephrodium pandum (C.B.Clarke) C.Hope; J. Bombay Nat. Hist. Soc. 12: 623 (1899)

= Polystichum bonatianum Brause; Hedwigia 54: 200, t. 4B (1914) (fide Fraser-Jenkins 1997)

species :

Dryopteris parabissetiana H.-J.Wei & Z.Y.Zuo; Taxon 73(1): 134 (2024)

Distribution: China (Anhui)

36 CHI

species :

Dryopteris paomowanensis Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(4): 10 (1984)

Distribution: China (Guangxi, Guizhou, Sichuan, Yunnan); Japan

36 CHI 38 JAP

Remarks: taxonomic rank uncertain, often treated as D. simasakii var. paleacea

Synonyms: hidden

= Dryopteris indusiata var. paleacea H.Itô; J. Jap. Bot. 9: 57 (1933)

= Dryopteris labordei f. paleacea (H.Itô) H.Itô; Nov. Fl. Jap. [Nakai & Honda] 4: 46 (1939)

= Dryopteris rufosquamosa Ching & P.S.Chiu; Bot. Res. Acad. Sin. 1987(2): 6 (1987)

= Dryopteris simasakii var. paleacea (H.Itô) Sa.Kurata; J. Geobot. 18: 5 (1970)

species :

Dryopteris papuae-novae-guineae Li Bing Zhang; Taxon 61(6): 1209 (2012), nom. nov.

Distribution: New Guinea (Papua New Guinea)

42 NWG

Synonyms: hidden

= Acrophorus stipellatus var. montanus Rosenst.; Repert. Spec. Nov. Regni Veg. 12: 166 (1913)

species :

Dryopteris papuana C.Chr.; Index Filic. 282 (1905)

Distribution: New Guinea (Papua New Guinea); Bismarck Arch. (New Ireland); Solomon Isl. (Kolombangara, Rendova)

42 NWG 43 BIS SOL

species :

Dryopteris patula (Sw.) Underw.; Native Ferns, ed. 4: 177 (1893)

Distribution: Mexico (Chiapas, Chihuahua, Colima, Ciudad de Mexico, Durango, Guanajuato, Hidalgo, Jalisco, Mexico State, Michoacan, Morelos, Nayarit, Nuevo Leon, Oaxaca, Puebla, Queretaro, Sinaloa, Veracruz); Guatemala; El Salvador; Honduras; Nicaragua; Costa Rica; Panama; Jamaica; Hispaniola (Haiti, Dominican Republic); Colombia (Antioquia, Boyacá, Cundinamarca, Guainía, Magdalena, Nariño); Venezuela (Aragua, Lara, Merida, Miranda, Portuguesa, Sucre, Tachira); Ecuador; Galapagos; Peru; Bolivia (Chuquisaca, Cochabamba, La Paz, Santa Cruz, Tarija); Brazil (Acre, Goiás, Maranhão, Minas Gerais, Mato Grosso, Pará, Rio de Janeiro, Rondônia, Rio Grande do Sul, São Paulo)

79 MEX 80 COS ELS GUA HON NIC PAN 81 HIS JAM 82 CLM VEN 83 BOL ECU GAL PER 84 BRA

Synonyms: hidden

= Aspidium apertum Fée; Mém. Foug. 8 (Icon. Esp. Nouv.): 106 (1857)

= Aspidium chaerophylloides Moritz ex Mett.; Abh. Senckenberg. Naturf. Ges. 2(2) [Pheg. u. Asp.]: 349 (1858)

= Aspidium diastematocarpon (Fée) Fée; Mém. Foug. 8 (Icon. Esp. Nouv.): 107 (1857)

= Aspidium imbecile Kuntze ex Ettingsh.; Farnkr. Jetztw. [Ettingshausen]: 176 (1865 publ. 1864)

= Aspidium leptorachis Kunze; Linnaea 18: 346 (1844)

= Aspidium mexicanum var. serratum Mett.; Abh. Senckenberg. Naturf. Ges. 2(2) [Pheg. u. Asp.]: 349 (1858)

= Aspidium patulum Sw.; Kongl. Vetensk. Acad. Handl. 1817: 64 (1817)

= Aspidium patulum var. chaerophylloides (Baker ex Mart.) J.Bommer & Christ; Bull. Soc. Roy. Bot. Belgique 35(1): 214 (1896)

= Aspidium paupertinum Roem.; Kunze, Linnaea 18: 345 (1844)

= Aspidium polystichoides (C.Presl) Mett.; Salomon, Nomencl. Gefäßkrypt. 67 (1883)

= Dryopteris aperta (Fée) C.Chr.; Index Filic. 252 (1905)

= Dryopteris patula var. chaerophylloides (Baker ex Mart.) Hieron.; Hedwigia 46: 345 (1907)

= Dryopteris patula var. serrata (Mett.) C.V.Morton; Contr. U. S. Natl. Herb. 38(6): 219 (1973)

= Dryopteris simplicior Mickel & Beitel; Mem. New York Bot. Gard. 46 [Pterid. Fl. Oaxaca]: 166 (1988)

= Filix patula (Sw.) Farw.; Rep. (Annual) Michigan Acad. Sci. 18: 81-83 (1916)

= Filix-mas patula (Sw.) Farw.; Amer. Midl. Naturalist 12: 234-251 (1931)

= Filix-mas patula var. chaerophylloides (Baker ex Mart.) Farw.; Amer. Midl. Naturalist 12: 256 (1931)

= Lastrea aperta (Fée) T.Moore; Index Fil. [T. Moore] 85 (1858)

= Lastrea diastematocarpa (Fée) T.Moore; Index Fil. [T. Moore] 90 (1858)

= Lastrea leptorachis (Kunze) Liebm.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 5, 1: 272 (seors. 120) (1849)

= Lastrea patula (Sw.) T.Moore; Index Fil. [T. Moore] 99 (1858)

= Lastrea paupertina (Roem.) Liebm.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 5, 1: 272 (seors. 120) (1849)

= Lastrea polystichoides C.Presl; Abh. Königl. Böhm. Ges. Wiss., ser. 5, 6 [Epim. Bot.]: 38 (1851)

= Nephrodium patulum (Sw.) Baker; Fl. Bras. [Martius] 1(2): 480 (1870)

= Nephrodium patulum f. robustior Donn.Sm.; Enum. Pl. Guatem. 3: 101 (1893), nom. nud.

= Nephrodium patulum var. chaerophylloides Baker; Fl. Bras. [Martius] 1(2): 481 (1870)

= Nephrodium patulum var. robustior Donn.Sm.; Enum. Pl. Guatem. 3: 101 (1893), nom. nud.

species :

Dryopteris paucisora Copel.; Philipp. J. Sci. 6: 136 (1911)

Distribution: Borneo (Mt. Kinabalu, Sabah, Sarawak)

42 BOR

Remarks: Subgen. Dryopsis

Synonyms: hidden

= Ctenitis paucisora (Copel.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 124 (1947)

= Dryopsis paucisora (Copel.) Holttum & P.J.Edwards; Kew Bull. 41(1): 183 (1986)

species :

Dryopteris pauliae Fraser-Jenk.; New Species Syndr. Indian Pteridol. 140 (1997)

Distribution: Nepal; India (Himachal Pradesh, Uttarakhand)

40 IND NEP

species :

Dryopteris peninsulae Kitag.; Rep. First Sci. Exped. Manchoukuo 4(2): 54 (1935)

Distribution: China (Gansu, Guizhou, Henan, Hubei, Jiangxi, Liaoning, Shaanxi, Shandong, Sichuan, NE-Yunnan)

36 CHI

Synonyms: hidden

= Dryopteris apicisora Ching & Y.T.Hsieh; Fl. Tsinling. 2: 228 (1974)

= Dryopteris cathayana Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(4): 1 (1984)

= Dryopteris lacera var. peninsulae Tagawa

= Dryopteris neolacera Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 440 (1938)

= Dryopteris shensicola Ching & Y.T.Hsieh; Fl. Tsinling. 2: 229 (1974)

species :

Dryopteris pentheri (Krasser) C.Chr.; Index Filic. 284 (1905)

Distribution: Cape Verde Isl. (Ilha de Sao Nicolau); Guinea-Bissau; Guinea; Sierra Leone; Liberia; Nigeria; Cameroon; Bioko Isl. [Fernando Poo]; São Tomé; D.R.Congo [Zaire]; Rwanda; Burundi; Sudan; Ethiopia; Uganda; Kenya; Tanzania; Angola; Zambia; Malawi; Zimbabwe; Mozambique; South Africa (Gauteng, Limpopo, Mpulamanga, KwaZulu-Natal, E-Cape Prov., W-Cape Prov., Free State); Lesotho; Eswatini [Swaziland]; Madagascar; La Réunion

21 CVI 22 GNB GUI LBR NGA SIE 23 BIO BUR CMN RWA SAO ZAI 24 ETH SUD 25 KEN TAN UGA 26 ANG MLW MOZ ZAM ZIM 27 LES SAF SWZ 29 MDG REU

Synonyms: hidden

= Dryopteris elongata Sim; Ferns S. Afr. [Sim], ed. 2: 104 (1916) (non Dryopteris elongata (Wall. ex Hook.) Kuntze)

= Nephrodium pentheri Krasser; Ann. Hofmus. Wien 16: 5, t. 2 t 1-5 (1900)

species :

Dryopteris peranema Li Bing Zhang; Taxon 61(6): 1211 (2012), nom. nov.

Distribution: China (Guizhou, Guangxi, Hubei, Hunan, Sichuan, Yunnan); Tibet; Taiwan; Nepal; Bhutan; India (Arunachal Pradesh, Manipur, Meghalaya, Nagaland, Sikkim, Uttarakhand, West Bengal); Myanmar (Chin, etc.); Vietnam; Philippines (Benguet: Mt. Pulag)

36 CHI TIB 38 TAI 40 BHU IND NEP 41 MYA VIE 42 PHI

Remarks: subgen. Nothoperanema. Inclusion of D. zhuweimingii fide Fraser-Jenkins et al. (2018)

Synonyms: hidden

= Dryopteris zhuweimingii Li Bing Zhang; Taxon 61(6): 1211 (2012), nom. nov. for Peranema luzonicum Copel.

= Peranema cyatheoides D.Don; Prodr. Fl. Nepal. 12 (1825)

= Peranema cyatheoides var. luzonicum (Copel.) Ching & S.H.Wu; Acta Phytotax. Sin. 21(4): 373-374 (1983)

= Peranema formosanum Hayata; Bot. Mag. (Tokyo) 26: 110 (1912), non Dryopteris formosana (Christ) C. Chr. 1905

= Peranema luzonicum Copel.; Philipp. J. Sci. 4(2): 111 (1909)

= Sphaeropteris barbata Wall.; Numer. List. [Wallich] no. 183 (1829), nomen, et Pl. Asiat. Rar. 1: 42 (1830)

species :

Dryopteris peranemiformis C.Chr.; Index Filic. 284 (1905), nom. nov.

Distribution: S-India (Kerala, Tamil Nadu: Nilgiri Hills, Annamalai Hills)

40 IND

Remarks: Subgen. Nothoperanema, sect. Shiehia

Conservation Status: IUCN CR for India (Fraser-Jenkins 2012)

Synonyms: hidden

= Ctenitis ferruginea (T.Moore ex Baker) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 296 (1938)

= Dryopsis ferruginea (T.Moore ex Baker) Holttum & P.J.Edwards; Kew Bull. 41(1): 201 (1986)

= Dryopteris ferruginea (T.Moore ex Baker) Kuntze; Revis. Gen. Pl. 2: 812 (1891), nom. inval.

= Lastrea ferruginea (T.Moore ex Baker) Bedd.; Ferns S. India t. 100 (1863) (non Aspidium ferrugineum Fée (1850))

= Nephrodium ferrugineum T.Moore ex Baker; Syn. Fil. [W. J. Hooker & Baker] 283 (1867)

species :

Dryopteris permagna M.G.Price; Gard. Bull. Singapore 30: 249 (1977)

Distribution: Philippines (Kalinga, Benguet, Mountain Province, South Cotabato)

42 PHI

Conservation Status: EN in the Philippines

species :

Dryopteris podophylla (Hook.) Kuntze; Revis. Gen. Pl. 2: 813 (1891)

Distribution: China (Fujian, Guangdong, Guangxi, Hainan, Yunnan)

36 CHI

Synonyms: hidden

= Aspidium podophyllum Hook.; J. Bot. 6: 236, t. 1 (1863)

= Dryopteris podophylla f. auriculata Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 400 (1938)

= Lastrea podophylla (Hook.) Hook.; Hooker´s J. Bot. Kew Gard Misc. 9: 339 (1857), non J. Sm. (1857)

= Nephrodium podophyllum (Hook.) Hook.; Sp. Fil. [W. J. Hooker] 4: 87 (1862)

species :

Dryopteris poilanei Tardieu; Bull. Soc. Bot. France 87: 368 (1941)

Distribution: Vietnam

41 VIE

Remarks: provisionally accepted name

species :

Dryopteris polita Rosenst.; Repert. Spec. Nov. Regni Veg. 13: 218 (1914)

Distribution: China (Hainan, Yunnan); S-Japan (Yakushima Isl.); Taiwan; Thailand; Cambodia; Vietnam; peninsular Malaysia (Cameron Highlands, Pahang); Sumatra; Borneo (Mt. Kinabalu, Sabah); Sulawesi; Philippines; New Guinea (Papua New Guinea)

36 CHI 38 JAP NNS TAI 41 CBD THA VIE 42 BOR MLY NWG PHI SUL SUM

Remarks: subgen. Acrorumohra

Synonyms: hidden

= Dryopteris chapaensis C.Chr. & Ching ex Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 474 (1938)

= Dryopteris exindusiata Ching & P.S.Chiu; Bot. Res. Acad. Sin. 1987(2): 32 (1987)

= Dryopteris laurisilvicola T.Suzuki; J. Jap. Bot. 11: 646, f. 2 (1935)

= Dryopteris lividis Ching & Chu H.Wang; Acta Phytotax. Sin. 8: 168, t.22(26) (1959)

= Dryopteris taitoensis Tagawa; Acta Phytotax. Geobot. 4: 134 (1935)

= Dryopteris viridis Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 474 (1938)

= Phegopteris polita (Rosenst.) Alderw.; Malayan Ferns Fern Allies, Suppl. 309 (1917)

species :

Dryopteris polylepis (Franch. & Sav.) C.Chr.; Index Filic. 285 (1905)

Distribution: South Korea; Japan

38 JAP KOR

Remarks: not in China

Synonyms: hidden

= Aspidium polylepis Franch. & Sav.; Enum. Pl. Jap. 2: 236, 631 (1877)

= Nephrodium filix-mas var. polylepis (Franch. & Sav.) Makino; Bot. Mag. (Tokyo) 15: 66 (1901)

= Nephrodium polylepis (Franch. & Sav.) Baker; Ann. Bot. (Oxford) 5: 320 (1891)

species :

Dryopteris pontica (Fraser-Jenk.) Fraser-Jenk.; Fern Gaz. 18(1): 7 (2007)

Distribution: ?Armenia; Turkey (N-Anatolia); NW-Iran

34 IRN TUR

Remarks: affinis agg.

Synonyms: hidden

= Dryopteris affinis subsp. pontica Fraser-Jenk.; Ann. Bot. Fenn. 33: 73 (1996)

species :

Dryopteris porosa Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 460 (1938)

Distribution: China (Guizhou, Sichuan, Yunnan); ?Bhutan; N-Thailand

36 CHI 41 THA

species :

Dryopteris protobissetiana K.Hori & N.Murak.; Acta Phytotax. Geobot. 66(1): 54 (2015)

Distribution: Japan (Kyushu)

38 JAP

species :

Dryopteris pseudocaenopteris (Kunze) Li Bing Zhang; Taxon 61(6): 1209 (2012)

Distribution: China (Sichuan, Hainan, Yunnan); Tibet; Taiwan; Nepal; Bhutan; India (Arunachal Pradesh, ?Assam, Manipur, Meghalaya, Mizoram, Nagaland, Sikkim, Tripura, West Bengal); Sri Lanka; Myanmar (Kachin, Chin, etc.); Thailand (widespread); Laos; Vietnam; peninsular Malaysia; Sumatra; Borneo (Sabah); Sulawesi; Philippines; Java; Lesser Sunda Isl. (Sumbawa); New Guinea (Irian Jaya, Papua New Guinea)

36 CHI TIB 38 TAI 40 BHU IND NEP SRL 41 LAO MYA THA VIE 42 BOR JAW LSI MLY NWG PHI SUL SUM

Remarks: subgen. Nothoperanema. Taxonomy fide Fraser-Jenkins et al. (2018)

Synonyms: hidden

= Cyathea aspidioides (Blume) Moritz; Syst. Verz. (Moritzi et al.) 1842-1844: 108 (1854)

= Diacalpe adscendens Ching ex S.H.Wu; Acta Phytotax. Sin. 21(4): 378 (1983), nom. illeg.

= Diacalpe aspidioides Blume; Enum. Pl. Javae 2: 241 (1828)

= Diacalpe aspidioides var. himalayensis Wall. ex Ching & S.K.Wu; S. H. Wu, Acta Phytotax. Sin. 21(4): 376 (1983), nom. nud.

= Diacalpe aspidioides var. hookeriana Wall. ex Ching & S.K.Wu; Fl. Xizangica 1: 194, t. 43, f. 1-3, t. 5, f. 7-8 (1983), nom. nud.

= Diacalpe aspidioides var. minor Ching ex S.K.Wu; Acta Phytotax. Sin. 21(4): 375, t. 1, f. 1 (1983)

= Diacalpe chinensis Ching & S.H.Wu; Acta Phytotax. Sin. 21(4): 376 (1983)

= Diacalpe hookeriana Wall. ex T.Moore; Gard. Chron. 1854: 135 (1854), nom. nud.

= Diacalpe laevigata Ching & S.H.Wu; Acta Phytotax. Sin. 21(4): 377 (1983)

= Diacalpe medogensis Ching & S.K.Wu; Fl. Xizangica 1: 196 (1983)

= Diacalpe omeiensis Ching; Bull. Fan Mem. Inst. Biol. Bot., ser. 2, 1: 293 (1949) (fide Fraser-Jenkins et al. 2018)

= Diacalpe pseudocaenopteris Kunze; Bot. Zeitung (Berlin) 4: 457 (1846)

= Dryopteris adscendens (Ching ex S.H.Wu) Li Bing Zhang; Taxon 61(6): 1210 (2012)

= Dryopteris diacalpe Li Bing Zhang; Taxon 61(6): 1210 (2012), nom. nov. for Diacalpe chinensis Ching & S. H. Wu, non Dryopteris chinensis (Baker) Koidz.

= Dryopteris hookeriana (Wall. ex T.Moore) Li Bing Zhang; Taxon 61(6): 1210 (2012)

= Dryopteris kungiana Li Bing Zhang; Taxon 61(6): 1210 (2012), nom. nov. for Diacalpe omeiensis Ching

= Dryopteris medogensis (Ching & S.K.Wu) Li Bing Zhang; Taxon 61(6): 1210 (2012)

= Peranema aspidioides (Blume) Mett.; Fil. Lechl. 2: 33 (1850)

= Peranema pseudocaenopteris (Kunze) Panigrahi; Pl. Sci. Res. (India) 20(1&2): 43 (1998), as ´pseudocystopteris´

= Physematium aspidioides (Blume) Kunze; Analecta Pteridogr. 43-44 (1837)

= Sphaeropteris hookeriana Wall.; Numer. List. [Wallich] no. 248 (1829), nomen (incorrect number, nom. nud.)

species :

Dryopteris pseudocomplexa (Fraser-Jenk.) P.D.Sell; Fl. Gr. Brit. Ireland 1: 671 (2018)

Distribution: England (Outer Hebrides, Scotland); Ireland; Norway; Germany (Sachsen-Anhalt); ?Austria; France (Vosges)

10 GRB IRE NOR 11 GER 12 FRA

Remarks: affinis agg., a fertile tetraploid probably deserving species rank

Synonyms: hidden

= Dryopteris affinis subsp. pseudocomplexa (Fraser-Jenk.) Fraser-Jenk.; Indian Fern J. 39(1): 120 (2022)

= Dryopteris cambrensis subsp. pseudocomplexa Fraser-Jenk.; Fern Gaz. 18(1): 6 (2007)

= Dryopteris pseudocomplexa (Fraser-Jenk.) Fraser-Jenk.; Indian J. Forest. 43(1): 83 (2020), isonym

species :

Dryopteris pseudodisjuncta (Tavel ex Fraser-Jenk.) Fraser-Jenk.; Fern Gaz. 18(1): 7 (2007)

Distribution: Norway; Belgium; Luxembourg; Germany (Baden-Württemberg, Bayern, Hessen, Nordrhein-Westfalen); Switzerland; England (SW-Scotland); Austria

10 GRB NOR 11 AUT BGM GER LUX SWI

Remarks: affinis agg.

Synonyms: hidden

= Aspidium filix-mas var. setosum Christ; Farnkr. Schweiz 1: 137 (1900)

= Dryopteris affinis subsp. pseudodisjuncta (Tavel ex Fraser-Jenk.) Fraser-Jenk.; Nordic J. Bot. 16(1): 4 (1996)

= Dryopteris affinis var. pseudodisjuncta Tavel ex Fraser-Jenk.; Ill. Fl. Mitt.-Eur., ed. 3, l(1): 147 (1984 publ. 1983)

= Dryopteris affinis var. setosa (Christ) Fraser-Jenk.; Sommerfeltia 6: xi (1987)

species :

Dryopteris pseudofilix-mas (Fée) Rothm.; Candollea 10: 96 (1945)

Distribution: Mexico (Chiapas, Ciudad de Mexico, Hidalgo, Mexico State, Michoacan, Morelos, Nuevo Leon, Oaxaca, Puebla, Queretaro, San Luis Potosi, Sonora, Tamaulipas, Veracruz); Guatemala

79 MEX 80 GUA

Synonyms: hidden

= Aspidium chrysocarpon Fée; Mém. Foug. 8 (Icon. Esp. Nouv.): 103 (1857)

= Aspidium filix-mas f. chrysocarpon (Fée) Milde; Fil. Eur. 120 (1867)

= Aspidium paleaceum var. majus E.Fourn.; Mexic. Pl. 1: 93 (1872)

= Aspidium pseudofilix-mas Fée; Mém. Foug. 8 (Icon. Esp. Nouv.): 103 (1857)

= Dryopteris ×chrysocarpa (Fée) Rothm.; Candollea 10: 95 (1945)

= Lastrea chrysocarpa (Fée) T.Moore; Index Fil. [T. Moore] 88 (1858)

species :

Dryopteris pseudolunanensis Tagawa; Acta Phytotax. Geobot. 9: 145 (1940)

Distribution: Taiwan

38 TAI

Conservation Status: very rare

Synonyms: hidden

= Dryopteris kuratae Nakaike ex Hoshiz. & K.A.Wilson; Amer. Fern J. 89(1): 10 (1999), nom. superfl.

species :

Dryopteris pseudoparasitica Alderw.; Nova Guinea 14: 19 (1924)

Distribution: New Guinea (Papua New Guinea)

42 NWG

Synonyms: hidden

= Cyclosorus pseudoparasiticus (Alderw.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 248 (1941)

= Thelypteris pseudoparasitica (Alderw.) C.F.Reed; Phytologia 17: 306 (1968)

species :

Dryopteris pseudosieboldii Hayata; Icon. Pl. Formosan. 4: 171-172, f. 111 (1914)

Distribution: Taiwan

38 TAI

Remarks: treated as full species by Knapp (2013), included as variety in D. enneaphylla by Flora of China

Synonyms: hidden

= Dryopteris enneaphylla var. pseudosieboldii (Hayata) Tagawa & K.Iwats.; Amer. Fern J. 50: 102 (1960)

= Dryopteris heteroneura (Tagawa) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 398 (1938)

= Dryopteris sieboldii var. heteroneura Tagawa; J. Jap. Bot. 12(7): 487 (1936)

species :

Dryopteris pseudosparsa Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 489 (1938)

Distribution: China (Yunnan, Guizhou, Guangxi, Sichuan); India (Meghalaya); Myanmar (Kachin, etc.); N-Thailand

36 CHI 40 IND 41 MYA THA

Synonyms: hidden

= Dryopteris cuneatiloba Ching & P.S.Chiu; Bot. Res. Acad. Sin. 1987(2): 11 (1987)

species :

Dryopteris pulvinulifera (Bedd.) Kuntze; Revis. Gen. Pl. 2: 213 (1891)

Distribution: China (NW-Yunnan); Nepal; Bhutan; India (Uttarakhand, Sikkim, Darjeeling, Meghalaya, Nagaland, Arunachal Pradesh, Assam State, Mizoram, West Bengal); Sri Lanka; Philippines

36 CHI 40 BHU IND NEP SRL 42 PHI

Synonyms: hidden

= Dryopteris harae H.Itô; Fl. E. Himal.: 476 (1966)

= Dryopteris reholttumii M.G.Price; Fiddlehead Forum 8(3): 19 (1981), nom. nud.

= Lastrea pulvinulifera Bedd.; Ferns Brit. India t. 333 (1870)

= Lastrea pulvinulifera var. zeylanica Bedd.; Ferns Brit. India Suppl. 17 (1876)

= Lastrea sparsa var. zeylanica (Bedd.) Bedd.; Ferns Brit. India 254 (1883)

= Nephrodium pulvinuliferum (Bedd.) Baker; Syn. Fil. [W. J. Hooker & Baker], ed. 2: 500 (1874) (p. p.)

= Nephrodium sparsum var. squamulosum C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 524 (1880) (p. p.)

= Polypodium marginatum T.Moore; Index Fil. [T. Moore] 297 (1861)

species :

Dryopteris purpurascens (Blume) Christ; Philipp. J. Sci. 2: 213 (1907)

Distribution: Borneo (Mt. Kinabalu, Sabah); Sulawesi; Philippines; New Guinea (Papua New Guinea)

42 BOR NWG PHI SUL

Synonyms: hidden

= Aspidium purpurascens Blume; Enum. Pl. Javae 2: 169 (1828)

= Nephrodium purpurascens (Blume) Hook.; Sp. Fil. [W. J. Hooker] 4: 132, t. 262 (1862)

species :

Dryopteris pycnopteroides (Christ) C.Chr.; Index Filic., Suppl. 2 (1906-1912): 38 (1913)

Distribution: China (Sichuan, Guizhou, Yunnan, Hubei); ?Japan

36 CHI

Synonyms: hidden

= Aspidium pycnopteroides Christ; Bull. Acad. Int. Géogr., sér. 3, 16: 116 (1906)

= Dryopteris sunii Ching; Bull. Fan Mem. Inst. Biol. Bot., ser. 2, 1: 295 (1949) (fide Fraser-Jenkins (1986))

species :

Dryopteris raiateensis (J.W.Moore) Li Bing Zhang; Taxon 61(6): 1209 (2012)

Distribution: Society Isl. (Raiatea)

61 SCI

Remarks: sometimes included in D. leucorhachis from Rarotonga

Synonyms: hidden

= Acrophorus raiateensis J.W.Moore; Bull. Bernice P. Bishop Mus. 102: 6 (1933)

species :

Dryopteris ramosa (C.Hope) C.Chr.; Index Filic. 266 (1905)

Distribution: E-Afghanistan; Pakistan (Kurram, Chitral, Swat, Hazara, Gilgit); Nepal; Bhutan; India (Himachal Pradesh, Jammu & Kashmir, Uttarakhand)

34 AFG 40 BHU IND NEP PAK

Synonyms: hidden

= Lastrea rigida Bedd.; Ferns Brit. India 251 (1883)

= Nephrodium ramosum C.Hope; J. Bot. 126 (1896)

species :

Dryopteris redactopinnata Basu & Panigrahi; Indian J. Forest. 3(3): 270, t. 2 (1980)

Distribution: China (Yunnan, Sichuan); Tibet; Taiwan; Pakistan (Hazara); Nepal; Bhutan; India (Jammu & Kashmir, Sikkim, Himachal Pradesh, Uttarakhand, Arunachal Pradesh)

36 CHI TIB 38 TAI 40 BHU IND NEP PAK

Synonyms: hidden

= Dryopteris pseudofibrillosa Ching; Fl. Xizangica 1: 252 (1983)

species :

Dryopteris reflexosquamata Hayata; Icon. Pl. Formosan. 4: 176, f. 114 (1914)

Distribution: China (Sichuan, Guizhou, Hunan, NE-Yunnan); Taiwan; ?India (?Sikkim)

36 CHI 38 TAI

Synonyms: hidden

= Dryopteris mingetsuensis Hayata; Icon. Pl. Formosan. 6: 281, f. 109 (1916)

= Dryopteris paludicola Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(3): 21 (1984)

species :

Dryopteris remota (A.Braun ex Döll) Druce; List Brit. Pl. 87 (1908)

Name: Entferntfiedriger Wurmfarn, Entferntfiedriger Dornfarn (DE); Scaly Buckler-fern (EN)

Distribution: +England (+Scotland); +Ireland (+W-Ireland); Germany (Baden-Württemberg, Bayern); Switzerland; ?Liechtenstein; ?Austria; Czech Republic; Spain; France; Slovenia; Croatia; European Russia; Northern Caucasus; Georgia [Caucasus]; Turkey (Anatolia)

10 GRB IRE 11 CZE GER SWI 12 FRA SPA 13 CRO SLV 14 RUS 33 GEO NCS 34 TUR

Remarks: filix-mas agg., derived from D. carthusiana and a diploid sexual ancestor of D. affinis.

Synonyms: hidden

= Aspidium filix-mas subsp. remotum (A.Braun ex Döll) Nyman; Consp. Fl. Eur. Suppl. 1: 865 (1884)

= Aspidium filix-mas var. remotum (A.Braun ex Döll) Jess.; Deut. Excurs.-Fl. 613 (1879)

= Aspidium rigidum var. remotum A.Braun ex Döll; Rhein. Fl. 16 (1843)

= Dryopteris kemulariae Mikheladze; Zametki Sist. Geogr. Rast. 23: 56 (1963)

= Dryopteris ×remota (A.Braun ex Döll) Hayek; Fl. Steiermark 56 (1908)

= Lastrea remota (A.Braun ex Döll) T.Moore; Index Fil. [T. Moore] 102 (1858)

= Nephrodium borbasio E.Walter; Guétrot, 104 (1929), var. orthogr.

= Nephrodium remoti Guétrot; 37 (1927), var. orthogr.

= Nephrodium remotum (A.Braun ex Döll) Murr; (1923), nom. illeg., non Heward nec Hook.

= Nephrodium spinulosum (A.Braun) Baker; (1867)

= Nephrodium spinulosum proles jordanii Rouy; Fl. France [Rouy & Foucaud] 14: 411 (1913)

= Nephrodium spinulosum subsp. jordanii (Rouy) P.E.Fourn.; Fl. Compl. Plaine Franç.: 595 (1928)

= Nephrodium spinulosum subsp. remotum (A.Braun ex Döll) G.Hensl.; Cat. Brit. Pl.: 44 (1879)

= Polystichum ×remotum (A.Braun ex Döll) P.E.Fourn.; Quatre Fl. France: 17 (1934)

species :

Dryopteris remotissima (Christ) Koidz.; Bot. Mag. (Tokyo) 43: 386 (1929)

Distribution: Japan

38 JAP

Synonyms: hidden

= Aspidium spinulosum var. remotissimum Christ; Bot. Mag. (Tokyo) 24: 240 (1910)

species :

Dryopteris renchangiana Z.Y.Zuo & D.Z.Li; Pl. Diversity 44(2): 185 (2021)

Distribution: China (Chongqing, Sichuan)

36 CHI

Remarks: sect. Diclisodon, see Zuo et al. 2022

Synonyms: hidden

= Dryopteris glabrior Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(4): 11 (1984), nom. illeg.

species :

Dryopteris rhomboideo-ovata H.Itô; J. Jap. Bot. 9: 55 (1933)

Distribution: Japan

38 JAP

species :

Dryopteris rodolfii J.P.Roux; Webbia 59(1): 143 (2004)

Distribution: Ethiopia; Kenya

24 ETH 25 KEN

species :

Dryopteris rossii C.Chr.; Mem. Revista Acad. Nac. Ci. Antonio Alzate 32: 178 (1912)

Name: Ross´ woodfern (EN)

Distribution: USA (Arizona); Mexico (Aguascalientes, Baja California Norte, Baja California Sur, Chihuahua, Colima, Ciudad de Mexico, Durango, Guanajuato, Guerrero, Hidalgo, Jalisco, Mexico State, Michoacan, Morelos, Nayarit, Oaxaca, Queretaro, San Luis Potosi, Sinaloa, Zacatecas); Nicaragua; Costa Rica

76 USA 79 MEX 80 COS NIC

Conservation Status: IUCN V (Arizona: E)

Synonyms: hidden

= Dryopteris patula var. rossii C.Chr.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 7, 10(2): 72 (1912 publ. 1913)

species :

Dryopteris rosthornii (Diels) C.Chr.; Index Filic. 289 (1905)

Distribution: China (Gansu, Guizhou, Hubei, Shanxi, Sichuan, Yunnan)

36 CHI

Remarks: not in India

Synonyms: hidden

= Aspidium filix-mas var. omeiense Christ; Bull. Acad. Int. Géogr., sér. 3, 16: 1906: 117 (1906)

= Dryopteris centrochinensis Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 423 (1938)

= Dryopteris filix-mas var. rosthornii (Diels) C.Chr.; Acta Horti Gothob. 1: 58 (1924)

= Dryopteris hupehensis Ching; Fl. Tsinling. 2: 226 (1974)

= Dryopteris omeicola Ching; Acta Phytotax. Sin. 10: 303 (1965)

= Nephrodium rosthornii Diels; Bot. Jahrb. Syst. 29: 190 (1900)

species :

Dryopteris rubiginosa (Brack.) Kuntze; Revis. Gen. Pl. 2: 813 (1891)

Distribution: Hawaii (Kauai, Oahu, Molokai, Lanai, Maui, Hawaii Isl.)

63 HAW

Remarks: subgen. Nothoperanema

Synonyms: hidden

= Aspidium rubiginosum (Brack.) Mann; Proc. Amer. Acad. Arts 7: 217 (1868)

= Ctenitis rubiginosa (Brack.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 125 (1947)

= Lastrea rubiginosa Brack.; U.S. Expl. Exped., Filic. 16: 201 (1854)

= Nephrodium rubiginosum (Brack.) Hook.; Sp. Fil. [W. J. Hooker] 4: 143 (1862)

= Nothoperanema rubiginosum (Brack.) A.R.Sm. & D.D.Palmer; Amer. Fern J. 85(2): 63 (1995)

species :

Dryopteris rubrobrunnea W.M.Chu; Acta Bot. Yunnan., Suppl. 5: 46 (1992)

Distribution: China (Yunnan); NE-India (Arunachal Pradesh); Myanmar (Kachin, etc.)

36 CHI 40 IND 41 MYA

Conservation Status: IUCN EN for India (Fraser-Jenkins 2012)

Synonyms: hidden

= Dryopteris rubripes Ching & Chu ex Fraser-Jenk.; (1986)

species :

Dryopteris ruwenzoriensis C.Chr. ex Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.), Bot. 14(3): 204 (1986)

Distribution: D.R.Congo [Zaire]; Rwanda; Uganda

23 RWA ZAI 25 UGA

species :

Dryopteris ryo-itoana Sa.Kurata; J. Geobot. 15: 84 (1967)

Distribution: China (Jiangxi, Zhejiang); Japan; Taiwan

36 CHI 38 JAP TAI

Remarks: new record for Taiwan see Knapp (2013)

Synonyms: hidden

= Dryopteris dispar Ching & C.F.Zhang; Bull. Bot. Res., Harbin 3(3): 17 (1983)

= Dryopteris gutishanensis Ching & C.F.Zhang; Bull. Bot. Res., Harbin 3(3): 29 (1983)

= Dryopteris kaihuaensis Ching & C.F.Zhang; Bull. Bot. Res., Harbin 3(3): 24 (1983)

= Dryopteris lushanensis Ching & P.S.Chiu; Bot. Res. Acad. Sin. 1987(2): 20 (1987)

= Dryopteris pseudofuscipes Ching & P.S.Chiu; Bot. Res. Acad. Sin. 1987(2): 3 (1987)

= Dryopteris triangularifrons Ching; Bull. Bot. Res., Harbin 2(2): 64 (1982)

species :

Dryopteris sabaei (Franch. & Sav.) C.Chr.; Index Filic. 290 (1905)

Distribution: S-Japan (incl. Yakushima Isl.)

38 JAP NNS

Remarks: sect. Diclisodon

Synonyms: hidden

= Aspidium sabaei Franch. & Sav.; Enum. Pl. Jap. 2: 259, 632 (1877)

= Nephrodium sabaei (Franch. & Sav.) Hand.-Mazz.; Symb. Sin. 6: 25 (1929)

species :

Dryopteris sacrosancta Koidz.; Bot. Mag. (Tokyo) 88: 108 (1924)

Distribution: China (Liaoning, Shandong, Zhejiang); North Korea; South Korea (incl. Cheju Isl.); Japan

36 CHI 38 JAP KOR NKO

Synonyms: hidden

= Dryopteris bissetiana var. sacrosancta (Koidz.) H.Itô; Bot. Mag. (Tokyo) 50: 36 (1936)

= Dryopteris kobayashii Kitag.; Rep. First Sci. Exped. Manchoukuo 4(2): 56 (1935) (fide Fraser-Jenkins (1986))

= Dryopteris tieanzuensis Ching & P.S.Chiu; Bot. Res. Acad. Sin. 1987(2): 24 (1987)

= Dryopteris varia subsp. sacrosancta (Koidz.) Sugim.; Keys Herb. Pl. Jap. Pterid. 281, 405 (1966)

= Dryopteris varia subsp. tenerifrons H.Itô; Bot. Mag. (Tokyo) 50: 37 (1936)

= Dryopteris varia var. sacrosancta (Koidz.) Ohwi; Fl. Jap. [Ohwi] 88 (1957)

= Polystichum bissetianum var. sacrosanctum (Koidz.) Nakai; Bot. Mag. (Tokyo) 45: 103 (1931)

= Polystichum sacrosanctum (Koidz.) Koidz.; Bot. Mag. (Tokyo) 43: 388 (1929)

species :

Dryopteris saffordii C.Chr.; Amer. Fern J. 1: 94 (1911)

Distribution: Peru; Bolivia (La Paz, Tarija)

83 BOL PER

species :

Dryopteris sandwicensis (Hook. & Arn.) C.Chr.; Index Filic. 290 (1905)

Distribution: Hawaii (Kauai, Oahu, Molokai, Lanai, Maui, Hawaii Isl.)

63 HAW

Conservation Status: IUCN E

Synonyms: hidden

= Phegopteris sandwicensis (Hook. & Arn.) Mann; Proc. Amer. Acad. Arts 7: 218 (1868)

= Polypodium sandwicense Hook. & Arn.; Bot. Beechey Voy. 105 (1832)

= Thelypteris sandwicensis (Hook. & Arn.) Fosberg; Occas. Pap. Bernice P. Bishop Mus. 23: 30 (1962)

species :

Dryopteris saxifraga H.Itô; Bot. Mag. (Tokyo) 60: 125 (1936)

Distribution: China (Jilin, Liaoning); North Korea (incl. Cheju Isl.); South Korea; Japan

36 CHI 38 JAP KOR NKO

Synonyms: hidden

= Dryopteris varia subsp. saxifraga (H.Itô) Sugim.; Keys Herb. Pl. Jap. Pterid. 282, 405 (1966)

= Dryopteris varia var. saxifraga (H.Itô) H.Ohba; Sci. Rep. Tohoku Imp. Univ., Ser. 4, Biol. 36: 111 (1971)

species :

Dryopteris saxifragivaria Nakai; J. Jap. Bot. 18: 286 (1942), pro hybr.

Distribution: South Korea (incl. Cheju Isl.)

38 KOR

Remarks: stabilized hybrid derived from Dryopteris saxifraga x bissetiana

Synonyms: hidden

= Dryopteris saxifraga var. deltoidea H.Itô; Bot. Mag. (Tokyo) 50: 126 (1936)

= Dryopteris saxifragivaria f. cristata Nakai

= Dryopteris varia f. cristata (Nakai) S.Y.Oh

species :

Dryopteris scabrosa (Kunze) Kuntze; Revis. Gen. Pl. 2: 813 (1891)

Distribution: S-India (Kerala, Tamil Nadu)

40 IND

Remarks: Subgen. Nothoperanema, sect. Shiehia

Conservation Status: IUCN VU for India (Fraser-Jenkins 2012)

Synonyms: hidden

= Aspidium scabrosum Kunze; Linnaea 24: 286 (1851)

= Ctenitis scabrosa (Kunze) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 292 (1938)

= Dryopsis scabrosa (Kunze) Holttum & P.J.Edwards; Kew Bull. 41(1): 199 (1986)

= Lastrea scabrosa (Kunze) T.Moore; Index Fil. [T. Moore] 103 (1858)

= Nephrodium scabrosum (Kunze) Baker; Syn. Fil. [W. J. Hooker & Baker] 283 (1867)

= Polypodium marginale Wall.; Numer. List. [Wallich] no. 322 (1829), nomen

= Polypodium nigrocarpum Bedd.; Ferns S. India t. 169 (1863) non Blume 1828

species :

Dryopteris schimperiana (Hochst. ex A.Braun) C.Chr.; Index Filic. 291 (1905)

Distribution: D.R.Congo [Zaire]; Rwanda; Burundi; Sudan; Ethiopia (Ahmar Mts.); Eritrea; Uganda; Kenya; Tanzania; Malawi; N-Yemen

23 BUR RWA ZAI 24 ERI ETH SUD 25 KEN TAN UGA 26 MLW 35 YEM

Synonyms: hidden

= Aspidium schimperianum Hochst. ex A.Braun; Flora 708 (1841)

= Lastrea filix-mas var. schimperiana (Hochst. ex A.Braun) Bedd.; Ferns Brit. India Suppl. 58 (1892)

= Nephrodium filix-mas var. schimperianum (Hochst. ex A.Braun) C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 520, t. 69, f. 2 (1880)

= Nephrodium schimperianum (Hochst. ex A.Braun) C.Hope; J. Bombay Nat. Hist. Soc. 14: 733 (1903) (non Aspidium schimperianum Hochst. (1841))

= Polystichum schimperianum (Hochst. ex A.Braun) Keyserl.; Polyp. Herb. Bunge 44 (1873)

species :

Dryopteris schizopaleata Eb.Fisch. & Lobin; Phytotaxa 587(2): 138 (2023)

Distribution: Rwanda

23 RWA

species :

Dryopteris schorapanensis Askerov; Izv. Akad. Nauk Azerbaidzhansk. SSR Biol. Nauk. 1978(4): 4 (1978 publ. 1979), ex Fraser-Jenkins, Fern Gaz. 18(1): 7 (2007), pro hybr.

Distribution: Northern Caucasus (Krasnodar, Daghestan); Georgia [Caucasus] (incl. Abkhazia); Azerbaijan; Turkey (Anatolia); NW-Iran

33 AZE GEO NCS 34 IRN TUR

Remarks: affinis agg.

Synonyms: hidden

= Dryopteris affinis subsp. persica Fraser-Jenk.; Willdenowia 10(1): 113 (1980)

species :

Dryopteris scottii (Bedd.) Ching; Bull. Biol. Dept. Sci. Coll. Sun Yatsen Univ. 6: 3 (1933)

Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Jiangsu, Jiangxi, Sichuan, Yunnan, Zhejiang); Japan; Taiwan; Nepal; Bhutan; India (Arunachal Pradesh, Assam State, Manipur, Meghalaya, Mizoram, Nagaland, Sikkim, West Bengal, Jammu & Kashmir); Myanmar (Kachin, Chin, etc.); N-Thailand; Vietnam; Sumatra; Borneo (Mt. Kinabalu, Sabah)

36 CHI 38 JAP TAI 40 BHU IND NEP 41 MYA THA VIE 42 BOR SUM

Synonyms: hidden

= Dryopteris grossa (Christ) C.Chr.; Index Filic. 269 (1905)

= Dryopteris hirtipes var. exinvolucrata (C.B.Clarke) Sledge ex Panigrahi & Basu; Proc. Indian Sci. Congr. Assoc. 69(3/6): 71 (1982) (fide Fraser-Jenkins 1997)

= Dryopteris nigrisquama f. subdecipiens (Hayata) H.Itô; Nov. Fl. Jap. [Nakai & Honda] 4: 16 (1938 publ. 1939)

= Dryopteris nigrisquama Hayata; Icon. Pl. Formosan. 4: 167, f. 106 (1914)

= Dryopteris subdecipiens Hayata; Icon. Pl. Formosan. 4: 181, f. 119 (1914)

= Nephrodium hirtipes var. exinvolucrata C.B.Clarke; J. Linn. Soc., Bot. 25: 93 (1889)

= Phegopteris grossa Christ; Bull. Herb. Boissier, sér. 1, 7: 13 (1899)

= Phegopteris melanolepis Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 16: 25 (1914)

= Phegopteris scottii (Bedd.) Bedd.; Ferns Brit. India Suppl. 19 (1876)

= Polypodium scottii Bedd.; Ferns Brit. India t. 345 (1870)

species :

Dryopteris sericea C.Chr.; Bot. Gaz. 66: 336 (1913)

Distribution: China (Gansu, Hubei, Shaanxi, Shanxi)

36 CHI

species :

Dryopteris serratodentata (Bedd.) Hayata; Icon. Pl. Formosan. 4: 179, f. 116 (1914)

Distribution: China (W-Sichuan, NW-Yunnan); Tibet; Taiwan; Pakistan (Hazara); Nepal; Bhutan; India (Jammu & Kashmir, Himachal Pradesh, Uttarakhand, Sikkim, Darjeeling, Arunachal Pradesh, West Bengal); Myanmar

36 CHI TIB 38 TAI 40 BHU IND NEP PAK 41 MYA

Synonyms: hidden

= Dryopteris filix-mas subsp. serratodentata (Bedd.) C.Chr.; Index Filic. 265 (1905)

= Dryopteris filix-mas var. serratodentata (Bedd.) Hayata; J. Coll. Sci. Imp. Univ. Tokyo 30: 416 (1911)

= Lastrea filix-mas var. odontoloma Baker; Syn. Fil. [W. J. Hooker & Baker] 498 (1874) (excl. fig. of Bedd. cited and the var. nidus)

= Lastrea filix-mas var. serratodentata Bedd.; Suppl. Ferns Brit. Ind. 55 (1892)

= Lastrea odontoloma (Baker) Bedd.; Ferns Brit. India t. 115 (1865)

= Nephrodium filix-mas var. odontoloma Baker; Syn. Fil. [W. J. Hooker & Baker], ed. 2: 498 (1874)

= Nephrodium odontoloma (Baker) C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 521 (1880)

= Nephrodium serratodentatum (Bedd.) C.Hope; J. Bombay Nat. Hist. Soc. 12: 622, t. 10 (1899)

= Woodsia veitchii Christ; Bull. Acad. Int. Géogr., sér. 3, 16: 121 (1906)

species :

Dryopteris shiakeana H.Shang & Y.H.Yan; Phytotaxa 218(2): 158 (2015)

Distribution: China (Guangdong)

36 CHI

species :

Dryopteris shikokiana (Makino) C.Chr.; Index Filic. 292 (1905)

Distribution: China (Guangxi, Guizhou, Hunan, Sichuan, Yunnan); Japan

36 CHI 38 JAP

Remarks: not in India. subgen. Nothoperanema

Synonyms: hidden

= Aspidium shikokianum Makino; Bot. Mag. (Tokyo) 6(60): 46 (1892)

= Ctenitis shikokiana (Makino) H.Itô; Nov. Fl. Jap. [Nakai & Honda] 4: 88 (1939)

= Dryopteris hirtosparsa Christ; Bull. Acad. Int. Géogr. Bot. 20: 176 (1909)

= Nephrodium shikokianum Makino; Bot. Mag. (Tokyo) 18: 62 (1899)

= Nothoperanema shikokianum (Makino) Ching; Acta Phytotax. Sin. 11: 28 (1966)

= Nothoperanema shikokianum (Makino) Nakaike; Enum. Pterid. Jap., Filic. 240 (1975), isonym

species :

Dryopteris shiroumensis Sa.Kurata & Nakam.; J. Geobot. 9: 54 (1960)

Distribution: Japan (Honshu)

38 JAP

Conservation Status: IUCN R

species :

Dryopteris sichotensis Kom.; Izv. Imp. Bot. Sada Petra Velikago 16: 146 (1916)

Distribution: Russian Far East (Sakhalin, Amur, Khabarovsk, Primorye); China (Heilongjiang, Jilin); North Korea; Japan

31 RFE 36 CHI 38 JAP NKO

Remarks: D. coreanomontana is sometimes kept separate

Synonyms: hidden

= Dryopteris barbellata Fomin; Fl. Sibir. Orient. Extremi 5: 59, fig. (1930)

= Dryopteris coreanomontana Nakai; Bot. Mag. (Tokyo) 36: 132 (1921)

= Dryopteris crassirhizoma var. setosa (Christ) Miyabe & Kudô; Fl. Hokkaido 1: 15 (1930)

= Dryopteris filix-mas var. setosa Christ; Bull. Acad. Int. Géogr. Bot. 1909: 163 (1909)

= Dryopteris setosa (Christ) Kudô; Medical Fl. Hokkaido: no. 1 (1922), nom. illeg., non Kuntze 1891

= Dryopteris sichotensis subsp. coreano-montana (Nakai) Vorosch.; Florist. Issl. Razn. Raionakh SSSR [A. K. Skvortsov] 144 (1985)

species :

Dryopteris sieboldii (T.Moore) Kuntze; Revis. Gen. Pl. 2: 813 (1891)

Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hunan, Jiangxi, Zhejiang); Japan

36 CHI 38 JAP

Synonyms: hidden

= Aspidium podophyllum E.J.Lowe; Ferns 6: t. 34-35 (1857) (non Hook. (1853))

= Aspidium sieboldii (T.Moore) Van Houtte ex Mett.; Fil. Hort. Bot. Lips. 87, t. 20, f. 1-4 (1856)

= Lastrea podophylla J.Sm.; Cat. Cult. Ferns: 57 (1857)

= Lastrea sieboldii (T.Moore) T.Moore; Index Fil. [T. Moore] 88 (1857)

= Nephrodium sieboldii (T.Moore) Hook.; Fil. Exot.: t. 31 (1859)

= Polystichum sieboldii (T.Moore) Keyserl.; Polyp. Herb. Bunge 41 (1873)

= Pycnopteris sieboldii T.Moore; Gard. Chron. 1855: 468 (1855)

species :

Dryopteris sikkimensis (Bedd.) Kuntze; Revis. Gen. Pl. 2: 813 (1891)

Distribution: China (Yunnan); Tibet; Bhutan; India (Sikkim, Arunachal Pradesh); Myanmar (Kachin, etc.)

36 CHI TIB 40 BHU IND 41 MYA

Conservation Status: IUCN VU for India (Fraser-Jenkins 2012)

Synonyms: hidden

= Aspidium sikkimense (Bedd.) Baker; Syn. Fil. [W. J. Hooker & Baker] 256 (1867)

= Dryopteris pseudosikkimensis Ching & S.K.Wu; Fl. Xizangica 1: 267 (1983)

= Lastrea sikkimensis (Bedd.) Bedd.; Ferns Brit. India 259 (1883), isonym

= Lastrea sikkimensis (Bedd.) C.B.Clarke; J. Linn. Soc., Bot. 17: 403 (1879)

= Nephrodium sikkimense (Bedd.) C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 525 (1880)

= Polystichum sikkimense Bedd.; Ferns Brit. India t. 1270 (1866)

= Thelypteris sikkimensis (Bedd.) C.F.Reed; Phytologia 17: 313 (1968)

species :

Dryopteris simasakii (H.Itô) Sa.Kurata; J. Geobot. 18: 5 (1970)

Distribution: China (Guangxi, Guizhou, Sichuan, Yunnan, Zhejiang); Japan

36 CHI 38 JAP

Synonyms: hidden

= Dryopteris erythrochlamys Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(4): 5 (1984)

= Dryopteris excelsior Ching & P.S.Chiu; Bot. Res. Acad. Sin. 1987(2): 8 (1987)

= Dryopteris indusiata var. simasakii H.Itô; J. Jap. Bot. 9: 57, f. 6 (1933)

= Dryopteris labordei var. simasakii (H.Itô) H.Itô

= Dryopteris laodianensis Ching & P.S.Chiu; Bot. Res. Acad. Sin. 1987(2): 16 (1987)

= Dryopteris occidentalizhejiangensis Ching & P.S.Chiu; Bot. Res. Acad. Sin. 1987(2): 17 (1987)

species :

Dryopteris sinonepalensis Z.Y.Zhuo & Fraser-Jenk.; Pl. Diversity 44(2): 185 (2021), nom. nov.

Distribution: China (Chongqing, Sichuan, Yunnan, ?Guangxi, ?Guizhou); Tibet; Nepal; Bhutan

36 CHI TIB 40 BHU NEP

Remarks: sect. Diclisodon, see Zuo et al. 2022. Often confused with D. yoroii

Synonyms: hidden

= Aspidium nitidulum Wall.; Numer. List. [Wallich] no. 392 (1829), nomen

= Aspidium nitidulum Wall. ex Kuhn; Linnaea 36: 117 (1869), nom. illeg., non (C. Presl) Kunze ex Mett.

= Dryopteris nitidula (Wall. ex Kuhn) Mitsuta & S.K.Wu; Vasc. Pl. Hengduan Mount. 1: 131 (1993), nom. illeg., non (C. Presl) Kuntze 1891

= Dryopteris sparsa subsp. nitidula (Bedd.) C.Chr.; Index Filic. 293 (1905)

= Dryopteris sparsa var. nitidula (Bedd.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 472 (1938)

= Lastrea sparsa var. nitidula Bedd.; Suppl. Ferns S. India 17, t. 374 (1876)

= Nephrodium sparsum var. nitidulum (Bedd.) C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 524 (1880)

species :

Dryopteris sledgei Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.), Bot. 18(5): 361 (1989)

Distribution: S-India (Tamil Nadu); +Sri Lanka

40 IND SRL

Conservation Status: IUCN EN for India (Fraser-Jenkins 2012)

species :

Dryopteris sordidipes Tagawa; Acta Phytotax. Geobot. 3: 29 (1934)

Distribution: S-Japan (incl. Yakushima Isl.); Ryukyu Isl.; Taiwan; Vietnam

38 JAP NNS TAI 41 VIE

species :

Dryopteris sororia (Maxon) M.A.McHenry, Sundue & Barrington; Taxon 62(6): 1158 (2013)

Distribution: Mexico (Chiapas); Guatemala

79 MEX 80 GUA

Synonyms: hidden

= Adenoderris sororia Maxon; Bot. Gaz. 39: 368 (1905)

species :

Dryopteris sparsa (D.Don) Kuntze; Revis. Gen. Pl. 2: 613 (1891)

subspecies :

subsp. sparsa

Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Jiangxi, Shanxi, Sichuan, Yunnan, Zhejiang); Tibet; Japan; Ryukyu Isl.; Taiwan; Bhutan; India (Arunachal Pradesh, Himachal Pradesh, Meghalaya, Sikkim, Uttarakhand, West Bengal); Sri Lanka; Myanmar (Kachin, etc.)

36 CHI TIB 38 JAP NNS TAI 40 BHU IND SRL 41 MYA

Remarks: sect. Diclisodon, see Zuo et al. 2022

Synonyms: hidden

= Aspidium densum Wall.; Numer. List. [Wallich] no. 390 (1829), nomen

= Aspidium densum Wall. ex Mett.; Abh. Senckenberg. Naturf. Ges. 2(2) [Pheg. u. Asp.]: 349 (1858)

= Aspidium oppositum Wall.; Numer. List. [Wallich] no. 7080 (1832), nomen, non (Vahl) Sw. (1829)

= Aspidium sparsum (D.Don) Spreng.; Syst. Veg., ed. 16 [Sprengel] 4(1): 106 (1827)

= Dryopteris parasparsa Ching & S.K.Wu; Fl. Xizangica 1: 267 (1983)

= Dryopteris sinosparsa Ching & K.H.Shing; Fl. Fujianica 1: 601 (1982)

= Dryopteris sparsa var. ryukyuensis Seriz.; J. Jap. Bot. 46: 279 (1971)

= Lastrea densa (Wall.) C.Presl; Tent. Pterid. 76 (1836)

= Lastrea sparsa (D.Don) Bedd.; Ferns S. India t. 103 (1863), isonym

= Lastrea sparsa (D.Don) T.Moore; Index Fil. [T. Moore] 87, 104 (1858)

= Nephrodium sparsum D.Don; Prodr. Fl. Nepal. 6 (1825)

= Polypodium oppositum Wall.; Numer. List. [Wallich] no. 7080 (1832), nomen

= Polystichum sparsum (D.Don) Keyserl.; Polyp. Herb. Bunge 43 (1873)

subspecies :

subsp. lehalii Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 322 (2008) [nothossp.]

Distribution: India (Darjeeling)

40 IND

Remarks: Dryopteris sparsa ssp. sparsa x sparsa ssp. rectipinnula

subspecies :

subsp. rectipinnula Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 320 (2008)

Distribution: China (Yunnan, Sichuan, Guizhou); Tibet; S-Japan; Nepal; Bhutan; India (Arunachal Pradesh, Assam State, Chhattisgarh, Karnataka, Kerala, Madhya Pradesh, Maharashtra, Manipur, Meghalaya, Mizoram, Nagaland, Tamil Nadu, Sikkim, Tripura, West Bengal); Myanmar; Thailand; ?Laos; ?Cambodia; Vietnam; peninsular Malaysia (Maxwell Hill, Perak, Fraser´s Hill & Cameron Highlands, Pahang); Sumatra; Borneo (Mt. Kinabalu, Sabah); Sulawesi; Palawan; Philippines; Java; Lesser Sunda Isl. (Bali, Flores); New Guinea (Irian Jaya, Papua New Guinea); ?Solomon Isl.; Australia (NE-Queensland); Fed. States of Micronesia (Pohnpei, Kosrae)

36 CHI TIB 38 JAP 40 BHU IND NEP 41 MYA THA VIE 42 BOR JAW LSI MLY NWG PHI PLW SUL SUM 50 AUS 62 MIC

Conservation Status: IUCN R (Queensland)

Synonyms: hidden

= Aspidium catophoron Kunze; Bot. Zeitung (Berlin) 6: 262 (1848)

= Aspidium viridescens (Baker) Miq.; Cat. Mus. Bot. Lugd.-Bat. 127 (1870)

= Aspidium weigleanum Kunze; Linnaea 24: 284 (1851)

= Dryopteris rheophila Mitsuta ex Darnaedi; M. Kato & K. Iwats., J. Jap. Bot. 64(10): 302 (1989) (fide Fraser-Jenkins et al. 2018)

= Dryopteris sparsa subsp. viridescens (Baker) Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 321 (2008)

= Dryopteris sparsa var. viridescens (Baker) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 473 (1938)

= Dryopteris viridescens (Baker) Kuntze; Revis. Gen. Pl. 2: 814 (1891)

= Nephrodium sparsum var. latisquamum C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 524 (1880)

= Nephrodium viridescens Baker; Syn. Fil. [W. J. Hooker & Baker] 275 (1867)

species :

Dryopteris speciosissima Copel.; Univ. Calif. Publ. Bot. 18: 219. (1942)

Distribution: New Guinea (Irian Jaya, Papua New Guinea)

42 NWG

Remarks: Subgen. Nothoperanema, sect. Stenolepia

Synonyms: hidden

= Ctenitis speciosissima (Copel.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 125 (1947)

= Stenolepia speciosissima (Copel.) Holttum ex P.J.Edwards; Guide Alpine Subalpine Fl. Mount Jaya [R. J. Johns et al.] 88 (2006)

species :

Dryopteris sphaeropteroides (Baker) C.Chr.; Index Filic. 293 (1905)

Distribution: China (Sichuan, Yunnan)

36 CHI

Remarks: Subgen. Dryopsis

Synonyms: hidden

= Aspidium sphaeropteroides (Baker) Christ; Bull. Acad. Int. Géogr. Bot. 119 (1900)

= Athyrium sphaeropteroides (Baker) C.Chr.; Acta Horti Gothob. 1: 77 (1924)

= Ctenitis sphaeropteroides (Baker) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 295 (1938)

= Dryopsis sphaeropteroides (Baker) Holttum & P.J.Edwards; Kew Bull. 41(1): 199-200 (1986)

= Phegopteris sphaeropteroides (Baker) Christ; Bull. Herb. Boissier, sér. 1, 7: 14 (1899)

= Polypodium sphaeropteroides Baker; Bull. Misc. Inform. Kew 1895: 55 (1895)

species :

Dryopteris splendens (Desv.) Kuntze; Revis. Gen. Pl. 2: 813 (1891)

Distribution: China (Yunnan); Tibet; Nepal; Bhutan; N-India (Arunachal Pradesh, Sikkim, West Bengal); peninsular Malaysia (Malacca)

36 CHI TIB 40 BHU IND NEP 42 MLY

Synonyms: hidden

= Aspidium splendens (Desv.) Christ; Farnkr. Erde: 259 (1897)

= Lastrea splendens (Desv.) Hook. ex Bedd.; Ferns Brit. India t. 42 (1865)

= Nephrodium splendens Hook.; Sp. Fil. [W. J. Hooker] 4: 126 (1862), nom. illeg.

species :

Dryopteris squamiseta (Hook.) Kuntze; Revis. Gen. Pl. 2: 813 (1891)

Distribution: Cameroon; Bioko Isl. [Fernando Poo]; Equatorial Guinea (Rio Muni); Rwanda; Sudan; Ethiopia; Somalia; Uganda; Kenya; Tanzania; Zambia; Malawi; E-Zimbabwe; South Africa (Limpopo, Mpulamanga, KwaZulu-Natal, E-Cape Prov.); Eswatini [Swaziland]; Madagascar; La Réunion; China (Yunnan); Tibet; Taiwan; Nepal; Bhutan; N-India (Arunachal Pradesh, Sikkim, West Bengal)

23 BIO CMN EQG RWA 24 ETH SOM SUD 25 KEN TAN UGA 26 MLW ZAM ZIM 27 SAF SWZ 29 MDG REU 36 CHI TIB 38 TAI 40 BHU IND NEP

Remarks: subgen. Nothoperanema

Conservation Status: IUCN R (Fernando Poo)

Synonyms: hidden

= Aspidium squamisetum (Hook.) Kuhn; Filic. Decken. 24 (1867)

= Ctenitis trichorhachis (Hayata) H.Itô; Nov. Fl. Jap. [Nakai & Honda] 4: 90 (1939)

= Dryopteris atrosetosa Rosenst.; Hedwigia 66: 342 (1915)

= Dryopteris buchananii (Baker) Kuntze; Revis. Gen. Pl. 2: 812 (1891)

= Dryopteris trichorhachis Hayata; Icon. Pl. Formosan. 4: 185, f. 123 (1914) (thrichorhachis)

= Lastrea buchananii (Baker) Bedd.; Ferns Brit. India 255 (1883)

= Nephrodium buchananii Baker; Syn. Fil. [W. J. Hooker & Baker], ed. 2: 498 (1874)

= Nephrodium squamisetum Hook.; Sp. Fil. [W. J. Hooker] 4: 140, t. 268 (1862)

= Nothoperanema squamisetum (Hook.) Ching; Acta Phytotax. Sin. 11: 27 (1966)

species :

Dryopteris stenolepis (Baker) C.Chr.; Index Filic. 294 (1905)

Distribution: China (Gansu, Guangxi, Sichuan, Yunnan); Tibet; Taiwan; Nepal; Bhutan; India (Arunachal Pradesh, Uttarakhand); Myanmar (Kachin, etc.); ?Laos; Vietnam

36 CHI TIB 38 TAI 40 BHU IND NEP 41 MYA VIE

Synonyms: hidden

= Aspidium stenolepis (Baker) Christ; Bull. Herb. Boissier, sér. 1, 7: 22 (1899)

= Dryopteris atrata var. stenolepis (Baker) Tagawa; Acta Phytotax. Geobot. 10(4): 281 (1941)

= Dryopteris hirtipes var. stenolepis (Baker) C.Chr.; Contr. U. S. Natl. Herb. 26(6): 279 (1931)

= Lastrea hirtipes (Hook.) Bedd.; Ferns Brit. India 232 (1883) (excl syn.)

= Nephrodium hirtipes Hook.; Sp. Fil. [W. J. Hooker] 4: 115, t. 2,49 (1862) (excl syn. Aspidium hirtipes Blume)

= Polypodium stenolepis Baker; Bull. Misc. Inform. Kew 1898: 231 (1898)

= Thelypteris stenolepis (Baker) C.F.Reed; Phytologia 17: 315 (1968)

species :

Dryopteris stewartii Fraser-Jenk.; Kalikasan 7(3): 272 (1978 publ. 1979)

Distribution: E-Afghanistan; Pakistan (Quetta, Kurram, Chitral, Swat, Pakistani Punjab, Hazara, Gilgit); W-Nepal; India (Jammu & Kashmir, Himachal Pradesh, Uttarakhand)

34 AFG 40 IND NEP PAK

Synonyms: hidden

= Dryopteris odontoloma f. brevifolia Mehra & Khullar; (1974 publ. 1980) (nom. nud., fide Fraser-Jenkins (1997))

species :

Dryopteris subarborea (Baker) C.Chr.; Index Filic. 295 (1905)

Distribution: peninsular Malaysia (Cameron Highlands, Pahang); Sumatra; Borneo (Mt. Kinabalu, Sabah); Sulawesi; Philippines; New Guinea (Irian Jaya, Papua New Guinea); Vanuatu; Fiji (Viti Levu)

42 BOR MLY NWG PHI SUL SUM 60 FIJ VAN

Synonyms: hidden

= Alsophila warburgii Christ; Ann. Jard. Bot. Buitenzorg 15: 80, t. 13, f. 3 (1897)

= Aspidium filix-mas var. megaphyllum (Baker) C.Chr.; Bull. Herb. Boissier, sér. 1, 6: 194 (1898) (non Mett. (1864))

= Dryopteris tabacicoma Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 16: 11 (1914)

= Lastrea megaphylla (Baker) Bedd.; Suppl. Ferns Brit. Ind. 63 (1892)

= Nephrodium megaphyllum Baker; J. Linn. Soc., Bot. 22: 227 (1886)

= Polypodium subarboreum Baker; J. Linn. Soc., Bot. 24: 259 (1887)

species :

Dryopteris subatrata Tagawa; Acta Phytotax. Geobot. 9: 207 (1940)

Distribution: Taiwan

38 TAI

species :

Dryopteris subbipinnata W.H.Wagner & Hobdy; Contr. Univ. Michigan Herb. 19: 69 (1993)

Distribution: Hawaii (Maui, Hawaii Isl.)

63 HAW

Synonyms: hidden

= Dryopteris palikuensis Herat ex Fraser-Jenk.; Thaiszia 4(1): 24 (1994)

species :

Dryopteris subcochleata Fraser-Jenk. & Katri Chhetri; Ferns and Fern-allies of Nepal 1: 27 (2015)

Distribution: India (Meghalaya, Manipur, Mizoram, Nagaland); Myanmar

40 IND 41 MYA

species :

Dryopteris subexaltata (Christ) C.Chr.; Index Filic. 295 (1905)

Distribution: South Korea (Cheju Isl.); Japan; Ryukyu Isl.; Taiwan

38 JAP KOR NNS TAI

Remarks: sect. Diclisodon, see Zuo et al. 2022

Synonyms: hidden

= Aspidium subexaltatum Christ; Bull. Herb. Boissier, sér. 2, 4: 616 (1904)

= Dryopteris hayatae Tagawa; Acta Phytotax. Geobot. 1: 156 (1932) (as hayatai)

species :

Dryopteris subhikonensis K.Hori & N.Murak.; Acta Phytotax. Geobot. 69(2): 99 (2018)

Distribution: Japan (Honshu)

38 JAP

species :

Dryopteris subimpressa Loyal; Nova Hedwigia 16: 467, t. 177, 178, f. 9-16 (1969)

Distribution: China (W-Yunnan); Nepal; Bhutan; India (Himachal Pradesh, Arunachal Pradesh, Sikkim, Uttarakhand); Myanmar

36 CHI 40 BHU IND NEP 41 MYA

Synonyms: hidden

= Dryopteris lancipinnula Ching; Sporae Pterid. Sin. 326 (1976), nom. nud.

= Dryopteris submarginata Loyal; Nova Hedwigia 16: 465, t. 175, 176, f. 1-8 (1969)

= Dryopteris subodontoloma Ching ex Loyal; (1961) (nom. nud., fide Fraser-Jenkins (1997))

species :

Dryopteris sublacera Christ; Notul. Syst. (Paris) 1: 43 (1909)

Distribution: China (Hubei, Shaanxi, Sichuan, Yunnan); Tibet; C-Taiwan; Nepal; Bhutan; India (Himachal Pradesh, Uttarakhand, ?Arunachal Pradesh)

36 CHI TIB 38 TAI 40 BHU IND NEP

Synonyms: hidden

= Dryopteris blepharolepis C.Chr.; H. Lév., Cat. Pl. Yun-Nan 103 (1916)

= Dryopteris juxtaposita f. multica H.Lév.; Cat. Pl. Yun-Nan 104 (1916)

= Dryopteris minjiangensis H.S.Kung; Acta Bot. Yunnan. 4(4): 341 (1982)

= Dryopteris schneideriana Hand.-Mazz.; Anz. Akad. Wiss. Wien, Math.-Naturwiss. Kl. no. 7.1 (1922)

= Nephrodium sublacerum (Christ) Hand.-Mazz.; Symb. Sin. 6: 21-25 (1929)

species :

Dryopteris submarginata Rosenst.; Repert. Spec. Nov. Regni Veg. 13: 132 (1914)

Distribution: China (Fujian, Guangxi, Guizhou, Hunan, Jiangxi, Sichuan, Zhejiang); Vietnam

36 CHI 41 VIE

Synonyms: hidden

= Dryopteris nudistipes Ching & P.S.Chiu; Bot. Res. Acad. Sin. 1987(2): 30 (1987)

= Dryopteris sessilipinna Ching & P.S.Chiu; Bot. Res. Acad. Sin. 1987(2): 29 (1987), nom. illeg., non Copel.

species :

Dryopteris subpycnopteroides Ching ex Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.), Bot. 14(3): 199 (1986)

Distribution: China (NW-Yunnan)

36 CHI

species :

Dryopteris subreflexipinna M.Ogata; J. Jap. Bot. 11: 30, f. 2 (1935)

Distribution: Taiwan; N-Thailand

38 TAI 41 THA

Remarks: status uncertain, possible syn. of A. diffracta

Conservation Status: IUCN R

Synonyms: hidden

= Acrorumohra subreflexipinna (M.Ogata) H.Itô; Nov. Fl. Jap. [Nakai & Honda] 4: 102 (1939)

= Arachniodes subreflexipinna (M.Ogata) Ching; Acta Bot. Sin. 10: 260 (1962)

= Rumohra subreflexipinna (M.Ogata) H.Itô; J. Jap. Bot. 11: 583 (1935)

species :

Dryopteris subtriangularis (C.Hope) C.Chr.; Index Filic. 296 (1905)

Distribution: China (Guangxi, Guizhou, Hainan, Sichuan, Yunnan); Tibet; Ryukyu Isl.; Taiwan; India (Assam, Arunachal Pradesh, Meghalaya); Myanmar; Thailand; Vietnam; Philippines

36 CHI TIB 38 NNS TAI 40 IND 41 MYA THA VIE 42 PHI

Remarks: often confused with D. tenuicula

Synonyms: hidden

= Dryopteris acutidens (C.Chr.) Ching; Acta Phytotax. Sin. 8: 168, t. 2 (24) (1959), not C. Chr. (1905)

= Dryopteris caudifrons Ching; Acta Phytotax. Sin. 8: 168, t. 22(25) (1959)

= Dryopteris subassamensis Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 451 (1938)

= Dryopteris uropinna M.G.Price; Gard. Bull. Singapore 30: 244, f. 1A-C (1977)

= Lastrea filix-mas var. subtriangularis (C.Hope) Bedd.; Ferns Brit. India Suppl. 56 (1892)

= Nephrodium subtriangulare C.Hope; J. Bot. 28: 327 (1890)

species :

Dryopteris subtsushimensis K.Hori & N.Murak.; Acta Phytotax. Geobot. 69(3): 154 (2018)

Distribution: China (Jiangsu); Japan (Kyushu)

36 CHI 38 JAP

species :

Dryopteris sukungiana Z.Y.Zuo; Phytotaxa 533(5): 260 (2022)

Distribution: China (Yunnan)

36 CHI

Remarks: sect. Diclisodon

species :

Dryopteris sweetiorum Lorence & W.L.Wagner; PhytoKeys 4: 19 (2011)

Distribution: Marquesas Isl. (Fatu Hiva)

61 MRQ

species :

Dryopteris tahmingensis Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 480 (1938)

Distribution: China (Fujian, Guangdong, Guangxi, Yunnan)

36 CHI

Synonyms: hidden

= Dryopteris hwangii Ching; Fl. Fujianica 1: 600 (1982)

species :

Dryopteris takeuchiana Koidz.; Acta Phytotax. Geobot. 1: 30 (1932)

Distribution: Japan

38 JAP

species :

Dryopteris tenuicula Matthew & Christ; Notul. Syst. (Paris) 1: 56 (1909)

Distribution: China (Guangdong, Guangxi, Guizhou, Hunan, Sichuan, Zhejiang); Japan; Ryukyu Isl. (Okinawa); Taiwan; Myanmar

36 CHI 38 JAP NNS TAI 41 MYA

Synonyms: hidden

= Dryopteris erythrosora var. purpurascens H.Itô

= Dryopteris indusiata var. purpurascens (H.Itô) Sa.Kurata; (1962)

= Dryopteris jiulungshanensis P.S.Chiu & G.H.Yao ex Ching; Bull. Bot. Res., Harbin 2(2): 62 (1982)

= Dryopteris labordei var. purpurascens (H.Itô) S.Sherizawa; J. Jap. Bot. 52: 17 (1977)

= Dryopteris neglecta Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(4): 4 (1984)

= Dryopteris purpurella Tagawa; Acta Phytotax. Geobot. 1: 307 (1932)

= Dryopteris rubristipes Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 3(4): 19 (1983) (fide Fraser-Jenkins 1997)

= Dryopteris subchampionii Ching; Bot. Res. Acad. Sin. 1987(2): 14 (1987)

= Dryopteris subtenuicula Ching & P.S.Chiu; Bot. Res. Acad. Sin. 1987(2): 29 (1987)

= Dryopteris yaoi Ching; Bull. Bot. Res., Harbin 2(2): 64 (1982)

= Dryopteris zhangii Ching; Bull. Bot. Res., Harbin 3(3): 26 (1983) (fide Fraser-Jenkins (1986)

= Nephrodium tenuiculum (Matthew & Christ) Tutcher; Bull. Misc. Inform. Kew, Addit. Ser. 10: 348 (1912)

species :

Dryopteris tenuipes (Rosenst.) Seriz.; J. Jap. Bot. 46: 19 (1971)

Distribution: Taiwan

38 TAI

Remarks: Subgen. Dryopsis

Synonyms: hidden

= Dryopteris erythrosora var. tenuipes Rosenst.; Hedwigia 56: 341 (1915)

species :

Dryopteris tetrapinnata W.H.Wagner & Hobdy; Contr. Univ. Michigan Herb. 22: 173 (1999)

Name: Wakamoi Woodfern

Distribution: Hawaii (Maui)

63 HAW

Synonyms: hidden

= Dryopteris unidentata var. ukelelensis Fraser-Jenk.; Thaiszia 4(1): 33 (1994)

species :

Dryopteris tingiensis Ching & S.K.Wu ex Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.), Bot. 14(3): 202 (1986)

Distribution: Tibet; Nepal

36 TIB 40 NEP

species :

Dryopteris tokyoensis (Matsum.) C.Chr.; Index Filic. 298 (1905)

Distribution: China (Fujian, Hubei, Hunan, Jiangxi, Zhejiang); South Korea; Japan

36 CHI 38 JAP KOR

Synonyms: hidden

= Aspidium transitorium Christ; Bull. Herb. Boissier, sér. 1, 7: 822 (1899)

= Dryopteris sanmingensis Ching; Fl. Fujianica 1: 600 (1982)

= Nephrodium tokyoense Matsum. ex Makino; Bot. Mag. (Tokyo) 12: 87 (1898)

species :

Dryopteris toyamae Tagawa; Acta Phytotax. Geobot. 8: 167 (1939), pro hybr.

Distribution: Japan; Taiwan

38 JAP TAI

Remarks: possibly a stabilized hybrid D. lacera x sieboldii

Synonyms: hidden

= Dryopteris enneaphylla var. toyamae (Tagawa) W.C.Shieh; J. Sci. Engin. (Natl. Chung-Hsing Univ.) 12: 305 (1975)

species :

Dryopteris transmorrisonensis (Hayata) Hayata; Icon. Pl. Formosan. 4: 187 (1914)

Distribution: S-China; Taiwan; Nepal; India (Sikkim)

36 CHI 38 TAI 40 IND NEP

Remarks: subgen. Dryopsis. Synonymized into D. nidus by Flora of China, but Fraser-Jenkins et al. (2018) and Knapp (Ferns of Taiwan) keep the species separate

Synonyms: hidden

= Ctenitis transmorrisonensis (Hayata) Tagawa; Acta Phytotax. Geobot. 8: 95 (1939)

= Dryopsis transmorrisonensis (Hayata) Holttum & P.J.Edwards; Kew Bull. 41(1): 191 (1986)

= Polystichum transmorrisonense Hayata; J. Coll. Sci. Imp. Univ. Tokyo 30: 427 (1911)

species :

Dryopteris tricellularis J.P.Roux; Kew Bull. 57(3): 735 (2002)

Distribution: Ethiopia; Uganda; Kenya; Tanzania

24 ETH 25 KEN TAN UGA

species :

Dryopteris tsoongii Ching; Bot. Res. Acad. Sin. 1987(2): 14 (1987)

Distribution: China (Anhui, Jiangxi, Zhejiang)

36 CHI

Synonyms: hidden

= Dryopteris wuyuanensis Ching; Bot. Res. Acad. Sin. 2: 15-16, pl. 6, f. 2 (1987)

species :

Dryopteris tsugiwoi Sa.Kurata; J. Geobot. 17: 34 (1969)

Distribution: Japan

38 JAP

Conservation Status: IUCN R. CR in Japan

species :

Dryopteris tsushimensis K.Hori & N.Murak.; Acta Phytotax. Geobot. 69(3): 149 (2018)

Distribution: Japan (Kyushu)

38 JAP

species :

Dryopteris tsutsuiana Sa.Kurata; J. Geobot. 17: 35 (1969)

Distribution: Japan (Kyushu)

38 JAP

Conservation Status: CR in Japan. IUCN V

species :

Dryopteris tyrrhena Fraser-Jenk. & Reichst.; Fern Gaz. 11: 180 (1975), nom. nov.

Distribution: Spain (S-Spain: Sa. Nevada); France; Corsica; Sardinia; Italy

12 COR FRA SAR SPA 13 ITA

Remarks: tyrrhena agg.

Synonyms: hidden

= Aspidium nevadense Boiss.; Elench. Pl. Nov. 93 (1838) (non Dryopteris nevadensis (Baker) Underw.)

= Dryopteris ×cebennae Fraser-Jenk.; Candollea 32(2): 318 (1977) [fide Jeßen (in litt.)]

= Nephrodium filix-mas proles rigidiformis Rouy; Fl. France [Rouy & Foucaud] 14: 408 (1913)

= Nephrodium rigidum subsp. nevadense (Boiss.) Rouy; Fl. France [Rouy & Foucaud] 14: 409 (1913)

species :

Dryopteris unidentata (Hook. & Arn.) C.Chr.; Index Filic. 299 (1905)

Distribution: Hawaii (Kauai, Oahu, Molokai, Lanai, Maui, Hawaii Isl.)

63 HAW

Synonyms: hidden

= Dryopteris acutidens C.Chr.; Index Filic. 250 (1905)

= Dryopteris unidentata var. acutidens (C.Chr.) Fraser-Jenk.; Thaiszia 4(1): 32 (1994)

= Nephrodium unidentatum (J.Sm.) Diels; Nat. Pflanzenfam. [Engler & Prantl] I, 4: 174 (1899)

= Phegopteris spinulosa Hillebr.; Fl. Hawaiian Isl. [Hillebrand] 566 (1888)

= Phegopteris unidentata J.Sm.; Ferns Brit. Foreign 170 (1866)

= Polypodium spinulosum (Hillebr.) Baker; Ann. Bot. (Oxford) 5: 459 (1891)

= Polypodium unidentatum (J.Sm.) Hook. & Arn.; Bot. Beechey Voy. 105 (1832)

variant :

var. paleacea (Hillebr.) Herat ex Fraser-Jenk.; Thaiszia 4(1): 31 (1994)

Distribution: Hawaii (Kauai, Oahu, Molokai, Maui, Hawaii Isl.)

63 HAW

Synonyms: hidden

= Phegopteris unidentata var. paleacea Hillebr.

species :

Dryopteris uniformis (Makino) Makino; Bot. Mag. (Tokyo) 23: 145 (1909)

Distribution: China (Anhui, Fujian, Jiangsu, Jiangxi, Zhejiang); South Korea (incl. Cheju Isl.); Japan

36 CHI 38 JAP KOR

Synonyms: hidden

= Dryopteris decurrentiloba Ching & C.F.Zhang; Bull. Bot. Res., Harbin 3(3): 31 (1983)

= Dryopteris dentipalea Nakai; Bot. Mag. (Tokyo) 34: 36 (1920)

= Dryopteris jiangshanensis Ching & P.S.Chiu; Bot. Res. Acad. Sin. 1987(2): 34 (1987)

= Dryopteris pseudouniformis Ching; Fl. Fujianica 1: 601 (1982)

= Dryopteris uniformis f. coreana (Nakai & H.Itô) H.Itô; Nov. Fl. Jap. [Nakai & Honda] 4: 14 (1939)

= Dryopteris uniformis var. coreana Nakai ex H.Itô; Nov. Fl. Jap. [Nakai & Honda] 4: 14 (1939)

= Dryopteris uniformis var. rufomarginata K.H.Shing; Jiangxi Sci. 8(3): 47 (1990)

= Nephrodium lacerum var. uniforme Makino; Bot. Mag. (Tokyo) 17: 79 (1903)

species :

Dryopteris varia (L.) Kuntze; Revis. Gen. Pl. 2: 814 (1891)

Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Henan, Hubei, Hunan, Jiangsu, Jiangxi, Sichuan, Yunnan, Zhejiang); South Korea (Cheju Isl.); Japan; Ryukyu Isl.; Bonin Isl. (Chichijima); Volcano Isl. (Kita-Iwojima, Minami-Iwojima); Taiwan; India; NE-India (Assam, Meghalaya); Myanmar; Vietnam; Sulawesi; Philippines

36 CHI 38 JAP KOR KZN NNS OGA TAI 40 IND 41 MYA VIE 42 PHI SUL

Conservation Status: IUCN CR for India (Fraser-Jenkins 2012)

Synonyms: hidden

= Aspidium lacerum var. subtripinnatum Franch. & Sav.; Enum. Pl. Jap. 2: 238 (1877)

= Aspidium opacum (Hook.) Benth.; Fl. Hongkong [Bentham] 456 (1861)

= Aspidium varium (L.) Sw.; J. Bot. (Schrader) 1800(2): 35 (1801)

= Dryopteris caudifolia Ching & P.S.Chiu; Bot. Res. Acad. Sin. 1987(2): 21 (1987)

= Dryopteris glabrescens Ching & P.S.Chiu ex K.H.Shing & J.F.Cheng; Jiangxi Sci. 8(3): 48 (1990)

= Dryopteris hololepis (Hayata) Tagawa; Acta Phytotax. Geobot. 4: 133 (1935)

= Dryopteris lingii Ching; Bot. Res. Acad. Sin. 1987(2): 23 (1987)

= Dryopteris matsuzoana Koidz.; Bot. Mag. (Tokyo) 39: 15 (1925)

= Dryopteris nanchuanensis Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 3(4): 20 (1983)

= Dryopteris ogawae H.Itô; Iconogr. Pl. Asiae Orient. 1: 18 (1935)

= Dryopteris pseudobissetiana Ching ex K.H.Shing & J.F.Cheng; Jiangxi Sci. 8(3): 49 (1990)

= Dryopteris sinovaria Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 6(1): 179, nom. nov.,

= Dryopteris yabei f. ogawae (H.Itô) H.Itô; Nov. Fl. Jap. [Nakai & Honda] 4: 59 (1938 publ. 1939)

= Dryopteris yabei Hayata; J. Coll. Sci. Imp. Univ. Tokyo 30: 424 (1911)

= Dryopteris yabei var. hololepis (Hayata) H.Itô; Bot. Mag. (Tokyo) 50(591): 128 (1936)

= Dryopteris yabei var. matsuzoana (Koidz.) H.Itô; Bot. Mag. (Tokyo) 50(591): 128 (1936)

= Dryopteris yabei var. ogawae (H.Itô) H.Itô; Bot. Mag. (Tokyo) 50: 128 (1936)

= Lastrea opaca Hook.; J. Bot. 9: 339 (1857)

= Lastrea varia (L.) T.Moore; Index Fil. [T. Moore] 107 (1858)

= Nephrodium coriaceum C.Hope; J. Bot. 28: 328 (1890)

= Nephrodium filix-mas f. subtripinnatum (Franch. & Sav.) Matsum.; Index Pl. Jap. 1: 319 (1904)

= Nephrodium opacum (Hook.) J.H.Veitch; Hort. Veitch.: 325 (1906)

= Nephrodium varium (L.) C.Presl; Reliq. Haenk. 1: 36 (1826)

= Nephrodium varium (L.) Desv.; Mem. Soc. Linn. Paris 6(2): 269 (1827), isonym

= Nephrodium villosum var. opacum (Hook.) Hieron.; Bot. Jahrb. Syst. 34: 446 (1904)

= Polypodium varium L.; Sp. Pl. 2: 1090 (1753)

= Polystichum hololepis Hayata; Icon. Pl. Formosan. 5: 332 (1915)

= Polystichum varium (L.) C.Presl; Abh. Königl. Böhm. Ges. Wiss., ser. 5, 6 [Epim. Bot.]: 57 (1851)

= Polystichum varium var. eurylepidota Rosenst.; Hedwigia 56(5): 340 (1915)

= Polystichum varium var. subtripinnatum (Franch. & Sav.) Nakai; Bot. Mag. (Tokyo) 39(461): 119 (1925)

species :

Dryopteris vescoi (Drake) C.Chr.; Index Filic. 300 (1905)

Distribution: Society Isl. (Tahiti)

61 SCI

Synonyms: hidden

= Nephrodium vescoi Drake; Fl. Polynésie Franç. [Drake] 301 (1893)

species :

Dryopteris vidyae Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.), Bot. 18(5): 466 (1989), pro hybr.

Distribution: N-India (West Bengal, Sikkim, Arunachal Pradesh); Myanmar (Kachin, etc.)

40 IND 41 MYA

Conservation Status: IUCN EN for India (Fraser-Jenkins 2012)

species :

Dryopteris villarii (Bellardi) Woyn. ex Schinz & Thell.; Vierteljahrsschr. Naturf. Ges. Zürich 60: 339 (1915)

Name: Starrer Wurmfarn (DE)

Distribution: Germany (Bayern); Switzerland; Liechtenstein; Austria; Baleares; France; Corsica; Sardinia; Italy; Sicily; Slovenia; Croatia; Bosnia & Hercegovina; Montenegro; Serbia; Kosovo; North Macedonia; Albania; Bulgaria; Greece (mountains in mainland); Northern Caucasus; Georgia [Caucasus]

11 AUT GER LIE SWI 12 BAL COR FRA SAR 13 ALB BIH BUL CRO GRC ITA KOS MNT NMA SIC SLV SRB 33 GEO NCS

Synonyms: hidden

= Aspidium fragrans (Vill.) Gray; Nat. Arr. Brit. Pl. 2: 9 (1821)

= Aspidium rigidum (Hoffm.) Sw.; J. Bot. (Schrader) 1800(2): 37 (1801)

= Aspidium rigidum f. fallax Milde

= Aspidium rigidum var. germanicum Christ; Burnat, Foug. Alpes Maritimes: 24 (1900)

= Dryopteris rigida (Sw.) A.Gray; Manual (Gray): 631 (1848)

= Dryopteris rigida (Sw.) Underw.; Native Ferns, ed. 4: 116 (1893), isonym

= Dryopteris villarii f. fallax (Milde) Niketic; Fl. Serbia 1: 155 (1992)

= Dryopteris villarii var. australis (Ten.) Diklic & V.Nikolic; Fl. SR Srbije 10: 10 (1986)

= Filix rigida (Sw.) Farw.; Rep. (Annual) Michigan Acad. Sci. 18: 81-83 (1916)

= Filix-mas rigida (Sw.) Farw.; Amer. Midl. Naturalist 12: 234-251 (1931)

= Hypodematium nivale Fée; Mém. Foug. 5 (Gen. Filic.): 298 (1852)

= Lastrea rigida (Sw.) C.Presl; Tent. Pterid. 77 (1836)

= Lophodium fragrans Newman; xxi (1871)

= Lophodium rigidum (Sw.) Newman; Phytologist 4: app. 21 (1851)

= Nephrodium meridionale (Lojac.) Rouy; Fl. France [Rouy & Foucaud] 14: 409 (1913)

= Nephrodium rigidum (Hoffm.) Desv.; Mem. Soc. Linn. Paris 6(2): 261 (1827)

= Nephrodium rigidum var. germanicum (Christ) Rouy; Fl. France [Rouy & Foucaud] 14: 409 (1913)

= Nephrodium villarii (Bellardi) Beck; Oesterr. Bot. Z. 67: 118 (1918)

= Nephrodium villarii subsp. rigidum (Hoffm.) Hayek; Repert. Spec. Nov. Regni Veg. Beih. 30(1): 26 (1924)

= Polypodium asplenioides Scop.; Fl. Carniol., ed. 2, 2: 298, t. 62 (1772)

= Polypodium fragrans Vill.; Hist. Pl. Dauphiné [Villars] 3(2): 843 (1789), nom. illeg.

= Polypodium odoratum Poir.; Encycl. [J. Lamarck et al.] 5: 541 (1804)

= Polypodium rigidum (Sw.) Hoffm.; Deutschl. Fl., Theil 2 (Hoffm.) 2: 6 (1795), nom. illeg.

= Polypodium villarii Bellardi; App. Fl. Pedem. 49 (1792)

= Polystichum meridionale Lojac.; Fl. Sicul. (Lojacono) 3: 405 (1909)

= Polystichum rigidum (Sw.) DC.; Fl. Franç. [Lam. & A. DC.], éd. 3,560 (1805)

= Polystichum strigosum Roth; Tent. Fl. Germ. 3(1.1): 86 (1799), nom. illeg.

species :

Dryopteris wallichiana (Spreng.) Hyl.; Bot. Not. 1963: 352 (1953)

subspecies :

subsp. wallichiana

Distribution: Turkey (Anatolia); China (Yunnan, Guizhou, Sichuan, Fujian, Jiangxi, Shanxi); Tibet; Japan; Taiwan; Nepal; Bhutan; India (Arunachal Pradesh, Himachal Pradesh, Manipur, Meghalaya, Nagaland, Sikkim, ?Tamil Nadu, Uttarakhand, West Bengal); Sri Lanka; Myanmar (Kachin, Chin, etc.); Vietnam; Borneo (Mt. Kinabalu, Sabah); Sulawesi; Philippines; Java; New Guinea (Irian Jaya, Papua New Guinea); Hawaii (Kauai, Oahu, Molokai, Lanai, Maui, Hawaii Isl.); Mexico (Chiapas, Chihuahua, Colima, Ciudad de Mexico, Guanajuato, Guerrero, Hidalgo, Jalisco, Mexico State, Michoacan, Morelos, Oaxaca, Puebla, Queretaro, San Luis Potosi, Tamaulipas, Tlaxcala, Veracruz); Guatemala; El Salvador; Honduras; Costa Rica; Panama; Jamaica; Hispaniola (Haiti, Dominican Republic); Colombia (Antioquia, Boyacá, Caldas, Cauca, Cesar, Cundinamarca, Magdalena, Meta, Nariño, Norte de Santander, Quindío, Risaralda, Santander, Tolima); Venezuela (Lara, Merida, Tachira, Trujillo); Ecuador; Galapagos; Peru; Bolivia (Chuquisaca, Cochabamba, La Paz, Santa Cruz, Tarija); Brazil (Espírito Santo, Minas Gerais, Paraná, Rio de Janeiro, Rio Grande do Sul, Santa Catarina, São Paulo); NE-Argentina (Catamarca, Jujuy, Salta, Tucuman); Gough Isl.

34 TUR 36 CHI TIB 38 JAP TAI 40 BHU IND NEP SRL 41 MYA VIE 42 BOR JAW NWG PHI SUL 63 HAW 79 MEX 80 COS ELS GUA HON PAN 81 HIS JAM 82 CLM VEN 83 BOL ECU GAL PER 84 BRA 85 ARG 91 TDC

Synonyms: hidden

= Aspidium crinitum M.Martens & Galeotti; Nouv. Mém. Acad. Roy. Sci. Bruxelles 16: 66, t. 17, f. 2 (1842)

= Aspidium donianum Spreng.; Syst. Veg., ed. 16 [Sprengel] 4(1): Suppl. 320 (1827), nom. superfl. nov. for Aspidium paleaceum D. Don

= Aspidium filix-mas f. paleaceum (Lag. ex Sw.) Asch.; Syn. Mitteleur. Fl. [Ascherson & Graebner] 1: 28 (1896)

= Aspidium filix-mas var. paleaceum (T.Moore) Mett.; Abh. Senckenberg. Naturf. Ges. 2(2) [Pheg. u. Asp.]: 339 (1858)

= Aspidium filix-mas var. parallelogrammum (Kunze) Hillebr.; Fl. Hawaiian Isl. [Hillebrand] 574 (1888)

= Aspidium paleaceum (T.Moore) D.Don; Prodr. Fl. Nepal. 4-5 (1825), non Sw. (1806), nom. illeg., non Aspidium paleaceum Lag. ex Sw. 1806

= Aspidium paleaceum (T.Moore) Dalla Torre & Sarnth.; Fl. Tirol 6(1): 46 (1906), non Sw. 1806

= Aspidium paleaceum D.Don; Prodr. Fl. Nepal. 4 (1824) non Sw. 1806

= Aspidium paleaceum Lag. ex Sw.; Syn. Fil. [Swart] 52 (1806)

= Aspidium parallelogrammum Kunze; Linnaea 13: 146 (1839)

= Aspidium patentissimum Wall.; Numer. List. [Wallich] no. 340 (1829), nomen

= Aspidium patentissimum Wall. ex Kunze; Linnaea 13: 146 (1839), nom. superfl.

= Aspidium wallichianum Spreng.; Syst. Veg., ed. 16 [Sprengel] 4(1): 104 (1827), non C. Presl ex Kunze (1851)

= Dichasium parallelogrammum (Kunze) Fée; Mém. Foug. 5 (Gen. Filic.): 302, t. 23 B, f. 1 (1852)

= Dichasium patentissimum (Wall. ex Kunze) Fée; Mém. Foug. 5 (Gen. Filic.): 302, t. 23 B, f. 2 (1852)

= Dryopteris cyrtolepis Hayata; Icon. Pl. Formosan. 4: 149 t 89 (1914)

= Dryopteris cyrtolepis var. doiana (Tagawa) H.Itô; Nov. Fl. Jap. [Nakai & Honda] 4: (1938 publ. 1939)

= Dryopteris doiana Tagawa; Acta Phytotax. Geobot. 5: 253 (1936)

= Dryopteris doniana (Spreng.) Ching; Sunyatsenia 6: 3 (1941)

= Dryopteris filix-mas subsp. paleacea (Lag. ex Sw.) W.Koch ex Braun-Blanq.; Veröff. Geobot. Inst. Rübel Zürich 7: 34 (1932), isonym

= Dryopteris filix-mas subsp. paleacea (Sw.) C.Chr.; Rosenvinge, Biol. Arbejd. Til. Eug. Warming 76 (1911)

= Dryopteris filix-mas subsp. parallelogramma (Kunze) Christ; Annuaire Conserv. Jard. Bot. Genève 15-16: 187 (1912)

= Dryopteris filix-mas subsp. patentissima (Wall. ex Kunze) C.Chr.; Index Filic. 5: 265 (1905) (p. p., excl. Nephrodium filix-mas var. fibrillosa C. B. Clarke (1880)) (fide Fraser-Jenkins 2008)

= Dryopteris filix-mas var. crinita (Wall.) Rosenst.; (1910)

= Dryopteris filix-mas var. paleacea (T.Moore) Druce; List Brit. Pl. 87 (1908), nom. illeg.

= Dryopteris filix-mas var. parallelogramma (Kunze) Christ; Philipp. J. Sci. 2: 212 (1907)

= Dryopteris filix-mas var. parallelogramma (Kunze) Conz.; Fl. Taxon Mex. (Conzatti) 1(2): 19 (1939), isonym

= Dryopteris himalaica (Ching & S.K.Wu) S.G.Lu; Acta Bot. Yunnan. 13(1): 40 (1991)

= Dryopteris kweichowicola Ching ex P.S.Wang; J. Arnold Arbor. 71(1): 119 (1990)

= Dryopteris mediterranea Fomin; Komarov, Fl. URSS 1: 35 (1934), nom. nov. for Lastrea filix-mas var. paleacea T. Moore

= Dryopteris paleacea (Lag. ex Sw.) C.Chr.; Amer. Fern J. 1: 94 (1911), non Hand.-Mazz. (1908)

= Dryopteris paleacea (T.Moore) Hand.-Mazz.; Verh. K. K. Zool.-Bot. Ges. Wien 58 (100) (1908), nom. illeg., non Aspidium paleaceum Lag. ex Sw. 1806

= Dryopteris parallelogramma (Kunze) Alston; Amer. Fern J. 47: 92 (1957)

= Dryopteris patentissima (Wall. ex Kunze) Fée; Mém. Foug. 5 (Gen. Filic.): 303, pl. 23B, f. 2 (1852)

= Dryopteris patentissima (Wall. ex Kunze) N.C.Nair; Indian Forester 94: 169 (1968), isonym

= Dryopteris patentissima (Wall. ex Kunze) Panigrahi & Basu; Indian J. Forest. 3(3): 266, f. 1, t. 2, 3 & 4 (1980), isonym

= Dryopteris quatanensis Ching; Wuyi Sci. J. 1: 6 (1981)

= Dryopteris reichsteinii Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.), Bot. 14: 191 (1986) (nom. nov., syn. fide Fraser-Jenkins (1997))

= Dryopteris ursipes Hayata; Icon. Pl. Formosan. 5: 291, f. 116 (1915)

= Dryopteris wallichiana (Spreng.) Alston & Bonner; Candollea 15(7): 216 (1956), nom. superfl.

= Dryopteris wallichiana subsp. reichsteinii (Fraser-Jenk.) Fraser-Jenk. ex Khullar; Fern Fl. W. Himalaya 5: 245 (2024)

= Dryopteris wallichiana var. himalaica Ching & S.K.Wu; Fl. Xizangica 1: 257 (1983)

= Dryopteris wallichiana var. kweichowicola (Ching ex P.S.Wang) S.K.Wu; Fl. Reipubl. Popularis Sin. [Fl. China Ed. Committee] 5(1): 149 (2000)

= Filix filix-mas var. paleacea (Sw.) Farw.; Amer. Midl. Naturalist 12: 254 (1931)

= Lastrea filix-mas var. paleacea T.Moore; Nat. Print. Brit. Ferns, 8vo. ed. 1: 178, 193, t. 33(A) (1859), nom. illeg., non Aspidium paleaceum Lag. ex Sw. 1806

= Lastrea filix-mas var. parallelogrammum (Kunze) Bedd.; Ferns Brit. India 249 (1883) (p. p. quoad. var. clarkei Bedd.) (fide Fraser-Jenkins 2008)

= Lastrea filix-mas var. patentissima (Wall. ex Kunze) Bedd.; Suppl. Ferns S. India 29 (1876)

= Lastrea paleacea (Sw.) T.Moore; Index Fil. [T. Moore] 99 (1858)

= Lastrea parallelogramma (Kunze) Liebm.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 5, 1: 271 (seors. 119) (1849)

= Lastrea patentissima (Wall. ex Kunze) J.Sm.; J. Bot. (Hooker) 4: 193 (1842)

= Nephrodium filix-mas f. paleaceum (T.Moore) Fiori; Fl. Anal. Ital. 1: 8 (1896)

= Nephrodium filix-mas subsp. paleaceum (T.Moore) Brause & Andres; Magyar Fl. Veg. 1: 545 (1964)

= Nephrodium filix-mas subsp. patentissimum (Wall. ex Kunze) C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 520 (1880)

= Nephrodium filix-mas var. paleaceum (T.Moore) Hook.; Fil. Exot.: t. 98 (1859), nom. illeg., non Aspidium paleaceum Lag. ex Sw. 1806

= Nephrodium filix-mas var. parallelogrammum (Kunze) Hook.; Sp. Fil. [W. J. Hooker] 4: 116 (1862)

= Nephrodium filix-mas var. patentissimum (Wall. ex Kunze) C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 520 (1880 (pro parte quoad sp.))

= Nephrodium parallelogrammum (Kunze) C.Hope; J. Bombay Nat. Hist. Soc. 14: 728 (1903)

= Nephrodium parallelogrammum f. patentissimum (Wall. ex Kunze) C.Hope; J. Bombay Nat. Hist. Soc. 14: 728 (1903)

= Nephrodium patentissimum (Wall. ex Kunze) C.B.Clarke; J. Linn. Soc., Bot. 15: 156 (1876) (incl. descr. in part, excl. type)

= Polystichum filix-mas var. paleaceum (T.Moore) Fiori; Nuov. Fl. Anal. Ital. 1: 23 (1923)

= Polystichum filix-mas var. paleaceum (T.Moore) P.E.Fourn.; Quatre Fl. France: 16 (1934), isonym

= Polystichum ursipes Hayata; Icon. Pl. Formosan. 5: 291 (1915), pro syn.

subspecies :

subsp. convexa Fraser-Jenk.; Pterid. Bhutan 317 (2023)

Distribution: India (West Bengal)

40 IND

subspecies :

subsp. himalaica Fraser-Jenk.; New Species Syndr. Indian Pteridol. 134 (1997)

Distribution: Nepal; Bhutan; India (Arunachal Pradesh, Nagaland, Sikkim, Uttarakhand, West Bengal)

40 BHU IND NEP

subspecies :

subsp. madrasensis (Fraser-Jenk.) Fraser-Jenk.; Thaiszia 4(1): 22 (1994)

Distribution: S-India (Tamil Nadu); Sri Lanka

40 IND SRL

Synonyms: hidden

= Dryopteris madrasensis Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.), Bot. 18(5): 359 (1989)

subspecies :

subsp. nepalensis Fraser-Jenk.; New Species Syndr. Indian Pteridol. 134 (1997)

Distribution: China (Yunnan, Sichuan); Tibet; C-Nepal; Bhutan; India (Uttarakhand, Sikkim, Arunachal Pradesh, Mizoram, Nagaland); Myanmar

36 CHI TIB 40 BHU IND NEP 41 MYA

subspecies :

subsp. madagascariensis (C.Chr.) J.P.Roux; Strelitzia 23: 124 (2009)

Distribution: Madagascar

29 MDG

Synonyms: hidden

= Dryopteris paleacea var. madagascariensis C.Chr.; Dansk Bot. Ark. 7: 53, t. 11, f. 5-9 (1932)

subspecies :

subsp. pachyphylla Fraser-Jenk. & Ralf Knapp; Annot. Checkl. Ind. Pterid. 2: 276 (2018), nom. nov.

Distribution: China; Taiwan; Bhutan; India (Arunachal Pradesh, Meghalaya, Nagaland); Myanmar

36 CHI 38 TAI 40 BHU IND 41 MYA

Synonyms: hidden

= Dryopteris neorosthornii Ching; Bull. Fan Mem. Inst. Biol. Bot. 11: 62 (1941)

= Dryopteris pachyphylla Hayata; Icon. Pl. Formosan. 4: 168, f. 108 (1914), non (Kunze) Kuntze (1891)

subspecies :

subsp. parrisiae (Fraser-Jenk.) Fraser-Jenk.; Indian Fern J. 39(1): 119 (2022)

Distribution: Moluccas (Seram); New Guinea (Papua New Guinea)

42 MOL NWG

Synonyms: hidden

= Dryopteris parrisiae Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.), Bot. 14(3): 199 (1986)

subspecies :

subsp. reichsteinii Fraser-Jenk.; Ann. Bot. Fenn. 33: 71 (1996), non D. reichsteinii Fraser-Jenk. (1986)

Distribution: Zimbabwe

26 ZIM

species :

Dryopteris wardii (Baker) Kuntze; Revis. Gen. Pl. 2: 814 (1891)

Distribution: Seychelles

29 SEY

Synonyms: hidden

= Araiostegia wardii (Baker) Tardieu; Adansonia, sér. 2, 5: 493 (1965)

= Aspidium wardii (Baker) Kuhn; Reisen Ost-Afr. [Decken] 3(3): Bot. 69 (1879)

= Nephrodium wardii Baker; Syn. Fil. [W. J. Hooker & Baker], ed. 2: 500 (1874)

= Polystichopsis wardii (Baker) Tardieu; Notul. Syst. (Paris) 15: 176 (1956)

species :

Dryopteris wattsii McKeown, Sundue & Barrington; PhytoKeys 14: 43-56 (2012), nom. nov.

Distribution: Australia (NE-Queensland)

50 AUS

Conservation Status: very rare, known only from 2 localities

Synonyms: hidden

= Polystichum fragile Watts; Trans. Linn. Soc. New South Wales 39: 775, t. 88, f. 9 (1915)

= Revwattsia fragilis (Watts) D.L.Jones; Fl. Australia 48: 711 (1998)

species :

Dryopteris whangshanensis Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 421 (1938)

Distribution: China (Anhui, Fujian, Hubei, Jiangxi, Zhejiang)

36 CHI

Synonyms: hidden

= Dryopteris crassirhizoma subsp. whangshangensis (Ching) Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.), Bot. 14(3): 191 (1986) (whangshangensis)

= Dryopteris huangangshanensis Ching; Fl. Fujianica 1: 602 (1982)

= Dryopteris submarginalis Ching, Boufford & K.H.Shing; J. Arnold Arbor. 64: 30. (1983), not (Langsdorf & Fisch.) C. Chr. (1905)

species :

Dryopteris wideniana Fraser-Jenk.; Ann. Bot. Fenn. 52(1-2): 63 (2015)

Distribution: Zimbabwe

26 ZIM

species :

Dryopteris woodsiisora Hayata; Icon. Pl. Formosan. 6: 158 (1916)

Distribution: China (Guangdong, Guizhou, Jiangxi, Liaoning, Shandong, Sichuan, Yunnan); Tibet; North Korea; Taiwan; Nepal; Bhutan; India (Himachal Pradesh, Arunachal Pradesh, ?Manipur, Meghalaya, Nagaland, Sikkim, Uttarakhand, West Bengal); Myanmar (Kachin, etc.); N-Thailand; Vietnam

36 CHI TIB 38 NKO TAI 40 BHU IND NEP 41 MYA THA VIE

Synonyms: hidden

= Dryopteris neochrysocoma Ching; Lingnan Sci. J. 21: 31 (1945)

= Dryopteris taishanensis F.Z.Li; Acta Phytotax. Sin. 20(3): 344 (1982)

= Dryopteris tenuissima f. serrata (Tagawa) H.Itô; Nov. Fl. Jap. [Nakai & Honda] 4: 71 (1938 publ. 1939)

= Dryopteris tenuissima Tagawa; Acta Phytotax. Geobot. 1: 308 (1932)

= Dryopteris tenuissima var. serrata Tagawa; J. Jap. Bot. 13(3): 185 (1937)

= Dryopteris woodsiisora var. tenuissima (Tagawa) Li & J.Z.Wang; Bull. Bot. Res., Harbin 9(4): 80 (1989)

= Dryopteris zinongii Z.R.Wang & Fraser-Jenk.; Acta Phytotax. Sin. 23(5). 345 (1985)

species :

Dryopteris wulingshanensis J.P.Shu, Y.H.Yan & R.J.Wang; PhytoKeys 185: 21, figs. 3-4 (2021)

Distribution: China (Hunan)

36 CHI

species :

Dryopteris wuyishanica Ching & P.S.Chiu; Bot. Res. Acad. Sin. 1987(2): 34 (1987)

Distribution: China (Fujian)

36 CHI

Synonyms: hidden

= Dryopteris wuyishanicola Ching & P.C.Chiu; Bot. Res. Acad. Sin. 1987(2): 31 (1987)

species :

Dryopteris wuzhaohongii Li Bing Zhang; Taxon 61(6): 1208 (2012), nom. nov.

Distribution: China (Yunnan); Tibet; Taiwan

36 CHI TIB 38 TAI

Remarks: subgen. Nothoperanema. Probably to be included in D. paleolatus fide Fraser-Jenkins et al. (2018)

Synonyms: hidden

= Acrophorus macrocarpus Ching & S.H.Wu; S. H. Wu, Acta Phytotax. Sin. 21(4): 381 (1983)

species :

Dryopteris xanthomelas (Christ) C.Chr.; Index Filic., Suppl. 2 (1906-1912): 41 (1913)

Distribution: China (Anhui, Gansu, Hebei, Henan, Shanxi, Sichuan); Tibet; Taiwan; Pakistan (Swat, Baltistan, Gilgit); Nepal; Bhutan; India (Jammu & Kashmir, Himachal Pradesh, Uttarakhand, Sikkim, Darjeeling, Arunachal Pradesh, West Bengal)

36 CHI TIB 38 TAI 40 BHU IND NEP PAK

Remarks: D. sinofibrillosa is kept separate by Flora of China but sunk into D. xanthomelas by Fraser-Jenkins (2008) and Knapp (2012)

Synonyms: hidden

= Aspidium xanthomelas Christ; Bull. Acad. Int. Géogr., sér. 3, 16: 117 (1906)

= Dryopteris canaliculata Ching; Fl. Xizangica 1: 251 (1983)

= Dryopteris chingii N.C.Nair; Indian Forester 94: 169 (1968) for D. fibrillosa (C. B. Clarke) Hand.-Mazz., nom. superfl. (fide Fraser-Jenkins 1997)

= Dryopteris fibrillosa (C.B.Clarke) Hand.-Mazz.; Anz. Akad. Wiss. Wien, Math.-Naturwiss. Kl. no. 7: 2 (1922), non Christ (1905)

= Dryopteris fibrillosissima Ching; Fl. Xizangica 1: 254 (1983) (fide Fraser-Jenkins et al. 2018)

= Dryopteris filix-mas subsp. fibrillosa (C.B.Clarke) C.Chr.; Acta Horti Gothob. 1: 57 (1924)

= Dryopteris filix-mas var. fibrillosa (C.B.Clarke) C.Chr.; Index Filic. 5: 265 (1905) (fide Fraser-Jenkins 1997)

= Dryopteris pulcherrima Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 422 (1938) (fide Fraser-Jenkins et al. 2018)

= Dryopteris sinofibrillosa Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 180 (1940) for D. fibrillosa (C. B. Clarke) Hand.-Mazz.

= Lastrea filix-mas subvar. fibrillosa (C.B.Clarke) Bedd.; Ferns Brit. India 250 (1883)

= Nephrodium clarkei var. fibrillosum (C.B.Clarke) Hand.-Mazz.; Symb. Sin. 6: 24 (1929)

= Nephrodium filix-mas var. fibrillosum C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 520, t. 70 (1880)

= Nephrodium parallelogrammum f. fibrillosum (C.B.Clarke) C.Hope; J. Bombay Nat. Hist. Soc. 14: 731 (1903)

species :

Dryopteris xunwuensis Ching & K.H.Shing; Jiangxi Sci. 8(3): 48 (1990)

Distribution: China (Guangxi, Jiangxi)

36 CHI

species :

Dryopteris yigongensis Ching; Fl. Xizangica 1: 253 (1983)

Distribution: China (Yunnan); Tibet; Bhutan

36 CHI TIB 40 BHU

Remarks: accepted as species separate from D. lepidopoda by Fraser-Jenkins (2023)

species :

Dryopteris yongdeensis W.M.Chu ex S.G.Lu; Acta Phytotax. Sin. 28(6): 473 (1990)

Distribution: China (Yunnan)

36 CHI

species :

Dryopteris yoroii Seriz.; J. Jap. Bot. 46: 20 (1971)

Distribution: China (Sichuan, Yunnan); SE-Tibet; Taiwan; India (Sikkim)

36 CHI TIB 38 TAI 40 IND

Remarks: sect. Diclisodon, see Zuo et al. 2022. Some material from India, Bhutan & Myanmar might refer to this species

Synonyms: hidden

= Acrorumohra yoroii (Seriz.) W.C.Shieh; J. Sci. Engin. (Natl. Chung-Hsing Univ.) 12: 301 (1975)

species :

Dryopteris zayuensis Ching & S.K.Wu; Fl. Xizangica 1: 255 (1983)

Distribution: China (Qinghai, Sichuan, Yunnan); Tibet; Taiwan; Pakistan (Azad Kashmir); Nepal; Bhutan; India (Jammu & Kashmir, Uttarakhand, Arunachal Pradesh, Himachal Pradesh, Sikkim, West Bengal)

36 CHI TIB 38 TAI 40 BHU IND NEP PAK

Remarks: D. zayuensis is the correct name fide Fraser-Jenkins et al. (2015)

Synonyms: hidden

= Dryopteris discreta Ching & S.K.Wu; Fl. Xizangica 1: 251 (1983)

= Dryopteris incisolobata Ching; Fl. Xizangica 1: 249 (1983) (fide Fraser-Jenkins 1997)

= Dryopteris nyalamensis Ching & S.K.Wu; Fl. Xizangica 1: 256 (1983)

= Dryopteris nyalamensis var. angustipinna Ching & S.K.Wu; Fl. Xizangica 1: 256 (1983)

= Dryopteris qamdoensis Ching; Fl. Xizangica 1: 252 (1983)

= Dryopteris squamifera Ching & S.K.Wu; Fl. Xizangica 1: 250 (1983)

= Dryopteris tsangpoensis Ching; Fl. Xizangica 1: 250 (1983)

species :

Dryopteris zhenfengensis P.S.Wang & X.Y.Wang; Pterid. Fl. Guizhou: 337 (2001)

Distribution: China (Guizhou)

36 CHI

Remarks: provisionally accepted name, status unconfirmed

species :

Dryopteris ×alejandrei Pérez Carro & Fern.Areces; Flora Montiber. 37: 33 (2007)

Distribution: Spain

12 SPA

Remarks: Dryopteris mindshelkensis x oreades

species :

Dryopteris ×algonquinensis D.M.Britton; Canad. Field-Naturalist 89(2): 165 (1975)

Name: Algonquin woodfern

Distribution: Canada (Ontario)

72 CDN

Remarks: Dryopteris fragrans x marginalis

species :

Dryopteris ×alpirsbachensis Freigang, Zenner, Bujnoch, S.Jess. & Magauer; Kochia 10: 27 (2017)

Distribution: Germany (Baden-Württemberg, Bayern); Austria; SW-France (Haute-Garonne); Italy

11 AUT GER 12 FRA 13 ITA

Remarks: Dryopteris carthusiana x remota

species :

Dryopteris ×ambroseae Fraser-Jenk. & Jermy; Fern Gaz. 11(5): 338 (1977)

Distribution: Europe; England; Germany (Baden-Württemberg, Bayern, Hessen, Nordrhein-Westfalen, Sachsen); Czech Republic; European Russia

10 GRB 11 CZE GER 14 RUS

Remarks: Dryopteris dilatata x expansa

species :

Dryopteris ×apuana Gibby, S.Jess. & Marchetti; Mem. Accad. Lunig. Sci. 66 [Atti Convegn. Internaz. 1994 - Pteridofite Lunigiana storica]: 92 (1996)

Distribution: Italy

13 ITA

Remarks: Dryopteris mindshelkensis x villarii

species :

Dryopteris ×arecesiae Pérez Carro & T.E.Díaz; Anales Jard. Bot. Madrid 47(1). 239 (1989 publ. 1990)

Distribution: Spain

12 SPA

Remarks: Dryopteris aemula x corleyi

species :

Dryopteris ×asturiensis Fraser-Jenk. & Gibby; Fern Gaz. 13(2): 113 (1986)

Distribution: Spain

12 SPA

Remarks: Dryopteris affinis x corleyi

species :

Dryopteris ×australis (Wherry) Small; Ferns S. E. States [Small] 279 cum tab., 477 (1938)

Distribution: USA (Alabama, Arkansas, Georgia, Louisiana, Maryland, Mississippi, North Carolina, South Carolina, Tennessee, Virginia)

78 USA

Remarks: Dryopteris celsa x Dryopteris ludoviciana

Synonyms: hidden

= Dryopteris clintoniana f. australis (Wherry) Wherry; Amer. Fern J. 40: 120 (1950)

= Dryopteris clintoniana var. australis Wherry; Amer. Fern J. 27: 2 (1937)

= Dryopteris cristata var. australis (Wherry) Blomq. & Correll

species :

Dryopteris ×benedictii (Farw.) Wherry; Amer. Fern J. 51: 34 (1961)

Distribution: Canada (Ontario); USA (Connecticut, Massachusetts, New Hampshire, New Jersey, New York, Ohio, Pennsylvania, Vermont)

72 CDN 75 USA

Remarks: Dryopteris carthusiana x clintoniana

Synonyms: hidden

= Filix benedictii Farw.; Pap. Michigan Acad. Sci. 2: 16 (1922)

= Filix-mas benedictii (Farw.) Farw.; Amer. Midl. Naturalist 12: 234-251 (1931)

= Filix ×benedictii (Farw.) Farw.; Pap. Michigan Acad. Sci. 2: 15 (1923)

species :

Dryopteris ×boottii (Tuck.) Underw.; Our nat. Ferns ed. 4: 117 (1893)

Distribution: Canada (New Brunswick, Newfoundland, Nova Scotia, Ontario, Québec); USA (Connecticut, Delaware, Iowa, Illinois, Indiana, Massachusetts, Maryland, Maine, Michigan, Minnesota, New Hampshire, New Jersey, New York, Ohio, Pennsylvania, Rhode Island, Tennessee, Virginia, Vermont, Wisconsin, West Virginia)

72 CDN 74 USA 75 USA 78 USA

Remarks: Dryopteris cristata x Dryopteris intermedia

Synonyms: hidden

= Aspidium spinulosum var. boottii (Tuck.) A.Gray; Manual (Gray), ed. 2: 598 (1856)

= Aspidium ×boottii Tuck.; Mag. Hort. Bot. 9: 145 (1843)

= Aspidium ×uliginosum (Newman) Nyman; Consp. Fl. Eur. Suppl. 1: 866 (1884), nom. illeg.

= Dryopteris ×boottii f. multiflora (Gilbert) M.Broun; Index No. Amer. Ferns: 63 (1938)

= Dryopteris ×rigida (Hoffm.) Underw.; Native Ferns, ed. 4: 116 (1893), nom. illeg.

= Dryopteris ×rigida A.Gray; Manual (Gray) ed. 1: 631 (1848)

= Filix ×boottii (Tuck.) Farw.; Rep. (Annual) Michigan Acad. Sci. 18: 81 (1916)

= Filix-mas ×boottii (Tuck.) Farw.; Amer. Midl. Naturalist 12: 253 (1931)

= Lastrea ×bootii (Tuck.) Nieuwl.; Midl. Naturalist (Notre Dame) 2: 278 (1912)

= Lastrea ×uliginosa Newman; Phytologist 3: 679 (1849)

= Lophodium ×uliginosum (Newman) Newman; Phytologist 4: 371 (1851)

= Nephrodium cristatum var. uliginosum (Newman) Baker; Syn. Fil. [W. J. Hooker & Baker] 273 (1867)

= Nephrodium spinulosum var. bootii (Tuck.) Baker; Syn. Fil. [W. J. Hooker & Baker] 275 (1867)

= Nephrodium ×boottii (Tuck.) Baker; Syn. Fil. [W. J. Hooker & Baker] 275 (1867)

= Nephrodium ×boottii f. multiflorum Gilbert; List N. Amer. Pterid.: 17, 36 (1901)

= Nephrodium ×uliginosum (Newman) Baker; Syn. Fil. [W. J. Hooker & Baker] 273 (1867)

= Nephrodium ×uliginosum (Newman) Rouy; Fl. France [Rouy & Foucaud] 14: 414 (1913), isonym

= Polystichum ×boottii (Tuck.) Gand.; Fl. Eur. [Gand.] 27: 168 (1891)

= Polystichum ×uliginosum (Newman) P.E.Fourn.; Quatre Fl. France: 17 (1934)

= Thelypteris ×boottii (Tuck.) Nieuwl.; Midl. Naturalist (Notre Dame) 1: 226 (1910)

= Thelypteris ×boottii f. multiflora (Gilbert) Dole; Fl. Vermont, ed. 3: 10 (1937)

= Thelypteris ×boottii var. multiflora (Gilbert) Mousley; Canad. Field-Naturalist 34-36: 149 (1920)

species :

Dryopteris ×borbasii Litard.; Bull. Soc. Bot. Deux-Sèvres 1909-1910: 85 (1910)

Distribution: France

12 FRA

Remarks: provisionally accepted name, status uncertain

Synonyms: hidden

= Aspidium ×subalpinum Hand.-Mazz.; Oesterr. Bot. Z. 53: 291 (1903)

= Nephrodium ×subalpinum Rouy; Fl. France [Rouy & Foucaud] 14: 415 (1913)

species :

Dryopteris ×brathaica Fraser-Jenk. & Reichst.; Fern Gaz. 11(5): 337 (1977)

Distribution: +England; Germany (Baden-Württemberg, Bayern, Niedersachsen); Austria; North Macedonia; European Russia; Canada (Québec)

10 GRB 11 AUT GER 13 NMA 14 RUS 72 CDN

Remarks: Dryopteris carthusiana x filix-mas

species :

Dryopteris ×burgessii B.Boivin; Naturaliste Canad. 93: 272 (1966)

Distribution: Canada (Ontario, Québec); USA (Connecticut, Massachusetts, Maryland, New Hampshire, New Jersey, New York, Pennsylvania, Vermont)

72 CDN 75 USA 78 USA

Remarks: Dryopteris clintoniana x marginalis

species :

Dryopteris ×burnatii Christ & Wilczek; Annuaire Conserv. Jard. Bot. Genève 15-16: 345 (1913)

Distribution: SE-France

12 FRA

species :

Dryopteris ×cantabrica Alejandre, Pérez Carro & Fern.Areces; Flora Montiber. 63: 69 (2016)

Distribution: Spain

12 SPA

Remarks: Dryopteris corleyi x guanchica

species :

Dryopteris ×cedroensis Gibby & Widén; Fern Gaz. 12(5): 267 (1983)

Distribution: Canary Isl. (La Gomera)

21 CNY

Remarks: Dryopteris guanchica x oligodonta

species :

Dryopteris ×complanata Fraser-Jenk.; Fern Gaz. 18(1): 11 (2007)

Distribution: Germany (Baden-Württemberg, Nordrhein-Westfalen)

11 GER

Remarks: Dryopteris filix-mas x pseudodisjuncta

species :

Dryopteris ×complexa Fraser-Jenk.; Sommerfeltia 6: xi (1987) [nothossp.]

subspecies :

subsp. complexa

Distribution: England (England, SW-Scotland, Wales); Ireland (Ireland, Northern Ireland); Belgium; Luxembourg; Germany (Baden-Württemberg, Nordrhein-Westfalen, Saarland, Sachsen-Anhalt); Switzerland; Austria; Portugal; Spain; France; Italy; Romania

10 GRB IRE 11 AUT BGM GER LUX SWI 12 FRA POR SPA 13 ITA ROM

Remarks: affinis agg., Dryopteris affinis ssp. affinis x filix-mas

subspecies :

subsp. contorta Fraser-Jenk.; Sommerfeltia 6: xii (1987) [nothossp.]

Distribution: England (England, Wales)

10 GRB

Remarks: affinis agg., Dryopteris affinis ssp. paleaceolobata x filix-mas

subspecies :

subsp. eschelmuelleri Fraser-Jenk.; Fern Gaz. 18(1): 10 (2007) [nothossp.]

Distribution: Germany (Bayern); Switzerland; Croatia; Turkey (Anatolia)

11 GER SWI 13 CRO 34 TUR

Remarks: affinis agg., Dryopteris affinis ssp. punctata x filix-mas

Synonyms: hidden

= Dryopteris ×complexa var. eschelmuelleri (Fraser-Jenk.) Fraser-Jenk. & Parris; Indian J. Forest. 43(3): 279 (2021)

subspecies :

subsp. transsilvanica Tlalka, S.Jess., A.Rostañski & Rojek; Fern Gaz. 21(1): 23-34 (2019) [nothossp.]

Distribution: S-Poland; North Macedonia; C-Romania

11 POL 13 NMA ROM

Remarks: affinis agg., Dryopteris affinis ssp. jessenii x filix-mas

Synonyms: hidden

= Dryopteris ×transsilvanica (Tlalka, S.Jess., A.Rostañski & Rojek) Tlalka; Ann. Bot. Fenn. 61: 17-23 (2024)

species :

Dryopteris ×convoluta (Fraser-Jenk.) Fraser-Jenk.; Fern Gaz. 18(1): 11 (2007) [nothossp.]

subspecies :

subsp. convoluta

Distribution: Germany (Baden-Württemberg, Bayern); Switzerland; France; Corsica

11 GER SWI 12 COR FRA

Remarks: Dryopteris filix-mas x cambrensis ssp. insubrica

subspecies :

subsp. inconspicua Fraser-Jenk.; Fern Gaz. 18(1): 11 (2007) [nothossp.], non D. inconspicua Copel. 1917

Distribution: England (Scotland); France

10 GRB 12 FRA

Remarks: Dryopteris filix-mas x cambrensis ssp. pseudocomplexa

Synonyms: hidden

= Dryopteris ×neoinconspicua P.D.Sell; Fl. Gr. Brit. Ireland 1: 671 (2018), nom. nov.

= Dryopteris ×complexa nothosubsp. inconspicua (Fraser-Jenk.) Fraser-Jenk.; Indian Fern J. 39(1): 121 (2022)

subspecies :

subsp. occidentalis Fraser-Jenk.; Fern Gaz. 18(1): 11 (2007) [nothossp.]

Distribution: England (England, Scotland, Wales); Ireland (Ireland); France; Corsica

10 GRB IRE 12 COR FRA

Remarks: Dryopteris filix-mas x cambrensis ssp. cambrensis

species :

Dryopteris ×correllii W.H.Wagner & A.V.Gilman; Amer. Fern J. 91: 11 (2001)

Distribution: USA (Vermont)

75 USA

species :

Dryopteris ×critica (Fraser-Jenk.) Fraser-Jenk.; Fern Gaz. 18(1): 11 (2007)

Distribution: England (England, Scotland, Wales); Norway; Belgium; Luxembourg; Germany (Baden-Württemberg, Bayern, Hessen, Niedersachsen, Saarland, Sachsen); Switzerland; Austria; Czech Republic; Hungary; ?Spain; France; Italy; Croatia; Romania; Bulgaria; European Turkey

10 GRB NOR 11 AUT BGM CZE GER HUN LUX SWI 12 FRA 13 BUL CRO ITA ROM TUE

Remarks: Dryopteris borreri x filix-mas

Synonyms: hidden

= Dryopteris ×complexa nothosubsp. critica Fraser-Jenk.; Sommerfeltia 6 (1987)

species :

Dryopteris ×deweveri (Jansen) Jansen & Wacht.; Geillustreerde Schoolflora voor Nederland [Heukels & Wachter], ed. II: 93 (1934)

Distribution: England (England, Wales, S-Scotland, Orkney Isl., Isle of Man); Ireland (Ireland, Northern Ireland); Germany (Brandenburg, Berlin, Baden-Württemberg, Bayern, Mecklenburg-Vorpommern, Rheinland-Pfalz, Sachsen); Switzerland; Czech Republic; Slovakia; etc.

10 GRB IRE 11 CZE GER SLO SWI

Remarks: Dryopteris carthusiana x dilatata

Synonyms: hidden

= Aspidium ×deweveri Jansen; Ned. Kruidk. Arch. 1932: 298 (1932)

species :

Dryopteris ×doeppii Rothm.; Candollea 10: 93 (1945)

Distribution: Germany (Baden-Württemberg)

11 GER

Remarks: Dryopteris affinis x carthusiana

species :

Dryopteris ×doluchanovii Askerov; Izv. Akad. Nauk Azerbaidzhansk. SSR Biol. Nauk. 1978(4): 4 (1978 publ. 1979)

Distribution: Northern Caucasus

33 NCS

Remarks: Dryopteris dilatata x affinis (pseudomas)

species :

Dryopteris ×dowellii (Farw.) Wherry; Amer. Fern J. 51: 34 (1961)

Distribution: Canada (Ontario); USA (Connecticut, Massachusetts, Maine, Michigan, New Hampshire, New Jersey, New York, Pennsylvania, Vermont)

72 CDN 75 USA

Remarks: Dryopteris clintoniana x Dryopteris intermedia

Synonyms: hidden

= Filix dowellii Farw.; Pap. Michigan Acad. Sci. 2: 16 (1922)

= Filix-mas dowellii (Farw.) Farw.; Amer. Midl. Naturalist 12: 234-251 (1931)

species :

Dryopteris ×euxinensis Fraser-Jenk. & Corley; Brit. Fern Gaz. 10: 225 (1972)

Distribution: Northern Caucasus; Turkey (Anatolia)

33 NCS 34 TUR

Remarks: Dryopteris caucasica x filix-mas

species :

Dryopteris ×fraser-jenkinsii Gibby & Widén; Fern Gaz. 12(5): 268 (1983)

Distribution: Spain

12 SPA

Remarks: Dryopteris affinis ssp. affinis x dilatata or D. guanchica

species :

Dryopteris ×fujipedis Sa.Kurata; J. Geobot. 14: 35 (1965)

Distribution: South Korea; Japan

38 JAP KOR

Remarks: Dryopteris crassirhizoma x lacera

species :

Dryopteris ×furadensis Bennert, Rasbach, K.Rasbach & Viane; Fern Gaz. 14(1): 10 (1991)

Distribution: Madeira (Madeira Isl.)

21 MDR

Remarks: Dryopteris aitoniana x intermedia ssp. maderensis

species :

Dryopteris ×ghatakii Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.), Bot. 18(5): 460 (1989)

Distribution: S-India (Tamil Nadu)

40 IND

Remarks: Dryopteris cochleata x sparsa ssp. rectipinnula

species :

Dryopteris ×gomerica Gibby & Widén; Fern Gaz. 12(5): 267 (1983)

Distribution: Canary Isl. (La Gomera)

21 CNY

Remarks: Dryopteris aemula x guanchica

species :

Dryopteris ×gotenbaensis Nakaike; J. Geobot. 17: 36, fig. (1969)

Distribution: Japan (Honshu)

38 JAP

Remarks: Dryopteris hondoensis x uniformis

species :

Dryopteris ×graeca Fraser-Jenk. & Gibby; Candollea 35(1): 309 (1980)

Distribution: Greece

13 GRC

Remarks: Dryopteris mindshelkensis x pallida

species :

Dryopteris ×haganecola Sa.Kurata; J. Geobot. 18: 4 (1970)

Distribution: Japan

38 JAP

Remarks: Dryopteris cycadina x uniformis

species :

Dryopteris ×hakonecola Sa.Kurata; J. Geobot. 7: 13 (1958)

Distribution: Japan

38 JAP

Remarks: Dryopteris dickinsii x uniformis

species :

Dryopteris ×hisatsuana Sa.Kurata; J. Geobot. 13: 42 (1964)

Distribution: Japan

38 JAP

Remarks: Dryopteris handeliana x lacera

species :

Dryopteris ×holttumii Li Bing Zhang; Fl. China 2-3: 616 (2013), nom. nov.

Distribution: Taiwan

38 TAI

Remarks: Dryopteris apiciflora x maximowicziana

Synonyms: hidden

= Dryopsis ×fauriei Holttum & P.J.Edwards; Kew Bull. 41(1): 198 (1986)

= Dryopteris splendens var. formosana Rosenst.; Hedwigia 56(5): 343 (1915)

= Dryopteris ×fauriei (Holttum & P.J.Edwards) Li Bing Zhang; Phytotaxa 71: 17-27 (2012), nom. illeg., non Kodama 1914

species :

Dryopteris ×initialis Fraser-Jenk. & Corley; Brit. Fern Gaz. 10: 230 (1972)

Distribution: European Russia; Northern Caucasus; Turkey (Anatolia)

14 RUS 33 NCS 34 TUR

Remarks: Dryopteris caucasica x oreades

species :

Dryopteris ×kominatoensis Tagawa; Acta Phytotax. Geobot. 6: 154 (1937)

Distribution: South Korea; Japan

38 JAP KOR

Remarks: Dryopteris monticola x tokyoensis

species :

Dryopteris ×kouzaii Akasawa; Bull. Kochi Women´s Univ., Ser. Nat. Sci. 27: 3 (1979)

Distribution: Japan (Shikoku)

38 JAP

Remarks: Dryopteris dickinsii x handeliana

species :

Dryopteris ×lawalreei Janch.; Cat. Fl. Austr., I. Pteridophyt. & Anthophyt.: 121 (1963)

Distribution: Austria

11 AUT

Remarks: Dryopteris borreri x carthusiana x filix-mas

species :

Dryopteris ×leedsii Wherry; Bartonia 21: 2, t. 1(1) (1942)

Distribution: USA (Arkansas, Maryland, Michigan, New Jersey, New York, Pennsylvania, Tennessee, Virginia)

75 USA 78 USA

Remarks: Dryopteris celsa x Dryopteris marginalis

species :

Dryopteris ×liddarensis Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.), Bot. 18(5): 460 (1989)

Distribution: NW-India (Jammu & Kashmir)

40 IND

Remarks: Dryopteris barbigera ssp. barbigera x serratodentata

species :

Dryopteris ×ligustica Gibby, S.Jess. & Marchetti; Fern Gaz. 22(3): 138 (2024), nom. nov.

Distribution: France; Italy

12 FRA 13 ITA

Remarks: Dryopteris filix-mas x tyrrhena

Synonyms: hidden

= Dryopteris ×lunensis Gibby, S.Jess. & Marchetti; Mem. Accad. Lunig. Sci. 66 [Atti Convegn. Internaz. 1994 - Pteridofite Lunigiana storica]: 93 (1996), nom. illeg., non (Christ) C. Chr.

species :

Dryopteris ×litardierei Rothm.; Candollea 10: 94 (1945)

Distribution: Corsica

12 COR

Remarks: Dryopteris paleacea x pallida

species :

Dryopteris ×loyalii Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.), Bot. 18(5): 462 (1989)

Distribution: India (Meghalaya)

40 IND

Remarks: Dryopteris carolihopei x marginata

species :

Dryopteris ×macdonellii Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.), Bot. 18(5): 464 (1989)

Distribution: Pakistan (Hazara); NW-India (Jammu & Kashmir)

40 IND PAK

Remarks: Dryopteris filix-mas x ramosa

species :

Dryopteris ×madalenae Fraser-Jenk.; Arquipelago, Sér. Ci. Nat. 2: 96 (1981)

Distribution: Azores (Pico)

21 AZO

Remarks: Dryopteris intermedia var. azorica x crispifolia

species :

Dryopteris ×makabensis Kaor.Yamam. & N.Murak.; Acta Phytotax. Geobot. 75(1): 5 (2024)

Distribution: Japan

38 JAP

Remarks: Dryopteris caudipinna x tokyoensis

species :

Dryopteris ×mantoniae Fraser-Jenk. & Corley; Brit. Fern Gaz. 10: 230 (1972)

Distribution: England (England, Scotland, Wales); Spain; Northern Caucasus

10 GRB 12 SPA 33 NCS

Remarks: Dryopteris filix-mas x oreades

species :

Dryopteris ×martinsiae Fraser-Jenk.; Arquipelago, Sér. Ci. Nat. 2: 95 (1981)

Distribution: Azores (Pico)

21 AZO

Remarks: Dryopteris aemula x crispifolia

species :

Dryopteris ×mayebarae Tagawa; Acta Phytotax. Geobot. 7: 185 (1938)

Distribution: South Korea; Japan

38 JAP KOR

Remarks: Dryopteris commixta x uniformis

species :

Dryopteris ×mickelii J.H.Peck; Amer. Fern J. 91(1): 36 (2001)

Distribution: Canada (Ontario); USA (Michigan, New Jersey, New York, Ohio, Pennsylvania, Vermont)

72 CDN 75 USA

Remarks: Dryopteris clintoniana x goldieana

species :

Dryopteris ×mituii Seriz.; J. Jap. Bot. 46: 19 (1971)

Distribution: South Korea; Japan

38 JAP KOR

Remarks: Dryopteris lacera x uniformis

species :

Dryopteris ×miyazakiensis Miyam.; J. Jap. Bot. 59(8): 243 (1984)

Distribution: Japan

38 JAP

Remarks: Dryopteris commixta x hangchowensis

species :

Dryopteris ×montgomeryi Fraser-Jenk. & Widén; Advances Forest. Res. India 29: 153 (2006)

Distribution: Canada (Ontario)

72 CDN

Remarks: Dryopteris filix-mas subsp. brittonii x marginalis

species :

Dryopteris ×neowherryi W.H.Wagner; W. & F. Wagner, Castanea 31: 136, f. 4E-H, 5, 6, 8G, H (1966)

Distribution: Canada (Ontario); USA (Connecticut, District of Columbia, Georgia, Illinois, Indiana, Massachusetts, Maryland, North Carolina, New Jersey, New York, Ohio, Pennsylvania, Virginia, Vermont, West Virginia)

72 CDN 74 USA 75 USA 78 USA

Remarks: Dryopteris goldieana x Dryopteris marginalis

species :

Dryopteris ×orexpansa Pérez Carro & Fern.Areces; Flora Montiber. 54: 152 (2013)

Distribution: Spain

12 SPA

Remarks: Dryopteris expansa x oreades

species :

Dryopteris ×otomasui Sa.Kurata; J. Geobot. 15: 83 (1967)

Distribution: Japan

38 JAP

Remarks: Dryopteris commixta x polylepis

species :

Dryopteris ×picoensis Fraser-Jenk. & Gibby; Arquipelago, Sér. Ci. Nat. 2: 94 (1981)

Distribution: Azores (Pico)

21 AZO

Remarks: Dryopteris affinis ssp. affinis x azorica

species :

Dryopteris ×pittsfordensis Sloss.; Rhodora 6: 75 (1904)

Distribution: Canada (Ontario); USA (Connecticut, Massachusetts, New Hampshire, New Jersey, New York, Ohio, Pennsylvania, Virginia, Vermont, Wisconsin, West Virginia)

72 CDN 74 USA 75 USA 78 USA

Remarks: Dryopteris carthusiana x marginalis

Synonyms: hidden

= Aspidium pittsfordense (Sloss.) Eastman; Ferns & Fern Allies New England 36 (1904)

= Filix spinulosa var. pittsfordensis (Sloss.) Farw.; Rep. (Annual) Michigan Acad. Sci. 18: 83 (1916)

= Nephrodium pittsfordensis (Sloss.) Woolson; Fern Bull. 12: 106 (1904)

= Thelypteris pittsfordensis (Sloss.) Vict.; Contr. Lab. Bot. Univ. Montréal 2: 51 (1923)

species :

Dryopteris ×pseudoabbreviata Jermy; Brit. Fern Gaz. 10: 12 (1968)

Distribution: England (Scotland: Mid Ebudes)

10 GRB

Remarks: Dryopteris aemula x oreades

species :

Dryopteris ×pseudocommixta Sa.Kurata; J. Geobot. 13: 43 (1964)

Distribution: Japan

38 JAP

Remarks: Dryopteris commixta x kuratae

species :

Dryopteris ×pseudohangchowensis Miyam.; J. Jap. Bot. 59(8): 242 (1984)

Distribution: Japan

38 JAP

Remarks: Dryopteris hangchowensis x kuratae

species :

Dryopteris ×pteridiiformis Christ; Bull. Acad. Int. Géogr. Bot. 17: 137-138 (1907)

Distribution: China (Yunnan, Guizhou); N-Myanmar

36 CHI 41 MYA

Remarks: obviously a hybrid D. carolihopei x camusiae fide Fraser-Jenkins et al. (2018)

species :

Dryopteris ×rarissima Sa.Kurata; J. Geobot. 13: 43 (1964)

Distribution: Japan

38 JAP

Remarks: Dryopteris commixta x sparsa

species :

Dryopteris ×ronald-vianensis Pérez Carro & Fern.Areces; Flora Montiber. 63: 74 (2016)

Distribution: Spain

12 SPA

Remarks: Dryopteris dilatata x guanchica

species :

Dryopteris ×sardoa Fraser-Jenk. & Reichst.; Fraser-Jenkins, Reichst. & G. Vida, Fern Gaz. 11: 193 (1975)

Distribution: Sardinia

12 SAR

Remarks: Dryopteris oreades x tyrrhena

species :

Dryopteris ×sarvelae Fraser-Jenk. & Jermy; Fern Gaz. 11(5): 339 (1977)

Distribution: England (England, Scotland); Finland; Germany (Mecklenburg-Vorpommern); Poland; Slovakia; European Russia

10 FIN GRB 11 GER POL SLO 14 RUS

Remarks: Dryopteris carthusiana x expansa

species :

Dryopteris ×satsumana Sa.Kurata; J. Geobot. 17: 89 (1969)

Distribution: Japan

38 JAP

Remarks: Dryopteris commixta x dickinsii

species :

Dryopteris ×separabilis Small; Ferns S. E. States [Small] 284, cum tab. (1938)

Distribution: USA (Maryland, Michigan, North Carolina, New Jersey, New York, Tennessee, Virginia)

75 USA 78 USA

Remarks: Dryopteris celsa x Dryopteris intermedia

species :

Dryopteris ×shibipedis Sa.Kurata; J. Geobot. 14: 84 (1966)

Distribution: Japan (Kyushu)

38 JAP

Remarks: Dryopteris kinkiensis x pacifica

Conservation Status: EW in Japan (listed as EX)

species :

Dryopteris ×shibisanensis Sa.Kurata; J. Geobot. 14: 37 (1965)

Distribution: Japan

38 JAP

Remarks: Dryopteris atrata x commixta

species :

Dryopteris ×sjoegrenii Fraser-Jenk.; Arquipelago, Sér. Ci. Nat. 2: 96 (1981)

Distribution: Azores (Pico)

21 AZO

Remarks: Dryopteris intermedia ssp. azorica x dilatata

species :

Dryopteris ×slossoniae (Davenp.) Wherry ex Lellinger; Amer. Fern J. 74(2): 62 (1984)

Distribution: Canada (New Brunswick, Ontario); USA (Connecticut, Delaware, Indiana, Massachusetts, Maryland, Maine, Michigan, New Hampshire, New Jersey, New York, Ohio, Pennsylvania, Rhode Island, South Carolina, Virginia, Vermont, Wisconsin)

72 CDN 74 USA 75 USA 78 USA

Remarks: Dryopteris cristata x Dryopteris marginalis

Synonyms: hidden

= Dryopteris clintoniana f. slossoniae (Davenp.) Clute; Ferns in Their Haunts 178, 385 (1938)

= Dryopteris clintoniana var. slossoniae (Davenp.) Burnham; Amer. Fern J. 6: 103 (1916)

= Dryopteris cristata f. slossoniae (Davenp.) Weath.; Amer. Fern J. 26: 65 (1936)

= Dryopteris ×slossoniae (Davenp.) Wherry; Bartonia 21: 15 (1942)

= Nephrodium cristatum var. slossoniae Davenp.; Rhodora 4: 52 (1902)

= Nephrodium ×slossoniae (Davenp.) Hahne; Allg. Bot. Z. Syst. 10: 103 (1904)

species :

Dryopteris ×subaustriaca Rothm.; Candollea 10: 100 (1945)

Distribution: Germany (Baden-Württemberg, Niedersachsen); etc.

11 GER

Remarks: Dryopteris dilatata x filix-mas

Synonyms: hidden

= Aspidium remotum var. subalpinum Borbás; Math. Term. Közlem. 8: 36 (1876)

= Polystichum ×subalpinum (Borbás) P.E.Fourn.; Quatre Fl. France: 17 (1934)

species :

Dryopteris ×sugino-takaoi Sa.Kurata; J. Geobot. 14: 36 (1965)

Distribution: Japan

38 JAP

Remarks: Dryopteris lacera x polylepis

species :

Dryopteris ×takachihoensis Miyam.; J. Jap. Bot. 59(8): 240 (1984)

Distribution: Japan

38 JAP

Remarks: Dryopteris hangchowensis x uniformis

species :

Dryopteris ×telesii Fraser-Jenk.; Arquipelago, Sér. Ci. Nat. 2: 97 (1981)

Distribution: Azores (Pico)

21 AZO

Remarks: Dryopteris crispifolia x dilatata

species :

Dryopteris ×tetsu-yamanakae Sa.Kurata; J. Geobot. 10: 100 (1962)

Distribution: Japan

38 JAP

Remarks: Dryopteris commixta x sieboldii

species :

Dryopteris ×tiantangzhaiensis H.-J.Wei & B.Chen; Taxon 73(1): 136 (2024)

Distribution: China

36 CHI

Remarks: Dryopteris gymnophylla x tsoongii

species :

Dryopteris ×tokudae Sugim.; J. Geobot. 7: 97 (1958)

Distribution: Japan

38 JAP

Remarks: Dryopteris crassirhizoma x polylepis

species :

Dryopteris ×triploidea Wherry; Amer. Fern J. 50: 90 (1960)

Distribution: Canada (New Brunswick, Nova Scotia, Ontario, Prince Edward Isl., Québec); USA (Connecticut, Delaware, Iowa, Illinois, Indiana, Kentucky, Massachusetts, Maryland, Maine, Michigan, Minnesota, North Carolina, New Hampshire, New Jersey, New York, Ohio, Pennsylvania, Rhode Island, Tennessee, Virginia, Vermont, Wisconsin, West Virginia)

72 CDN 74 USA 75 USA 78 USA

Remarks: Dryopteris intermedia ssp. intermedia x carthusiana

Synonyms: hidden

= Dryopteris austriaca var. fructuosa (Gilbert) C.V.Morton; Amer. Fern J. 40: 222 (1950)

= Dryopteris intermedia f. fructuosa (Gilbert) Clute; Ferns in Their Haunts 183, 385 (1938)

= Dryopteris intermedia var. fructuosa (Gilbert) Wherry

= Dryopteris spinulosa var. fructuosa (Gilbert) Trudell; Amer. Fern J. 19: 136, et 20: 39 (1930)

= Lastrea dilatata var. glandulosa T.Moore; Ferns Gr. Brit. (Moore): t. 23 (1856)

= Nephrodium spinulosum f. fructuosum (Gilbert) Clute; Fern Bull. 1907: 23 (1907)

= Nephrodium spinulosum var. fructuosum Gilbert; List N. Amer. Pterid.: 17, 37 (1901)

= Thelypteris spinulosa var. fructuosa (Gilbert) Fernald; Rhodora 28: 146 (1926)

species :

Dryopteris ×uliginosa (A.Braun ex Döll) Kuntze ex Druce; List Brit. Pl. 87 (1908)

Distribution: England; Germany (Brandenburg, Berlin, Baden-Württemberg, Bayern, Hessen, Hamburg, Mecklenburg-Vorpommern, Niedersachsen, Schleswig-Holstein, Sachsen, Thüringen); Czech Republic; Slovakia; European Russia; Canada (New Brunswick, Newfoundland, Ontario); USA (Connecticut, Delaware, Iowa, Massachusetts, Maryland, Maine, Minnesota, North Carolina, North Dakota, New Hampshire, New Jersey, New York, Ohio, Pennsylvania, Virginia, Vermont, Wisconsin, West Virginia)

10 GRB 11 CZE GER SLO 14 RUS 72 CDN 74 USA 75 USA 78 USA

Remarks: Dryopteris carthusiana x cristata

Synonyms: hidden

= Aspidium cristatum subsp. uliginosum (A.Braun ex Döll) Nyman; Consp. Fl. Eur. Suppl. 1: 866 (1884)

= Aspidium spinulosum subsp. tauschii Celak.; Prodr. Fl. Böhmen 1: 10 (1867)

= Aspidium spinulosum var. uliginosum A.Braun ex Döll; Rhein. Fl. 17 (1843)

= Dryopteris ×laschii E.Walter; Pterid. Exsicc. 1(2): p. 36 (1939)

= Dryopteris ×tauschii (Celak.) Domin; Preslia 13-15: 8 (1936)

= Lastrea cristata subsp. uliginosa (A.Braun ex Döll) Hayw.; Bot. Pocket-Book 180 (1872)

= Lastrea spinulosa var. uliginosa (A.Braun ex Döll) H.C.Watson & Dennes; London Cat. Brit. Pl. [H. C. Watson], ed. 3: 14 (1850)

= Lastrea ×uliginosa (A.Braun ex Döll) Syme & N.E.Br.; Engl. Bot., 3rd ed., vol. XII: 73 (1886)

= Lophodium ×uliginosum (A.Braun ex Döll) Newman; 371 (1851)

= Nephrodium cristatum var. uliginosum (A.Braun ex Döll) Hook. & Baker; 273 (1867)

species :

Dryopteris ×uralensis Gureeva & Moczalov; Sist. Zametki Mater. Gerb. Krylova Tomsk. Gosud. Univ. 110: 32 (2014)

Distribution: E-European Russia

14 RUS

Remarks: Dryopteris expansa x filix-mas

species :

Dryopteris ×urbachensis S.Jess. & Bujnoch; Fern Gaz. 22(3): 129 (2024)

Distribution: Switzerland (Bern)

11 SWI

Remarks: Dryopteris filix-mas x lacunosa

species :

Dryopteris ×vidae Fraser-Jenk. & Gibby; Candollea 35(1): 306 (1980)

Distribution: Montenegro; etc.

13 MNT

Remarks: Dryopteris pallida x vilarii

species :

Dryopteris ×watanabei Sa.Kurata; J. Geobot. 14: 36 (1965)

Distribution: South Korea; Japan

38 JAP KOR

Remarks: Dryopteris crassirhizoma x uniformis

species :

Dryopteris ×wechteriana Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.), Bot. 18(5): 466 (1989)

Distribution: India (Himachal Pradesh, Uttarakhand)

40 IND

Remarks: Dryopteris chrysocoma x nigropaleacea

species :

Dryopteris ×woynarii Rothm.; Candollea 10: 94 (1946)

Distribution: England; etc.

10 GRB

Remarks: Dryopteris affinis x dilatata

species :

Dryopteris ×yamashitae Sa.Kurata; J. Geobot. 17: 4 (1969)

Distribution: Japan

38 JAP

Remarks: Dryopteris hayatae x sparsa

species :

Dryopteris ×yuyamae Sa.Kurata; J. Geobot. 15: 8 (1966)

Distribution: Japan

38 JAP

Remarks: Dryopteris kuratae x uniformis

species :

Dryopteris ×zygoparentalis Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.), Bot. 18(5): 467 (1989)

Distribution: India (West Bengal)

40 IND

Remarks: Dryopteris gamblei x scottii

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A complete, synonymic checklist of the Higher Plants of the World

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A complete, synonymic checklist of the Higher Plants of the World

Version 24.11 (November 5th, 2024)

Covering 355,576 accepted species,
8,715 hybrids, 47,863 accepted infraspecific taxa and 1,000,581 synonyms,
in total 1,412,785* names

[about 25,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus*" are deliberately omitted]

With complete, detailed distribution data (countries and regions) and literature references

Arranged in linear phylogenetic sequence reflecting true relationships

Fully searchable and continously updated

With 38,209 pictures of 15,787 species

Assembled through 40 years independently from other databases

In cooperation with Catalogue of Life

Contact

How to Cite:

Search

+ Family

Filter

Order

Family (28)

Genus (27)

Display options

A complete, synonymic checklist of the Higher Plants of the World

Version 24.11 (November 5th, 2024)

Covering 355,576 accepted species, 8,715 hybrids, 47,863 accepted infraspecific taxa and 1,000,581 synonyms, in total 1,412,785* names

[*about 25,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus" are deliberately omitted]

With complete, detailed distribution data (countries and regions) and literature references

Arranged in linear phylogenetic sequence reflecting true relationships

Fully searchable and continously updated

With 38,209 pictures of 15,787 species

Assembled through 40 years independently from other databases

In cooperation with Catalogue of Life

Covering 355,576 accepted species,
8,715 hybrids, 47,863 accepted infraspecific taxa and 1,000,581 synonyms,
in total 1,412,785* names

[about 25,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus*" are deliberately omitted]