134.0000 Family Orchidaceae Juss.; Gen. Pl. [Jussieu] 64 (1789), nom. cons.

over 800 genera, more than 30.000 spp. worldwide, mostly tropical

Species listed in database: 30538; hybrids: 1461

134.4327 Genus Dactylorhiza Neck. ex Nevski; Trudy Bot. Inst. Akad. Nauk S.S.S.R., Ser. 1, Fl. Sist. Vyssh. Rast. 4: 332 (1937), nom. cons.

Distribution: Atlantic Islands, Europe, European Russia, few to Russian Far East and Alaska. The species concept in this genus is highly controversial and fluctuating, depending on author, leading to countless synonyms. Recent molecular studies point to relatively few highly variable species and a certain number of polyploid hybridogenous (but stabilized) taxa. Coeloglossum is phylogenetically embedded in Dactylorhiza.

Species listed in database: 46; hybrids: 83

species :

Dactylorhiza aristata (Fisch. ex Lindl.) Soó; Nom. Nov. Gen. Dactylorhiza 5 (1962)

Name: Fischer´s Orchid

Distribution: Russian Far East (Kamchatka, Khabarovsk, Kuril Isl., Sakhalin); China (Hebei, Henan, Shandong, Shanxi); North Korea; Japan; Aleutes; SW-Alaska

31 RFE 36 CHI 38 JAP NKO 70 ASK

Remarks: sambucina agg.

Synonyms: hidden

= Dactylorchis aristata (Fisch.) Verm.; Stud. Dactylorch. 64 (1947)

= Dactylorhiza aristata f. alba P.M.Br.; N. Amer. Native Orchid J. 1(1): 10 (1995)

= Dactylorhiza aristata f. albomaculata P.M.Br.; N. Amer. Native Orchid J. 10: 36 (2004 publ. 2005)

= Dactylorhiza aristata f. perbracteata (Lepage) Catling; Naturaliste Canad. 109: 278 (1982)

= Dactylorhiza aristata f. perbracteata (Lepage) P.M.Br.; N. Amer. Native Orchid J. 2(1): 68 (1996), isonym

= Dactylorhiza aristata f. rosea P.M.Br.; N. Amer. Native Orchid J. 1(3): 199 (1995)

= Dactylorhiza aristata var. kodiakensis Luer & G.M.Luer; Amer. Orchid Soc. Bull. 41: 207 (1972)

= Dactylorhiza aristata var. perbracteata (Lepage) Catling; Naturaliste Canad. 109(2): 278 (1982)

= Orchis aristata f. perbracteata Lepage; Amer. Midl. Naturalist 46: 757 (1952)

= Orchis aristata f. punctata Tatew.; Trans. Sapporo Nat. Hist. Soc. 9: 10 (1927)

= Orchis aristata Fisch. ex Lindl.; Gen. Sp. Orchid. Pl. 262 (1835)

= Orchis aristata var. albiflora Koidz.; Bot. Mag. (Tokyo) 31: 34 (1917)

= Orchis aristata var. immaculata Makino; J. Jap. Bot. 8: 17 (1932)

= Orchis aristata var. maculata Makino; J. Jap. Bot. 8: 6 (1932)

= Orchis beeringiana (Cham.) Kudô; J. Coll. Agric. Hokkaido Imp. Univ. 11: 94 (1922)

= Orchis latifolia var. beeringiana Cham.; Linnaea 3: 26 (1828)

species :

Dactylorhiza armeniaca Hedrén; Pl. Syst. Evol. 229(1-2): 42 (2001)

Distribution: Northern Caucasus (Tuapse, Krasnodar); Georgia [Caucasus]; Azerbaijan; NE-Turkey

33 AZE GEO NCS 34 TUR

Synonyms: hidden

= Dactylorhiza euxina subsp. armeniaca (Hedrén) Kreutz; Kompend. Eur. Orchid. 37 (2004)

species :

Dactylorhiza baumanniana J.Hölz. & Künkele; Mitt. Arbeitskreises Heimische Orchideen 15(4): 514 (1983)

subspecies :

subsp. baumanniana

Name: Baumanns Knabenkraut (DE)

Distribution: Albania; Bulgaria; Greece (mountains of NW-, NC-Greece, C-Greece around Mt. Giona, Mt. Erymanthos)

13 ALB BUL GRC

Remarks: traunsteineri agg. Included by Pedersen et al. 2019 in D. majalis subsp. pindica

subspecies :

subsp. smolikana (B.Willing & E.Willing) H.Baumann & R.Lorenz; J. Eur. Orch. 37(3): 607 (2005)

Distribution: N-Greece (Mt. Smolikas etc.)

13 GRC

Synonyms: hidden

= Dactylorhiza smolikana B.Willing & E.Willing; Ber. Arbeitskreis. Heimische Orchid. 6(1): 33 (1989)

species :

Dactylorhiza cantabrica H.A.Pedersen; Bot. J. Linn. Soc. 152(4): 428 (2006)

Distribution: Spain (Galicia: Léon, Lugo, Palencia, Cantabria)

12 SPA

Remarks: a tetraploid species derived from D. insularis x D. sambucina

Synonyms: hidden

= Dactylorhiza sambucina subsp. cantabrica (H.A.Pedersen) Kreutz; Ber. Arbeitskreis. Heimische Orchid. 24(1): 155 (2007)

species :

Dactylorhiza cordigera (Fr.) Soó; Nom. Nov. Gen. Dactylorhiza 5 (1962)

subspecies :

subsp. cordigera

Name: Herz-Knabenkraut

Distribution: Croatia; Bosnia & Hercegovina; Serbia; Kosovo; North Macedonia; Albania; Romania; Bulgaria; Greece (mountains of NW-, NC-, NE-Greece); European Russia

13 ALB BIH BUL CRO GRC KOS NMA ROM SRB 14 RUS

Remarks: majalis agg., possibly only a subspecis of D. majalis

Synonyms: hidden

= Dactylorchis cordigera (Fr.) Verm.; Stud. Dactylorch. 67 (1947)

= Dactylorhiza cordigera f. albiflora Landwehr; Wilde Orchid. Eur. 2: 557 (1977)

= Dactylorhiza cordigera var. grisebachii (Pant.) Soó; Nom. Nov. Gen. Dactylorhiza 5 (1962)

= Dactylorhiza cordigera var. rhodopeia Presser; J. Eur. Orch. 36(4): 982 (2004)

= Dactylorhiza cordigera var. rochelii (Griseb. & Schenk) Soó; Nom. Nov. Gen. Dactylorhiza 5 (1962)

= Dactylorhiza cordigera var. vermionica B.Willing & E.Willing; Ber. Arbeitskreis. Heimische Orchid. 4(1): 36 (1987)

= Dactylorhiza majalis subsp. cordigera (Fr.) E.Nelson; Monogr. & Ikonogr. Orchid.-Gattung Dactylorhiza 82 (1976) [without basionym page]

= Dactylorhiza majalis subsp. cordigera (Fr.) H.Sund.; Europ. Medit. Orchid., ed. 2: 45 (1975)

= Dactylorhiza richteri B.Bock; Bull. Soc. Bot. Centre-Ouest 42: 263 (2012)

= Orchis cordigera lusus immaculata Klinge; Revis. Orchis cordigera & O. angustifolia 19 (1893)

= Orchis cordigera proles rochelii (Griseb. & Schenk) Asch. & Graebn.; Syn. Mitteleur. Fl. [Ascherson & Graebner] 3: 742 (1907)

= Orchis cordigera f. banatica Asch. & Graebn.; Syn. Mitteleur. Fl. [Ascherson & Graebner] 3: 741 (1907)

= Orchis cordigera f. grisebachii (Pant.) Soó; Notizbl. Bot. Gart. Berlin-Dahlem 9: 910 (1926)

= Orchis cordigera f. macrobracteata (Schur) Soó; Bot. Arch. 23: 72 (1928)

= Orchis cordigera f. rivularis (Heuff.) Pauca; Fl. Reipubl. Popul. Roman. (Fl. Republ. Socialist. Romania) 12: 702 (1972)

= Orchis cordigera f. rochelii (Griseb. & Schenk) Klinge; Revis. Orchis cordigera & O. angustifolia 23 (1893)

= Orchis cordigera f. simonkaiana Soó; Repert. Spec. Nov. Regni Veg. 24: 31 (1927)

= Orchis cordigera f. subtriloba Soó; Bot. Arch. 23: 71 (1928)

= Orchis cordigera Fr.; Novit. Fl. Suec. Mant. 3: 130 (1845)

= Orchis cordigera var. foliosa Klinge; Revis. Orchis cordigera & O. angustifolia 37 (1893)

= Orchis cordigera var. grisebachii (Pant.) Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 33 (1899)

= Orchis cordigera var. klingei E.G.Camus; Monogr. Orchid. 174 (1908)

= Orchis cordigera var. macrobracteata Schur; Verh. Mitth. Siebenbürg. Vereins Naturwiss. Hermannstadt 4: 72 (1853)

= Orchis cordigera var. rocheliana Klinge; Revis. Orchis cordigera & O. angustifolia 20 (1893)

= Orchis cruenta Rochel; Pl. Banat. Rar. 34, t. 1, f. 1 (1828) non O. F. Müll.

= Orchis foliosa Schur; Enum. Pl. Transsilv. 642 (1866)

= Orchis grisebachii Pant.; Verh. Vereins Natur- Heilk. Presburg 2: 27 (1871-72)

= Orchis hungarica Pant. ex G.Keller & Soó; Repert. Spec. Nov. Regni Veg. Sonderbeih. A: 2: 227 (1933)

= Orchis latifolia f. gracilis Schur; Verh. Mitth. Siebenbürg. Vereins Naturwiss. Hermannstadt 4: 72 (1853)

= Orchis latifolia var. conica Lindl.; Gen. Sp. Orchid. Pl. 260 (1835)

= Orchis latifolia var. cordigera (Fr.) Rchb.f.; Icon. Fl. Germ. Helv. [H.G.L.Reichenbach] 13-14: 62 (1851)

= Orchis latifolia var. orbiculatus Zapal.; Consp. Fl. Galic. Crit. 209 (1906)

= Orchis latifolia var. rochelii Griseb. & Schenk; Arch. Naturgesch. (Berlin) 18: 355 (1852)

= Orchis monticola Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(1): 178 (1898), nom. superfl.

= Orchis monticola subsp. bosniaca (Beck) Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(1): 180 (1898)

= Orchis monticola subsp. cordigera (Fr.) Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(1): 179 (1898), nom. superfl.

= Orchis rivularis Heuff. ex Schur; Enum. Pl. Transsilv. 642 (1866)

= Orchis rocheliana (Klinge) Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(1): 179 (1898), nom. inval.

= Orchis traunsteineri var. grisebachii (Pant.) E.G.Camus; Monogr. Orchid. 183 (1908)

subspecies :

subsp. bosniaca (Beck) Soó; Nom. Nov. Gen. Dactylorhiza 5 (1962)

Distribution: Bosnia & Hercegovina; Montenegro; Serbia; Kosovo; Albania; North Macedonia; Bulgaria

13 ALB BIH BUL KOS MNT NMA SRB

Synonyms: hidden

= Dactylorhiza bosniaca (Beck) Aver.; Bot. Zhurn. (Moscow & Leningrad) 69(6): 875 (1984)

= Dactylorhiza lagotis (Rchb.f.) H.Baumann; Mitt. Arbeitskreises Heimische Orchideen 15(1): 50 (1983)

= Orchis bosniaca Beck; Ann. K.K. Naturhist. Hofmus. 2: 53 (1887)

= Orchis cordigera subsp. bosniaca (Beck) Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(1): 34 (1898)

= Orchis cordigera var. bosniaca (Beck) Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 28 (1899)

= Orchis lagotis Rchb.f.; Icon. Fl. Germ. Helv. [H.G.L.Reichenbach] 13: t. 403 (1851)

= Orchis latifolia var. lagotis Rchb.f.; Icon. Fl. Germ. Helv. [H.G.L.Reichenbach] 13-14: 58 (1851)

subspecies :

subsp. graeca (H.Baumann) Kreutz; Kompend. Eur. Orchid. 34 (2004)

Name: Griechisches Knabenkraut (DE)

Distribution: N-Greece (Greek Macedonia, Thracia, Rodopi, Mt. Varvousia)

13 GRC

Synonyms: hidden

= Dactylorhiza graeca H.Baumann; Mitt. Arbeitskreises Heimische Orchideen 15(1): 59 (1983)

= Dactylorhiza cordigera var. graeca (H.Baumann) Presser; J. Eur. Orch. 36(4): 981 (2004)

subspecies :

subsp. pindica (B.Willing & E.Willing) H.Baumann & R.Lorenz; J. Eur. Orch. 37(3): 706 (2005)

Name: Pindos-Knabenkraut (DE)

Distribution: Bulgaria; Greece (mountains of NW-, NC-Greece, C-Greece around Mt. Giona)

13 BUL GRC

Synonyms: hidden

= Dactylorhiza pindica B.Willing & E.Willing; Ber. Arbeitskreis. Heimische Orchid. 3(2): 215 (1986 publ. 1987)

= Dactylorhiza majalis subsp. pindica (B.Willing & E.Willing) H.A.Pedersen, P.J.Cribb & Rolf Kühn; Field Guide Orchids Europe Medit. 161 (2019)

subspecies :

subsp. siculorum (Soó) Soó; Nom. Nov. Gen. Dactylorhiza 5 (1962)

Distribution: N-Romania; SW-Ukraine

13 ROM 14 UKR

Synonyms: hidden

= Dactylorhiza majalis subsp. siculorum (Soó) E.Nelson; Monogr. & Ikonogr. Orchid.-Gattung Dactylorhiza 81 (1976)

= Dactylorhiza siculorum (Soó) Kreutz & Bobocea; Ber. Arbeitskreis. Heimische Orchid. 38(2): 109 (2021)

= Orchis cordigera subsp. siculorum Sóo; Notizbl. Bot. Gart. Berlin-Dahlem 9(89): 910 (1926), et Magyar Bot. Lapok 25: 274 (1927)

= Orchis cordigera var. siculorum (Soó) Pauca; Fl. Reipubl. Popul. Roman. (Fl. Republ. Socialist. Romania) 12: 701 (1972)

= Orchis siculorum (Soó) Soó; Repert. Spec. Nov. Regni Veg. 24: 31 (1927)

species :

Dactylorhiza elata (Poir.) Soó; Nom. Nov. Gen. Dactylorhiza 7 (1962)

Name: Hohes Knabenkraut (DE)

Distribution: Portugal; Spain; France; Corsica; Sardinia; Morocco; Algeria; Tunisia

12 COR FRA POR SAR SPA 20 ALG MOR TUN

Synonyms: hidden

= Dactylorchis elata (Poir.) Verm.; Stud. Dactylorch. 68 (1947)

= Dactylorhiza elata f. alba (Soó) Raynaud; Orchid. Maroc 103 (1985)

= Dactylorhiza elata f. cordata Landwehr; Wilde Orchid. Eur. 2: 557 (1977)

= Dactylorhiza elata f. leucantha (Maire & Weiller) Raynaud; Orchid. Maroc 104 (1985)

= Dactylorhiza elata f. pallida (Maire & Weiller) Raynaud; Orchid. Maroc 104 (1985)

= Dactylorhiza elata f. peltieri (Soó) Raynaud; Orchid. Maroc 103 (1985)

= Dactylorhiza elata f. speciosissima (Soó) Raynaud; Orchid. Maroc 103 (1985)

= Dactylorhiza elata subvar. maroccanica (Soó) Raynaud; Orchid. Maroc 104 (1985)

= Dactylorhiza elata subvar. speciosissima (Soó) Raynaud; Orchid. Maroc 103 (1985)

= Dactylorhiza elata subvar. stephensonii (Maire & Weiller) Raynaud; Orchid. Maroc 103 (1985)

= Dactylorhiza elata var. algerica (Rchb.f.) P.Delforge; Naturalistes Belges 96(Orchid. 28): 14 (2015)

= Dactylorhiza elata var. corsica (E.Rev. ex E.G.Camus) Soó; Nom. Nov. Gen. Dactylorhiza 7 (1962)

= Dactylorhiza elata var. stephensonii (Maire & Weiller) Raynaud; Orchid. Maroc 103 (1985)

= Dactylorhiza incarnata subsp. elata (Poir.) H.Sund.; Trop. & Mediterr. Orchideen 45 (1975)

= Gymnadenia elata (Poir.) Lindl.; Gen. Sp. Orchid. Pl. 277 (1835)

= Orchis elata f. alba Soó; Bull. Soc. Hist. Nat. Afrique N. 24: 173 (1933)

= Orchis elata f. albiflora Maire; Bull. Soc. Hist. Nat. Afrique N. 31: 42 (1940)

= Orchis elata f. klingeana Soó; Repert. Spec. Nov. Regni Veg. 24: 31 (1927)

= Orchis elata f. leucantha Maire & Weiller; Fl. Afrique N. 6: 314 (1959 publ. 1960)

= Orchis elata f. mairei Soó; Bull. Soc. Hist. Nat. Afrique N. 24: 173 (1933)

= Orchis elata f. maroccanica Soó; Repert. Spec. Nov. Regni Veg. 24: 32 (1927)

= Orchis elata f. pallida Maire & Weiller; Fl. Afrique N. 6: 313 (1959 publ. 1960)

= Orchis elata f. peltieri Soó; Bull. Soc. Hist. Nat. Afrique N. 24: 173 (1933)

= Orchis elata f. speciosissima Soó; Bull. Soc. Hist. Nat. Afrique N. 24: 173 (1933)

= Orchis elata f. willdenowiana Soó; Repert. Spec. Nov. Regni Veg. 24: 31 (1927)

= Orchis elata Poir.; Voy. Barbarie 2: 248 (1789)

= Orchis elata subvar. stephensonii Maire & Weiller; Fl. Afrique N. 6: 343 (1959 publ. 1960)

= Orchis elata var. algerica (Rchb.f.) Maire & Weiller; Fl. Afrique N. 6: 312 (1960)

= Orchis elata var. corsica (E.Rev. ex E.G.Camus) Soó; Repert. Spec. Nov. Regni Veg. 24: 31 (1927)

= Orchis elata var. poiretiana Maire & Weiller; Fl. Afrique N. 6: 312 (1959 publ. 1960)

= Orchis incarnata var. algerica Rchb.f.; Icon. Fl. Germ. Helv. [H.G.L.Reichenbach] 13-14: 53, t. CCCXCVI (1851)

= Orchis incarnata var. elatior E.G.Camus; Monogr. Orchid. 178 (1908)

= Orchis latifolia subsp. elata (Poir.) Maire; Cat. Pl. Maroc 1: 151 (1931)

= Orchis latifolia var. corsica E.Rev. ex E.G.Camus; J. Bot. [Morot] 6: 158 (1892)

= Orchis latifolia var. elata (Poir.) Rchb.f.; Icon. Fl. Germ. Helv. [H.G.L.Reichenbach] 13-14: 59 (1851)

= Orchis latifolia var. elatior Batt.; Fl. Alger [Battandier & Trabut] 196 (1884), nom. illeg.

= Orchis latifolia var. labrovaria Brot.; Phytogr. Lusitan. Select. 2: 25 (1827 publ. 1835)

= Orchis sesquipedalis var. algerica (Rchb.f.) Briq.; Prodr. Fl. Corse 1: 369 (1910)

= Orchis sesquipedalis var. corsica (E.Rev. ex E.G.Camus) Briq.; Prodr. Fl. Corse 1: 369 (1910)

= Orchis vestita Cav.; Anales Ci. Nat. 6: 142 (1803)

variant :

var. atlantica (Kreutz & Vlaciha) comb.ined.

Distribution: S-Morocco (High Atlas)

20 MOR

Synonyms: hidden

= Dactylorhiza elata subsp. atlantica (Kreutz & Vlaciha) Eccarius; Orchideengattung Dactylorhiza 446 (2016)

= Dactylorhiza atlantica Kreutz & Vlaciha; Orchidee (Hamburg) 57(5): 584 (2006)

variant :

var. durandii (Boiss. & Reut.) Landwehr; Wilde Orchid. Eur. 1: 210 (1977)

Distribution: S-Spain; N-Morocco; NW-Algeria

12 SPA 20 ALG MOR

Synonyms: hidden

= Dactylorchis durandii (Boiss. & Reut.) Verm.; Stud. Dactylorch. 68 (1947)

= Dactylorhiza durandii (Boiss. & Reut.) Aver.; Bot. Zhurn. (Moscow & Leningrad) 69(6): 875 (1984), isonym

= Dactylorhiza durandii (Boiss. & Reut.) M.Laínz; Aport. Conoc. Fl. Gallega 7: 31 (1971)

= Dactylorhiza elata subsp. durandii (Boiss. & Reut.) Molero Mesa & Pérez Raya; Fl. Sierra Nevada 365 (1987), isonym

= Dactylorhiza elata subsp. durandii (Boiss. & Reut.) Soó; Nom. Nov. Gen. Dactylorhiza 7 (1962)

= Orchis durandii Boiss. & Reut.; Pugill. Pl. Afr. Bor. Hispan. 111 (1852)

= Orchis elata subsp. durandii (Boiss. & Reut.) Soó; Repert. Spec. Nov. Regni Veg. 24: 32 (1927)

= Orchis elata subvar. durandiana Maire & Weiller; Fl. Afrique N. 6: 313 (1959 publ. 1960)

= Orchis incarnata subsp. durandii (Boiss. & Reut.) E.G.Camus; Monogr. Orchid. 180 (1908)

= Orchis incarnata subsp. durandii (Boiss. & Reut.) Gand.; Nov. Consp. Fl. Eur. 461 (1910), isonym

= Orchis incarnata var. durandii (Boiss. & Reut.) Willk.; Prodr. Fl. Hispan. [Willkomm & Lange] 1(1): 170 (1861)

= Orchis latifolia subsp. durandii (Boiss. & Reut.) Batt.; Fl. Algérie [Battandier et al.] 30 (1895)

= Orchis latifolia subsp. durandii (Boiss. & Reut.) Malag.; Acta Phytotax. Barcinon. 1: 64 (1968), isonym

= Orchis latifolia var. durandii (Boiss. & Reut.) Ball; J. Linn. Soc., Bot. 16: 672 (1878)

= Orchis sesquipedalis var. durandii (Boiss. & Reut.) Briq.; Prodr. Fl. Corse 1: 369 (1910)

variant :

var. elongata (Maire) Maire; Bull. Soc. Hist. Nat. Afrique N. 23: 216 (1932)

Distribution: Morocco

20 MOR

Synonyms: hidden

= Dactylorhiza elata subsp. elongata (Maire) Kreutz, Rebbas & Véla; Ber. Arbeitskreis. Heimische Orchid. 31(2): 165 (2015)

= Dactylorhiza elata var. elongata (Maire) Raynaud; Orchid. Maroc 103 (1985), isonym

= Orchis elata f. elongata (Maire) Soó; Bull. Soc. Hist. Nat. Afrique N. 24: 173 (1933)

= Orchis latifolia var. elongata Maire; Cat. Pl. Maroc 1: 151 (1931)

variant :

var. iberica (T.Stephenson) Soó; Nom. Nov. Gen. Dactylorhiza 7 (1962)

Distribution: NW-France; NW-Spain; N-Portugal

12 FRA POR SPA

Synonyms: hidden

= Orchis sesquipedalis var. iberica T.Stephenson; Bull. Soc. Bot. France 492 (1928)

= Dactylorhiza asturiensis Kreutz; Guide Orchids Europe N. Afr. Middle E. 399 (2024), nom. nov. for O. sesquipedalis var. iberica T. Stephenson

= Dactylorhiza elata subsp. iberica (T.Stephenson) Kreutz; Eurorchis 17: 104 (2005) [basion. illeg.]

variant :

var. kabyliensis (G.Keller) comb.ined.

Distribution: N-Algeria

20 ALG

Remarks: possibly of hybridogenous origin

Synonyms: hidden

= Orchis rassautae (Alleiz.) Alleiz.; Bull. Soc. Bot. France 109: 94 (1962), no exact basionym page.

= xOrchidactyla ×kabyliensis (G.Keller) Aver.; Bot. Zhurn. (Moscow & Leningrad) 71: 93 (1986)

= Orchis kabyliensis G.Keller; Repert. Spec. Nov. Regni Veg. Sonderbeih. A: 2: 260 (1933)

= Orchis kabyliensis var. rassautae Alleiz.; Bull. Soc. Bot. France 101: 115 (1954)

variant :

var. mauritanica (B.Baumann & H.Baumann) comb.ined.

Distribution: Morocco (Middle Atlas)

20 MOR

Synonyms: hidden

= Dactylorhiza elata subsp. mauritanica B.Baumann & H.Baumann; J. Eur. Orch. 37(4): 919 (2005)

= Dactylorhiza mauritanica (B.Baumann & H.Baumann) Kreutz; Guide Orchids Europe N. Afr. Middle E. 396 (2024)

variant :

var. munbyana (Boiss. & Reut.) Soó; Nom. Nov. Gen. Dactylorhiza 7 (1962)

Distribution: N-Algeria; NW-Tunisia

20 ALG TUN

Synonyms: hidden

= Dactylorchis munbyana (Boiss. & Reut.) Verm.; Stud. Dactylorch. 69, t. 3, fig. 3 (1947)

= Dactylorhiza elata subsp. africana (Klinge) H.Sund.; Europ. Medit. Orchid., ed. 3: 40 (1980)

= Dactylorhiza elata subsp. munbyana (Boiss. & Reut.) Kreutz; Ber. Arbeitskreis. Heimische Orchid. 31(2): 161 (2015)

= Dactylorhiza elata subvar. munbyana (Boiss. & Reut.) Raynaud; Orchid. Maroc 103 (1985)

= Dactylorhiza incarnata subsp. africana (Klinge) H.Sund.; Europ. Medit. Orchid., ed. 3: 40 (1980), isonym

= Dactylorhiza incarnata subsp. africana (Klinge) P.D.Sell; Watsonia 6: 317 (1967)

= Dactylorhiza munbyana (Boiss. & Reut.) Aver.; Bot. Zhurn. (Moscow & Leningrad) 69(6): 875 (1984)

= Dactylorhiza munbyana (Boiss. & Reut.) Holub; Folia Geobot. Phytotax. 19(2): 214 (1984)

= Orchis africana (Klinge) A.W.Hill; Index Kew., Suppl. 9: 193 (1938)

= Orchis elata f. munbyana (Boiss. & Reut.) Soó; Repert. Spec. Nov. Regni Veg. 24: 32 (1927)

= Orchis elata subsp. munbyana (Boiss. & Reut.) E.G.Camus; Iconogr. Orchid. Europe 221 (1928)

= Orchis elata var. munbyana (Boiss. & Reut.) Soó; Bull. Soc. Hist. Nat. Afrique N. 24: 173 (1933)

= Orchis incarnata subsp. munbyana (Boiss. & Reut.) E.G.Camus; Monogr. Orchid. 178 (1908)

= Orchis latifolia var. munbyana (Boiss. & Reut.) Batt.; Fl. Alger [Battandier & Trabut] 196 (1884)

= Orchis munbyana Boiss. & Reut.; Pugill. Pl. Afr. Bor. Hispan. 112 (1852)

= Orchis orientalis subsp. africana Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17: 1. 156, in clavi, et 186 (1898)

variant :

var. occitanica (Geniez, Melki, Pain & Soca) P.Delforge; Naturalistes Belges 92(Orchid. 24): 21 (2011)

Distribution: S-France

12 FRA

Synonyms: hidden

= Dactylorhiza elata subsp. ambigua (J.A.Guim.) Kreutz; Kompend. Eur. Orchid. 35 (2004)

= Dactylorhiza elata var. ambigua (J.A.Guim.) Soó; Nom. Nov. Gen. Dactylorhiza 7 (1962)

= Dactylorhiza occitanica Geniez, Melki, Pain & Soca; Orchidophile (Paris) 115: 25 (1995)

= Orchis ambigua Martrin-Donos; Fl. Tarn 705 (1864), nom. illeg.

= Orchis elata subsp. ambigua (J.A.Guim.) Soó; Repert. Spec. Nov. Regni Veg. 24: 31 (1927)

= Orchis elata var. ambigua (Martrin-Donos) Schltr.; Repert. Spec. Nov. Regni Veg. Sonderbeih. A: 24: 181 (1927)

= Orchis incarnata var. ambigua J.A.Guim.; Bol. Soc. Brot. 5: 77 (1887)

= Orchis latifolia var. ambigua (Martrin-Donos) Rouy; Fl. France [Rouy & Foucaud] 13: 151 (1912)

variant :

var. sesquipedalis (Willd.) Landwehr; Wilde Orchid. Eur. 1: 204 (1977)

Distribution: S-France; Spain; N-Portugal; +Corsica

12 COR FRA POR SPA

Synonyms: hidden

= Dactylorchis sesquipedalis (Willd.) Verm.; Stud. Dactylorch. 68 (1947)

= Dactylorhiza elata subsp. sesquipedalis (Willd.) Soó; Nom. Nov. Gen. Dactylorhiza 7 (1962)

= Dactylorhiza majalis subsp. sesquipedalis (Willd.) H.A.Pedersen & Hedrén; Bot. J. Linn. Soc. 168(2): 191 (2012)

= Dactylorhiza sesquipedalis (Willd.) Aver.; Bot. Zhurn. (Moscow & Leningrad) 68(7): 895 (1983), isonym

= Dactylorhiza sesquipedalis (Willd.) M.Laínz; Aport. Conoc. Fl. Gallega 7: 31 (1971), isonym

= Dactylorhiza sesquipedalis (Willd.) Verm.; Bol. Soc. Brot., Sér. 2, 44: 89 (1970)

= Dactylorhiza vestita (Lag. & Rodr.) Aver.; Bot. Zhurn. (Moscow & Leningrad) 71: 92 (1986)

= Orchis elata subsp. sesquipedalis (Willd.) P.Fourn.; Quatre Fl. France 208 (1935), isonym

= Orchis elata subsp. sesquipedalis (Willd.) Soó; Repert. Spec. Nov. Regni Veg. 24: 31 (1928)

= Orchis elata var. sesquipedalis (Willd.) Schltr.; Monogr. Iconogr. Orchid. Eur. 1: 180 (1927)

= Orchis elata var. vestita (Lag. & Rodr.) Losa & Rivas Goday; Arch. Inst. Aclim. Cons. Super. Invest. Ci. 13(1): 79 (1968)

= Orchis incarnata subsp. sesquipedalis (Willd.) E.G.Camus; J. Bot. [Morot] 6: 156 (1892)

= Orchis incarnata subsp. sesquipedalis (Willd.) Gand.; Nov. Consp. Fl. Eur. 461 (1910), isonym

= Orchis incarnata var. sesquipedalis (Willd.) Rchb.f.; Icon. Fl. Germ. Helv. [H.G.L.Reichenbach] 13/14: 53, tab. 48 (1851)

= Orchis latifolia proles sesquipedalis (Willd.) Rouy; Fl. France [Rouy & Foucaud] 13: 151 (1912)

= Orchis latifolia subsp. lusitanica (Steud.) Gand.; Nov. Consp. Fl. Eur. 461 (1910)

= Orchis latifolia subsp. sesquipedalis (Willd.) Maire; Cat. Pl. Maroc 1: 151 (1931)

= Orchis lusitanica Steud.; Nomencl. Bot. [Steudel], ed. 2, 2: 224 (1841)

= Orchis sesquipedalis Willd.; Sp. Pl., ed. 4 [Willdenow] 4: 30 (1805)

= Orchis vestita Lag. & Rodr.; Anales Ci. Nat. 6: 142 (1803)

species :

Dactylorhiza euxina (Nevski) Czerep.; Sosud. Rast. S.S.S.R. 308 (1981)

Name: Schwarzmeer-Knabenkraut (DE)

Distribution: Northern Caucasus; Georgia [Caucasus] (Svaneti, Racha-Lechkhumi, Guria, Adzharia, Shiga Kartli, Kartli, Mtiuleti, Kvemo Kartli, Javakheti); Armenia; Azerbaijan; NE-Turkey

33 ARM AZE GEO NCS 34 TUR

Remarks: incarnata agg.

Synonyms: hidden

= Dactylorhiza cataonica subsp. caucasica (Klinge) Soó; Nom. Nov. Gen. Dactylorhiza 5 (1962) [nom. illeg.], et Ann. Univ. Sci. Budapest. Rolando Eötvös, Sect. Biol., 11: 69 (1969)

= Dactylorhiza caucasica (Klinge) Soó; Nom. Nov. Gen. Dactylorhiza 5 (1962) [nom. inval. illeg.]

= Dactylorhiza euxina (Nevski) H.Baumann & Künkele; Mitt. Arbeitskreises Heimische Orchideen 13(2): 235 (1981), isonym

= Dactylorhiza euxina (Nevski) Soó; [nom. inval.]

= Dactylorhiza majalis subsp. caucasica (Klinge) H.Sund.; Europ. Medit. Orchid., ed. 2: 45 (1975)

= Orchis angustifolia var. caucasica (Klinge) E.G.Camus; Monogr. Orchid. 183 (1908)

= Orchis caucasica (Klinge) Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17 1: 155, in clavi, 181 (1898)

= Orchis caucasica (Klinge) Medw.; Vestn. Tiflissk. Bot. Sada 18: 476 (1919), nom. illeg.

= Orchis cordigera Kraenzl.; Repert. Spec. Nov. Regni Veg., Beih. 65: 39 (1931)

= Orchis cordigera var. caucasica (Klinge) Klinge; V. I. Lipsky, Fl. Cauc. Impr. Colch. 306 (1899)

= Orchis euxina Nevski; Fl. URSS [Komarov et al., eds.] 4: 709 (1935)

= Orchis monticola subsp. caucasica (Lipsky) Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17: 35 (1898)

variant :

var. markowitschii (Soó) Renz & Taubenheim; Notes Roy. Bot. Gard. Edinburgh 41(2): 270 (1983)

Distribution: Northern Caucasus; Georgia [Caucasus]; Armenia; Azerbaijan; NE-Turkey

33 ARM AZE GEO NCS 34 TUR

Synonyms: hidden

= Dactylorhiza caucasica var. markowitschii Soó; Nom. Nov. Gen. Dactylorhiza 5 (1962)

= Dactylorhiza markowitschii (Soó) Aver.; Bot. Zhurn. (Moscow & Leningrad) 68(7): 893 (1983)

= Orchis caucasica var. markowitschii Soó; Notizbl. Bot. Gart. Berlin-Dahlem 9: 909 (1926)

species :

Dactylorhiza foliosa (Rchb.f.) Soó; Nom. Nov. Gen. Dactylorhiza 7 (1962)

Name: Madeira-Knabenkraut

Distribution: Madeira

21 MDR

Remarks: maculata agg.

Synonyms: hidden

= Dactylorchis foliosa (Rchb.f.) Verm.; Stud. Dactylorch. 69 (1947)

= Dactylorchis orientalis subsp. foliosa (Rchb.f.) Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(1): 157, 189 (1898)

= Dactylorhiza incarnata subsp. foliosa (Rchb.f.) H.Sund.; Europ. Medit. Orchid., ed. 2: 45 (1975)

= Orchis foliosa Sol. ex Lowe; Trans. Cambridge Philos. Soc. 4: 13 (1831), nom. illeg.

= Orchis incarnata var. foliosa Rchb.f.; Icon. Fl. Germ. Helv. [H.G.L.Reichenbach] 13-14: 52 (1851)

= Orchis latifolia var. foliosa (Rchb.f.) Rouy; Fl. France [Rouy & Foucaud] 13: 151 (1912), nom. illeg.

= Orchis maderensis Summerh.; Kew Bull. 3: 123 (1947 publ. 1948)

= Orchis orientalis subsp. foliosa (Rchb.f.) Klinge; Trudy Imp. St.-Peterburgsk. Bot. Sada 17(1): 11, 43 (1898)

species :

Dactylorhiza fuchsii (Druce) Soó; Nom. Nov. Gen. Dactylorhiza 8 (1962)

subspecies :

subsp. fuchsii

Name: Fuchs-Fingerwurz, Fuchs-Fingerknabenkraut (DE), Common Spotted-orchid (EN)

Distribution: Iceland; England (England, Wales, Isle of Man, Scotland, Outer Hebrides, Shetland Isl., +Isles of Scilly); Ireland (Ireland, Northern Ireland); Denmark; Norway; Sweden; Finland; Netherlands; Belgium; Luxembourg; Germany (Brandenburg, Berlin, Baden-Württemberg, Bayern, Hessen, Hamburg, Mecklenburg-Vorpommern, Niedersachsen, Rheinland-Pfalz, Schleswig-Holstein, Sachsen, Sachsen-Anhalt); Switzerland; Liechtenstein; Austria; Poland; Czech Republic; Slovakia; Hungary; France; Andorra; ?Spain; Channel Isl. (Alderney, Guernsey, Jersey); Italy; San Marino; Slovenia; Croatia; Bosnia & Hercegovina; Montenegro; Serbia; Kosovo; North Macedonia; Romania; Estonia; Latvia; Lithuania; Belarus; Ukraine; European Russia (N-European Russia, C-European Russia, W-European Russia, E-European Russia); Siberia (Altai, Buryatia, Chita, Irkutsk, Krasnoyarsk, Tuva, W-Siberia, Yakutia); Kazakhstan; China (N-Xinjiang); Mongolia

10 DEN FIN GRB ICE IRE NOR SWE 11 AUT BGM CZE GER HUN LIE LUX NET POL SLO SWI 12 ADO CIS FRA 13 BIH CRO ITA KOS MNT NMA ROM SLV SMA SRB 14 BLR EST LAT LIT RUS UKR 30 SIB 32 KAZ 36 CHI 37 MON

Remarks: maculata agg. Separate from D. maculata based on cytological and genetic grounds. In Central Europa, D. fuchsii is much more abundant than D. maculata, and most records for ´D. maculata´ refer to D. fuchsii

Synonyms: hidden

= Dactylorchis fuchsii (Druce) Verm.; Stud. Dactylorch. 69, 147 (1947)

= Dactylorhiza fuchsii f. alpina Landwehr; Orchideeën 37(3): 78 (1975), et Wilde Orchid. Eur. 2: 556 (1977)

= Dactylorhiza fuchsii f. meyeri (Rchb.f.) P.Delforge; Naturalistes Belges 92: 202 (2011)

= Dactylorhiza fuchsii f. rhodochila (Turner Ettl.) P.Delforge; Naturalistes Belges 88(Orchid. 20): 246 (2007)

= Dactylorhiza fuchsii subsp. meyeri (Rchb.f.) Kulikov & E.G.Philippov; Orchid Memories 107 (2004)

= Dactylorhiza fuchsii var. alpina (Landwehr) R.M.Bateman & Denholm; Watsonia 17(3): 341 (1989)

= Dactylorhiza fuchsii var. longibracteata (Schur) Vöth; Linzer Biol. Beitr. 10(1): 189 (1978)

= Dactylorhiza fuchsii var. magnocalcarata Landwehr; Orchideeën 37(3): 79 (1975), et Wilde Orchid. Eur. 2: 556 (1977)

= Dactylorhiza fuchsii var. meyeri (Rchb.f.) Soó; Nom. Nov. Gen. Dactylorhiza 8 (1962)

= Dactylorhiza fuchsii var. rhodochila Turner Ettl.; Watsonia 18: 308 (1991)

= Dactylorhiza fuchsii var. tatrensis Borsos ex Soó; Ann. Univ. Sci. Budapest. Rolando Eötvös, Sect. Biol., 3: 351 (1960)

= Dactylorhiza fuchsii var. trilobata (Bréb.) P.D.Sell; Fl. Gr. Brit. Ireland 5: 364 (1996)

= Dactylorhiza longibracteata (F.W.Schmidt) Holub; Folia Geobot. Phytotax. 18(2): 204 (1983) [p. p.]

= Dactylorhiza maculata subsp. austriaca Vöth; Linzer Biol. Beitr. 10(1): 190 (1978)

= Dactylorhiza maculata subsp. fuchsii (Druce) Hyl.; Nordisk Kärlväxtfl. 2: 238 (1966) [nom. inval.]

= Dactylorhiza maculata subsp. meyeri (Rchb.f.) Tournay; Fl. Belg. Nord France Rég. Vois. 691 (1967)

= Dactylorhiza meyeri (Rchb.f.) Aver.; Bot. Zhurn. (Moscow & Leningrad) 67(3): 307 (1982)

= Orchis comosa F.W.Schmidt; Samml. Phys.-Oekon. Aufsätze 1: 233 (1795), nom. illeg.

= Orchis fuchsii Druce; Rep. Bot. Soc. Exch. Club Brit. Isles 4: 105 (1914 publ. 1915)

= Orchis fuchsii f. lancefolia Snarskis; Lietuvos T.S.R. Fl. 2: 679 (1963)

= Orchis fuchsii f. maxima Snarskis; Lietuvos T.S.R. Fl. 2: 679 (1963)

= Orchis longibracteata F.W.Schmidt; Samml. Phys.-Oekon. Aufsätze 1: 233. t. 2 (1795) [would have priority, but conservation of D. fuchsii proposed]

= Orchis maculata f. alba Zapal.; Consp. Fl. Galic. Crit. 216 (1906)

= Orchis maculata f. minor Zapal.; Consp. Fl. Galic. Crit. 211 (1906)

= Orchis maculata f. paluster E.G.Camus; Monogr. Orchid. 50 (1908)

= Orchis maculata f. recurvifolia Brenner; Acta Soc. Fauna Fl. Fenn. 34(4): 10 (1911)

= Orchis maculata f. trilobata (Bréb.) Soó; Bot. Arch. 23: 78 (1928)

= Orchis maculata subsp. meyeri (Rchb.f.) E.G.Camus & A.Camus; isonym

= Orchis maculata subsp. meyeri (Rchb.f.) K.Richt.; Pl. Eur. 1: 272 (1890)

= Orchis maculata subsp. meyeri (Rchb.) O.Bolòs & Vigo; Fl. Països Catalans 4: 629 (2001), isonym

= Orchis maculata var. curvifolia E.G.Camus; Monogr. Orchid. 194 (1908)

= Orchis maculata var. elongata Gadeceau; Mem. Soc. Sci. Nat. Ouest 2: 10 (1892)

= Orchis maculata var. media Bréb.; 313 (1836)

= Orchis maculata var. meyeri Rchb.f.; Icon. Fl. Germ. Helv. [H.G.L.Reichenbach] 13-14: 67 (1851)

= Orchis maculata var. slandzinskii Zapal.; Consp. Fl. Galic. Crit. 211 (1906)

= Orchis maculata var. submontana Zapal.; Consp. Fl. Galic. Crit. 211 (1906)

= Orchis maculata var. triloba E.G.Camus; Monogr. Orchid. 50 (1908)

= Orchis maculata var. trilobata Bréb.; Fl. Normandie, éd. 5: 387 (1879)

= Orchis mascula var. triloba Bréb.; Fl. Normandie, éd. 5: 387 (1879)

variant :

var. maculatensis Lachmair; Ber. Arbeitskreis. Heimische Orchid. 40(2): 84 (2024)

Distribution: Austria

11 AUT

variant :

var. sacciferoides (Klinge) Landwehr; Orchideeën 31: 78 (1975), et Wilde Orchid. Eur. 1: 114 (1977)

Distribution: Netherlands; Germany; etc.

11 GER NET

Synonyms: hidden

= Dactylorhiza maculata var. sacciferoides (Klinge ex Schulze) Soó; Nom. Nov. Gen. Dactylorhiza 7 (1962)

= Orchis maculata f. sacciferoides (Klinge ex Schulze) Soó; Bot. Arch. 23: 76 (1928)

= Orchis maculata var. sacciferoides Klinge ex Schultze; Mitth. Thüring. Bot. Vereins 4 (1902)

variant :

var. punicea (J.Essink, L.Essink & Kreutz) Kreutz; Guide Orchids Europe N. Afr. Middle E. 440 (2024)

Distribution: Norway

10 NOR

Synonyms: hidden

= Dactylorhiza fuchsii subsp. punicea J.Essink, L.Essink & Kreutz; Ber. Arbeitskreis. Heimische Orchid. 24(1): 79 (2007)

= Dactylorhiza andoeyana Perko; Ber. Arbeitskreis. Heimische Orchid. 24(1): 40 (2007)

= Dactylorhiza fuchsii f. andoeyana (Perko) P.Delforge; Naturalistes Belges 89(Orchid. 21): 250 (2008)

= Dactylorhiza fuchsii var. punicea (J.Essink, L.Essink & Kreutz) Kreutz; Guide Orchids Europe N. Afr. Middle E. 440 (2024)

variant :

var. sooana (Borsos) Kreutz; Guide Orchids Europe N. Afr. Middle E. 439 (2024)

Distribution: Poland; Czech Republic; Slovakia; Hungary; W-Ukraine

11 CZE HUN POL SLO 14 UKR

Conservation Status: IUCN V

Synonyms: hidden

= Dactylorhiza fuchsii subsp. sooana (Borsos) Borsos; Ann. Univ. Sci. Budapest. Rolando Eötvös, Sect. Biol. 4: 76 (1961)

= Dactylorhiza fuchsii subsp. sooana (Borsos) Soó; Nom. Nov. Gen. Dactylorhiza 8 (1962), isonym

= Dactylorhiza maculata subsp. sooana Borsos ex Batou?ek; Pl. Syst. Evol. 307(4)-51: 16 (2021)

subspecies :

subsp. carpatica (Batousek & Kreutz) Kreutz; Kompend. Eur. Orchid. 38 (2004)

Distribution: Czech Repubplic; Slovakia (Bile Karpaty); Romania

11 SLO 13 ROM

Synonyms: hidden

= Dactylorhiza carpatica (Batousek & Kreutz) P.Delforge; Naturalistes Belges 81(4): 396 (2000)

= Dactylorhiza traunsteineri subsp. carpatica Batousek & Kreutz; J. Eur. Orch. 31(3): 684, 683 (1999)

subspecies :

subsp. hebridensis (Wilmott) Soó; Nom. Nov. Gen. Dactylorhiza 8 (1962)

Distribution: England (Scotland, Outer Hebrides); Ireland (NW-Ireland)

10 GRB IRE

Remarks: frequently treated as bona sp.

Synonyms: hidden

= Dactylorchis fuchsii subsp. hebridensis (Wilmott) Hesl.-Harr.; Ber. Geobot. Inst. ETH Stiftung Rübel 1953: 55 (1954)

= Dactylorchis maculata subsp. hebridensis (Wilmott) Verm.

= Dactylorhiza fuchsii var. cornubiensis (Pugsley) Soó; Nom. Nov. Gen. Dactylorhiza 8 (1962)

= Dactylorhiza fuchsii var. hebridensis (Wilmott) R.M.Bateman & Denholm; Watsonia 17(3): 340 (1989)

= Dactylorhiza hebridensis (Wilmott) Aver.; Bot. Zhurn. (Moscow & Leningrad) 71(1): 92 (1986), et Spis. Rast. Gerb. Fl. S.S.S.R. 25 (127130): 26 (1986), et Novosti Sist. Vyssh. Rast. 26: 50 (1989)

= Dactylorhiza maculata subsp. hebridensis (Wilmott) E.Nelson; Monogr. & Ikonogr. Orchid.-Gattung Dactylorhiza 102 (1976) [without basionym page]

= Dactylorhiza maculata subsp. hebridensis (Wilmott) H.Baumann & Künkele; Mitt. Arbeitskreises Heimische Orchideen 20(3): 631 (1988)

= Orchis fuchsii subsp. hebridensis (Wilmott) Clapham; Fl. Brit. Isles [Clapham et al.] 1317 (1952)

= Orchis hebridensis Wilmott; J. Bot. 77: 192 (1939)

= Orchis maculata var. cornubiensis Pugsley; J. Bot. 78: 178 (1940)

subspecies :

subsp. okellyi (Druce) Soó; Nom. Nov. Gen. Dactylorhiza 8 (1962)

Distribution: England (Scotland); Ireland (W-Ireland, Northern Ireland)

10 GRB IRE

Synonyms: hidden

= Dactylorchis fuchsii subsp. okellyi (Druce) Verm.; Stud. Dactylorch. 69 (1947)

= Dactylorhiza fuchsii var. okellyi (Druce) R.M.Bateman & Denholm; Watsonia 17(3): 342 (1989)

= Dactylorhiza maculata subsp. okellyi (Druce) H.Baumann & Künkele; Mitt. Arbeitskreises Heimische Orchideen 20(3): 632 (1988)

= Dactylorhiza okellyi (Druce) Aver.; Bot. Zhurn. (Moscow & Leningrad) 69(6): 875 (1984)

= Orchis maculata subsp. okellyi (Druce) Druce; Bot. Pocket-book ed. 15: 277 (1917)

= Orchis maculata var. okellyi Druce; Irish Naturalist 1909: 211 (1909)

= Orchis okellyi (Druce) Druce; Rep. Bot. Soc. Exch. Club Brit. Isles 4: 108 (1914 publ. 1915)

subspecies :

subsp. psychrophila (Schltr.) Holub; Preslia 36: 252 (1964)

Distribution: Norway; Sweden; Finland; N-European Russia (Kola peninsula)

10 FIN NOR SWE 14 RUS

Synonyms: hidden

= Dactylorchis fuchsii subsp. psychrophila (Schltr.) Verm.; Stud. Dactylorch. 69 (1947)

= Dactylorhiza fuchsii var. psychrophila (Schltr.) B.Bock; Bull. Soc. Bot. Centre-Ouest 43: 214 (2012)

= Dactylorhiza fuchsii var. psychrophila (Schltr.) Soó; Nom. Nov. Gen. Dactylorhiza 8 (1962)

= Dactylorhiza maculata subsp. psychrophila (Schltr.) Soó; Nom. Nov. Gen. Dactylorhiza 7 (1962)

= Dactylorhiza maculata var. psychrophila (Schltr.) Vöth; Linzer Biol. Beitr. 12(2): 429 (1980)

= Dactylorhiza psychrophila (Schltr.) Aver.; Bot. Zhurn. (Moscow & Leningrad) 67(3): 308 (1982)

= Orchis maculata var. psychrophila Schltr.; Monogr. Iconogr. Orchid. Eur. 1: 183 (1927)

subspecies :

subsp. sudetica (Poech ex Rchb.f.) Verm.; Orchideeën 37(3): 78 (1975), et Wilde Orchid. Eur. 1: 106 (1977)

Distribution: Germany (Sachsen); ?Austria; Poland; Czech Republic; European Russia

11 CZE GER POL 14 RUS

Remarks: possibly a synonym of ssp. psychrophila or a nothosubspecies fuchsii - psychrophila

Synonyms: hidden

= Dactylorchis fuchsii subsp. sudetica (Poech ex Rchb.f.) Verm.; Wilde Orchid. Eur. 1: 106 (1977)

= Dactylorchis maculata var. sudetica (Poech ex Rchb.f.) Verm.; Ned. Kruidk. Arch. 56: 236 (1949)

= Dactylorhiza fuchsii var. sudetica (Poech ex Rchb.f.) H.Baumann, Künkele & R.Lorenz; J. Eur. Orch. 36: 771 (2004)

= Dactylorhiza longibracteata subsp. sudetica (Poech ex Rchb.f.) Holub; Folia Geobot. Phytotax. 18: 205 (1983)

= Dactylorhiza maculata subsp. sudetica (Poech ex Rchb.f.) Vöth; Linzer Biol. Beitr. 12(2): 430 (1980)

= Dactylorhiza maculata var. sudetica (Poech ex Rchb.f.) Baumann, Künkele & R.Lorenz; J. Eur. Orch. 34(1): 144 (2002), isonym

= Dactylorhiza sudetica (Poech ex Rchb.f.) Aver.; Bot. Zhurn. (Moscow & Leningrad) 67(3): 310 (1982)

= Orchis maculata f. sudetica (Poech ex Rchb.f.) Neuman; Sver. Fl. 631 (1901)

= Orchis maculata subsp. sudetica (Poech ex Rchb.f.) K.Richt.; Pl. Eur. 1: 272 (1890)

= Orchis maculata var. sudetica Poech ex Rchb.f.; Icon. Fl. Germ. Helv. [H.G.L.Reichenbach] 13-14: 66 (1850)

= Orchis maculata var. sudetica Rchb.f.; Icon. Fl. Germ. Helv. [H.G.L.Reichenbach] 13-14: 66 (1851)

species :

Dactylorhiza hatagirea (D.Don) Soó; Ann. Univ. Sci. Budapest. Rolando Eötvös, Sect. Biol. 3: 341 (1960), et Nom. Nov. Gen. Dactylorhiza 4 (1962)

Distribution: China (Gansu, Heilongjiang, Jilin, Nei Mongol, Ningxia, Qinghai, W-Sichuan, Xinjiang); E-Tibet; Mongolia; Pakistan (Gilgit, Karakorum, Kagan, Astor, Hazara, Swat); Nepal; Bhutan; India (Himachal Pradesh, Jammu & Kashmir, Sikkim, Uttarakhand)

36 CHI TIB 37 MON 40 BHU IND NEP PAK

Synonyms: hidden

= Dactylorhiza graggeriana (Soó) Soó; Nom. Nov. Gen. Dactylorhiza 4 (1962)

= Dactylorhiza hatagirea var. doniana (Soó) Soó; Nom. Nov. Gen. Dactylorhiza 4 (1962)

= Dactylorhiza hatagirea var. schlaginweitii (Soó) Soó; Nom. Nov. Gen. Dactylorhiza 4 (1962)

= Dactylorhiza umbrosa var. afghanica (Soó) Soó; Nom. Nov. Gen. Dactylorhiza 4 (1962)

= Orchis graggeriana Soó; J. Bot. 66: 15 (1928)

= Orchis hatagirea D.Don; Prodr. Fl. Nepal. 23 (1825)

= Orchis hatagirea var. afghanica Soó; J. Bot. 66: 19 (1928)

= Orchis hatagirea var. doniana Soó; J. Bot. 66: 19 (1928)

= Orchis hatagirea var. schlaginweitii Soó; J. Bot. 66: 19 (1928)

= Orchis latifolia var. indica Lindl.; Gen. Sp. Orchid. Pl. 260 (1835)

species :

Dactylorhiza iberica (M.Bieb. ex Willd.) Soó; Nom. Nov. Gen. Dactylorhiza 3 (1962)

Distribution: Greece (mountains of NW-, C-Greece, N-Peloponnisos, NW-Evvoia); European Turkey; Crimea; Northern Caucasus; Georgia [Caucasus] (Shiga Kartli, Kartli); Armenia; Azerbaijan; Turkey; NE-Iraq; Iran; Cyprus; Lebanon; Syria

13 GRC TUE 14 KRY 33 ARM AZE GEO NCS 34 CYP IRN IRQ LEB SYR TUR

Remarks: maculata agg.

Synonyms: hidden

= Dactylorchis iberica (M.Bieb. ex Willd.) Verm.; Stud. Dactylorch. 65 (1947)

= Gymnadenia angustifolia (M.Bieb.) Spreng.; Syst. Veg., ed. 16 [Sprengel] 3: 693 (1826)

= Orchis angustifolia M.Bieb.; Fl. Taur.-Caucas. 2: 368 (1808)

= Orchis angustifolia var. longifolia (K.Koch) Rchb.f.; Icon. Fl. Germ. Helv. [H.G.L.Reichenbach] 13-14: 34 (1851)

= Orchis angustifolia var. stevenii Rchb.f.; Icon. Fl. Germ. Helv. [H.G.L.Reichenbach] 13-14: 34 (1851)

= Orchis iberica f. leptophylla E.G.Camus; Monogr. Orchid. 165 (1908)

= Orchis iberica M.Bieb. ex Willd.; Sp. Pl., ed. 4 [Willdenow] 4: 25 (1805)

= Orchis iberica var. longifolia (K.Koch) Rchb.f.; Icon. Fl. Germ. Helv. [H.G.L.Reichenbach] 13-14: 34 (1851)

= Orchis iberica var. stevenii Rchb.f.; Icon. Fl. Germ. Helv. [H.G.L.Reichenbach] 13-14: 34 (1851)

= Orchis leptophylla K.Koch; Linnaea 22: 282 (1850)

= Orchis leptophylla var. laxa K.Koch; Linnaea 22: 282 (1850)

= Orchis leptophylla var. longifolia K.Koch; Linnaea 22: 282 (1850)

= Orchis natolica Fisch. & C.A.Mey.; Ann. Sci. Nat., Bot., Sér. 4, 1: 30 (1854)

species :

Dactylorhiza incarnata (L.) Soó; Nom. Nov. Gen. Dactylorhiza 3 (1962)

subspecies :

subsp. incarnata

Name: Fleischrote Fingerwurz (DE), Early Marsh-orchid (EN)

Distribution: Faeroe Isl.; England (England, Wales, Isle of Man, Scotland, Outer Hebrides, Orkney Isl., Shetland Isl.); Ireland (Ireland, Northern Ireland); Denmark; Norway; Sweden; Finland; Netherlands; Belgium; Luxembourg; Germany (Brandenburg, Berlin, Baden-Württemberg, Bayern, Hessen, Hamburg, Mecklenburg-Vorpommern, Niedersachsen, Nordrhein-Westfalen, Rheinland-Pfalz, Schleswig-Holstein, Saarland, Sachsen, Sachsen-Anhalt, Thüringen); Switzerland; Liechtenstein; Austria; Poland; Czech Republic; Slovakia; Hungary; +Portugal; Spain; Andorra; France; Channel Isl.; Italy; Slovenia; Croatia; Bosnia & Hercegovina; Montenegro; Serbia; Kosovo; North Macedonia; Albania; Romania; Bulgaria; N-Greece (Rodopi, Mt. Falakron, very rare NW-Greece); Estonia; Latvia; Lithuania; Belarus; Moldova; Ukraine; Crimea; European Russia (N-European Russia, C-European Russia, W-European Russia, E-European Russia); Siberia (Altai, Buryatia, Irkutsk, Krasnoyarsk, W-Siberia, Yakutia); Russian Far East (Khabarovsk); Kazakhstan; Kyrgyzstan; ?Northern Caucasus; N-Turkey; Mongolia

10 DEN FIN FOR GRB IRE NOR SWE 11 AUT BGM CZE GER HUN LIE LUX NET POL SLO SWI 12 ADO CIS FRA POR SPA 13 ALB BIH BUL CRO GRC ITA KOS MNT NMA ROM SLV SRB 14 BLR EST KRY LAT LIT MLD RUS UKR 30 SIB 31 RFE 32 KAZ KGZ 34 TUR 37 MON

Remarks: incarnata agg.

Synonyms: hidden

= Dactylorchis incarnata (L.) Verm.; Stud. Dactylorch. 65, 101 (1947)

= Dactylorchis incarnata f. multiangulata Verm.; Ned. Kruidk. Arch. 56: 210 (1949)

= Dactylorchis incarnata f. punctata Verm.; Ned. Kruidk. Arch. 56: 209 (1949)

= Dactylorchis incarnata var. angustispicata Verm.; Ned. Kruidk. Arch. 56: 209 (1949)

= Dactylorchis incarnata var. beckii Verm.; Ned. Kruidk. Arch. 56: 211 (1949)

= Dactylorchis incarnata var. brachyphylla Verm.; Ned. Kruidk. Arch. 56: 207 (1949)

= Dactylorchis incarnata var. brevicalcarata (Rchb.f.) Verm.; Ned. Kruidk. Arch. 56: 210 (1949)

= Dactylorchis incarnata var. divaricata (Rich. ex Loisel.) Verm.; Ned. Kruidk. Arch. 56: 208 (1949)

= Dactylorchis incarnata var. leucantha Verm.; Ned. Kruidk. Arch. 56: 212 (1949)

= Dactylorchis incarnata var. longibracteata Verm.; Ned. Kruidk. Arch. 56: 209 (1949)

= Dactylorchis incarnata var. noncircumscripta Verm.; Ned. Kruidk. Arch. 56: 211 (1949)

= Dactylorchis incarnata var. remota Verm.; Ned. Kruidk. Arch. 56: 207 (1949)

= Dactylorchis incarnata var. rhombeilabia Verm.; Ned. Kruidk. Arch. 56: 209 (1949)

= Dactylorchis incarnata var. sublatifolia (Rchb.f.) Verm.; Ned. Kruidk. Arch. 56: 205 (1949)

= Dactylorchis incarnata var. triangularis Verm.; Ned. Kruidk. Arch. 56: 210 (1949)

= Dactylorchis incarnata var. vulgaris Verm.; Ned. Kruidk. Arch. 56: 206 (1949)

= Dactylorhiza comosa (Scop.) P.D.Sell; Fl. Gr. Brit. Ireland 5: 364 (1996)

= Dactylorhiza incarnata lusus albida (Rchb.f.) Soó; Acta Bot. Acad. Sci. Hung. 16: 368 (1970 publ. 1971), nom. inval.

= Dactylorhiza incarnata f. ochrantha Landwehr; Orchideeën 37: 80 (1975)

= Dactylorhiza incarnata f. punctata (Verm.) R.M.Bateman & Denholm; Watsonia 15: 346 (1985)

= Dactylorhiza incarnata subsp. punctata R.Doll; Feddes Repert. 89(56): 346 (1978)

= Dactylorhiza incarnata subsp. pyrenaica Balayer; Bull. Soc. Bot. France, Lett. Bot. 133(3): 280 (1986)

= Dactylorhiza incarnata var. brachyphylla (Verm.) Soó; Nom. Nov. Gen. Dactylorhiza 3 (1962)

= Dactylorhiza incarnata var. macrophylla (Schur) Soó; Nom. Nov. Gen. Dactylorhiza 3 (1962)

= Dactylorhiza incarnata var. seemenii (Asch. & Graebn.) Soó; Nom. Nov. Gen. Dactylorhiza 3 (1962)

= Dactylorhiza incarnata var. sublatifolia (Rchb.f.) Soó; Nom. Nov. Gen. Dactylorhiza 3 (1962)

= Dactylorhiza incarnata var. trifurca (Rchb.f.) Aver.; Konsp. Sosud. Rast. Zakaz. Mshinskoe Boloto Leningrad. Oblasti 38 (1988)

= Dactylorhiza intermedia (Serg.) Kulikov & E.G.Philippov; Byull. Moskovsk. Obshch. Isp. Prír., Otd. Biol., n. s. 104(2): 31 (1999)

= Dactylorhiza strictifolia (Opiz) Rauschert ex Hudziok; Wiss. Z. Martin-Luther-Univ. Halle-Wittenberg, Math.-Naturwiss. Reihe 14: 492 (1965)

= Orchis altaica (Rchb.f.) Soó; Repert. Spec. Nov. Regni Veg. Sonderbeih. A: 2: 214 (1933)

= Orchis angustifolia Loisel. ex Rchb.; Iconogr. Bot. Pl. Crit. 9: 17 (1831), nom. inval.

= Orchis angustifolia var. divaricata Nyman; Consp. Fl. Eur. 692 (1882)

= Orchis angustifolia var. fraasii Rchb.f.; Icon. Fl. Germ. Helv. [H.G.L.Reichenbach] 13-14: 34 (1851)

= Orchis angustifolia Wimm. & Grab.; Fl. Siles. 2(1): 252 (1829), nom. illeg.

= Orchis comosa Scop.; Fl. Carniol., ed. 2, 2: 198 (1771)

= Orchis comosa var. angustifolia Ambrosi; Fl. Tirolo Mer. 1: 794 (1854)

= Orchis condensa Schur; Enum. Pl. Transsilv. 642 (1866)

= Orchis iberica var. fraasii Rchb.f.; Icon. Fl. Germ. Helv. [H.G.L.Reichenbach] 13-14: 34 (1851)

= Orchis incarnata f. ampla Höppner; Verh. Naturhist. Vereines Preuss. Rheinl. Westphalens 81: 268 (1924 publ. 1925)

= Orchis incarnata f. brevicalcarata (Rchb.f.) Seeland; 51 (1940)

= Orchis incarnata f. elatior Serg.; Sist. Zametki Mater. Gerb. Krylova Tomsk. Gosud. Univ. Kuybysheva 1: 4 (1929), nom. illeg.

= Orchis incarnata f. lanceata (A.Dietr.) Snarskis; Lietuvos T.S.R. Fl. 2: 679 (1963)

= Orchis incarnata f. macrophylla (Asch. & Graebn.) Seeland; 87 (1929)

= Orchis incarnata f. macrophylla Schur; Enum. Pl. Transsilv. 641 (1866)

= Orchis incarnata f. praecox Engenst.; Allg. Bot. Z. Syst. 18: 110 (1912)

= Orchis incarnata f. retusa Soó; Bot. Arch. 23: 66 (1928)

= Orchis incarnata f. rostriformis (Zapal.) Soó; Bot. Arch. 23: 65 (1928)

= Orchis incarnata f. stenophylla (Asch. & Graebn.) Seeland; 87 (1929)

= Orchis incarnata f. stenophylla Asch. & Graebn.; Syn. Mitteleur. Fl. [Ascherson & Graebner] 3: 717 (1907), nom. illeg.

= Orchis incarnata f. subfoliosa M.Schulze ex Asch. & Graebn.; Syn. Mitteleur. Fl. [Ascherson & Graebner] 3: 718 (1907)

= Orchis incarnata L.; Fl. Suec., ed. 2 [Linnaeus] 312 (1755)

= Orchis incarnata subsp. fraasii (Rchb.f.) K.Richt.; Pl. Eur. 1: 270 (1890)

= Orchis incarnata subsp. fraasii K.Richt.; Pl. Eur. 1: 270 (1890)

= Orchis incarnata subsp. lanceata (A.Dietr.) E.G.Camus; Monogr. Orchid. 177 (1908)

= Orchis incarnata subsp. lanceata (A.Dietr.) Gsell; 8 (1936)

= Orchis incarnata var. albida Rchb.f.; Icon. Fl. Germ. Helv. [H.G.L.Reichenbach] 13-14: 183 (1851), nom. inval.

= Orchis incarnata var. altaica Rchb.f.; Icon. Fl. Germ. Helv. [H.G.L.Reichenbach] 13-14: 53 (1851)

= Orchis incarnata var. angustifolia Blytt; Enum. Pl. Vasc. 54 (1844)

= Orchis incarnata var. brevicalcarata Rchb.f.; Icon. Fl. Germ. Helv. [H.G.L.Reichenbach] 13-14: 53 (1851)

= Orchis incarnata var. brevifolia A.Fuchs; Ber. Naturwiss. Vereins Schwaben Neuburg 42: 69 (1919)

= Orchis incarnata var. divaricata (Rich. ex Loisel.) St.-Lag.; Étude Fl. [A.Cariot], éd. 8, 2: 806 (1889)

= Orchis incarnata var. fraasii (K.Richt.) Soó; Bot. Arch. 23: 66 (1928)

= Orchis incarnata var. incisiloba Zapal.; Consp. Fl. Galic. Crit. 207 (1906)

= Orchis incarnata var. intermedia Serg.; Sist. Zametki Mater. Gerb. Krylova Tomsk. Gosud. Univ. Kuybysheva 3: 685 (1929)

= Orchis incarnata var. lanceata (A.Dietr.) Rchb.f.; Icon. Fl. Germ. Helv. [H.G.L.Reichenbach] 13-14: 51 (1851)

= Orchis incarnata var. lanceata-fraasii Rchb.f.; Icon. Fl. Germ. Helv. [H.G.L.Reichenbach] 13-14: 52 (1851)

= Orchis incarnata var. lanceolata Rchb.f.; Icon. Fl. Germ. Helv. [H.G.L.Reichenbach] 13-14: 176 (1851)

= Orchis incarnata var. latifolia (L.) Blytt; Enum. Pl. Vasc. 54 (1844)

= Orchis incarnata var. maxima Cuatrec.; Treb. Inst. Catalana Hist. Nat. 12: 253 (1929)

= Orchis incarnata var. pallida Brenner; Florist. Handb. 185 (1886)

= Orchis incarnata var. platyphylla Snarskis; Lietuvos T.S.R. Fl. 2: 679 (1963)

= Orchis incarnata var. rhombeilabia-acroglossa Rchb.f.; Icon. Fl. Germ. Helv. [H.G.L.Reichenbach] 13-14: 53 (1851)

= Orchis incarnata var. rostriformis Zapal.; Consp. Fl. Galic. Crit. 207 (1906)

= Orchis incarnata var. seemenii Asch. & Graebn.; Syn. Mitteleur. Fl. [Ascherson & Graebner] 3: 721 (1907)

= Orchis incarnata var. subextensa Neuman; Bot. Not. 1909: 159 (1909)

= Orchis incarnata var. sublatifolia Rchb.f.; Icon. Fl. Germ. Helv. [H.G.L.Reichenbach] 13-14: 51 (1851)

= Orchis incarnata var. trifurca Rchb.f.; Icon. Fl. Germ. Helv. [H.G.L.Reichenbach] 13-14: 53 (1851)

= Orchis incarnata var. triloba-retusa Rchb.f.; Icon. Fl. Germ. Helv. [H.G.L.Reichenbach] 13-14: 53 (1851)

= Orchis lanceata A.Dietr.; Fl. Boruss. t. 5 (1833)

= Orchis latifolia L.; Sp. Pl. [Linnaeus] 2: 941 (1753), nom. rej.

= Orchis latifolia subsp. impudica Soó; Repert. Spec. Nov. Regni Veg. 24: 30 (1927), nom. inval.

= Orchis latifolia subsp. incarnata (L.) Corb.; Nouv. Fl. Normandie 557 (1894), isonym

= Orchis latifolia subsp. incarnata (L.) Hook.f.; Students Fl. Brit. Isl. 353 (1870)

= Orchis latifolia var. angustifolia Bab.; Man. Brit. Bot. [Babington] 291 (1843), nom. illeg.

= Orchis latifolia var. angustifolia Wimm. & Grab. ex Döll; Rhein. Fl. 225 (1843)

= Orchis latifolia var. cambrica Pugsley; Bot. J. Linn. Soc. 49: 579 (1935)

= Orchis latifolia var. comosa Harz; Fl. Deutschl. [Schlechtendal], ed. 5, Neuer Abdruck, 4: 242 (1895 publ. 1896)

= Orchis latifolia var. divaricata (Rich. ex Loisel.) Nyman; Consp. Fl. Eur. 692 (1882)

= Orchis latifolia var. ecalcarata Lej.; Comp. Fl. Belg. 3: 185 (1836), nom. superfl.

= Orchis latifolia var. hocquartiana Dumort.; Fl. Belg. [Dumortier] 132 (1827)

= Orchis latifolia var. incarnata (L.) Bab.; 310 (1847)

= Orchis latifolia var. incarnata (L.) Coss. & Germ.; Fl. Descr. Anal. Paris, éd. 2: 684 (1861)

= Orchis latifolia var. lanceata Brügger; Jahresber. Naturf. Ges. Graubündens 17: 58 (1874), nom. inval.

= Orchis latifolia var. lineare Tinant; Fl. Luxemb. 441 (1836)

= Orchis latifolia var. longibracteata Neilr.; Fl. Wien 129 (1846)

= Orchis latifolia var. maculata Tinant; Fl. Luxemb. 441 (1836)

= Orchis latifolia var. nemorosa Westerl.; Bot. Not. 1863: 147 (1863)

= Orchis latifolia var. pallida Tinant; Fl. Luxemb. 441 (1836)

= Orchis latifolia var. rubra Gray; Nat. Arr. Brit. Pl. 2: 202 (1821 publ. 1822)

= Orchis latifolia var. tharandina Rchb.f.; Icon. Fl. Germ. Helv. [H.G.L.Reichenbach] 13-14: 58 (1851)

= Orchis macra Schur; Enum. Pl. Transsilv. 641 (1866), nom. illeg.

= Orchis maculata subsp. incarnata (L.) Douin; Fl. Ill. France 11: 30 (1931)

= Orchis matodes Rchb.f.; Icon. Fl. Germ. Helv. [H.G.L.Reichenbach] 13: t. 398 (1851)

= Orchis militaris Puccin. ex Parl.; Fl. Ital. [Parlatore] 3: 521 (1858)

= Orchis mixta Retz.; Fl. Scand. Prodr. 167 (1779), nom. superfl.

= Orchis mixta var. incantata (L.) Retz.; Fl. Scand. Prodr. 167 (1779)

= Orchis palmata Gilib.; Exerc. Phyt. 2: 479 (1792), opus utique oppr.

= Orchis palmata subsp. incarnata (L.) Syme; 100 (1869), nom. illeg.

= Orchis richardii Tratt.; Freye Auswahl Pflanzenabbild.: no. 517 (1816)

= Orchis sambucina var. incarnata (L.) Hampe; 55 (1836)

= Orchis strictifolia Opiz; Naturalientausch 10: 217 (1825)

= Orchis tharandina Rchb.f.; Icon. Fl. Germ. Helv. [H.G.L.Reichenbach] 13-14: t. 403 (1851)

variant :

var. baumgartneriana (B.Baumann, H.Baumann, R.Lorenz & Ruedi Peter) P.Delforge; Naturalistes Belges 85 (Orchid. 17): 250 (2004)

Distribution: Azerbaijan

33 AZE

Synonyms: hidden

= Dactylorhiza baumgartneriana (B.Baumann, H.Baumann, R.Lorenz & Ruedi Peter) Kreutz; Guide Orchids Europe N. Afr. Middle E. 357 (2024)

= Dactylorhiza incarnata subsp. baumgartneriana B.Baumann, H.Baumann, R.Lorenz & Ruedi Peter; J. Eur. Orch. 35(1): 165 (2003)

= Dactylorhiza kafiriana subsp. baumgartneriana (B.Baumann, H.Baumann, R.Lorenz & Ruedi Peter) Sczep. & Kreutz; Ber. Arbeitskreis. Heimische Orchid. 24(2): 103 (2007)

variant :

var. borealis (Neuman) Hyl.; Nordisk Kärlväxtfl. 2: 387 (1966)

Distribution: Norway; Sweden; Finland; Estonia; N-European Russia

10 FIN NOR SWE 14 EST RUS

Synonyms: hidden

= Orchis incarnata var. borealis Neuman; Bot. Not. 1909: 229 (1909)

variant :

var. cungsii (Kreutz) comb.ined.

Distribution: Luxembourg (Differdange)

11 LUX

Synonyms: hidden

= Dactylorhiza incarnata subsp. cungsii Kreutz; Ber. Arbeitskreis. Heimische Orchid. 32(2): 158

= Dactylorhiza cungsii (Kreutz) Kreutz; Guide Orchids Europe N. Afr. Middle E. 350 (2024)

variant :

var. dunensis (Druce) Hyl.; Nordisk Kärlväxtfl. 2: 387 (1966)

Distribution: England (England, Scotland); Ireland; Denmark; Norway; Netherlands; Belgium; N-Germany (Borkum)

10 DEN GRB IRE NOR 11 BGM GER NET

Synonyms: hidden

= Dactylorchis incarnata f. dunensis (Druce) Verm.; Ned. Kruidk. Arch. 56: 207 (1949)

= Dactylorchis incarnata var. dunensis (Druce) Verm.; Ned. Kruidk. Arch. 56: 207 (1949)

= Orchis incarnata var. dunensis Druce; Rep. Bot. Soc. Exch. Club Brit. Isles 4: 212 (1916)

variant :

var. haematodes (Rchb.) Verm.; Ned. Kruidk. Arch. 56: 208 (1949)

Distribution: Netherlands; Germany; Hungary; Austria; Switzerland; France; etc.

11 AUT GER HUN NET SWI 12 FRA

Synonyms: hidden

= Dactylorchis incarnata var. extensa (Hartm.) Verm.; Ned. Kruidk. Arch. 56: 208 (1949)

= Dactylorchis incarnata var. haematodes (Rchb.) Verm.; Stud. Dactylorch. 117 (1947)

= Dactylorhiza haematodes (Rchb.) G.H.Loos; Jahrb. Bochum. Bot. Vereins 1: 120 (2010)

= Dactylorhiza incarnata f. haematodes (Rchb.) P.Delforge; Naturalistes Belges 87(Orchids. 19): 259 (2006)

= Dactylorhiza incarnata subsp. haematodes (Rchb.) Soó; Nom. Nov. Gen. Dactylorhiza 4 (1962)

= Dactylorhiza incarnata var. extensa (Hartm.) Soó; Nom. Nov. Gen. Dactylorhiza 3 (1962)

= Dactylorhiza incarnata var. haematodes (Rchb.) Soó; Magyar Fl. Veg., Rnd. Kézik. 171 (1973), isonym

= Dactylorhiza incarnata var. reichenbachii Gathoye & D.Tyteca; Lejeunia 143: 51 (1994)

= Orchis angustifolia var. extensa Hartm.; Handb. Skand. Fl., ed. 4: 281 (1843)

= Orchis cruenta var. haematodes (Rchb.) Neuman; Bot. Not. 1909: 157 (1909)

= Orchis extensa (Hartm.) Pritz.; Icon. Bot. Index [Pritzel] 2: 768 (1855)

= Orchis haematodes Rchb.f.; Fl. Germ. Excurs. 126 (1830), nom. illeg.

= Orchis incarnata lusus haematodes (Rchb.) Soó; Bot. Arch. 23: 66 (1928)

= Orchis incarnata f. haematodes (Rchb.) M.Schulze; Syn. Mitteleur. Fl. [Ascherson & Graebner] 3: 717 (1907)

= Orchis incarnata var. extensa (Hartm.) Rchb.f.; Icon. Fl. Germ. Helv. [H.G.L.Reichenbach] 13-14: 51 (1851)

= Orchis incarnata var. haematodes (Rchb.f.) M.Schultze; Syn. Mitteleur. Fl. [Ascherson & Graebner] 3(5): 717 (1907)

= Orchis incarnata var. haematodes (Rchb.f.) Pauca; Fl. Reipubl. Popul. Roman. (Fl. Republ. Socialist. Romania) 12: 692 (1972), isonym

= Orchis incarnata var. haematodes (Rchb.) Nyman; Consp. Fl. Eur. 692 (1882)

= Orchis strictifolia var. extensa (Hartm.) Verm.; Hyl., Uppsala Univ. Årsskr. 7: 117 (1945)

variant :

var. hyphaematodes (Neuman) Løjtnant; Feddes Repert. 89(1): 13 (1978)

Distribution: Ireland; Denmark; Sweden; Norway; Finland; N-Germany; Austria; Switzerland; N-Italy; Slovenia; Czech Republic; Slovakia; Bulgaria; Romania; Estonia

10 DEN FIN IRE NOR SWE 11 AUT CZE GER SLO SWI 13 BUL ITA ROM SLV 14 EST

Synonyms: hidden

= Dactylorhiza incarnata subsp. hyphaematodes (Neuman) Landwehr; Orchideeën 37: 80 (1975)

= Orchis incarnata var. hyphaematodes Neuman; Bot. Not. 1909: 244 (1909)

variant :

var. immaculata Romolini & Sodi; GIROS Notiz. 25: 21 (2004)

Distribution: Italy

13 ITA

variant :

var. jugicrucis (Akhalk., R.Lorenz & Mosul.) P.Delforge; Naturalistes Belges 96(Orchid. 28): 15 (2015)

Distribution: Georgia [Caucasus] (Guria, Adzharia, Kartli, Meskheti)

33 GEO

Synonyms: hidden

= Dactylorhiza incarnata subsp. jugicrucis Akhalk., R.Lorenz & Mosul.; Jahresber. Naturwiss. Vereins Wuppertal 12: 36 (2007)

= Dactylorhiza jugicrucis (Akhalk., R.Lorenz & Mosul.) Kreutz; Guide Orchids Europe N. Afr. Middle E. 349 (2024)

variant :

var. latissima (Zapal.) Verm.; Ned. Kruidk. Arch. 56: 205 (1949)

Distribution: S-Sweden; Netherlands; Belgium; France; Germany; Switzerland; Austria; N-Italy; Czech Republic; Slovakia; Hungary; Denmark; England; Ireland; Poland; Belarus; Estonia; Crimea; Moldova; Turkey; etc.

10 DEN GRB IRE SWE 11 AUT BGM CZE GER HUN NET POL SLO SWI 12 FRA 13 ITA 14 BLR EST KRY MLD 34 TUR

Synonyms: hidden

= Dactylorhiza incarnata var. latissima (Zapal.) Hyl.; Nordisk Kärlväxtfl. 2: 387 (1966)

= Orchis incarnata var. latissima Zapal.; Consp. Fl. Galic. Crit. 207 (1906)

= Dactylorhiza gemmana (Pugsley) Aver.; Bot. Zhurn. (Moscow & Leningrad) 69(6): 875 (1984)

= Dactylorhiza incarnata subsp. gemmana (Pugsley) Hesl.-Harr.; Ber. Geobot. Inst. ETH Stiftung Rübel 1953: 55 (1954)

= Dactylorhiza incarnata subsp. gemmana (Pugsley) P.D.Sell; Watsonia 6: 317 (1967), isonym

= Orchis latifolia var. gemmana Pugsley; J. Linn. Soc., Bot. 49: 578 (1935)

variant :

var. lobelii Verm.; Ned. Kruidk. Arch. 56: 206 (1949)

Distribution: England (Wales); Norway; Netherlands; Belgium

10 GRB NOR 11 BGM NET

Remarks: often included in var. dunensis

Synonyms: hidden

= Dactylorhiza incarnata subsp. lobelii (Verm.) H.A.Pedersen; Nordic J. Bot. 21(2): 184 (2001)

= Dactylorhiza incarnata var. lobelii (Verm.) Soó; Nom. Nov. Gen. Dactylorhiza 3 (1962)

= Dactylorhiza lobelii (Verm.) Kreutz; Guide Orchids Europe N. Afr. Middle E. 352 (2024)

subspecies :

subsp. coccinea (Pugsley) Soó; Nom. Nov. Gen. Dactylorhiza 4 (1962)

Distribution: England (N-England, Wales, Scotland, Outer Hebrides, Shetland Isl.); Ireland (Ireland, Northern Ireland)

10 GRB IRE

Synonyms: hidden

= Dactylorchis incarnata subsp. coccinea (Pugsley) Hesl.-Harr.; Ber. Geobot. Inst. ETH Stiftung Rübel 1953: 54 (1954)

= Dactylorhiza coccinea (Pugsley) Aver.; Bot. Zhurn. (Moscow & Leningrad) 69(6): 875 (1984)

= Orchis incarnata f. atrirubra Godfery; Monogr. Icon. Brit. Nat. Orchid. 187 (1933)

= Orchis latifolia var. coccinea Pugsley; Bot. J. Linn. Soc. 49: 579 (1935)

= Orchis strictifolia subsp. coccinea (Pugsley) Clapham; Fl. Brit. Isles [Clapham et al.] 1319 (1952) [nom. inval.]

subspecies :

subsp. cruenta (O.F.Müll.) P.D.Sell; Watsonia 6: 317 (1967)

Name: Blutrote Fingerwurz, Blutrotes Fingerknabenkraut (DE)

Distribution: England (N-Scotland, Orkney Isl.); Ireland (NW- & W-Ireland); +Denmark; Norway; Sweden; Finland; Slovakia; France; Italy; Croatia; Bosnia & Hercegovina; Estonia; Latvia; Lithuania; +Belarus; Siberia (Altai, Buryatia, Irkutsk, Krasnoyarsk, Tuva, W-Siberia, Yakutia); China (NW-Xinjiang)

10 DEN FIN GRB IRE NOR SWE 11 SLO 12 FRA 13 BIH CRO ITA 14 BLR EST LAT LIT 30 SIB 36 CHI

Synonyms: hidden

= Dactylorchis cruenta (O.F.Müll.) Verm.; Stud. Dactylorch. 66, 121 (1947)

= Dactylorchis incarnata subsp. cruenta (O.F.Müll.) Verm.; Stud. Dactylorch. 66, 124 (1947)

= Dactylorhiza cruenta (O.F.Müll.) Soó; Nom. Nov. Gen. Dactylorhiza 4 (1962)

= Dactylorhiza cruenta f. ochrantha Wief.; Wilde Orchid. Eur. 2: 557 (1977)

= Dactylorhiza cruenta var. lanceolata (Neuman) Landwehr; 80 (1975)

= Dactylorhiza incarnata var. cruenta (O.F.Müll.) Hyl.; Nordisk Kärlväxtfl. 2: 387 (1966) [nom. inval.]

= Orchis angustifolia subsp. cruenta (O.F.Müll.) C.Hartm.; Handb. Skand. Fl., ed. 5: 225 (1849)

= Orchis cruenta O.F.Müll.; Fl. Dan. [Oeder et al.] t. 876 (1782)

= Orchis cruenta var. lanceolata Neuman; Bot. Not. 1909: 157 (1909)

= Orchis cruenta var. subelliptica Neuman; Bot. Not. 1909: 151 (1909)

= Orchis cruenta var. subtriangularis Neuman; Bot. Not. 1909: 156 (1909)

= Orchis cruentiformis Neuman; Bot. Not. 243 (1909)

= Orchis incarnata f. cruenta (O.F.Müll.) Cajander; Melan. Suom. Kasvio [Cajander], ed. 5: 171 (1906)

= Orchis incarnata subsp. cruenta (O.F.Müll.) Asch. & Graebn.; Syn. Mitteleur. Fl. [Ascherson & Graebner] 3(5): 561 (1907), isonym

= Orchis incarnata subsp. cruenta (O.F.Müll.) Hegi; 352 (1909), isonym

= Orchis incarnata subsp. cruenta (O.F.Müll.) Nyman; Consp. Fl. Eur. 692 (1882)

= Orchis incarnata var. rhombeilabia-cruenta Rchb.f.; Icon. Fl. Germ. Helv. [H.G.L.Reichenbach] 13-14: 53 (1851)

= Orchis latifolia var. cruenta (O.F.Müll.) Lindl.; Gen. Sp. Orchid. Pl. 260 (1835)

= Orchis latifolia var. subincarnata Rchb.f.; Icon. Fl. Germ. Helv. [H.G.L.Reichenbach] 13-14: 59 (1851)

subspecies :

subsp. ochroleuca (Wüstnei ex Boll) P.F.Hunt & Summerh.; Watsonia 6: 130 (1965)

Name: Hellgelbe Fingerwurz, Blassgelbes Fingerknabenkraut (DE)

Distribution: England (E-England: Suffolk, +Cambridgshire); Denmark; Sweden; Germany (Brandenburg, Baden-Württemberg, Bayern, Mecklenburg-Vorpommern, +Niedersachsen, +Schleswig-Holstein, +Sachsen-Anhalt); Switzerland; Austria; Poland; Slovakia; Hungary; Croatia; Romania; Estonia; Latvia; Lithuania; Belarus; European Russia

10 DEN GRB SWE 11 AUT GER HUN POL SLO SWI 13 CRO ROM 14 BLR EST LAT LIT RUS

Synonyms: hidden

= Dactylorhiza incarnata subsp. ochroleuca (Wüstnei ex Boll) Hesl.-Harr.f.; Ber. Geobot. Inst. ETH Stiftung Rübel 1953: 55 (1954), nom. inval. (basionymo non cit.)

= Dactylorhiza incarnata var. ochroleuca (Wüstnei ex Boll) Hyl.; Nordisk Kärlväxtfl. 2: 387 (1966)

= Dactylorhiza incarnata var. straminea (Rchb.f.) Soó; Nom. Nov. Gen. Dactylorhiza 3 (1962)

= Dactylorhiza ochroleuca (Wüstnei ex Boll) Aver.; Bot. Zhurn. (Moscow & Leningrad) 68(7): 893 (1983), isonym

= Dactylorhiza ochroleuca (Wüstnei ex Boll) Holub; Folia Geobot. Phytotax. 9(3): 272 (1974)

= Orchis incarnata subsp. ochroleuca (Wüstnei ex Boll) Hegi; Ill. Fl. Mitt.-Eur. [Hegi] 2: 352 (1909)

= Orchis incarnata subsp. ochroleuca (Wüstnei ex Boll) O.Schwarz; Mitt. Thüring. Bot. Ges. 1: 94 (1949), isonym

= Orchis incarnata var. ochroleuca Wüstnei ex Boll; Arch. Vereins Freunde Naturgesch. Mecklenburg 14: 307 (1860)

= Orchis incarnata var. straminea Rchb.f.; Icon. Fl. Germ. Helv. [H.G.L.Reichenbach] 13-14: 183 (1851)

= Orchis latifolia var. ochroleuca (Wüstnei ex Boll) Pugsley; J. Linn. Soc., Bot. 49: 578 (1935)

= Orchis strictifolia var. ochroleuca (Wüstnei ex Boll) Nannf.; Festschrift The Svedberg 601-622 (1944)

= Orchis traunsteineri subsp. ochroleuca (Wüstnei ex Boll) O.Schwarz; Mitt. Thüring. Bot. Ges., n. s. 1: 94 (1949)

subspecies :

subsp. pulchella (Druce) Hesl.-Harr.f.; Ber. Geobot. Inst. ETH Stiftung Rübel 1953: 55 (1954)

Distribution: England (England, Wales, Scotland, Outer Hebrides, Shetland Isl.); Ireland (Ireland, Northern Ireland)

10 GRB IRE

Synonyms: hidden

= Dactylorhiza incarnata subsp. pulchella (Druce) Soó; Nom. Nov. Gen. Dactylorhiza 4 (1962)

= Dactylorhiza majalis var. pulchella (Druce) R.M.Bateman & Denholm; Watsonia 14: 370 (1983)

= Dactylorhiza pulchella (Druce) Aver.; Bot. Zhurn. (Moscow & Leningrad) 69(6): 875 (1984)

= Dactylorhiza purpurella var. pulchella (Druce) Soó; Nom. Nov. Gen. Dactylorhiza 5 (1962)

= Orchis incarnata var. pulchella Druce; Rep. Bot. Soc. Exch. Club Brit. Isles 5: 167 (1918)

= Orchis latifolia var. pulchella (Druce) Pugsley; J. Linn. Soc., Bot. 49: 587 (1935)

= Orchis praetermissa var. pulchella Druce; Rep. Bot. Soc. Exch. Club Brit. Isles 5: 577 (1920)

= Orchis purpurella var. pulchella (Druce) Pugsley; Bot. J. Linn. Soc. 49: 583 (1935)

= Orchis strictifolia var. pulchella (Druce) Clapham; Fl. Brit. Isles [Clapham et al.] 1319 (1952)

subspecies :

subsp. serotina (Hausskn. ex M.Schulze) Soó & D.M.Moore; Bot. J. Linn. Soc. 76: 367 (1978)

Name: Späte Steifblättrige Fingerwurz, Spätes Steifblättriges Knabenkraut (DE)

Distribution: Denmark; Norway; Sweden; Germany; Switzerland; Austria; Czech Republic; Hungary; ?Spain; France; European Russia

10 DEN NOR SWE 11 AUT CZE GER HUN SWI 12 FRA 14 RUS

Synonyms: hidden

= Dactylorchis incarnata var. serotina (Hausskn. ex M.Schulze) Verm.; Stud. Dactylorch. 162 (1947)

= Dactylorhiza incarnata f. brevibracteata (Landwehr) P.Delforge; Naturalistes Belges 87(Orchid. 19): 258 (2006)

= Dactylorhiza incarnata var. brevibracteata Landwehr; Orchideeën 37(3): 78 (1975), et Wilde Orchid. Eur. 2: 556 (1977)

= Dactylorhiza incarnata var. haussknechtii (Klinge) Buttler; Bot. Naturschutz Hessen 17: 108 (2004)

= Dactylorhiza incarnata var. serotina (Hausskn. ex M.Schulze) Soó; Nom. Nov. Gen. Dactylorhiza 3 (1962)

= Dactylorhiza serotina (Hausskn. ex M.Schulze) G.H.Loos; Jahrb. Bochum. Bot. Vereins 1: 120 (2010), with incorrect basionym ref.

= Orchis angustifolia var. haussknechtii Klinge; Revis. Orchis cordigera & O. angustifolia 70 (1893)

= Orchis incarnata f. serotina (A.F.Schwarz) M.Goldschm.; 156 (1912)

= Orchis incarnata subsp. serotina (A.F.Schwarz) Hegi; 352 (1909)

= Orchis incarnata var. haussknechtii (Klinge) O.Schwarz; Mitt. Thüring. Bot. Ges. 1: 94 (1949)

= Orchis incarnata var. serotina Hausskn. & M.Schultze; Orch. Deutschl. Österr. Schweiz Nr. 19, 4 (1894), nom. illeg.

= Orchis serotina (Hausskn. ex M.Schulze) A.F.Schwarz; Fl. Nürnberg-Erlangen [Schwarz] 765 (1901)

= Orchis traunsteineri var. serotina Hausskn. ex M.Schulze; Orch. Deutschl. Österr. Schweiz 4 (1894)

subspecies :

subsp. versicolor (J.C.Schmidt ex Lüscher) Potucek; Klic. Urc. Ceskosl. Vstavac. Pardubice 29 (1969) [nothosubsp.]

Distribution: Germany; Austria

11 AUT GER

Remarks: Dactylorhiza incarnata ssp. incarnata x D. incarnata ssp. ochroleuca

Synonyms: hidden

= Dactylorhiza incarnata nothovar. versicolor (J.C.Schmidt ex Lüscher) Soó; Nom. Nov. Gen. Dactylorhiza 4 (1962)

= Dactylorhiza ×versicolor (Schmidt & Lüscher) Gathoye & D.Tyteca; Lejeunia 143: 60 (1994)

= Orchis incarnata lusus pulchella Höppner; Verh. Naturhist. Vereines Preuss. Rheinl. Westphalens 81: 270 (1924 publ. 1925)

= Orchis incarnata nothovar. versicolor J.C.Schmidt ex Lüscher; Fl. Aargau 150 (1918)

= Orchis ×pedemontana Gsell; Jahresber. Naturf. Ges. Graubündens, N. F. 74: 16 (1936 publ. 1936)

species :

Dactylorhiza insularis (Sommier) Landwehr; Orchidee (Hamburg) 20(3): 128 (1969)

Name: Insel-Knabenkraut

Distribution: Portugal (Estremadura); Spain; France (Aude, Corbières); Corsica; Sardinia; Italy (Toscana, Emilia Romagna, Elba); N-Morocco (Rif)

12 COR FRA POR SAR SPA 13 ITA 20 MOR

Synonyms: hidden

= Dactylorhiza iberica subsp. insularis (Sommier) Soó; Nom. Nov. Gen. Dactylorhiza 3 (1962)

= Dactylorhiza insularis (Sommier ex Martelli) E.Nelson; Monogr. & Ikonogr. Orchid.-Gattung Dactylorhiza 104 (1976), isonym

= Dactylorhiza insularis (Sommier) Ó.Sánchez & Herrero; Fl. Iber. [Castroviejo et al.] 21: 98 (2005), isonym

= Dactylorhiza insularis f. castellana (Rivas Goday) Bernardos; Belg. J. Bot. 135(1-2): 81 (2002)

= Dactylorhiza insularis subsp. castellana (Rivas Goday) D.Rivera & López Vélez; Orquid. Prov. Albacete 82 (1987)

= Dactylorhiza sambucina subsp. insularis (Sommier ex Martelli) Soó; Nom. Nov. Gen. Dactylorhiza 3 (1962)

= Orchis insularis Sommier; Bull. Soc. Bot. Ital. 1895: 248 (1895)

= Orchis insularis Sommier ex Martelli; Monocot. Sardoae, Fasc. 1, 58 (1896)

= Orchis pseudosambucina subsp. castellana Rivas Goday; Farmacognosia 6: 193 (1945)

= Orchis romana var. castellana Rivas Goday; Farmacognosia 6: 193, 197 (1945) [nom. altern.]

= Orchis romana var. insularis (Sommier) E.G.Camus; Monogr. Orchid. 172 (1908)

= Orchis sambucina proles insularis (Sommier) Rouy; Fl. France [Rouy & Foucaud] 13: 155 (1912)

= Orchis sambucina subsp. insularis (Sommier) Briq.; Prodr. Fl. Corse 1: 371 (1910)

= Orchis sambucina var. castellana Rivas Goday; Farmacognosia 6: 193, 197 (1945) [nom. altern.]

= Orchis sambucina var. insularis (Sommier) Fiori; Fl. Anal. Italia 1: 245 (1896)

= Orchis sulphurea subsp. castellana (Rivas Goday) Rivas Goday; Anales Jard. Bot. Madrid 21(2): 306 (1963)

form :

f. bartonii (Huxley & P.F.Hunt) Gathoye & D.Tyteca; Lejeunia 143: 52 (1994)

Distribution: Spain; France; Corsica

12 COR FRA SPA

Synonyms: hidden

= Dactylorhiza bartonii (Huxley & P.F.Hunt) Aver.; Bot. Zhurn. (Moscow & Leningrad) 69(6): 876 (1984)

= Dactylorhiza insularis var. bartonii (Huxley & P.F.Hunt) D.Rivera & Lopez Velez; Orquíd. Prov. Albacete 82 (1987), isonym

= Dactylorhiza insularis var. bartonii (Huxley & P.F.Hunt) Landwehr; Orchidee (Hamburg) 20(3): 128 (1969)

= Dactylorhiza romana subsp. bartonii Huxley & P.F.Hunt; J. Roy. Hort. Soc. 92: 309 (1967), et Jahresber. Naturwiss. Vereins Wuppertal 21/22: 97 (1968)

species :

Dactylorhiza kafiriana Renz; Fl. Iranica [Rechinger] 26: 125 (1978)

Distribution: Afghanistan (Badakshan, Wakhan); N-Pakistan (Chitral, Hunza, Gilgit, Karakorum); India (Jammu & Kashmir)

34 AFG 40 IND PAK

species :

Dactylorhiza kerryensis (Wilmott) P.F.Hunt & Summerh.; Watsonia 6: 131 (1965), nom. cons.

Distribution: Ireland (W-Ireland, Northern Ireland)

10 IRE

Remarks: for the status as full species see Bateman & Denholm 2011. A stabilized hybridogenous species derived from D. maculata and D. incarnata.

Synonyms: hidden

= Dactylorchis kerryensis (Wilmott) Verm.; Stud. Dactylorch. 67 (1947)

= Dactylorhiza comosa f. kerryensis (Roberts) P.D.Sell; Fl. Gr. Brit. Ireland 5: 365 (1996)

= Dactylorhiza latifolia var. kerryensis (Wilmott) Soó; Nom. Nov. Gen. Dactylorhiza 4 (1962)

= Dactylorhiza majalis subsp. kerryensis (Wilmott) Senghas; Jahresber. Naturwiss. Vereins Wuppertal 21/22: 53 (1968)

= Dactylorhiza majalis var. kerryensis (Wilmott) R.M.Bateman & Denholm; Watsonia 14(4): 368 (1983)

= Dactylorhiza occidentalis var. kerryensis (Wilmott) R.M.Bateman & Denholm; Watsonia 27(3): 247 (2009)

= Orchis kerryensis Wilmott; Proc. Linn. Soc. London Sess. 148, 126 (1936)

= Orchis occidentalis subsp. kerryensis (Wilmott) Clapham; Fl. Brit. Isles [Clapham et al.] 1321 (1952), nom. illeg.

variant :

var. occidentalis (Pugsley) Jebb; Watsonia 27(4): 371 (2009)

Distribution: Ireland (SW- & W-Ireland)

10 IRE

Synonyms: hidden

= Dactylorchis majalis subsp. occidentalis (Pugsley) Hesl.-Harr.f.; Ber. Geobot. Inst. ETH Stiftung Rübel 1953: 55 (1954)

= Dactylorchis occidentalis (Pugsley) Verm.; Stud. Dactylorch. 67 (1947)

= Dactylorhiza comosa f. occidentalis (Pugsley) P.D.Sell; Fl. Gr. Brit. Ireland 5: 365 (1996)

= Dactylorhiza comosa subsp. occidentalis (Pugsley) P.D.Sell; Fl. Gr. Brit. Ireland 5: 364 (1996)

= Dactylorhiza kerryensis subsp. occidentalis (Pugsley) H.Baumann & Künkele; Mitt. Arbeitskreises Heimische Orchideen 20(3): 629 (1988)

= Dactylorhiza latifolia subsp. occidentalis (Pugsley) Soó; Nom. Nov. Gen. Dactylorhiza 4 (1962)

= Dactylorhiza majalis subsp. occidentalis (Pugsley) E.Nelson; Monogr. & Ikonogr. Orchid.-Gattung Dactylorhiza 90 (1976), isonym

= Dactylorhiza majalis subsp. occidentalis (Pugsley) P.D.Sell; Acta Fac. Rerum Nat. Univ. Comen., Bot. 14: 19 (1968)

= Dactylorhiza majalis var. occidentalis (Pugsley) R.M.Bateman & Denholm; Watsonia 14(4): 368 (1983)

= Dactylorhiza occidentalis (Pugsley) P.Delforge; Naturalistes Belges 72(3): 100 (1991)

= Orchis majalis var. occidentalis Pugsley; Bot. J. Linn. Soc. 49: 586 (1935)

= Orchis occidentalis (Pugsley) Wilmott; Rep. Bot. Soc. Exch. Club Brit. Isles 10: 458, 551 (1937 publ. 1938), nom. rej.

species :

Dactylorhiza lapponica (Laest. ex Hartm.) Soó; Nom. Nov. Gen. Dactylorhiza 5 (1962)

Name: Lappland-Fingerwurz, Lappland-Fingerknabenkraut (DE)

Distribution: Norway; Sweden; Finland; Estonia; N-European Russia; ?Siberia (?W-Siberia)

10 FIN NOR SWE 14 EST RUS

Remarks: majalis agg. Distribution of synonyms between this species and D. traunsteineri is chaotic

Synonyms: hidden

= Dactylorchis lapponica (Laest. ex Hartm.) Verm.; Stud. Dactylorch. 67 (1947)

= Dactylorhiza angustifolia subsp. pycnantha (Neuman) Aver.; Bot. Not. 1909: 232 (1909)

= Dactylorhiza cruenta subsp. lapponica (Laest. ex Rchb.f.) E.Nelson; Monogr. & Ikonogr. Orchid.-Gattung Dactylorhiza 72 (1976) [comb. inval.]

= Dactylorhiza majalis subsp. lapponica (Laest. ex Hartm.) H.Sund.; Europ. Medit. Orchid., ed. 2: 45 (1975)

= Dactylorhiza pycnantha (Neuman) Aver.; Bot. Zhurn. (Moscow & Leningrad) 68(9): 1163 (1983)

= Dactylorhiza traunsteineri subsp. lapponica (Laest. ex Hartm.) Soó; Bot. J. Linn. Soc. 76(4): 367 (1978)

= Dactylorhiza traunsteineri subsp. pycnantha (Neuman) Soó; Nom. Nov. Gen. Dactylorhiza 6 (1962), isonym

= Dactylorhiza traunsteineri subsp. pycnantha (Neuman) Verm.; Stud. Dactylorch. 66 (1947)

= Dactylorhiza traunsteineri var. blyttii (Rchb.f.) Soó; Nom. Nov. Gen. Dactylorhiza 5 (1962)

= Dactylorhiza traunsteineri var. nylandri (Klinge) Soó; Nom. Nov. Gen. Dactylorhiza 5 (1962)

= Dactylorhiza traunsteineri var. recurva (Klinge) Soó; Nom. Nov. Gen. Dactylorhiza 6 (1962)

= Dactylorhiza traunsteineri var. tarbatonica (Klinge) Soó; Nom. Nov. Gen. Dactylorhiza 6 (1962)

= Orchis angustifolia subsp. pycnantha Neuman; Bot. Not. 1909: 232 (1909)

= Orchis angustifolia var. lapponica Laest. ex Hartm.; Handb. Skand. Fl., ed. 4: 281 (1843)

= Orchis angustifolia var. nylandri Klinge; Revis. Orchis cordigera & O. angustifolia 67 (1893)

= Orchis angustifolia var. sauteri (Klinge) Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 75 (1899)

= Orchis blyttii Soó; Bot. Arch. 23: 160 (1928), nomen

= Orchis cordigera subsp. blyttii (Rchb.f.) K.Richt.; Pl. Eur. 1: 271 (1890)

= Orchis cordigera var. blyttii Rchb.f.; Icon. Fl. Germ. Helv. [H.G.L.Reichenbach] 13/14: 60 (1851)

= Orchis cruenta subsp. blyttii (Rchb.f.) Soó; Repert. Spec. Nov. Regni Veg. 24: 30 (1927)

= Orchis lapponica (Laest. ex Hartm.) K.Richt.; Pl. Eur. 1: 271 (1890)

= Orchis latifolia subsp. lapponica (Laest. ex Rchb.f.) Soó; Repert. Spec. Nov. Regni Veg. 24: 25 (1927)

= Orchis latifolia var. blyttii Rchb.f.; Icon. Fl. Germ. Helv. [H.G.L.Reichenbach] 13-14: 60 (1851)

= Orchis latifolia var. lapponica (Laest. ex Hartm.) Rchb.f.; Icon. Fl. Germ. Helv. [H.G.L.Reichenbach] 13-14: 58 (1851)

= Orchis maculata subsp. lapponica (Laest. ex Hartm.) Nyman; Consp. Fl. Eur. 693 (1882)

= Orchis maculata var. recurva (F.Nyl. ex Fr.) Nyman; Consp. Fl. Eur. 693 (1882)

= Orchis sambucina var. subalpina Rchb.f.; Icon. Fl. Germ. Helv. [H.G.L.Reichenbach] 13-14: 52 (1851)

= Orchis traunsteineri var. lapponica Hartm. ex Rchb.f.; Icon. Fl. Germ. Helv. [H.G.L.Reichenbach] 13/14: 77 (1851)

species :

Dactylorhiza maculata (L.) Soó; Nom. Nov. Gen. Dactylorhiza 7 (1962)

subspecies :

subsp. maculata

Name: Gefleckte Fingerwurz, Geflecktes Fingerknabenkraut (DE)

Distribution: Denmark; Norway; Sweden; Finland; Netherlands; Belgium; Luxembourg; Germany (Niedersachsen, Nordrhein-Westfalen, ?Rheinland-Pfalz); Switzerland; Austria; Poland; Czech Republic; Slovakia; Hungary; Portugal; Spain; ?Baleares; France; Italy; Slovenia; Croatia; Bosnia & Hercegovina; Montenegro; Serbia; Kosovo; North Macedonia; Romania; Estonia; Latvia; Lithuania; Belarus; ?Moldova; Ukraine; European Russia (N-European Russia, W-European Russia, C-European Russia); Siberia (Krasnoyarsk, W-Siberia)

10 DEN FIN NOR SWE 11 AUT BGM CZE GER HUN LUX NET POL SLO SWI 12 FRA POR SPA 13 BIH CRO ITA KOS MNT NMA ROM SLV SRB 14 BLR EST LAT LIT RUS UKR 30 SIB

Remarks: maculata agg. Distinct from D. fuchsii and in Central Europe much less frequent

Synonyms: hidden

= Dactylorchis maculata (L.) Verm.; Stud. Dactylorch. 68, 130 (1947)

= Dactylorchis maculata var. angustiflora Verm.; Ned. Kruidk. Arch. 56: 235 (1949)

= Dactylorchis maculata var. compacta Verm.; Ned. Kruidk. Arch. 56: 232 (1949)

= Dactylorchis maculata var. concolor Verm.; Ned. Kruidk. Arch. 56: 232 (1949)

= Dactylorchis maculata var. densiflora Verm.; Ned. Kruidk. Arch. 56: 233 (1949)

= Dactylorchis maculata var. grandiflora Verm.; Ned. Kruidk. Arch. 56: 233 (1949)

= Dactylorchis maculata var. hunyadensis Borsos & Soó; Acta Bot. Acad. Sci. Hung. 5: 324 (1959)

= Dactylorchis maculata var. ibericoides Verm.; Ned. Kruidk. Arch. 56: 235 (1949)

= Dactylorchis maculata var. immaculata (Schur) Verm.; Ned. Kruidk. Arch. 56: 233 (1949)

= Dactylorchis maculata var. laxiflora Verm.; Ned. Kruidk. Arch. 56: 233 (1949)

= Dactylorchis maculata var. lineata Verm.; Ned. Kruidk. Arch. 56: 232 (1949)

= Dactylorchis maculata var. ovalifolia (Beck) Verm.; Ned. Kruidk. Arch. 56: 232 (1949)

= Dactylorchis maculata var. spathulata Verm.; Ned. Kruidk. Arch. 56: 232 (1949)

= Dactylorchis maculata var. traunsteineriifolius Verm.; Ned. Kruidk. Arch. 56: 231 (1949)

= Dactylorchis maculata var. vespertilio (Lindm.) Verm.; Ned. Kruidk. Arch. 56: 234 (1949)

= Dactylorchis maculata var. vexillum (Lindm.) Verm.; Ned. Kruidk. Arch. 56: 234 (1949)

= Dactylorchis maculata var. vulgaris Verm.; Ned. Kruidk. Arch. 56: 236 (1949)

= Dactylorhiza maculata lusus albicans (Marchand ex Tinant) Soó; Acta Bot. Acad. Sci. Hung. 16: 368 (1970 publ. 1971)

= Dactylorhiza maculata lusus carnea (Tinant) Soó; Acta Bot. Acad. Sci. Hung. 16: 368 (1970 publ. 1971)

= Dactylorhiza maculata f. angustiflora (Verm.) Soó; Acta Bot. Acad. Sci. Hung. 16: 368 (1970 publ. 1971)

= Dactylorhiza maculata f. brevicalcarata (Rohlena) Soó; Acta Bot. Acad. Sci. Hung. 16: 368 (1970 publ. 1971)

= Dactylorhiza maculata var. hunyadensis (Borsos & Soó) Soó; Nom. Nov. Gen. Dactylorhiza 8 (1962)

= Dactylorhiza maculata var. olgae Soó; Ann. Univ. Sci. Budapest. Rolando Eötvös, Sect. Biol., 3: 349 (1960)

= Dactylorhiza maculata var. pseudoelata Balayer; Bull. Soc. Bot. France, Lett. Bot. 133(3): 280 (1986)

= Dactylorhiza maculata var. submacrostachys Borsos ex Soó; Ann. Univ. Sci. Budapest. Rolando Eötvös, Sect. Biol., 3: 348 (1960)

= Dactylorhiza maculata var. traunsteineriifolius (Harz) Soó; Nom. Nov. Gen. Dactylorhiza 7 (1962)

= Orchis basilica L. ex Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(1): 190 (1898), nom. superfl.

= Orchis basilica subsp. maculata (L.) Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(1): 192 (1898), nom. superfl.

= Orchis biermannii Ortmann; Bot. Zeitung (Berlin) 8: 738 (1850)

= Orchis calvellii A.Terracc.; Annuario Reale Ist. Bot. Roma 4: 47 (1889-1890 publ. 1891)

= Orchis danguyana P.Fourn.; Bull. Soc. Bot. France 78: 434 (1931)

= Orchis latifolia var. immaculata Regel; Bull. Soc. Imp. Naturalistes Moscou 41(2): 429 (1868)

= Orchis longibracteata Schur; Enum. Pl. Transsilv. 643 (1866), non F. W. Schmidt

= Orchis macedonica Griseb.; Spic. Fl. Rumel. 2: 361 (1844)

= Orchis maculata lusus calvellii (A.Terracc.) Soó; Bot. Arch. 23: 76 (1928)

= Orchis maculata unranked immaculata Schur; Oesterr. Bot. Z. 20: 366 (1870)

= Orchis maculata f. bracteosa Harz; Fl. Deutschl. [Schlechtendal], ed. 5, Neuer Abdruck, 4: 261 (1895 publ. 1896)

= Orchis maculata f. brevicalcarata Rohlena; Preslia 10: 153 (1931)

= Orchis maculata f. carpatica Zapal.; Consp. Fl. Galic. Crit. 211 (1906)

= Orchis maculata f. depressa Engenst.; Allg. Bot. Z. Syst. 18: 110 (1912)

= Orchis maculata f. falcata A.Fuchs; Ber. Naturwiss. Vereins Schwaben Neuburg 42: 154 (1919)

= Orchis maculata f. lancifolia Harz; Fl. Deutschl. [Schlechtendal], ed. 5, Neuer Abdruck, 4: 258 (1895 publ. 1896)

= Orchis maculata f. leucantha Druce; Rep. Bot. Soc. Exch. Club Brit. Isles 4: 213 (1915 publ. 1916)

= Orchis maculata f. macroglossa Druce; Rep. Bot. Soc. Exch. Club Brit. Isles 5: 579 (1919)

= Orchis maculata f. media E.G.Camus; Monogr. Orchid. 50 (1908)

= Orchis maculata f. oblongifolia Harz; Fl. Deutschl. [Schlechtendal], ed. 5, Neuer Abdruck, 4: 258 (1895 publ. 1896)

= Orchis maculata f. obscura Neuman; Bot. Not. 1909: 153 (1909)

= Orchis maculata f. obtusifolia Schur; Verh. Mitth. Siebenbürg. Vereins Naturwiss. Hermannstadt 4: 72 (1853)

= Orchis maculata f. ovalifolia Beck; Fl. Nieder-Österreich 204 (1890)

= Orchis maculata f. subintegriflora Druce; Rep. Bot. Soc. Exch. Club Brit. Isles 1921: 396 (1921)

= Orchis maculata f. traunsteineriifolius Harz; Fl. Deutschl. [Schlechtendal], ed. 5, Neuer Abdruck, 4: 260 (1895 publ. 1896)

= Orchis maculata L.; Sp. Pl. [Linnaeus] 2: 942 (1753)

= Orchis maculata var. albicans Marchand ex Tinant; Fl. Luxemb. 441 (1836)

= Orchis maculata var. alpina Schur; Verh. Mitth. Siebenbürg. Vereins Naturwiss. Hermannstadt 4: 72 (1853)

= Orchis maculata var. angustata Harz; Fl. Deutschl. [Schlechtendal], ed. 5, Neuer Abdruck, 4: 261 (1896)

= Orchis maculata var. angustifolia Hartm.; Handb. Skand. Fl. 328 (1820)

= Orchis maculata var. angustifolia Tinant; Fl. Luxemb. 441 (1836)

= Orchis maculata var. biermannii (Ortm.) Nyman; Consp. Fl. Eur. 693 (1882)

= Orchis maculata var. carnea Tinant; Fl. Luxemb. 441 (1836)

= Orchis maculata var. comosa Schur; Verh. Mitth. Siebenbürg. Vereins Naturwiss. Hermannstadt 4: 72 (1853)

= Orchis maculata var. concolor Lange; Haandb. Danske Fl. 502 (1851)

= Orchis maculata var. elabiata R.Keller; Ber. Schweiz. Bot. Ges. 14: 116 (1904)

= Orchis maculata var. foliis-immaculatis Tinant; Fl. Luxemb. 441 (1836), nom. inval.

= Orchis maculata var. gruenii Van Heurck & Wesm.; Prodr. Fl. Brabant 79 (1861)

= Orchis maculata var. hunyadensis (Borsos & Soó) Pauca; Fl. Reipubl. Popul. Roman. (Fl. Republ. Socialist. Romania) 12: 698 (1972)

= Orchis maculata var. latifolia Hartm.; Handb. Skand. Fl. 328 (1820)

= Orchis maculata var. longibracteata Schur; Oesterr. Bot. Z. 20: 296 (1870)

= Orchis maculata var. lusitanica J.A.Guim.; Bol. Soc. Brot. 5: 79 (1887)

= Orchis maculata var. macrobracteata Borbás; Deutsche Bot. Monatsschr. 1883: 296 (1883)

= Orchis maculata var. minor Bréb.; Fl. Normandie, éd. 5: 389 (1879)

= Orchis maculata var. montana Marchand ex Tinant; Fl. Luxemb. 441 (1836)

= Orchis maculata var. olgae (Soó) Pauca; Fl. Reipubl. Popul. Roman. (Fl. Republ. Socialist. Romania) 12: 698 (1972)

= Orchis maculata var. paludosa Marchand ex Tinant; Fl. Luxemb. 441 (1836)

= Orchis maculata var. palustris Cortesi; 193 (1903)

= Orchis maculata var. pumila Neuman; Bot. Not. 1909: 245 (1909)

= Orchis maculata var. sandomiriensis Zapal.; Consp. Fl. Galic. Crit. 211 (1906)

= Orchis maculata var. vespertilio Lindm.; Kongl. Svenska Vetensk.-Akad. Handl., N. F. 23(3, 1): 6 (1897)

= Orchis maculata var. vexillum Lindm.; Kongl. Svenska Vetensk.-Akad. Handl., N. F. 23(3, 1): 6 (1897)

= Orchis nemorosa Montandon; Syn. Fl. Jura, ed. 2: 294 (1868)

= Orchis obtusifolia Schur; Verh. Mitth. Siebenbürg. Vereins Naturwiss. Hermannstadt 4: 72 (1853)

= Orchis solida Moench; Methodus [Moench] 713 (1794), nom. illeg.

= Orchis tetragona Heuff.; Flora (Regensburg) 16: 363 (1833)

variant :

var. arduennensis (Zadoks) Kreutz; Ber. Arbeitskreis. Heimische Orchid. 24(1): 153 (2007)

Distribution: Belgium; NE-France; Luxembourg

11 BGM LUX 12 FRA

Synonyms: hidden

= Dactylorchis arduennensis Zadoks; Bull. Soc. Naturalistes Luxemb. 59: 37 (1954)

= Dactylorchis maculata subsp. arduennensis Zadoks; Bull. Soc. Naturalistes Luxemb. 59: 125 (1954)

= Dactylorhiza maculata f. arduennensis (Zadoks) P.Delforge; Naturalistes Belges 92(Orchid. 24): 202 (2011)

= Dactylorhiza maculata subsp. arduennensis (Zadoks) Tournay; Fl. Belg. Nord France Rég. Vois. 691 (1967)

variant :

var. averyanovii Jagiello; Fragm. Florist. Geobot. 3132 (34): 369 (198687 publ. 1988)

Distribution: Poland; Czech Republic; Slovakia

11 CZE POL SLO

Synonyms: hidden

= Dactylorhiza averyanovii (Jagiello) Kreutz & Tenschert; Guide Orchids Europe N. Afr. Middle E. 459 (2024)

= Dactylorhiza maculata subsp. averyanovii (Jagiello) Jagiello; Acta Univ. Wratislav., Prace Bot. 1055, 45 (1990)

variant :

var. bucovina (Kreutz, Bobocea & Bradeanu) comb.ined.

Distribution: Romania (Bucovina, Maramures)

13 ROM

Synonyms: hidden

= Dactylorhiza bucovina Kreutz, Bobocea & Bradeanu; Ber. Arbeitskreis. Heimische Orchid. 38(2): 101 (2021)

variant :

var. nummularia Pikner; Ber. Arbeitskreis. Heimische Orchid. 30: 295 (2013)

Distribution: Estonia

14 EST

subspecies :

subsp. arcana Trávn., Taraska, Batousek & Lamla; Folia Geobot. 58(2): 177 (2024)

Distribution: S-Poland; Slovakia

11 POL SLO

Synonyms: hidden

= Dactylorhiza arcana (Trávn., Taraska, Batousek & Lamla) Kreutz; Guide Orchids Europe N. Afr. Middle E. 457 (2024)

subspecies :

subsp. caramulensis Verm.; Bol. Soc. Brot., Sér. 2, 44: 94 (1970)

Distribution: Portugal (C- & N-Portugal); Spain (Galicia, Extremadura, Castilla y Leon)

12 POR SPA

Synonyms: hidden

= Dactylorhiza caramulensis (Verm.) D.Tyteca; Orchidophile (Paris) 88: 155 (1989)

subspecies :

subsp. elodes (Griseb.) Soó; Nom. Nov. Gen. Dactylorhiza 7 (1962)

Name: Heidemoor-Fingerwurz, Heidemoor-Fingerknabenkraut (DE)

Distribution: Netherlands; NW-Germany; Belgium; NE-France

11 BGM GER NET 12 FRA

Synonyms: hidden

= Dactylorchis elodes (Griseb.) Verm.; Stud. Dactylorch. 137 (1947)

= Dactylorchis maculata subsp. elodes (Griseb.) Verm.; Stud. Dactylorch. 68 (1947)

= Dactylorhiza elodes (Griseb.) Aver.; Bot. Zhurn. (Moscow & Leningrad) 67(3): 309 (1982)

= Dactylorhiza elodes (Griseb.) Englmaier; Abh. Zool.-Bot. Ges. Österreich 22: 107 (1984), isonym

= Dactylorhiza maculata var. dalarnensis Landwehr; Orchideeën 37(3): 77 (1975), et Wilde Orchid. Eur. 2: 556 (1977)

= Dactylorhiza maculata var. elodes (Griseb.) Aver.; Turczaninowia 3(1): 49 (2000)

= Orchis elodes Griseb.; Bild. Torfs. 67 (1845)

= Orchis maculata proles elodes (Griseb.) Rouy; Fl. France [Rouy & Foucaud] 13: 154 (1912)

= Orchis maculata subsp. elodes (Griseb.) K.Richt.; Pl. Eur. 1: 272 (1890)

= Orchis maculata subsp. elodes (Griseb.) O.Bolòs & Vigo; Fl. Països Catalans 4: 629 (2001), isonym

= Orchis maculata subsp. elodes (Griseb.) P.Fourn.; Quatre Fl. France 206 (1935)

= Orchis maculata var. elodes (Griseb.) Jess.; Deut. Excurs.-Fl. 482 (1879), isonym

= Orchis maculata var. elodes (Griseb.) Rchb.f.; Icon. Fl. Germ. Helv. [H.G.L.Reichenbach] 13-14: 67 (1851)

subspecies :

subsp. ericetorum (E.F.Linton) P.F.Hunt & Summerh.; Watsonia 6: 132 (1965)

Name: Heath Spotted-orchid (EN)

Distribution: Faeroe Isl.; England (England, Wales, Isle of Man, Scotland, Outer Hebrides, Orkney Isl., Shetland Isl., St. Kilda Isl.); Ireland (Ireland, Northern Ireland); Denmark; Norway; Sweden; Finland; Netherlands; Belgium; ?Luxembourg; Germany; Slovakia; Portugal; Spain; France; Channel Isl. (Alderney, Guernsey, Jersey, Sark); ?Romania

10 DEN FIN FOR GRB IRE NOR SWE 11 BGM GER NET SLO 12 CIS FRA POR SPA

Synonyms: hidden

= Dactylorchis maculata subsp. ericetorum (E.F.Linton) Verm.; Stud. Dactylorch. 69 (1947)

= Dactylorchis maculata subsp. rhoumensis (Hesl.-Harr.f.) Hesl.-Harr.f.; Watsonia 4: 48 (1957)

= Dactylorchis maculata var. ericetorum (E.F.Linton) Verm.; Fl. Neerl. 1(5): 85 (1958)

= Dactylorhiza ericetorum (E.F.Linton) Aver.; Bot. Zhurn. (Moscow & Leningrad) 67(3): 309 (1981)

= Dactylorhiza fuchsii subsp. rhoumensis (Hesl.-Harr.f.) A.R.Clapham, Tutin & E.F.Warb.; Fl. Brit. Isles [Clapham et al.] ed. 2: 1046 (1962)

= Dactylorhiza maculata subsp. rhoumensis (Hesl.-Harr.f.) Soó; Nom. Nov. Gen. Dactylorhiza 7 (1962)

= Dactylorhiza maculata var. ericetorum (E.F.Linton) Soó; Ann. Univ. Sci. Budapest. Rolando Eötvös, Sect. Biol. 3: 349 (1960), nom. inval.

= Dactylorhiza maculata var. praecox (Webster) Soó; Nom. Nov. Gen. Dactylorhiza 7 (1962)

= Dactylorhiza maculata var. rhoumensis (Hesl.-Harr.f.) P.Delforge; Naturalistes Belges 96 (Orchid. 28): 15 (2015)

= Orchis ericetorum (E.F.Linton) A.Benn.; Ann. Scott. Nat. Hist. 1905: 39 (1905)

= Orchis fuchsii subsp. rhoumensis Hesl.-Harr.f.; Trans. Bot. Soc. Edinburgh 35: 53 (1948)

= Orchis maculata f. ericetorum (E.F.Linton) Hagerup; Dansk Bot. Ark. 11: 3 (1944)

= Orchis maculata subsp. ericetorum E.F.Linton; Fl. Bournemouth 208 (1900)

= Orchis maculata subsp. rhoumensis Hesl.-Harr.f.; Trans. Bot. Soc. Edinburgh 35: 64 (1948)

= Orchis maculata var. praecox Webster; Brit. Orchids 54 (1886)

variant :

var. terschellingensis (Verm.) comb.ined.

Distribution: England (Scotland); Norway; Denmark (Jutland); Netherlands (Terschelling Isl., Vlieland)

10 DEN GRB NOR 11 NET

Synonyms: hidden

= Dactylorhiza maculata subsp. podesta (Landwehr) Kreutz; Orchid. van Nederland [Kreutz & Dekker] 179 (2000)

= Dactylorhiza maculata var. podesta Landwehr; Wilde Orchid. Eur. 2: 556 (1977)

= Dactylorchis maculata var. terschellingensis Verm.; Ned. Kruidk. Arch. 56: 233 (1949)

= Dactylorhiza podesta (Landwehr) Kreutz; Guide Orchids Europe N. Afr. Middle E. 461 (2024)

subspecies :

subsp. islandica (Á.Löve & D.Löve) Soó; Nom. Nov. Gen. Dactylorhiza 7 (1962)

Distribution: Iceland

10 ICE

Synonyms: hidden

= Dactylorchis maculata subsp. islandica Á.Löve & D.Löve; Chrom. Numb. Nort. Pl. 106 (1948)

= Dactylorhiza islandica (Á.Löve & D.Löve) Aver.; Bot. Zhurn. (Moscow & Leningrad) 69(6): 875 (1984)

= Dactylorhiza maculata var. islandica (Á.Löve & D.Löve) Hyl.; Nordisk Kärlväxtfl. 2: 387 (1966)

subspecies :

subsp. kolaensis (Montell) Kreutz; Kompend. Eur. Orchid. 46 (2004)

Distribution: N-Norway; N-Finland; N-European Russia (Kola peninsula)

10 FIN NOR 14 RUS

Synonyms: hidden

= Dactylorchis maculata subsp. montellii Verm.; Stud. Dactylorch. 69, 141 (1947)

= Dactylorhiza kolaensis (Montell) Aver.; Bot. Zhurn. (Moscow & Leningrad) 69(6): 875 (1984)

= Dactylorhiza maculata subsp. deflexa Landwehr; Orchideeën 37(3): 77 (1975), et Wilde Orchid. Eur. 2: 556 (1977)

= Dactylorhiza maculata subsp. montellii (Verm.) Landwehr; Orchideeën 37(3): 79 (1975)

= Dactylorhiza maculata var. kolaensis (Montell) Soó; Nom. Nov. Gen. Dactylorhiza 7 (1962)

= Dactylorhiza montellii (Verm.) P.Delforge; Naturalistes Belges 71(3): 101 (1990)

= Orchis maculata var. kolaensis Montell; Memoranda Soc. Faun. Fl. Fenn. 23: 166 (1947)

subspecies :

subsp. pyrenaica Kreutz; Ber. Arbeitskreis. Heimische Orchid. 30(2): 279 (2013)

Distribution: Andorra; SW-France (Pyrénées)

12 ADO FRA

Synonyms: hidden

= Dactylorhiza savogiensis f. pyrenaica (Kreutz) P.Delforge; Naturalistes Belges 96 (Orchid. 28): 15 (2015)

= Dactylorhiza pyrenaica (Kreutz) Kreutz; Guide Orchids Europe N. Afr. Middle E. 468 (2024)

subspecies :

subsp. savogiensis (D.Tyteca & Gathoye) Kreutz; Kompend. Eur. Orchid. 46 (2004)

Distribution: France (Massif Central, W-Alps, Pyrenees); Andorra; Spain (Huesca); NW-Italy (Aosta); Switzerland (Wallis, Graubünden, Tessin, Uri)

11 SWI 12 ADO FRA SPA 13 ITA

Synonyms: hidden

= Dactylorhiza fuchsii var. brachystachys (E.G.Camus) Soó; Nom. Nov. Gen. Dactylorhiza 8 (1962)

= Dactylorhiza savogiensis D.Tyteca & Gathoye; Orchidophile (Paris) 92: 108 (1990)

= Orchis maculata var. brachystachys E.G.Camus; Iconogr. Orchid. Europe, Atlas 1: 4 (1921)

subspecies :

subsp. schurii (Klinge) Soó; Rev. Roumaine Biol., Sér. Bot. 12: 229 (1967)

Distribution: Romania (Carpathians)

13 ROM

Synonyms: hidden

= Dactylorhiza maculata var. schurii (Klinge) Soó; Nom. Nov. Gen. Dactylorhiza 7 (1962)

= Dactylorhiza schurii (Klinge) Aver.; Bot. Zhurn. (Moscow & Leningrad) 69(6): 875 (1984)

= Dactylorhiza traunsteineri subsp. schurii (Klinge) Kreutz; Ber. Arbeitskreis. Heimische Orchid. 31: 106 (2014 publ. 2015)

= Orchis angustifolia f. schurii Klinge; Revis. Orchis cordigera & O. angustifolia 63, 83 (1893)

= Orchis maculata f. schurii (Klinge) Soó; Monogr. Iconogr. Orchid. Eur. 2: 240, 243 (1933)

= Orchis maculata var. schurii (Klinge) Pauca; Fl. Reipubl. Popul. Roman. (Fl. Republ. Socialist. Romania) 12: 697 (1972)

= Orchis traunsteineri var. schurii (Klinge) Asch. & Graebn.; Syn. Mitteleur. Fl. [Ascherson & Graebner] 3: 729 (1907)

subspecies :

subsp. sooana Borsos ex Batousek; Pl. Syst. Evol. 307(4)-51: 16 (2021)

Distribution: Slovakia

11 SLO

subspecies :

subsp. transsilvanica (Schur) Soó; Nom. Nov. Gen. Dactylorhiza 7 (1962)

Name: Siebenbürger Fingerwurz, Siebenbürger Fingerknabenkraut (DE)

Distribution: Czech Republic; Slovakia; Slovenia; Croatia; Bosnia & Hercegovina; Serbia; Kosovo; Romania; Estonia; European Russia

11 CZE SLO 13 BIH CRO KOS ROM SLV SRB 14 EST RUS

Synonyms: hidden

= Dactylorchis maculata subsp. transsilvanica (Schur) Verm.; Stud. Dactylorch. 69 (1947)

= Dactylorchis maculata var. candidissima (Krock.) Verm.; Ned. Kruidk. Arch. 56: 235 (1949)

= Dactylorhiza fuchsii subsp. transsilvanica (Schur) S.E.Fröhner; Ber. Arbeitsgem. Sächs. Bot. N. F.8: 143 (1968)

= Dactylorhiza maculata var. transsilvanica (Schur) P.Delforge; Naturalistes Belges 81(4): 397 (2000)

= Dactylorhiza maculata var. wettsteinii (H.Fleischm.) Soó; Nom. Nov. Gen. Dactylorhiza 8 (1962)

= Dactylorhiza transsilvanica (Schur) Aver.; Bot. Zhurn. (Moscow & Leningrad) 67(3): 309 (1982)

= Orchis candidissima Krock.; Fl. Siles. [Krocker] 3: 16, t. 2 (1814)

= Orchis maculata f. wettsteinii (H.Fleischm.) Soó; Repert. Spec. Nov. Regni Veg. 24: 32 (1927)

= Orchis maculata subsp. transsilvanica (Schur) Soó; Notizbl. Bot. Gart. Berlin-Dahlem 9: 911 (1926)

= Orchis maculata var. candidissima (Krock.) Nyman; Consp. Fl. Eur. Suppl. 2: 291 (1890)

= Orchis maculata var. ochrantha Pancic; Verh. K.K. Zool.-Bot. Ges. Wien 6: 575 (1856)

= Orchis maculata var. transsilvanica (Schur) Nyman; Consp. Fl. Eur. Suppl. 2: 291 (1890)

= Orchis ochrantha (Pancic) H.Fleischm.; Mitt. Naturwiss. Vereines Steiermark 45: 176 (1908 publ. 1909)

= Orchis ochrantha var. wettsteinii H.Fleischm.; Mitt. Naturwiss. Vereines Steiermark 45: 178 (1909)

= Orchis transsilvanica Schur; Verh. Mitth. Siebenbürg. Vereins Naturwiss. Hermannstadt 4: 72 (1853)

species :

Dactylorhiza magna (Czerniak.) Ikonn.; Novosti Sist. Vyssh. Rast. 9: 303 (1972)

subspecies :

subsp. magna

Distribution: Uzbekistan; Tajikistan

32 TZK UZB

Synonyms: hidden

= Dactylorhiza baldshuanica Chernyak.; Bot. Zhurn. (Moscow & Leningrad) 68(4): 534 (1983) ex Aver.

= Dactylorhiza umbrosa subsp. magna (Czerniak.) Soó; Ann. Univ. Sci. Budapest. Rolando Eötvös, Sect. Biol., 11: 68 (1969)

= Orchis magna Czerniak.; Fl. Uzbekist. 1: 546 (1941)

subspecies :

subsp. czerniakowskae (Aver.) Eccarius; Orchideengattung Dactylorhiza 102 (2016)

Distribution: Kazakhstan; Uzbekistan; Tajikistan

32 KAZ TZK UZB

Synonyms: hidden

= Dactylorhiza czerniakowskae Aver.; Bot. Zhurn. (Moscow & Leningrad) 68(4): 536 (1983)

subspecies :

subsp. kulikalonica (Chernyak.) Eccarius; Orchideengattung Dactylorhiza 106 (2016)

Distribution: Kazakhstan; Uzbekistan; Tajikistan

32 KAZ TZK UZB

Synonyms: hidden

= Dactylorhiza kulikalonica Chernyak.; Bot. Zhurn. (Moscow & Leningrad) 68(4): 535 (1983) ex Aver.

species :

Dactylorhiza majalis (Rchb.) P.F.Hunt & Summerh.; Watsonia 6: 130 (1965), nom. cons.

subspecies :

subsp. majalis

Name: Breitblättrige Fingerwurz, Breitblättriges Fingerknabenkraut (DE)

Distribution: Denmark; Norway; Sweden; Netherlands; Belgium; Luxembourg; Germany (Brandenburg, Berlin, Baden-Württemberg, Bayern, Hessen, Hamburg, Mecklenburg-Vorpommern, Niedersachsen, Nordrhein-Westfalen, Rheinland-Pfalz, Schleswig-Holstein, Saarland, Sachsen, Sachsen-Anhalt, Thüringen); Switzerland; Liechtenstein; Austria; Poland; Czech Republic; Slovakia; Hungary; Spain; Andorra; France; Italy; Slovenia; Croatia; Bosnia & Hercegovina; Montenegro; Serbia; Kosovo; North Macedonia; Romania; Lithuania; Belarus; Ukraine; European Russia

10 DEN NOR SWE 11 AUT BGM CZE GER HUN LIE LUX NET POL SLO SWI 12 ADO FRA SPA 13 BIH CRO ITA KOS MNT NMA ROM SLV SRB 14 BLR LIT RUS UKR

Remarks: majalis agg.

Synonyms: hidden

= Dactylorchis latifolia (L.) Rothm.; Feddes Repert. Beih. 67 (1-3): 4 (1963) [nom. rej.]

= Dactylorchis latifolia subsp. brevifolia Bisse; Feddes Repert. Spec. Nov. Regni Veg. 67: 184 (1963)

= Dactylorchis majalis (Rchb.) Verm.; Stud. Dactylorch. 67 (1947)

= Dactylorchis majalis f. albolileacea Verm.; Ned. Kruidk. Arch. 56: 217 (1949)

= Dactylorchis majalis f. ampla (Asch. & Graebn.) Verm.; Ned. Kruidk. Arch. 56: 214 (1949)

= Dactylorchis majalis var. aequiloba (Harz) Verm.; Ned. Kruidk. Arch. 56: 216 (1949)

= Dactylorchis majalis var. albiflora (Tinant) Verm.; Ned. Kruidk. Arch. 56: 218 (1949)

= Dactylorchis majalis var. alborosacea (Erdner) Verm.; Ned. Kruidk. Arch. 56: 217 (1949)

= Dactylorchis majalis var. alpestroides Verm.; Ned. Kruidk. Arch. 56: 215 (1949)

= Dactylorchis majalis var. atrorubens Verm.; Ned. Kruidk. Arch. 56: 215 (1949)

= Dactylorchis majalis var. brachyphylla Verm.; Ned. Kruidk. Arch. 56: 213 (1949)

= Dactylorchis majalis var. brevicalcarata (Harz) Verm.; Ned. Kruidk. Arch. 56: 216 (1949)

= Dactylorchis majalis var. gracilis Verm.; Ned. Kruidk. Arch. 56: 213 (1949)

= Dactylorchis majalis var. immaculata (Regel) Verm.; Ned. Kruidk. Arch. 56: 215 (1949)

= Dactylorchis majalis var. integra (Harz) Verm.; Ned. Kruidk. Arch. 56: 216 (1949)

= Dactylorchis majalis var. laxa (Harz) Verm.; Ned. Kruidk. Arch. 56: 215 (1949)

= Dactylorchis majalis var. linguata (Harz) Verm.; Ned. Kruidk. Arch. 56: 216 (1949)

= Dactylorchis majalis var. macrobracteata (Schur) Verm.; Ned. Kruidk. Arch. 56: 215 (1949)

= Dactylorchis majalis var. magnicalcarata (Harz) Verm.; Ned. Kruidk. Arch. 56: 217 (1949)

= Dactylorchis majalis var. submaculata (Rchb.f.) Verm.; Ned. Kruidk. Arch. 56: 215 (1949)

= Dactylorhiza comosa subsp. majalis (Rchb.) P.D.Sell; Fl. Gr. Brit. Ireland 5: 365 (1996)

= Dactylorhiza fistulosa (Moench) H.Baumann & Künkele; Mitt. Arbeitskreises Heimische Orchideen 15(4): 456 (1983)

= Dactylorhiza latifolia (L.) H.Baumann & Künkele; Mitt. Arbeitskreises Heimische Orchideen 15(4): 444 (1983), isonym

= Dactylorhiza latifolia (L.) Soó; Nom. Nov. Gen. Dactylorhiza 4 (1962) [nom. illeg.]

= Dactylorhiza latifolia var. gracilis (Verm.) Soó; Nom. Nov. Gen. Dactylorhiza 4 (1962)

= Dactylorhiza latifolia var. limbata Hudziok; 707 (1963)

= Dactylorhiza latifolia var. pinguis (Asch. & Graebn.) Soó; Nom. Nov. Gen. Dactylorhiza 4 (1962)

= Dactylorhiza majalis (Rchb.) Rauschert ex Hudziok; Wiss. Z. Martin-Luther-Univ. Halle-Wittenberg, Math.-Naturwiss. Reihe 14: 492 (1965)

= Dactylorhiza majalis lusus albiflora (Thielens) Soó; Acta Bot. Acad. Sci. Hung. 16: 368 (1970 publ. 1971)

= Dactylorhiza majalis lusus alboliliacea (Verm.) Soó; Acta Bot. Acad. Sci. Hung. 16: 368 (1970 publ. 1971)

= Dactylorhiza majalis lusus alborosacea (Erdner) Soó; Acta Bot. Acad. Sci. Hung. 16: 368 (1970 publ. 1971)

= Dactylorhiza majalis lusus carnea (Neuman) Soó; Acta Bot. Acad. Sci. Hung. 16: 368 (1970 publ. 1971)

= Dactylorhiza majalis lusus pallida (Thielens) Soó; Acta Bot. Acad. Sci. Hung. 16: 368 (1970 publ. 1971)

= Dactylorhiza majalis f. alpestroides (Verm.) Soó; Acta Bot. Acad. Sci. Hung. 16: 368 (1970 publ. 1971)

= Dactylorhiza majalis f. ampla (Asch. & Graebn.) Landwehr; 79 (1975)

= Dactylorhiza majalis f. aspersa F.Proch.; Preslia 54: 290 (1982)

= Dactylorhiza majalis f. atrorubens (Verm.) Soó; Acta Bot. Acad. Sci. Hung. 16: 368 (1970 publ. 1971)

= Dactylorhiza majalis f. barlae (E.G.Camus) Soó; Acta Bot. Acad. Sci. Hung. 16: 368 (1970 publ. 1971)

= Dactylorhiza majalis f. brevicalcarata (Harz) Soó; Acta Bot. Acad. Sci. Hung. 16: 368 (1970 publ. 1971)

= Dactylorhiza majalis f. chwaszczyniensis Marg. & Tukallo; Richardiana 12(2): 53 (51-58

= Dactylorhiza majalis f. heikeana Gottschalk; 14 (2014)

= Dactylorhiza majalis f. immaculata (Harz) Soó; Acta Bot. Acad. Sci. Hung. 16: 368 (1970 publ. 1971)

= Dactylorhiza majalis f. laxa (Harz) Soó; Acta Bot. Acad. Sci. Hung. 16: 368 (1970 publ. 1971)

= Dactylorhiza majalis f. linearis (Thielens) Soó; Acta Bot. Acad. Sci. Hung. 16: 368 (1970 publ. 1971)

= Dactylorhiza majalis f. linguata (Harz) Soó; Acta Bot. Acad. Sci. Hung. 16: 368 (1970 publ. 1971)

= Dactylorhiza majalis f. maculata (Thielens) Soó; Acta Bot. Acad. Sci. Hung. 16: 368 (1970 publ. 1971)

= Dactylorhiza majalis f. magnicalcarata (Harz) Soó; Acta Bot. Acad. Sci. Hung. 16: 368 (1970 publ. 1971)

= Dactylorhiza majalis f. rubra (Winterl) D.Tyteca & Gathoye; Orchidophile (Paris) 92: 112 (1990)

= Dactylorhiza majalis f. tharandina (Rchb.f.) Soó; Acta Bot. Acad. Sci. Hung. 16: 368 (1970 publ. 1971)

= Dactylorhiza majalis subsp. brevifolia (Bisse) Senghas; Jahresber. Naturwiss. Vereins Wuppertal 21/22: 126 (1963)

= Dactylorhiza majalis var. druceoides Balayer; Bull. Soc. Bot. France, Lett. Bot. 133(3): 280 (1986)

= Dactylorhiza majalis var. pinguis (Asch. & Graebn.) Landwehr; 80 (1975)

= Dactylorhiza majalis var. semimaculata R.Doll; Feddes Repert. 89(56): 347 (1978)

= Orchis divaricata Rich. ex Loisel.; Fl. Gall. [Loiseleur] 2: 606 (1807)

= Orchis elata Hartm.; Handb. Skand. Fl., ed. 10: 202 (1870) non Poir. 1789

= Orchis fistulata Stokes; Bot. Mat. Med. 4: 296 (1812)

= Orchis fistulosa Moench; Methodus [Moench] 713 (1794), nom. illeg.

= Orchis incarnata subsp. latifolia (L.) Cajander; Melan. Suom. Kasvio [Cajander], ed. 5: 171 (1906)

= Orchis latifolia f. aequiloba (Harz) Soó; Bot. Arch. 23: 69 (1928)

= Orchis latifolia f. ampla Asch. & Graebn.; Syn. Mitteleur. Fl. [Ascherson & Graebner] 3: 735 (1907)

= Orchis latifolia f. brevicalcarata (Harz) Soó; Bot. Arch. 23: 68 (1928)

= Orchis latifolia f. densa (Harz) Soó; Bot. Arch. 23: 68 (1928)

= Orchis latifolia f. inaequiloba (Harz) Soó; Bot. Arch. 23: 69 (1928)

= Orchis latifolia f. laxa (Harz) Soó; Bot. Arch. 23: 68 (1928)

= Orchis latifolia f. laxiflora Zapal.; Consp. Fl. Galic. Crit. 1: 215 (1906)

= Orchis latifolia f. macrobracteata (Schur) Soó; Bot. Arch. 23: 68 (1928)

= Orchis latifolia f. macrochlamys (Asch. & Graebn.) M.Goldschm.; 285 (1908)

= Orchis latifolia f. macrophylla Zapal.; Consp. Fl. Galic. Crit. 1: 215 (1906)

= Orchis latifolia f. pinguis Asch. & Graebn.; Syn. Mitteleur. Fl. [Ascherson & Graebner] 3: 735 (1907)

= Orchis latifolia Scop.; Fl. Carniol., ed. 2, 2: 196 (1771), nom. illeg.

= Orchis latifolia subsp. majalis (Rchb.) Ces.; 315 (1844)

= Orchis latifolia subsp. majalis (Rchb.) Gand.; Nov. Consp. Fl. Eur. 461 (1910), isonym

= Orchis latifolia subsp. majalis (Rchb.) Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(1): 168 (1898), isonym

= Orchis latifolia var. abbreviata Harz; Fl. Deutschl. [Schlechtendal], ed. 5, Neuer Abdruck, 4: 242 (1895 publ. 1896)

= Orchis latifolia var. aequiloba Harz; Fl. Deutschl. [Schlechtendal], ed. 5, Neuer Abdruck, 4: 242 (1895 publ. 1896)

= Orchis latifolia var. aggregata Harz; Fl. Deutschl. [Schlechtendal], ed. 5, Neuer Abdruck, 4: 242 (1895 publ. 1896)

= Orchis latifolia var. albiflora Tinant; Fl. Luxemb. 441 (1836)

= Orchis latifolia var. albiflora Zapal.; Consp. Fl. Galic. Crit. 1: 214 (1906)

= Orchis latifolia var. ampla Asch. & Graebn.; Syn. Mitteleur. Fl. [Ascherson & Graebner] 3: 735 (1907)

= Orchis latifolia var. brevibracteata Harz; Fl. Deutschl. [Schlechtendal], ed. 5, Neuer Abdruck, 4: 242 (1895 publ. 1896)

= Orchis latifolia var. brevicalcarata Harz; Fl. Deutschl. [Schlechtendal], ed. 5, Neuer Abdruck, 4: 242 (1896)

= Orchis latifolia var. carnea R.Neumann; 9 (1905)

= Orchis latifolia var. densa Harz; Fl. Deutschl. [Schlechtendal], ed. 5, Neuer Abdruck, 4: 242 (1895 publ. 1896)

= Orchis latifolia var. densiflora Harz; Fl. Deutschl. [Schlechtendal], ed. 5, Neuer Abdruck, 4: 242 (1895 publ. 1896)

= Orchis latifolia var. elata Harz; Fl. Deutschl. [Schlechtendal], ed. 5, Neuer Abdruck, 4: 242 (1895 publ. 1896), nom. illeg.

= Orchis latifolia var. gracilis Warnst.; Deutsche Bot. Monatsschr. 1883: 169 (1883), nom. illeg.

= Orchis latifolia var. humilis Harz; Fl. Deutschl. [Schlechtendal], ed. 5, Neuer Abdruck, 4: 242 (1895 publ. 1896)

= Orchis latifolia var. inaequiloba Harz; Fl. Deutschl. [Schlechtendal], ed. 5, Neuer Abdruck, 4: 242 (1895 publ. 1896)

= Orchis latifolia var. integra Harz; Fl. Deutschl. [Schlechtendal], ed. 5, Neuer Abdruck, 4: 242 (1896)

= Orchis latifolia var. lanceata Harz; Fl. Deutschl. [Schlechtendal], ed. 5, Neuer Abdruck, 4: 242 (1895 publ. 1896)

= Orchis latifolia var. laxa Harz; Fl. Deutschl. [Schlechtendal], ed. 5, Neuer Abdruck, 4: 242 (1895 publ. 1896)

= Orchis latifolia var. laxiuscula Harz; Fl. Deutschl. [Schlechtendal], ed. 5, Neuer Abdruck, 4: 242 (1895 publ. 1896)

= Orchis latifolia var. linguata Harz; Fl. Deutschl. [Schlechtendal], ed. 5, Neuer Abdruck, 4: 242 (1895 publ. 1896)

= Orchis latifolia var. macilenta Harz; Fl. Deutschl. [Schlechtendal], ed. 5, Neuer Abdruck, 4: 242 (1895 publ. 1896)

= Orchis latifolia var. macrobracteata Schur; Mitt. Geogr. Ges. Thüringen (Jena) 2: 220 (1884 publ. 1885)

= Orchis latifolia var. macrochlamys Asch. & Graebn.; Syn. Mitteleur. Fl. [Ascherson & Graebner] 3: 735 (1907)

= Orchis latifolia var. magnicalcarata Harz; Fl. Deutschl. [Schlechtendal], ed. 5, Neuer Abdruck, 4: 242 (1895 publ. 1896)

= Orchis latifolia var. majalis (Rchb.) Kitt.; Taschenb. Fl. Deutschl., ed. 1: 137 (1837)

= Orchis latifolia var. majalis (Rchb.) Nyman; Consp. Fl. Eur. 692 (1882)

= Orchis latifolia var. obtusa Gray; Nat. Arr. Brit. Pl. 2: 202 (1822)

= Orchis latifolia var. obtusa Harz; Fl. Deutschl. [Schlechtendal], ed. 5, Neuer Abdruck, 4: 242 (1895 publ. 1896), nom. illeg.

= Orchis latifolia var. palustris Gray; Nat. Arr. Brit. Pl. 2: 202 (1822)

= Orchis latifolia var. parvicalcarata Harz; Fl. Deutschl. [Schlechtendal], ed. 5, Neuer Abdruck, 4: 242 (1895 publ. 1896)

= Orchis latifolia var. parvilabia Zapal.; Consp. Fl. Galic. Crit. 1: 215 (1906)

= Orchis latifolia var. pinguis (Asch. & Graebn.) Hegi; 355 (1909)

= Orchis latifolia var. praecox M.Loehr in Wirtg.; 164 (1842)

= Orchis latifolia var. pyramidalis Harz; Fl. Deutschl. [Schlechtendal], ed. 5, Neuer Abdruck, 4: 242 (1896)

= Orchis latifolia var. robusta Harz; Fl. Deutschl. [Schlechtendal], ed. 5, Neuer Abdruck, 4: 242 (1896)

= Orchis latifolia var. sparsiflora Harz; Fl. Deutschl. [Schlechtendal], ed. 5, Neuer Abdruck, 4: 242 (1895 publ. 1896)

= Orchis latifolia var. submaculata Rchb.f.; Icon. Fl. Germ. Helv. [H.G.L.Reichenbach] 13-14: 57 (1851)

= Orchis latifolia var. traunsteineriiformis Harz; Fl. Deutschl. [Schlechtendal], ed. 5, Neuer Abdruck, 4: 242 (1895 publ. 1896)

= Orchis latifolia var. trifida Harz; Fl. Deutschl. [Schlechtendal], ed. 5, Neuer Abdruck, 4: 242 (1895 publ. 1896)

= Orchis latifolia var. trilobata Harz; Fl. Deutschl. [Schlechtendal], ed. 5, Neuer Abdruck, 4: 242 (1895 publ. 1896)

= Orchis latifolia var. uncinata Rchb.f.; Icon. Fl. Germ. Helv. [H.G.L.Reichenbach] 13-14: 176 (1851)

= Orchis maculata subsp. latifolia Bonnier & Layens; Tabl. Syn. Pl. Vasc. France 311 (1894)

= Orchis maculata subsp. latifolia Douin; Fl. Ill. France 11: 30 (1931)

= Orchis majalis lusus albiflora Thielens; Fl. Luxemb. 60 (1875)

= Orchis majalis lusus alborosacea Erdner; Ber. Naturwiss. Vereins Schwaben Neuburg 39: 142 (1911)

= Orchis majalis lusus pallida Thielens; Fl. Luxemb. 60 (1875)

= Orchis majalis carnea Neuman; Mitt. Bad. Landesvereins Naturk. 5: 201 (1905)

= Orchis majalis f. barlae E.G.Camus; Iconogr. Orchid. Europe 4 (1921)

= Orchis majalis f. brevicalcarata Harz; Fl. Deutschl. [Schlechtendal], ed. 5, Neuer Abdruck, 4: 247 (1895 publ. 1896)

= Orchis majalis f. immaculata Harz; Fl. Deutschl. [Schlechtendal], ed. 5, Neuer Abdruck, 4: 247 (1895 publ. 1896)

= Orchis majalis f. laxa Harz; Fl. Deutschl. [Schlechtendal], ed. 5, Neuer Abdruck, 4: 247 (1895 publ. 1896)

= Orchis majalis f. linearis Thielens; Fl. Luxemb. 60 (1875)

= Orchis majalis f. linguata Harz; Fl. Deutschl. [Schlechtendal], ed. 5, Neuer Abdruck, 4: 247 (1895 publ. 1896)

= Orchis majalis f. maculata Thielens; Fl. Luxemb. 60 (1875)

= Orchis majalis f. magnicalcarata Harz; Fl. Deutschl. [Schlechtendal], ed. 5, Neuer Abdruck, 4: 247 (1895 publ. 1896)

= Orchis majalis Rchb.; Iconogr. Bot. Pl. Crit. 6: 7 (1828), nom. cons.

= Orchis majalis var. pinguis (Asch. & Graebn.) Pugsley; 585 (1935)

= Orchis majalis var. tharandina Rchb.f.; Icon. Fl. Germ. Helv. [H.G.L.Reichenbach] 13-14: 58 (1851)

= Orchis palmata Syme; 99 (1869), nom. illeg.

= Orchis tingitana Durand ex G.Keller & Soó; Repert. Spec. Nov. Regni Veg. Sonderbeih. A: 2: 230 (1933)

= Orchis ×ulehlae Podp.; Klíc Kvét. Ceskoslov. 796 (1928), et ex Soó, in Repert. Spec. Nov. Regni Veg. Sonderbeih. A: 2: 199 (1932)

variant :

var. alpestris (Pugsley) Wucherpf.; Ber. Arbeitskreis. Heimische Orchid. 19(2): 106 (2002 publ. 2003)

Distribution: Germany; Switzerland; Austria; NE-Spain; France; Italy

11 AUT GER SWI 12 FRA SPA 13 ITA

Remarks: status disputed, probably only a boreoalpine ecovariety

Synonyms: hidden

= Dactylorchis alpestris (Pugsley) Verm.; Stud. Dactylorch. 67 (1947)

= Dactylorhiza alpestris (Pugsley) Aver.; Bot. Zhurn. (Moscow & Leningrad) 68(9): 1164 (1983)

= Dactylorhiza fistulosa subsp. alpestris (Pugsley) H.Baumann & Künkele; Mitt. Arbeitskreises Heimische Orchideen 20(3): 628 (1988), nom. illeg.

= Dactylorhiza latifolia subsp. alpestris (Pugsley) Soó; Nom. Nov. Gen. Dactylorhiza 4 (1962)

= Dactylorhiza latifolia var. pumila (Freyn ex M.Schultze) Soó; Nom. Nov. Gen. Dactylorhiza 4 (1962)

= Dactylorhiza majalis subsp. alpestris (Pugsley) Senghas; Jahresber. Naturwiss. Vereins Wuppertal 21-22: 51 (1968)

= Dactylorhiza majalis var. pumila (Freyn ex M.Schultze) Landwehr; Orchideeën 37(3): 80 (1975)

= Dactylorhiza majalis var. pumila (Freyn ex M.Schulze) H.Baumann, Künkele & R.Lorenz; J. Eur. Orch. 34: 151 (2002), isonym

= Orchis alpestris Pugsley; Proc. Linn. Soc. London Sess. 146, 98 (1934), in clavi, et J. Linn. Soc., Lond., Bot. 49: 587 (1935) [descr. ampl.]

= Orchis impudica Crantz; Stirp. Austr. Fasc., ed. 2, 6: 497 (1769), nom. illeg.

= Orchis latifolia f. pumila Freyn ex M.Schulze; Mitth. Thüring. Bot. Vereins, N. F. 17: 51 (1902)

= Orchis latifolia subsp. alpestris (Pugsley) Janch.; 72 (1950)

= Orchis latifolia subsp. impudica (Crantz) Soó; Repert. Spec. Nov. Regni Veg. 24: 30 (1927)

= Orchis latifolia var. macrantha Zapal.; Consp. Fl. Galic. Crit. 210 (1906)

= Orchis latifolia var. pumila (Freyn ex M.Schultze) W.Zimmerm.; Formen Orch. Deutschl., Deutsch-Österr. & Schweiz 36 (1912)

= Orchis latifolius subsp. impudicus (Crantz) Soó; Repert. Spec. Nov. Regni Veg. 24: 30 (1927)

= Orchis latifolius var. impudicus (Crantz) Asch. & Graebn.; Syn. Mitteleur. Fl. [Ascherson & Graebner] 3(6): 736 (1907)

= Orchis majalis subsp. alpestris (Pugsley) Hellm.; Boissiera 7: 382 (1943)

variant :

var. brevifolia (Rchb.f.) Kreutz; Kompend. Eur. Orchid. 50 (2004)

Distribution: Belgium; Germany (Mecklenburg-Vorpommern, Brandenburg, Thüringen, Bayern); Czech Republic

11 BGM CZE GER

Synonyms: hidden

= Dactylorchis cruenta var. brevifolia (Rchb.f.) Verm.; Stud. Dactylorch. 159 (1947)

= Dactylorhiza cruenta var. brevifolia (Rchb.f.) Soó; Nom. Nov. Gen. Dactylorhiza 4 (1962)

= Dactylorhiza majalis var. gracilis (Warnst.) Landwehr; 79 (1975)

= Orchis angustifolia var. brevifolia Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 81 (1899)

= Orchis cruenta f. brevifolia (Rchb.f.) Neuman; Sver. Fl. 631 (1901)

= Orchis latifolia var. angustifolia W.Becker; 68 (1898), nom. illeg.

= Orchis latifolia var. brevifolia Rchb.f.; Icon. Fl. Germ. Helv. [H.G.L.Reichenbach] 13-14: 58 (1851)

variant :

var. parvimajalis (D.Tyteca & Gathoye) comb.ined.

Distribution: SE-France (Isère, Savoie, Haute-Savoie, Hautes-Alpes, Jura, Vosges)

12 FRA

Synonyms: hidden

= Dactylorhiza majalis subsp. parvimajalis (D.Tyteca & Gathoye) Kreutz; Kompend. Eur. Orchid. 50 (2004)

= Dactylorhiza parvimajalis D.Tyteca & Gathoye; J. Eur. Orch. 32 (3/4): 471-511 (2000)

subspecies :

subsp. baltica (Klinge) Senghas; Jahresber. Naturwiss. Vereins Wuppertal 21/22: 126 (1968)

Name: Baltisches Knabenkraut (DE)

Distribution: Denmark; Finland; Poland; Estonia; Latvia; Lithuania; Belarus; ?N-Ukraine; Siberia (Altai, Irkutsk, Krasnoyarsk, W-Siberia); Kazakhstan; ?China (?Xinjiang); European Russia

10 DEN FIN 11 POL 14 BLR EST LAT LIT RUS 30 SIB 32 KAZ

Remarks: status as subspecies confirmed by genetic analysis of Naczk et al. 2024

Synonyms: hidden

= Dactylorchis baltica (Klinge) Verm.; Stud. Dactylorch. 67 (1947)

= Dactylorchis longifolia (Neuman) Verm.; Stud. Dactylorch. 67 (1947)

= Dactylorchis traunsteineri var. dunensis (Rchb.f.) Verm.; Ned. Kruidk. Arch. 56: 226 (1949)

= Dactylorhiza baltica (Klinge) N.I.Orlova; Konspekt. Fl. Pskovsk. Obl. 57 (1970), isonym

= Dactylorhiza baltica (Klinge) N.I.Orlova ex Aver.; Bot. Zhurn. (Moscow & Leningrad) 68: 1163 (1983), isonym

= Dactylorhiza baltica (Klinge) Nevski; Fl. URSS [Komarov et al., eds.] 4: 713 (1935)

= Dactylorhiza baltica var. kuzkenembe Pikner; Ber. Arbeitskreis. Heimische Orchid. 28(2): 260 (2011), et 29(1): 278 (2012)

= Dactylorhiza incarnata f. dunensis (Rchb.f.) P.Delforge; Naturalistes Belges 92(Orchid. 24): 202 (2011), isonym

= Dactylorhiza latifolia subsp. baltica (Klinge) Soó; Nom. Nov. Gen. Dactylorhiza 5 (1962) [nom. inval.]

= Dactylorhiza latifolia var. longifolia (Neuman) Soó; Nom. Nov. Gen. Dactylorhiza 5 (1962)

= Dactylorhiza longifolia (Neuman) Aver.; Bot. Zhurn. (Moscow & Leningrad) 69(6): 875 (1984), isonym

= Dactylorhiza longifolia (Neuman) Verm.; Stud. Dactylorch. 67 (1947)

= Dactylorhiza majalis subsp. baltica (Klinge) E.Nelson; Monogr. & Ikonogr. Orchid.-Gattung Dactylorhiza 91 (1976), isonym

= Dactylorhiza majalis subsp. baltica (Klinge) H.Sund.; Europ. Medit. Orchid., ed. 3: 40 (1980), isonym

= Dactylorhiza traunsteineri var. dunensis (Rchb.f.) Soó; Nom. Nov. Gen. Dactylorhiza 6 (1962)

= Dactylorhiza vironii Kreutz; Ber. Arbeitskreis. Heimische Orchid. 24(1): 90 (2007)

= Orchis baltica (Klinge) A.Fuchs; Ber. Naturwiss. Vereins Schwaben Neuburg 42: 42 (1919), isonym

= Orchis baltica (Klinge) Nevski; Trudy Imp. S.-Peterburgsk. Bot. Sada 17 1: 153, in clavi, et 170 (1898)

= Orchis lapponica subsp. dunensis (Rchb.f.) K.Richt.; Pl. Eur. 1: 271 (1890)

= Orchis latifolia f. baltica (Klinge) M.Schulze; Mitth. Thüring. Bot. Vereins, N. F. 17: 51 (1902)

= Orchis latifolia subsp. baltica Klinge; Trudy Imp. St.-Peterburgsk. Bot. Sada 17(1): 7, 24 (1898), et ex Lehmann, Fl. Pol. Livl. 188 (1895)

= Orchis latifolia subsp. dunensis (Rchb.f.) Krysht.; 420 (1929)

= Orchis latifolia var. dunensis Rchb.f.; Icon. Fl. Germ. Helv. [H.G.L.Reichenbach] 13-14: 59 (1851)

= Orchis latifolia var. tenuior Neuman; Bot. Not. 1909: 240 (1909)

= Orchis longifolia Neuman; Bot. Not. 1909: 241 (1909)

= Orchis longifolia var. gracilis Neuman; Bot. Not. 1909: 241 (1909)

subspecies :

subsp. brennensis (E.Nelson) D.Tyteca & M.Hedrèn; J. Eur. Orch. 52(1): 59 (2020)

Distribution: C-France (Indre)

12 FRA

Synonyms: hidden

= Dactylorhiza brennensis (E.Nelson) D.Tyteca & Gathoye; Orchidophile (Paris) 81: 58 (1988)

= Dactylorhiza elata subsp. brennensis E.Nelson; Monogr. & Ikonogr. Orchid.-Gattung Dactylorhiza 50 (1976)

= Dactylorhiza elata var. brennensis (E.Nelson) P.Delforge; Naturalistes Belges 81(4): 397 (2000)

subspecies :

subsp. elatior (Fr.) M.Hedrén & H.A.Pedersen; Nordic J. Bot. 30: 271 (2012)

Distribution: Sweden (Gotland); Estonia (Saaremaa Isl.); Lithuania

10 SWE 14 EST LIT

Synonyms: hidden

= Dactylorhiza elatior (Fr.) Soó; Nom. Nov. Gen. Dactylorhiza 7 (1962)

= Dactylorhiza osiliensis Pikner; Naturalistes Belges 86(Orchid. 18): 77 (2005)

= Dactylorhiza praetermissa subsp. osiliensis (Pikner) Kreutz; Orchidee (Hamburg) 57(1): 99 (2006)

= Dactylorhiza ruthei subsp. osiliensis (Pikner) Eccarius; Orchideengattung Dactylorhiza 490 (2016)

= Orchis angustifolia subsp. elata (Hartm.) C.Hartm.; Handb. Skand. Fl., ed. 5: 225 (1849)

= Orchis angustifolia var. elata Hartm.; Handb. Skand. Fl., ed. 4: 281 (1843)

= Orchis elatior (Fr.) Afzel. ex Rchb.f.; Icon. Fl. Germ. Helv. [H.G.L.Reichenbach] 13-14: 51 (1851)

= Orchis latifolia subsp. elatior Fr.; Summa Veg. Scand. [Fries] 217 (1845)

= Orchis strictifolia var. elata (Hartm.) Hyl.; Uppsala Univ. Årsskr. 7: 117 (1945)

subspecies :

subsp. kalopissii (E.Nelson) H.A.Pedersen, P.J.Cribb & Rolf Kühn; Field Guide Orchids Europe Medit. 154 (2019)

Name: Kalopissis´ Knabenkraut (DE)

Distribution: SE-Albania; North Macedonia; Bulgaria; N-Greece (mountains from Pindos to Drama)

13 ALB BUL GRC NMA

Conservation Status: IUCN R

Synonyms: hidden

= Dactylorhiza kalopissii E.Nelson; Monogr. & Ikonogr. Orchid.-Gattung Dactylorhiza 43 (1976)

subspecies :

subsp. macedonica (J.Hölz. & Künkele) H.A.Pedersen, P.J.Cribb & Rolf Kühn; Field Guide Orchids Europe Medit. 158 (2019)

Distribution: North Macedonia; Greece (mountains of NC-Greece south to Mt. Olimbos, Rodopi, Mt. Falakron)

13 GRC NMA

Synonyms: hidden

= Dactylorhiza kalopissii f. macedonica (J.Hölz. & Künkele) P.Delforge; Naturalistes Belges 88(Orchid. 20): 246 (2007)

= Dactylorhiza macedonica J.Hölz. & Künkele; Mitt. Arbeitskreises Heimische Orchideen 20(1): 191 (1988)

= Dactylorhizia kalopissii subsp. macedonica (J.Hölz. & Künkele) Kreutz; Kompend. Eur. Orchid. 43 (2004)

subspecies :

subsp. pythagorae (Gölz & H.R.Reinhard) H.A.Pedersen, P.J.Cribb & Rolf Kühn; Field Guide Orchids Europe Medit. 165 (2019)

Distribution: East Aegaean Isl. (Samos)

34 EAI

Synonyms: hidden

= Dactylorhiza kalopissii subsp. pythagorae (Gölz & H.R.Reinhard) Kreutz; Kompend. Eur. Orchid. 43 (-44) (2004)

= Dactylorhiza pythagorae Gölz & H.R.Reinhard; Mitt. Arbeitskreises Heimische Orchideen 24(1): 58, 52 (1992)

subspecies :

subsp. ruthei (M.Schultze ex R.Ruthe) H.Kretzschmar; Orchideen Deutschlands [Kretzschmar] 16 (2008)

Name: Ruthe-Fingerwurz, Ruthe-Fingerknabenkraut (DE)

Distribution: NE-Germany (Mecklenburg-Vorpommern); +NW-Poland

11 GER POL

Remarks: status as subspecies confirmed by genetic analysis of Naczk et al. 2024

Synonyms: hidden

= Dactylorchis ruthei (M.Schultze ex R.Ruthe) Verm.; Stud. Dactylorch. 67 (1947)

= Dactylorhiza ruthei (M.Schultze ex R.Ruthe) Soó; Nom. Nov. Gen. Dactylorhiza 6 (1962)

= Orchis maculata subsp. ruthei (M.Schulze ex R.Ruthe) Hegi; 354 (1909)

= Orchis ruthei M.Schultze ex R.Ruthe; Deutsche Bot. Monatsschr. 15: 237 (1897)

species :

Dactylorhiza maurusia (Em. & Maire) Holub; Folia Geobot. Phytotax. 8(2): 176 (1973)

Distribution: N-Morocco (Rif Mts.); Algeria (Kabylie)

20 ALG MOR

Synonyms: hidden

= Dactylorchis maurusia (Emb. & Maire) Verm.; Stud. Dactylorch. 69 (1947)

= Dactylorhiza battandieri Raynaud; Orchid. Maroc 62 (1985)

= Dactylorhiza maculata subsp. baborica (Maire & Weiller) Soó; Acta Bot. Acad. Sci. Hung. 16 (3-4): 368 (1970 publ. 1971)

= Dactylorhiza maculata subsp. battandieri (Raynaud) H.Baumann & Künkele; Mitt. Arbeitskreises Heimische Orchideen 20: 631 (1988)

= Dactylorhiza maculata subsp. maurusia (Emb. & Maire) Soó; Nom. Nov. Gen. Dactylorhiza 8 (1962)

= Dactylorhiza maurusia (Emb. & Maire) Aver.; Bot. Zhurn. (Moscow & Leningrad) 69(6): 875 (1984), isonym

= Dactylorhiza maurusia (Emb. & Maire) Holub; Folia Geobot. Phytotax. 8: 176 (1973)

= Dactylorhiza maurusia (Emb. & Maire) Raynaud; Orchid. Maroc 59 (1985), isonym

= Orchis incarnata var. xauensis Pau & Font Quer; Iter Marocc. [Font Quer], Exsicc. 1928: no. 56 (1929)

= Orchis latifolia subsp. maurusia (Emb. & Maire) Maire; Cat. Pl. Maroc 1: 151 (1931)

= Orchis maculata subsp. baborica Maire & Weiller; Fl. Afrique N. 6: 309 (1959 publ. 1960), no Latin descr.

= Orchis maculata subsp. maurusia (Emb. & Maire) Maire & Weiller; Fl. Afrique N. 6: 309 (1960)

= Orchis maurusia Emb. & Maire; Arch. Sci. Maroc 3 (Pl. Marocc. Nov. 3): 7 (1930), et Bull. Soc. Sci. Nat. Maroc 11: Nos. 4-6, 109 (1931)

species :

Dactylorhiza nieschalkiorum H.Baumann & Künkele; Mitt. Arbeitskreises Heimische Orchideen 13(2): 259 (1981)

Name: Nieschalks Knabenkraut (DE)

Distribution: NE-Turkey (Bolu, Kastamonu, Bursa, Kütahya)

34 TUR

Remarks: praetermissa agg. Status doubtful, possibly in Anatolia the same as D. osmanica

Conservation Status: IUCN R

Synonyms: hidden

= Dactylorhiza majalis subsp. nieschalkiorum (H.Baumann & Künkele) H.A.Pedersen, P.J.Cribb & Rolf Kühn; Field Guide Orchids Europe Medit. 159 (2019)

species :

Dactylorhiza osmanica (Klinge) Soó; Ann. Univ. Sci. Budapest. Rolando Eötvös, Sect. Biol. 3: 341, 355 (1960)

Name: Osmanisches Knabenkraut

Distribution: Siberia (Altai, Buryatia, Chita, Tuva); Turkmenistan; Uzbekistan; Kyrgyzstan; Tajikistan; Northern Caucasus; Transcaucasus; Turkey; Iran; Lebanon; Syria

30 SIB 32 KGZ TKM TZK UZB 33 NCS 34 IRN LEB SYR TUR

Remarks: incarnata agg.

Synonyms: hidden

= Dactylorhiza cataonica (H.Fleischm.) Holub; Preslia 36: 252 (1964)

= Dactylorhiza caucasica var. cataonica (H.Fleischm.) Soó; Nom. Nov. Gen. Dactylorhiza 5 (1962)

= Dactylorhiza cilicica (Klinge) P.F.Hunt & Summerh.; Watsonia 6: 130 (1965), isonym

= Dactylorhiza cilicica (Klinge) Soó; Ann. Univ. Sci. Budapest. Rolando Eötvös, Sect. Biol., 3: 340 (1960), et Nom. Nov. Gen. Dactylorhiza 4 (1962)

= Dactylorhiza euxina var. cataonica (H.Fleischm.) Soó; [nom. inval.]

= Dactylorhiza incarnata subsp. cilicica (Klinge) H.Sund.; Europ. Medit. Orchid., ed. 2: 45 (1975)

= Dactylorhiza maculata subsp. osmanica (Klinge) E.Nelson; Monogr. & Ikonogr. Orchid.-Gattung Dactylorhiza 98 (1976)

= Dactylorhiza maculata subsp. osmanica (Klinge) H.Sund.; Europ. Medit. Orchid., ed. 3: 41 (1980)

= Dactylorhiza olocheilos (Boiss.) Aver.; Bot. Zhurn. (Moscow & Leningrad) 69(6): 875 (1984)

= Dactylorhiza osmanica (Klinge) P.F.Hunt & Summerh.; Watsonia 6: 130 (1965), isonym

= Orchis cataonica H.Fleischm.; Ann. K.K. Naturhist. Hofmus. 28: 34 (1914)

= Orchis cilicica (Klinge) Schltr.; Repert. Spec. Nov. Regni Veg. Sonderbeih. A: 1: 178 (1927)

= Orchis cilicica Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17 1: 156, in clavi, et 187 (1898)

= Orchis holocheilos (Boiss.) G.Keller & Soó; Repert. Spec. Nov. Regni Veg. Sonderbeih. A: 2: 228 (1933)

= Orchis incarnata var. latifolia Freyn; Bull. Herb. Boissier, Sér. 2, 1: 285 (1901), nom. illeg.

= Orchis incarnata var. olocheilos Boiss.; Fl. Orient. [Boissier] 5: 71 (1882)

= Orchis incarnata var. stenophylla Freyn; Bull. Herb. Boissier, Sér. 2, 1: 285 (1901)

= Orchis olocheilos (Boiss.) Soó; Repert. Spec. Nov. Regni Veg. 24: 30 (1927)

= Orchis olocheilos var. osmanica (Klinge) Soó; Bot. Arch. 23: 73 (1928)

= Orchis orientalis subsp. cilicica Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(1): 36, 37 (1898)

= Orchis orientalis subsp. osmanica Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(1): 157, 188 (1898)

= Orchis osmanica (Klinge) A.W.Hill; Index Kew., Suppl. 9: 194 (1938), isonym

= Orchis osmanica (Klinge) Lipsky; Trudy Imp. S.-Peterburgsk. Bot. Sada 17 1: 157, in clavi, et 188 (1898)

variant :

var. anatolica (E.Nelson) Renz & Taubenheim; Notes Roy. Bot. Gard. Edinburgh 41: 271 (1983)

Distribution: Turkey (SC-Anatolia)

34 TUR

Synonyms: hidden

= Dactylorhiza elata subsp. anatolica E.Nelson; Monogr. & Ikonogr. Orchid.-Gattung Dactylorhiza 45 (1976), et Taxon 28 (5-6): 593 (1979)

= Dactylorhiza anatolica (E.Nelson) Kreutz; Guide Orchids Europe N. Afr. Middle E. 359 (2024)

= Dactylorhiza osmanica subsp. anatolica (E.Nelson) Eccarius; Orchideengattung Dactylorhiza 232 (2016)

species :

Dactylorhiza praetermissa (Druce) Soó; Nom. Nov. Gen. Dactylorhiza 5 (1962)

subspecies :

subsp. praetermissa

Name: Übersehene Fingerwurz, Übersehenes Fingerknabenkraut (DE)

Distribution: England (England, Wales, Isles of Scilly); Denmark; Norway; Netherlands; Belgium; Luxembourg; Germany (Hessen [I], Hamburg, Mecklenburg-Vorpommern [I], Niedersachsen, Nordrhein-Westfalen, Rheinland-Pfalz, Schleswig-Holstein, Saarland); France; Channel Isl. (Alderney, Guernsey, Jersey); Estonia; NE-Canada [I] (Tilt Cove [I])

10 DEN GRB NOR 11 BGM GER LUX NET 12 CIS FRA 14 EST 72 cdn

Synonyms: hidden

= Dactylorchis praetermissa (Druce) Verm.; Stud. Dactylorch. 67 (1947)

= Dactylorchis praetermissa var. cuspidata Deinum ex Verm.; Ned. Kruidk. Arch. 56: 230 (1949)

= Dactylorchis praetermissa var. elata Verm.; Ned. Kruidk. Arch. 56: 229 (1949)

= Dactylorchis praetermissa var. longifolia Verm.; Ned. Kruidk. Arch. 56: 230 (1949)

= Dactylorchis praetermissa var. macrantha (Sipkes) Verm.; Ned. Kruidk. Arch. 56: 229 (1949)

= Dactylorchis praetermissa var. parvifructa Verm.; Ned. Kruidk. Arch. 56: 231 (1949)

= Dactylorchis praetermissa var. tenera Verm.; Ned. Kruidk. Arch. 56: 230 (1949)

= Dactylorchis praetermissa var. triangularis Verm.; Ned. Kruidk. Arch. 56: 231 (1949)

= Dactylorhiza incarnata subsp. praetermissa (Druce) H.Sund.; Europ. Medit. Orchid., ed. 3: 40 (1980)

= Dactylorhiza majalis subsp. praetermissa (Druce) D.M.Moore & Soó; J. Linn. Soc., Bot. 76: 367 (1978)

= Dactylorhiza majalis var. integrata (E.G.Camus) H.A.Pedersen; J. Eur. Orch. 43: 80 (2011)

= Dactylorhiza majalis var. macrantha (Sipkes) R.M.Bateman & Denholm; Watsonia 14(4): 372 (1983)

= Dactylorhiza majalis var. praetermissa (Druce) R.M.Bateman & Denholm; Watsonia 14(4): 371 (1983)

= Dactylorhiza praetermissa var. bracteosa Kreutz; J. Eur. Orch. 48: 321 (2016)

= Dactylorhiza praetermissa var. cuspidata (Deinum ex Verm.) Soó; Nom. Nov. Gen. Dactylorhiza 5 (1962)

= Dactylorhiza praetermissa var. elata (Verm.) Soó; Nom. Nov. Gen. Dactylorhiza 5 (1962)

= Dactylorhiza praetermissa var. robusta Bussche; Liparis 7: 32 (2001)

= Orchis incarnata var. albiflora Lecoq & Lamotte ex Thielens; Bull. Soc. Roy. Bot. Belgique 12: 82 (1873)

= Orchis praetermissa Druce; Rep. Bot. Soc. Exch. Club Brit. Isles 3: 341 (1913 publ. 1914)

= Orchis praetermissa f. macrantha (Sipkes) Soó; Bot. Arch. 23: 69 (1938)

= Orchis praetermissa var. albiflora Druce; Rep. Bot. Soc. Exch. Club Brit. Isles 5: 53 (1919)

= Orchis praetermissa var. macrantha Sipkes; Levende Natuur 26: 52 (1921)

variant :

var. integrata (E.G.Camus ex Fourcy) D.Tyteca & Gathoye; Belg. J. Bot. 126(1): 93 (1993)

Distribution: N-France; Belgium; Netherlands; Luxembourg

11 BGM LUX NET 12 FRA

Synonyms: hidden

= Dactylorhiza integrata (E.G.Camus) Aver.; Bot. Zhurn. (Moscow & Leningrad) 69(6): 875 (1984)

= Dactylorhiza majalis subsp. integrata (E.G.Camus) H.A.Pedersen; J. Eur. Orch. 41(3/4): 504 (2009)

= Dactylorhiza praetermissa subsp. integrata (E.G.Camus ex Fourcy) Soó; Nom. Nov. Gen. Dactylorhiza 5 (1962)

= Orchis incarnata subsp. integrata (E.G.Camus) E.G.Camus; J. Bot. [Morot] 4: 157 (1892)

= Orchis incarnata var. integrata E.G.Camus ex Fourcy; Vadem. Herb. Paris [de Fourcy], éd. 6: 325 (1891)

= Orchis integrata (E.G.Camus) E.G.Camus; Monogr. Orchid. France 48 (1894)

= Orchis praetermissa subsp. integrata (E.G.Camus) E.G.Camus & A.Camus; Iconogr. Orchid. Europe 1: 219 (1928)

variant :

var. junialis (Verm.) Senghas; Jahresber. Naturwiss. Vereins Wuppertal 21/22: 126 (1968)

Distribution: England; NW-Germany; NE-Canada [I] (St. Johns [I])

10 GRB 11 GER 72 cdn

Synonyms: hidden

= Dactylorchis praetermissa var. junialis (Verm.) Verm.; Stud. Dactylorch. 67 (1947), nom. inval., et Ned. Kruidk. Arch. 56: 229 (1949)

= Dactylorhiza majalis subsp. pardalina (Pugsley) E.Nelson; Monogr. & Ikonogr. Orchid.-Gattung Dactylorhiza 88 (1976) [without basionym page]

= Dactylorhiza majalis var. junialis (Verm.) H.A.Pedersen; Flor. Med. Fra Dansk Herb. I, URT 23(2): 64-71 (1999), isonym

= Dactylorhiza majalis var. junialis (Verm.) R.M.Bateman & Denholm; Watsonia 14: 371 (1983)

= Dactylorhiza praetermissa f. junialis (Verm.) P.D.Sell; Fl. Gr. Brit. Ireland 5: 365 (1996)

= Dactylorhiza praetermissa subsp. junialis (Verm.) Soó; Nom. Nov. Gen. Dactylorhiza 5 (1962)

= Dactylorhiza praetermissa var. junialis (Verm.) H.A.Pedersen; J. Eur. Orch. 41(3/4): 504 (2009), nom. illeg.

= Dactylorhiza ×pardalina (Pugsley) Aver.; Bot. Zhurn. (Moscow & Leningrad) 71(1): 93 (1986)

= Orchis latifolia var. junialis Verm.; Ned. Kruidk. Arch. 43: 404, 409 (1933)

= Orchis pardalina Pugsley; Proc. Linn. Soc. London Sess. 146, 97 (1934), in clavi, et J. Linn. Soc., Lond., Bot. 49: 581 (1935) [descr. ampl.]

= Orchis praetermissa subsp. junialis (Verm.) Verm.; Chron. Bot. 4(2): 108 (1934)

variant :

var. maculosa D.Tyteca & Gathoye; Orchidophile (Paris) 92: 112 (1990)

Distribution: Belgium

11 BGM

subspecies :

subsp. schoenophila R.M.Bateman & Denholm; New J. Bot. 2(1): 47 (2012), nom. nov.

Distribution: England (S- & E-England)

10 GRB

Synonyms: hidden

= Dactylorhiza majalis var. bowmannii Jenk.; (1995)

= Dactylorhiza schoenophila (R.M.Bateman & Denholm) Kreutz; Guide Orchids Europe N. Afr. Middle E. 407 (2024)

species :

Dactylorhiza purpurella (T.Stephenson & T.A.Stephenson) Soó; Nom. Nov. Gen. Dactylorhiza 5 (1962)

Name: Purpurblütige Fingerwurz, Purpurblütiges Fingerknabenkraut (DE), Northern Marsh-orchid (EN)

Distribution: Faeroe Isl.; England (N-England, Wales, Isle of Man, Scotland, Outer Hebrides, Orkney Isl., Shetland Isl.); Ireland (N- & SE-Ireland, Northern Ireland); S-Sweden; Denmark (Jutland); Norway; Germany [I] (Bayern [I])

10 DEN FOR GRB IRE NOR SWE 11 ger

Synonyms: hidden

= Dactylorchis purpurella (T.Stephenson & T.A.Stephenson) Verm.; Stud. Dactylorch. 67 (1947)

= Dactylorhiza incarnata var. obscura (Höppner) Soó; Nom. Nov. Gen. Dactylorhiza 3 (1962)

= Dactylorhiza majalis subsp. purpurella (T.Stephenson & T.A.Stephenson) D.M.Moore & Soó; Bot. J. Linn. Soc. 76(4): 367 (1978)

= Dactylorhiza majalis var. purpurella (T.Stephenson & T.A.Stephenson) R.M.Bateman & Denholm; Watsonia 14: 370 (1983)

= Orchis incarnata var. obscura Höppner; Fl. Niederrheins, ed. 3: 106 (1913)

= Orchis purpurella T.Stephenson & T.A.Stephenson; J. Bot. 58: 164 (1920)

variant :

var. atrata (A.J.Richards) A.J.Richards & L.Lewis; J. Eur. Orch. 50(1): 85 (2018)

Distribution: England (N-England, Wales)

10 GRB

Synonyms: hidden

= Dactylorhiza majalis var. atrata A.J.Richards; Watsonia 18(3): 308 (1991)

variant :

var. cambrensis (R.H.Roberts) R.M.Bateman & Denholm; Watsonia 14(4): 368 (1983), et Orchids Brit. Isles 373 (2005)

Distribution: England (N-England, Scotland, Wales); Ireland (Northern Ireland, Ireland)

10 GRB IRE

Synonyms: hidden

= Dactylorchis majalis subsp. cambrensis R.H.Roberts; Watsonia 5(1): 41 (1961)

= Dactylorhiza cambrensis (R.H.Roberts) Aver.; Bot. Zhurn. (Moscow & Leningrad) 69(6): 875 (1984)

= Dactylorhiza comosa subsp. cambrensis (R.H.Roberts) P.D.Sell; Fl. Gr. Brit. Ireland 5: 365 (1996)

= Dactylorhiza latifolia subsp. cambrensis (R.H.Roberts) Soó; Nom. Nov. Gen. Dactylorhiza 5 (1962)

= Dactylorhiza majalis subsp. cambrensis (R.H.Roberts) E.Nelson; Monogr. & Ikonogr. Orchid.-Gattung Dactylorhiza 89 (1976)

= Dactylorhiza majalis var. cambrensis (R.H.Roberts) H.A.Pedersen; J. Eur. Orch. 39(2): 350 (2007), isonym

= Dactylorhiza majalis var. eboracensis (Godfery) R.M.Bateman & Denholm; Watsonia 14: 373 (1983)

= Dactylorhiza purpurella var. cambrensis (R.H.Roberts) R.M.Bateman & Denholm; Orchids Brit. Isles [Foley] 373 (2005)

= Orchis latifolia var. eboracensis Godfery; Mitt. Geogr. Ges. Thüringen (Jena) 2: 220 (1885)

variant :

var. crassifolia (T.Stephenson & T.A.Stephenson) Landwehr; Wilde Orchideen Eur. 80 (1975)

Distribution: Faeroe Isl.; England (N-England, Wales, Scotland); Ireland; Norway; Denmark (Jutland)

10 DEN FOR GRB IRE NOR

Synonyms: hidden

= Orchis purpurella var. crassifolia T.Stephenson; Rep. Bot. Soc. Exch. Club Brit. Isles 11: 356 (1937)

= Dactylorhiza majalis var. crassifolia (T.Stephenson & T.A.Stephenson) Landwehr; Orchideeën 37: 80 (1975)

variant :

var. maculosa (T.Stephenson & T.A.Stephenson) Kreutz, H.Dekker & L.Lewis; J. Eur. Orch. 48(2-4): 304 (2016)

Distribution: England (Scotland); Netherlands

10 GRB 11 NET

Synonyms: hidden

= Dactylorhiza britannica Kreutz; Guide Orchids Europe N. Afr. Middle E. 382 (2024), nom. nov. for O. purpurella var. maculosa

= Dactylorhiza majalis var. maculosa (T.Stephenson) R.M.Bateman & Denholm; Watsonia 14(4): 370 (1983)

= Orchis purpurella var. maculosa T.Stephenson; Rep. Bot. Soc. Exch. Club Brit. Isles 11: 355 (1937)

variant :

var. majaliformis (E.Nelson) Kreutz; Eurorchis 18: 95 (2006)

Distribution: England (Scotland, Outer Hebrides, Orkney Isl.)

10 GRB

Synonyms: hidden

= Dactylorhiza purpurella subsp. majaliformis E.Nelson; Monogr. & Ikonogr. Orchid.-Gattung Dactylorhiza 77 (1976), without type, et Taxon 28(56): 593 (1979)

= Dactylorhiza purpurella subsp. majaliformis E.Nelson ex Løjtnant; Bot. Tidsskr. 74: 176 (1979), nom. illeg.

variant :

var. vadorum (Kreutz & H.Dekker) comb.ined.

Distribution: Denmark; ?Germany

10 DEN

Synonyms: hidden

= Dactylorhiza vadorum Kreutz & H.Dekker; Gorteria 38: 176 (2016)

= Dactylorhiza vadorum var. picturata Kreutz & H.Dekker; Gorteria 38: 184 (2016)

species :

Dactylorhiza romana (Sebast.) Soó; Nom. Nov. Gen. Dactylorhiza 3 (1962)

subspecies :

subsp. romana

Name: Römisches Knabenkraut (DE); Roman Spotted-orchid (EN)

Distribution: Italy; Sicily; Croatia; Montenegro; Serbia; Kosovo; North Macedonia; Albania; Bulgaria; Greece (scattered mainland, Thasos, Kerkyra, Ionian Isl., Sporades, Skiros, Kiklades); Crete; European Turkey; Crimea; East Aegaean Isl. (Lesvos, Chios, Samos, Kos, Rhodos); Turkey; Cyprus; N-Israel

13 ALB BUL CRO GRC ITA KOS KRI MNT NMA SIC SRB TUE 14 KRY 34 CYP EAI ISR TUR

Remarks: sambucina agg.

Synonyms: hidden

= Dactylorchis romana (Sebast.) Verm.; Stud. Dactylorch. 65 (1947)

= Dactylorhiza sambucina f. zimmermannii (A.Camus) P.Delforge; Naturalistes Belges 88(Orchid. 20): 246 (2007)

= Dactylorhiza sambucina subsp. pseudosambucina (Ten.) H.Sund.; Europ. Medit. Orchid., ed. 3: 40 (1980)

= Dactylorhiza sambucina subsp. romana (Sebast.) Bornm.; Beibl. Bot. Jahrb. Syst. 140: 126 (1928)

= Dactylorhiza sulphurea (Spreng.) Franco; J. Linn. Soc., Bot. 76: 336 (1978)

= Dactylorhiza sulphurea subsp. pseudosambucina (Ten.) Franco; Bot. J. Linn. Soc. 76(4): 366 (1978)

= Orchis bracteata Ten.; Fl. Napol. 1(Prodr.): LII (1812), nom. illeg.

= Orchis fasciculata var. obtusifolia Tineo; Fl. Sicul. Syn. 2: 875 (1845)

= Orchis lucana Spreng.; Pl. Min. Cogn. Pug. 2: 79 (1815)

= Orchis mediterranea Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(1): 163 (1898), nom. illeg.

= Orchis mediterranea subsp. pseudosambucina (Ten.) KIinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(1): 151, 164 (1898)

= Orchis natalis Tineo; Pl. Rar. Sicil. 8 (1846)

= Orchis ochroleuca Schur; Enum. Pl. Transsilv. 641 (1866), nom. illeg.

= Orchis pseudosambucina subsp. romana (Sebast.) Gand.; Nov. Consp. Fl. Eur. 462 (1910)

= Orchis pseudosambucina Ten.; Fl. Napol. 2: 284 (1820)

= Orchis pseudosambucina var. natalis (Tineo) Nyman; Consp. Fl. Eur. 693 (1882)

= Orchis romana lusus bicolor G.Keller; Repert. Spec. Nov. Regni Veg. Sonderbeih. A: 2: 206 (1933)

= Orchis romana Sebast.; Roman. Pl. Fasc. Prim. 1: 12 (1813)

= Orchis romana var. incarnata E.G.Camus & A.Camus; Iconogr. Orchid. Europe 214 (1928)

= Orchis romana var. lutea E.G.Camus & A.Camus; Iconogr. Orchid. Europe 214 (1928)

= Orchis sambucina Brot.; Fl. Lusit. 1: 21 (1800 publ. 1804)

= Orchis sambucina var. zimmermannii A.Camus; Iconogr. Orchid. Europe 212 (1928)

= Orchis sulphurea Spreng.; Syst. Veg., ed. 16 [Sprengel] 3: 688 (1826) [in syn.]

subspecies :

subsp. georgica (Klotzsch) Soó ex Renz & Taubenheim; Notes Roy. Bot. Gard. Edinburgh 41(2): 271 (1983)

Name: Georgisches Knabenkraut (DE)

Distribution: Turkmenistan; Northern Caucasus; Georgia [Caucasus] (Racha-Lechkhumi, Adzharia, Kartli, Kakheti, Trialeti); Armenia; Azerbaijan (incl. Talysh); E-Turkey; N-Iran; N-Iraq

32 TKM 33 ARM AZE GEO NCS 34 IRN IRQ TUR

Synonyms: hidden

= Dactylorchis flavescens (K.Koch) Verm.; Stud. Dactylorch. 65 (1947)

= Dactylorhiza flavescens (K.Koch) H.Baumann & Künkele; Mitt. Arbeitskreises Heimische Orchideen 13(2): 237 (1981), isonym

= Dactylorhiza flavescens (K.Koch) Holub; Folia Geobot. Phytotax. 11(1): 83 (1976)

= Dactylorhiza flavescens (Koch) É.M.Seifulin; Czerep., Opred. khvoshcheobraznykh. rastenii Turkmenistana 278 (1978) [isonym, isonym]

= Dactylorhiza ruprechtii Aver.; Bot. Zhurn. (Moscow & Leningrad) 68(4): 537 (1983)

= Dactylorhiza sambucina subsp. georgica (Klinge) H.Sund.; Europ. Medit. Orchid., ed. 3: 40 (1980)

= Dactylorhiza sulphurea subsp. georgica (Klinge) Herrero & Ó.Sánchez; Doc. Jard. Bot. Atlántico 7: 80 (2010)

= Orchis flavescens K.Koch; Linnaea 22: 281 (1850)

= Orchis georgica (Klinge) Lipsky; Trudy Imp. S.-Peterburgsk. Bot. Sada 17 1: 151, in clavi, et 166 (1898)

= Orchis georgica (Klinge) Medw.; Trudy Tiflissk. Bot. Sada 18: 473 (1919), in clavi

= Orchis mediterranea subsp. georgica Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(1): 19 (1898)

= Orchis pseudosambucina subsp. georgica Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(1): 151 (1898)

= Orchis romana lusus flavescens (K.Koch) Soó; Repert. Spec. Nov. Regni Veg. 24: 30 (1927)

= Orchis romana lusus tenuifolia Soó; Repert. Spec. Nov. Regni Veg. 24: 30 (1927)

= Orchis romana subsp. georgica (Klinge) E.G.Camus; Monogr. Orchid. 172 (1908)

= Orchis romana var. georgica (Klinge) Schltr.; Repert. Spec. Nov. Regni Veg. Sonderbeih. A: 1: 187 (1927)

= Orchis tenuifolia K.Koch; Linnaea 22: 281 (1850)

subspecies :

subsp. guimaraesii (E.G.Camus) H.A.Pedersen; Bot. J. Linn. Soc. 152(4): 426 (2006)

Distribution: N-Portugal; Spain (C- & S-Spain); Baleares; Sicily (Messina); Morocco; N-Algeria

12 BAL POR SPA 13 SIC 20 ALG MOR

Synonyms: hidden

= Dactylorhiza guimaraesii (E.G.Camus) P.Delforge; Naturalistes Belges 96(Orchid. 28): 15 (2015)

= Orchis guimaraesii (E.G.Camus) Rivas Goday; Farmacognosia 4(6): 193 (1945)

= Orchis romana lusus sulphurea (Link) Soó; Bot. Arch. 23: 63 (1929)

= Orchis romana var. guimaraesii E.G.Camus; Iconogr. Orchid. Europe 1: 216 (1928)

subspecies :

subsp. libanotica (Mouterde) Kreutz; Ber. Arbeitskreis. Heimische Orchid. 24(1): 154 (2007)

Distribution: Lebanon; N-Israel

34 ISR LEB

Synonyms: hidden

= Dactylorhiza libanotica (Mouterde) Aver.; Bot. Zhurn. (Moscow & Leningrad) 69(6): 876 (1984)

= Orchis romana subsp. libanotica Mouterde; Nouv. Fl. Liban Syrie 1: 342 (1966)

subspecies :

subsp. markusii (Tineo) Holub; Folia Geobot. Phytotax. 19(2): 214 (1984)

Name: Sizilianisches Knabenkraut (DE)

Distribution: S-Italy (Calabria); Sicily (Palermo)

13 ITA SIC

Remarks: records for the Baleares refer to ssp. guimaraesii, those for Sardinia to D. insularis

Synonyms: hidden

= Dactylorhiza markusii (Tineo) H.Baumann & Künkele; Mitt. Arbeitskreises Heimische Orchideen 13(4): 461 (1981)

= Dactylorhiza romana subsp. siciliensis (Klinge) Soó; Nom. Nov. Gen. Dactylorhiza 3 (1962)

= Dactylorhiza romana var. markusii (Tineo) Soó; Nom. Nov. Gen. Dactylorhiza 3 (1962)

= Dactylorhiza romana var. sicula (Tineo) Soó; Nom. Nov. Gen. Dactylorhiza 3 (1962)

= Dactylorhiza sambucina subsp. siciliensis (Klinge) H.Sund.; Europ. Medit. Orchid., ed. 3: 40 (1980)

= Dactylorhiza siciliensis (Klinge) E.Nelson; Monogr. & Ikonogr. Orchid.-Gattung Dactylorhiza 105 (1976) [without basionym page]

= Dactylorhiza sicula (Tineo ex Rchb.f.) Aver.; Bot. Zhurn. (Moscow & Leningrad) 69(6): 875 (1984)

= Dactylorhiza sulphurea (Link) Franco; Bot. J. Linn. Soc. 76: 366 (1978), nom. inval.

= Dactylorhiza sulphurea subsp. siciliensis (Klinge) Franco; J. Linn. Soc., Bot. 76(4): 367 (1978)

= Orchis markusii Tineo; Pl. Rar. Sicil. 9 (1846)

= Orchis mediterranea subsp. siciliensis Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(1): 165, 166 (1898)

= Orchis pseudosambucina subsp. markusii (Tineo) Gand.; Nov. Consp. Fl. Eur. 462 (1910), isonym

= Orchis pseudosambucina subsp. markusii (Tineo) Nyman; Consp. Fl. Eur. 4: 693 (1882)

= Orchis pseudosambucina subsp. sicula (Tineo) K.Richt.; Pl. Eur. 1: 271 (1890)

= Orchis pseudosambucina subsp. sulphurea Link; Nov. Consp. Fl. Eur. 462 (1910)

= Orchis romana f. markusii (Tineo) Soó; Repert. Spec. Nov. Regni Veg. 24: 30 (1927)

= Orchis romana var. markusii (Tineo) E.G.Camus, Bergon & A.Camus; Monogr. Orchid. 171 (1908)

= Orchis romana var. markusii (Tineo) Rivas Goday; Farmacognosia 4(6): 194, 195 (1945), isonym

= Orchis siciliensis (Klinge) A.W.Hill; Index Kew., Suppl. 9: 194 (1938)

= Orchis sicula Tineo; Pl. Rar. Sicil. 8 (1846)

= Orchis sulphurea Link; J. Bot. [Schrader] 1806: 1 (1806), et Nomencl. Bot. [Steudel], ed. 1: 225 (1821)

= Orchis sulphurea var. markusii (Tineo) Maire; Cat. Pl. Maroc 1: 151 (1931)

species :

Dactylorhiza saccifera (Brongn.) Soó; Nom. Nov. Gen. Dactylorhiza 8 (1962)

subspecies :

subsp. saccifera

Name: Langähriges Knabenkraut (DE)

Distribution: Croatia; Bosnia & Hercegovina; Albania; Romania; Bulgaria; Greece (widespread mountains of mainland, Thasos, Evvoia); European Turkey; Turkey (NE-Anatolia)

13 ALB BIH BUL CRO GRC ROM TUE 34 TUR

Remarks: maculata agg.

Synonyms: hidden

= Dactylorchis maculata subsp. saccifera (Brongn.) Verm.; Stud. Dactylorch. 68 (1947)

= Dactylorchis maculata var. heuffelii Borsos & Soó; Acta Bot. Acad. Sci. Hung. 5: 324 (1959)

= Dactylorchis saccifera (Brongn.) Verm.; Fl. Neerl. 1: Afl. 5, 87 (1958)

= Dactylorhiza fuchsii var. nesogenes (Briq.) Soó; Nom. Nov. Gen. Dactylorhiza 8 (1962)

= Dactylorhiza maculata subsp. saccifera (Brongn.) Diklic; Fl. Srbije 8: 77 (1976)

= Dactylorhiza maculata subsp. saccifera (Brongn.) H.Sund.; Europ. Medit. Orchid., ed. 3: 40 (1980)

= Dactylorhiza maculata var. orophila (Briq.) Soó; Nom. Nov. Gen. Dactylorhiza 7 (1962)

= Dactylorhiza saccifera subsp. macrostachys (Tineo) Niketic & Djordjevic; Bull. Nat. Hist. Mus. Belgrade 11: 104 (2018)

= Dactylorhiza saccifera var. heuffelii (Borsos & Soó) Soó; Nom. Nov. Gen. Dactylorhiza 8 (1962)

= Dactylorhiza saccifera var. lanceolata Landwehr; Wilde Orchid. Eur. 2: 556 (1977)

= Orchis basilica subsp. saccifera (Brongn.) Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(1): 157 (1898)

= Orchis bonanniana Tod.; Orchid. Sicul. 57 (1842)

= Orchis comosa Schur; Enum. Pl. Transsilv. 641 (1866)

= Orchis macrostachys Tineo; Pl. Rar. Sicil. 7 (1846)

= Orchis maculata proles macrostachys (Tineo) Asch. & Graebn.; Syn. Mitteleur. Fl. [Ascherson & Graebner] 3: 748 (1907)

= Orchis maculata proles nesogenes (Briq.) Rouy; Fl. France [Rouy & Foucaud] 13: 154 (1912)

= Orchis maculata subsp. macrostachys (Tineo) Soó; Repert. Spec. Nov. Regni Veg. 24: 32 (1927)

= Orchis maculata subsp. saccifera (Brongn.) E.G.Camus; Iconogr. Orchid. Europe Text(1): 242 (1928)

= Orchis maculata subsp. saccifera (Brongn.) K.Richt.; Pl. Eur. 1: 272 (1890)

= Orchis maculata var. bonanniana (Tod.) Nyman; Consp. Fl. Eur. 693 (1882)

= Orchis maculata var. cartalinoides Klinge ex H.Fleischm.; Mitt. Naturwiss. Vereines Steiermark 45: 176 (1908 publ. 1909)

= Orchis maculata var. heuffelii (Borsos & Soó) Pauca; Fl. Reipubl. Popul. Roman. (Fl. Republ. Socialist. Romania) 12: 698 (1972)

= Orchis maculata var. kortiathica Formánek; Verh. Naturf. Vereins Brünn 32: 26 (1894)

= Orchis maculata var. nesogenes Briq.; Prodr. Fl. Corse 1: 371 (1910)

= Orchis maculata var. orophila Briq.; Prodr. Fl. Corse 1: 370 (1912)

= Orchis maculata var. pyramidata Schur; Enum. Pl. Transsilv. 643 (1866)

= Orchis maculata var. saccifera (Brongn.) Rchb.f.; Icon. Fl. Germ. Helv. [H.G.L.Reichenbach] 13/14: 67 (1851)

= Orchis maculata var. turfosa Schur; Verh. Mitth. Siebenbürg. Vereins Naturwiss. Hermannstadt 4: 72 (1853)

= Orchis nesogenes (Briq.) Rouy; Fl. France [Rouy & Foucaud] 13: 154 (1912)

= Orchis saccifera Brongn.; Expéd. Sci. Morée, Bot. 259 (1832)

subspecies :

subsp. cyrnea (Foelsche & Cord-Landwehr) comb.ined.

Distribution: Corsica

12 COR

Synonyms: hidden

= Dactylorhiza cyrnea Foelsche & Cord-Landwehr; J. Eur. Orch. 44(1): 179 (2012)

subspecies :

subsp. gervasiana (Tod.) Kreutz; Kompend. Eur. Orchid. 54 (2004)

Distribution: Italy (C- & S-Italy); Sicily (N-Sicily)

13 ITA SIC

Synonyms: hidden

= Dactylorhiza gervasiana (Tod.) H.Baumann & Künkele; Mitt. Arbeitskreises Heimische Orchideen 13(3): 346 (1981)

= Orchis gervasiana Tod.; Orchid. Sicul. 57 (1842)

= Orchis maculata var. gervasiana (Tod.) Nyman; Consp. Fl. Eur. 693 (1882)

subspecies :

subsp. taurica Kreutz & Isler; Res. Türkiye Fl. 3a: 135 (2022)

Distribution: ?N-Greece; Turkey (SC-Anatolia)

34 TUR

Synonyms: hidden

= Dactylorhiza taurica (Kreutz & Isler) Kreutz; Guide Orchids Europe N. Afr. Middle E. 449 (2024)

species :

Dactylorhiza sajanensis Stepanov; Red Book Krasnoyarsk. Territ. [ed. 3] 304 (2022)

Distribution: Siberia (Krasnoyarsk)

30 SIB

Remarks: provisionally accepted name

species :

Dactylorhiza salina (Turcz. ex Lindl.) Soó; Nom. Nov. Gen. Dactylorhiza 4 (1962)

Distribution: Siberia (Altai, Buryatia, Chita, Krasnoyarsk); Russian Far East (Amur); Kazakhstan; Kyrgyzstan; Northern Caucasus; Georgia [Caucasus]; Armenia; Mongolia; Pakistan (Karakorum)

30 SIB 31 RFE 32 KAZ KGZ 33 ARM GEO NCS 37 MON 40 PAK

Synonyms: hidden

= Dactylorchis salina (Turcz. ex Lindl.) Verm.; Stud. Dactylorch. 66 (1947)

= Dactylorhiza cruenta subsp. salina (Turcz. ex Lindl.) E.Nelson; Monogr. & Ikonogr. Orchid.-Gattung Dactylorhiza 70 (1976)

= Dactylorhiza knorringiana (Kraenzl.) Dickoré; Stapfia 39: 221 (1995) isonym

= Dactylorhiza knorringiana (Kraenzl.) Ikonn.; Novosti Sist. Vyssh. Rast. 6: 267 (1970)

= Dactylorhiza umbrosa subsp. knorringiana (Kraenzl.) Soó; Ann. Univ. Sci. Budapest. Rolando Eötvös, Sect. Biol., 11: 68 (1969)

= Dactylorhiza umbrosa var. knorringiana (Kraenzl.) Soó; Ann. Univ. Sci. Budapest. Rolando Eötvös, Sect. Biol. 5: 4 (1962)

= Orchis incarnata var. knorringiana Kraenzl.; Repert. Spec. Nov. Regni Veg. 65: 34 (1931)

= Orchis knorringiana (Kraenzl.) Czerniak.; Fl. Kirgizsk. S.S.R. 3: 138 (1951)

= Orchis orientalis subsp. salina (Turcz. ex Lindl.) Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(1): 185 (1898), nom. superfl.

= Orchis salina Turcz. ex Lindl.; Gen. Sp. Orchid. Pl. 259 (1835)

species :

Dactylorhiza sambucina (L.) Soó; Nom. Nov. Gen. Dactylorhiza 3 (1962)

Name: Holunder-Knabenkraut

Distribution: Denmark; Norway; Sweden; Finland; Germany (Brandenburg, Baden-Württemberg, Bayern, +Hessen, Rheinland-Pfalz, Sachsen, Sachsen-Anhalt, Thüringen); Switzerland; Austria; Poland; Czech Republic; Slovakia; Hungary; Portugal; Spain; Andorra; France; Corsica; Sardinia; Italy; Sicily; Slovenia; Croatia; Bosnia & Hercegovina; Montenegro; Serbia; Kosovo; North Macedonia; Albania; Romania; Bulgaria; Greece (mountains of mainland incl. Peloponnisos); Estonia; +Belarus; Ukraine; European Russia (W-European Russia, C-European Russia)

10 DEN FIN NOR SWE 11 AUT CZE GER HUN POL SLO SWI 12 ADO COR FRA POR SAR SPA 13 ALB BIH BUL CRO GRC ITA KOS MNT NMA ROM SIC SLV SRB 14 BLR EST RUS UKR

Remarks: sambucina agg.

Synonyms: hidden

= Dactylorchis sambucina (L.) Verm.; Stud. Dactylorch. 65 (1947)

= Dactylorhiza fasciculata (Tineo) Aver.; Bot. Zhurn. (Moscow & Leningrad) 69(6): 875 (1984), isonym

= Dactylorhiza fasciculata (Tineo) H.Baumann & Künkele; Mitt. Arbeitskreises Heimische Orchideen 13: 470 (1981)

= Dactylorhiza latifolia f. rubra (Winterl) D.Tyteca & Gathoye; Orchidophile (Paris) 92: 112 (1990)

= Dactylorhiza sambucina lusus incarnata (Wahlenb.) Soó; Nom. Nov. Gen. Dactylorhiza 3 (1962)

= Dactylorhiza sambucina f. chusae C.E.Hermos.; Estud. Mus. Cienc. Nat. Álava 15: 190 (2000)

= Dactylorhiza sambucina f. hybrida (W.Zimm.) Kümpel; Ber. Arbeitskreis. Heimische Orchid. 18: 27 (2001 publ. 2002)

= Dactylorhiza sambucina f. robusta (Neuman) Soó; Acta Bot. Acad. Sci. Hung. 16: 368 (1970 publ. 1971)

= Dactylorhiza sambucina f. rubra (Winterl) Hyl.; Nordisk Kärlväxtfl. 2: 387 (1966)

= Dactylorhiza sambucina f. rubrobracteata (Harz) Soó; Acta Bot. Acad. Sci. Hung. 16: 368 (1970 publ. 1971)

= Dactylorhiza sambucina subsp. laurentina (R.Bolós ex Vayr.) Malag.; Acta Phytotax. Barcinon. 1: 64 (1968)

= Dactylorhiza sambucina var. hungarica (Soó) Soó; Nom. Nov. Gen. Dactylorhiza 3 (1962)

= Dactylorhiza sambucina var. hybrida (W.Zimm.) Peitz; Jahresber. Naturwiss. Vereins Wuppertal 25: 178 (1972)

= Dactylorhiza sambucina var. laurentina (R.Bolòs ex Vayr.) Soó; Nom. Nov. Gen. Dactylorhiza 3 (1962)

= Orchis fasciculata Tineo; Fl. Sicul. Syn. 2: 875 (1845)

= Orchis guffroyi P.Fourn.; Bull. Soc. Bot. France 78: 434 (1931)

= Orchis incarnata var. sambucina (L.) Lapeyr. ex Bubani; Fl. Pyren. [Bubani] 1: 125 (1897)

= Orchis incarnata Willd.; Sp. Pl., ed. 4 [Willdenow] 4: 80 (1805), nom. illeg.

= Orchis laurentina R.Bolòs ex Vayr.; Anales Soc. Esp. Hist. Nat. 9: 96, t. 1 (1880)

= Orchis lutea Dulac; Fl. Hautes-Pyrénées 125 (1867), nom. illeg.

= Orchis maculata subsp. sambucina (L.) Bonnier & Layens; Tabl. Syn. Pl. Vasc. France 1: 311 (1894)

= Orchis mixta var. sambucina (L.) Retz.; 167 (1779), nom. illeg.

= Orchis pallens Moritzi; Fl. Schweiz [Moritzi] 508 (1844), Puccin. Syn. Fl. Luc. 476

= Orchis pseudosambucina var. fasciculata (Tineo) Rchb.f.; Icon. Fl. Germ. Helv. [H.G.L.Reichenbach] 13-14: 176 (1851)

= Orchis saccata Rchb.; Fl. Germ. Excurs. 122 (1830), nom. illeg.

= Orchis salina Fronius; Verh. Mitth. Siebenbürg. Vereins Naturwiss. Hermannstadt 8: 102 (1857)

= Orchis sambucina lusus hybrida W.Zimm.; Allg. Bot. Z. Syst. 22: 129 (1916)

= Orchis sambucina lusus incarnata (Wahlenb.) Soó; Bot. Arch. 23: 62 (1928)

= Orchis sambucina lusus lutea W.Zimm.; Form. Orchid. Deutschl. 38 (1912)

= Orchis sambucina lusus luteopurpurea (Rouy) G.Keller & Soó; Repert. Spec. Nov. Regni Veg. Sonderbeih. A: 2: 203 (1933)

= Orchis sambucina lusus rubra (Winterl) Soó; Repert. Spec. Nov. Regni Veg. Sonderbeih. A: 2: 203 (1933)

= Orchis sambucina f. bracteata M.Schulze; Mitt. Geogr. Ges. Thüringen (Jena) 6(Gesamtthüringen): 26 (1889)

= Orchis sambucina f. hungarica Soó; Repert. Spec. Nov. Regni Veg. 24: 29 (1927)

= Orchis sambucina f. lanceolata W.Zimm.; Allg. Bot. Z. Syst. 22: 129 (1916)

= Orchis sambucina f. laurentina (R.Bolòs ex Vayr.) Soó; Repert. Spec. Nov. Regni Veg. 24: 29 (1927)

= Orchis sambucina f. obovata W.Zimm.; Allg. Bot. Z. Syst. 22: 129 (1916)

= Orchis sambucina f. purpurea (W.D.J.Koch) Neuman; Sver. Fl. 629 (1901)

= Orchis sambucina f. rubrobracteata Harz; Fl. Deutschl. [Schlechtendal], ed. 5, Neuer Abdruck, 4: 271 (1895 publ. 1896)

= Orchis sambucina f. tripartita A.Fuchs; Ber. Naturwiss. Vereins Schwaben Neuburg 1921: 177 (1921)

= Orchis sambucina L.; Fl. Suec., ed. 2 [Linnaeus] 312 (1755)

= Orchis sambucina unranked laurentina (R.Bolòs ex Vayr.) Willk.; Suppl. Prodr. Fl. Hispan. [Willkomm & Lange] 42 (1893)

= Orchis sambucina subsp. asturica (Gand.) Gand.; Nov. Consp. Fl. Eur. 462 (1910), nom. inval.

= Orchis sambucina subsp. incarnata (Willd.) Ces.in Car.Catt.; 315 (1844), nom. inval.

= Orchis sambucina subsp. laurentina (R.Bolòs ex Vayr.) Malag.; Acta Phytotax. Barcinon. 1: 64 (1969), nom. superfl.

= Orchis sambucina subsp. laurentina (R.Bolòs ex Vayr.) Willk.; Suppl. Prodr. Fl. Hispan. [Willkomm & Lange] 42 (1893), nom. alt.

= Orchis sambucina subvar. luteopurpurea Rouy; Fl. France [Rouy & Foucaud] 13: 155 (1912)

= Orchis sambucina subvar. purpurea (W.D.J.Koch) Rouy; Fl. France [Rouy & Foucaud] 13: 155 (1912)

= Orchis sambucina var. barlae Rouy; Fl. France [Rouy & Foucaud] 13: 155 (1912)

= Orchis sambucina var. bracteata (M.Schulze) Harz; Fl. Deutschl. [Schlechtendal], ed. 5, Neuer Abdruck, 4: 271 (1895 publ. 1896)

= Orchis sambucina var. candida St.-Lag.; Étude Fl. [A.Cariot], éd. 8, 2: 806 (1889)

= Orchis sambucina var. hungarica (Soó) A.Camus; Iconogr. Orchid. Europe 212 (1928)

= Orchis sambucina var. incarnata (Willd.) Amo; Fl. Fan. Penins. Iberica 1: 455 (1871)

= Orchis sambucina var. incarnata (Willd.) Mayrhofer; 23 (1832), isonym

= Orchis sambucina var. incarnata (Willd.) Rchb.; XIII (1833), isonym

= Orchis sambucina var. incarnata (Willd.) Wahlenb.; Fl. Suec. [Wahlenberg] 2: 554 (1826)

= Orchis sambucina var. laurentina (R.Bolòs ex Vayr.) Willk.; Prodr. Fl. Hispan. [Willkomm & Lange] 42: 356 (1893)

= Orchis sambucina var. lutea E.G.Camus; Iconogr. Orchid. Europe 212 (1928), nom. illeg.

= Orchis sambucina var. pallide-flava Schur; 295 (1870), nom. inval.

= Orchis sambucina var. purpurascens R.Hinterh. & J.Hinterh.; Prodr. Fl. Salzburg 192 (1851)

= Orchis sambucina var. purpurea W.D.J.Koch; Syn. Fl. Germ. Helv., ed. 2: 972 (1845)

= Orchis sambucina var. robusta Neuman; Sver. Fl. 629 (1901), nom. inval.

= Orchis sambucina var. rubra Huter; 48 (1872), nom. inval.

= Orchis sambucina var. rubra Winterl; Index Hort. Bot. Univ. Hung., s. p. (1788)

= Orchis sambucina var. rubriflora St.-Lag.; Étude Fl. [A.Cariot], éd. 8, 2: 806 (1889)

= Orchis schleicheri Sweet; Brit. Fl. Gard. [Sweet] 2: t. 199 (1827)

species :

Dactylorhiza sibirica Efimov; Phytotaxa 258(2): 111 (2016)

Distribution: Siberia (Krasnoyarsk)

30 SIB

species :

Dactylorhiza sphagnicola (Höppner ex Soó) Aver.; Bot. Zhurn. (Moscow & Leningrad) 69(6): 875 (1984)

Name: Torf-Fingerwurz, Torf-Fingerknabenkraut (DE)

Distribution: Denmark; Norway; Sweden; +Netherlands; Belgium; Germany (Hamburg, Niedersachsen, Nordrhein-Westfalen, Schleswig-Holstein); France

10 DEN NOR SWE 11 BGM GER NET 12 FRA

Remarks: praetermissa agg.

Synonyms: hidden

= Dactylorchis sphagnicola (Höppner) Verm.; Stud. Dactylorch. 66 (1947)

= Dactylorchis traunsteineri subsp. sphagnicola (Höppner) Verm.; Fl. Neerl. 1(5): 70 (1958)

= Dactylorhiza incarnata subsp. sphagnicola (Höppner) H.Sund.; Europ. Medit. Orchid., ed. 3: 40 (1980)

= Dactylorhiza majalis subsp. sphagnicola (Höppner) H.A.Pedersen & Hedrén; Nordic J. Bot. 22(6): 651 (2004)

= Dactylorhiza sphagnicola (Höppner) Aver.; Bot. Zhurn. (Moscow & Leningrad) 69: 875 (1984)

= Orchis angustifolia var. densiflora Neuman; Bot. Not. 1909: 234 (1909), nom. illeg.

= Orchis sphagnicola Höppner; Sitzungsber. Niederrhein. Ges. Natur- Heilk. Bonn 1926D: 23 (1927)

= Orchis ×sphagnicola Höppner ex G.Keller & Soó; Repert. Spec. Nov. Regni Veg. Sonderbeih. A: 2: 252, 259 (1933)

form :

f. calcifugiens (H.A.Pedersen) P.Delforge; Naturalistes Belges 88(Orchid. 20): 246 (2007)

Distribution: Denmark (Jutland)

10 DEN

Synonyms: hidden

= Dactylorhiza calcifugiens (H.A.Pedersen) Kreutz & Hennigs; Guide Orchids Europe N. Afr. Middle E. 420 (2024)

= Dactylorhiza majalis subsp. calcifugiens H.A.Pedersen; Nordic J. Bot. 22: 655 (2004)

= Dactylorhiza sphagnicola subsp. calcifugiens (H.A.Pedersen) Kreutz & Sczep.; Ber. Arbeitskreis. Heimische Orchid. 24(2): 97 (2007)

variant :

var. deweveri (Verm.) Kreutz; J. Eur. Orch. 48(2-4): 314 (2016)

Distribution: Netherlands (Limburg); ?SE-Belgium

11 NET

Synonyms: hidden

= Dactylorchis deweveri Verm.; Ned. Kruidk. Arch., Versl. 56: 227 (1949)

= Dactylorhiza deweveri (Verm.) Soó; Nom. Nov. Gen. Dactylorhiza 6 (1962)

= Dactylorhiza sphagnicola f. deweveri (Verm.) P.Delforge; Naturalistes Belges 88(Orchid. 20): 246 (2007)

= Dactylorhiza sphagnicola subsp. deweveri (Verm.) Wenker; Orchideen Nordrhein-Westfalens 105 (2001)

= Dactylorhiza traunsteineri subsp. deweveri (Verm.) Verm.; Fl. Neerl., I, 5, Orchidaceae 71 (1958)

variant :

var. hoeppneri (A.Fuchs) Kreutz; J. Eur. Orch. 48(2-4): 317 (2016)

Distribution: +Netherlands; NW-Germany (Nordrhein-Westfalen)

11 GER NET

Synonyms: hidden

= Dactylorchis hoeppneri (A.Fuchs) Verm.; Stud. Dactylorch. 66 (1947)

= Dactylorhiza hoeppneri (A.Fuchs) Soó; Ann. Univ. Sci. Budapest. Rolando Eötvös, Sect. Biol. 3: 346 (1960)

= Dactylorhiza sphagnicola f. hoeppneri (A.Fuchs) P.Delforge; Naturalistes Belges 88(Orchid. 20): 246 (2007)

= Dactylorhiza sphagnicola subsp. hoeppneri (A.Fuchs) Dalkowski, Wende & Wenker; Orchideen Nordrhein-Westfalens 105 (2001), nom. inval.

= Orchis hoeppneri (A.Fuchs) Höppner ex Verm.; Stud. Dactylorch. 66 (1947)

= Orchis traunsteineri f. hoeppneri (A.Fuchs) Soó; Monogr. Iconogr. Orchid. Eur. 2: 251 (1933)

= Orchis ×pseudotraunsteineri subsp. hoeppneri A.Fuchs; Ber. Naturwiss. Vereins Schwaben Neuburg 42: 149 (1919)

form :

f. sennia (Vollmar) P.Delforge; Naturalistes Belges 88(Orchid. 20): 246 (2007)

Distribution: Germany (E-Nordrhein-Westfalen)

11 GER

Synonyms: hidden

= Dactylorhiza maculata subsp. sennia (Vollmar) Kreutz; Kompend. Eur. Orchid. 47 (2004)

= Dactylorhiza sennia Vollmar; Ber. Arbeitskreis. Heimische Orchid. 18(2): 98 (2001 publ. 2002)

= Dactylorhiza sphagnicola subsp. sennia (Vollmar) Kreutz; Eurorchis 17: 105 (2005)

variant :

var. wirtgenii (Höppner) comb.ined.

Distribution: NW-Germany; France

11 GER 12 FRA

Synonyms: hidden

= Dactylorhiza traunsteineri subsp. wirtgenii (Höppner) Kreutz; Kompend. Eur. Orchid. 57 (2004)

= Dactylorhiza wirtgenii (Höppner) Soó; Nom. Nov. Gen. Dactylorhiza 6 (1962)

= Orchis wirtgenii Höppner; Abh. Vereins Naturwiss. Erforsch. Niederrheins 2: 57 (1916)

species :

Dactylorhiza stortonii Benigni, Mandozzi, Monaldi, Barigelli & Petroselli; Micol. Marche, n. s. 5(2): 43 (2021)

Distribution: Italy

13 ITA

Remarks: provisionally accepted name

species :

Dactylorhiza traunsteineri (Saut. ex Rchb.) Soó; Nom. Nov. Gen. Dactylorhiza 5 (1962)

subspecies :

subsp. traunsteineri

Name: Traunsteiners Knabenkraut (DE)

Distribution: +Netherlands; Germany (Baden-Württemberg, Bayern); Switzerland; Austria; France; Italy; Slovenia; Croatia

11 AUT GER NET SWI 12 FRA 13 CRO ITA SLV

Remarks: traunsteineri agg. This Central European, mostly alpine species is not identical with the nordic D. lapponica

Synonyms: hidden

= Dactylorchis gracilis Hesl.-Harr.; Vasculum 35: 19 (1950)

= Dactylorchis traunsteineri (Saut. ex Rchb.) Verm.; Stud. Dactylorch. 66 (1947)

= Dactylorchis traunsteineri var. klingei Verm.; Ned. Kruidk. Arch. 56: 225 (1949)

= Dactylorchis traunsteineri var. rhombilabris Verm.; Ned. Kruidk. Arch. 56: 225 (1949)

= Dactylorhiza devillersiorum P.Delforge; Naturalistes Belges 92(Orchid. 24): 79 (2011)

= Dactylorhiza majalis subsp. traunsteineri (Saut. ex Rchb.) H.Sund.; Europ. Medit. Orchid., ed. 3: 40 (1980)

= Dactylorhiza traunsteineri var. mielichhoferi (Klinge) Soó; Nom. Nov. Gen. Dactylorhiza 6 (1962)

= Dactylorhiza ×gracilis (Höppner) Hesl.-Harr.; Vasculum 35: 19 (1950)

= Dactylorhiza ×gracilis (Höppner) Soó; Nom. Nov. Gen. Dactylorhiza 6 (1962)

= Dactylorhiza ×rhenana (Höppner) Soó; Nom. Nov. Gen. Dactylorhiza 6 (1962)

= Dactylorhiza ×rigida (Höppner) Soó; Nom. Nov. Gen. Dactylorhiza 6 (1962)

= Dactylorhiza ×steegeri (Höppner) Soó; Nom. Nov. Gen. Dactylorhiza 6 (1962)

= Dactylorhiza ×suevica (A.Fuchs) Soó; Nom. Nov. Gen. Dactylorhiza 6 (1962)

= Orchis angustifolia subf. conferta Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 88 (1899)

= Orchis angustifolia subf. densata Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 78 (1899)

= Orchis angustifolia subf. dissita Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 88 (1899)

= Orchis angustifolia subf. elegantior Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 83 (1899)

= Orchis angustifolia subf. filiformis Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 84 (1899)

= Orchis angustifolia subf. intermedia Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 146 (1899)

= Orchis angustifolia subf. laxata Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 78 (1899)

= Orchis angustifolia subf. maculata Klinge; Revis. Orchis cordigera & O. angustifolia 68 (1893)

= Orchis angustifolia subf. strictior Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 83 (1899)

= Orchis angustifolia subf. subcurva (Klinge) Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 79 (1899)

= Orchis angustifolia subf. superans Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 84 (1899)

= Orchis angustifolia subf. venusta Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 79 (1899)

= Orchis angustifolia f. acuminata Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 76 (1899)

= Orchis angustifolia f. ammatica Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 79 (1899)

= Orchis angustifolia f. amplivaginata Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 78 (1899)

= Orchis angustifolia f. angustivaginata Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 78 (1899)

= Orchis angustifolia f. anziensis Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 80 (1899)

= Orchis angustifolia f. brevispicata Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 81 (1899)

= Orchis angustifolia f. comosa Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 84 (1899)

= Orchis angustifolia f. densiflora Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 86 (1899)

= Orchis angustifolia f. densissima Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 87 (1899)

= Orchis angustifolia f. densiuscula Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 77 (1899)

= Orchis angustifolia f. fichtenbergii Klinge; Revis. Orchis cordigera & O. angustifolia 68 (1893)

= Orchis angustifolia f. friesii Klinge; Revis. Orchis cordigera & O. angustifolia 67 (1893)

= Orchis angustifolia f. gracilis Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 84 (1899)

= Orchis angustifolia f. immaculata Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 78 (1899), nom. illeg.

= Orchis angustifolia f. laxa Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 85 (1899)

= Orchis angustifolia f. laxiflora Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 86 (1899)

= Orchis angustifolia f. laxiuscula Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 77 (1899)

= Orchis angustifolia f. lehnertii Klinge; Revis. Orchis cordigera & O. angustifolia 67 (1893)

= Orchis angustifolia f. levaschevoica Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 87 (1899)

= Orchis angustifolia f. longispicata Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 81 (1899)

= Orchis angustifolia f. oblonga Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 85 (1899)

= Orchis angustifolia f. obtusifolia Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 75 (1899)

= Orchis angustifolia f. paanajaervica Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 88 (1899)

= Orchis angustifolia f. papjerwica Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 83 (1899)

= Orchis angustifolia f. perhoica Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 82 (1899)

= Orchis angustifolia f. pulcherrima Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 87 (1899)

= Orchis angustifolia f. reichenbachii Klinge; Revis. Orchis cordigera & O. angustifolia 67 (1893)

= Orchis angustifolia f. remota Klinge; Revis. Orchis cordigera & O. angustifolia 67 (1893)

= Orchis angustifolia f. rosiensis Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 80 (1899)

= Orchis angustifolia f. sauteri Klinge; Revis. Orchis cordigera & O. angustifolia 67 (1893)

= Orchis angustifolia f. schmidtii Klinge; Revis. Orchis cordigera & O. angustifolia 68 (1893)

= Orchis angustifolia f. spathulata Klinge; Revis. Orchis cordigera & O. angustifolia 67 (1893), et Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 76 (1899)

= Orchis angustifolia f. subcurva Klinge; Revis. Orchis cordigera & O. angustifolia 69 (1893)

= Orchis angustifolia f. suberecta Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 87 (1899)

= Orchis angustifolia f. tiudiensis Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 88 (1899)

= Orchis angustifolia Rchb. ex Klinge; 291 (1893), nom. illeg.

= Orchis angustifolia subsp. friesii (Klinge) Neuman; Bot. Not. 1909: 231 (1909)

= Orchis angustifolia subsp. subcapitata Neuman; Bot. Not. 1909: 233 (1909)

= Orchis angustifolia subsp. traunsteineri (Saut. ex Rchb.) C.Hartm.; Handb. Skand. Fl., ed. 11: 388 (1879) [Unplaced]

= Orchis angustifolia subsp. traunsteineri (Saut. ex Rchb.) F.Aresch.; 409 (1881)

= Orchis angustifolia subsp. traunsteineri (Saut. ex Rchb.) Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(1): 175 (1898)

= Orchis angustifolia var. acuminata (Klinge) Neuman; Bot. Not. 1909: 232 (1909)

= Orchis angustifolia var. blyttii (Rchb.f.) Klinge; Revis. Orchis cordigera & O. angustifolia 67 (1893)

= Orchis angustifolia var. elongata Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 83 (1899)

= Orchis angustifolia var. erecta Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 81 (1899)

= Orchis angustifolia var. gracillima Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 86 (1899)

= Orchis angustifolia var. immaculata Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 81 (1899), nom. illeg.

= Orchis angustifolia var. mielichhoferi Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 75 (1899)

= Orchis angustifolia var. patens Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 82 (1899)

= Orchis angustifolia var. poenalica Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 86 (1899)

= Orchis angustifolia var. pusilla Neuman; Bot. Not. 1909: 233 (1909)

= Orchis angustifolia var. recurva Klinge; Revis. Orchis cordigera & O. angustifolia 68 (1893)

= Orchis angustifolia var. reolana Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 86 (1899)

= Orchis angustifolia var. rigidula Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 82 (1899)

= Orchis angustifolia var. sanionis Klinge; Revis. Orchis cordigera & O. angustifolia 67 (1893)

= Orchis angustifolia var. stricta Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 82 (1899), nom. illeg.

= Orchis angustifolia var. superba Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 80 (1899)

= Orchis angustifolia var. tarbatonica Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 78 (1899)

= Orchis angustifolia var. tenuifolia Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 81 (1899)

= Orchis angustifolia var. traunsteineri (Saut. ex Rchb.) Klinge; 329 (1893)

= Orchis incarnata subsp. traunsteineri (Saut. ex Rchb.) Cajander; Melan. Suom. Kasvio [Cajander], ed. 5: 171 (1906)

= Orchis incarnata var. angustifolia (Lindl.) Rchb.f.; Icon. Fl. Germ. Helv. [H.G.L.Reichenbach] 13-14: 52 (1851), nom. illeg.

= Orchis incarnata var. traunsteineri (Saut. ex Rchb.) Parl.; Fl. Ital. [Parlatore] 3: 521 (1858)

= Orchis latifolia var. traunsteineri (Saut. ex Rchb.) Godr.; Fl. Lorraine 3: 30 (1844)

= Orchis maculata subsp. traunsteineri (Saut. ex Rchb.) Douin; Fl. Ill. France 11: 30 (1931)

= Orchis sambucina var. subalpina Rchb.; 140[hochgestellt:]18 (1832)

= Orchis traunsteineri Saut. ex Koch; Syn. Fl. Germ. Helv. 688 (1837)

= Orchis traunsteineri Saut. ex Rchb.; Fl. Germ. Excurs. 140 (1831)

= Orchis ×gracilis Höppner; Sitzungsber. Naturhist. Vereins Preuss. Rheinl. 14 (1926 publ. 1927)

= Orchis ×gracilis Höppner ex G.Keller & Soó; Repert. Spec. Nov. Regni Veg. Sonderbeih. A: 2: 252 (1933)

= Orchis ×pseudotraunsteineri subsp. suevica A.Fuchs; Ber. Naturwiss. Vereins Schwaben Neuburg 42: 107 (1919)

= Orchis ×rhenana Höppner; Abk. Ver. Naturw. Erforsch. Niederrh. 2: 69 (1916)

= Orchis ×steegeri Höppner; Sitzungsber. Naturhist. Vereins Preuss. Rheinl. 16 (1926 publ. 1927)

= Orchis ×steegeri Höppner ex G.Keller & Soó; Repert. Spec. Nov. Regni Veg. Sonderbeih. A: 2: 252 (1933)

variant :

var. vosagiaca (Kreutz & P.Wolff) Sonnb.; Fl. German. [M. Hassler & T. Muer] 1675 (2022)

Name: Wasgau-Fingerwurz (DE)

Distribution: SW-Germany (Rheinland-Pfalz, +Saarland); NE-France (Vosges)

11 GER 12 FRA

Synonyms: hidden

= Dactylorhiza traunsteineri subsp. vosagiaca P.Wolff ex Kreutz & P.Wolff; Ber. Arbeitskreis. Heimische Orchid. 27(2): 196 (2010 publ. 2011)

= Dactylorhiza vosagiaca (P.Wolff ex Kreutz & P.Wolff) Herr-Heidtke & Heidtke; 11 (2012), nom. inval.

= Dactylorhiza vosagiaca (P.Wolff ex Kreutz & P.Wolff) Herr-Heidtke & Heidtke ex P.Wolff; Kochia 10: 57 (2017)

subspecies :

subsp. bohemica (Businský) Kreutz & Sczep.; Ber. Arbeitskreis. Heimische Orchid. 24(2): 94 (2007)

Distribution: Czech Republic (N-Bohemia)

11 CZE

Synonyms: hidden

= Dactylorhiza bohemica Businsky; Preslia 61(4): 296 (1989)

subspecies :

subsp. irenica (F.M.Vázquez) Kreutz; Ber. Arbeitskreis. Heimische Orchid. 28(2): 269 (2011)

Distribution: Spain (Cáceres)

12 SPA

Synonyms: hidden

= Dactylorhiza irenica F.M.Vázquez; J. Eur. Orch. 40(1): 32 (2008)

subspecies :

subsp. rhaetica (H.Baumann & R.Lorenz) F.Benoît; Orchidophile (Paris) 205: 228 (2015)

Distribution: S-Germany (Bayern); Switzerland; Austria; Slovakia (Mala Fatra); France; N-Italy; ?Ukraine (Carpathians)

11 AUT GER SLO SWI 12 FRA 13 ITA

Synonyms: hidden

= Dactylorhiza lapponica subsp. rhaetica H.Baumann & R.Lorenz; J. Eur. Orch. 37(4): 941 (2005)

= Dactylorhiza rhaetica (H.Baumann & R.Lorenz) Biagioli, Kreutz & De Simoni; GIROS Orch. Spont. Eur. 65(2): 149 (2022)

subspecies :

subsp. russowii (Klinge) Soó; Nom. Nov. Gen. Dactylorhiza 6 (1962)

Distribution: Sweden; Finland; NE-Germany (Mecklenburg-Vorpommern); Poland; Czech Republic; Estonia; Latvia; Lithuania; N-Belarus; European Russia (N-European Russia, C-European Russia); Siberia (Krasnoyarsk, W-Siberia); Kazakhstan

10 FIN SWE 11 CZE GER POL 14 BLR EST LAT LIT RUS 30 SIB 32 KAZ

Synonyms: hidden

= Dactylorchis curvifolia (F.Nyl.) Verm.; Stud. Dactylorch. 151 (1947)

= Dactylorchis pseudocordigera (Neuman) Lid; Norsk Fl. [Nordhagen], ed. 2, 210 (1952)

= Dactylorchis russowii (Klinge) Á.Löve & D.Löve; Bot. Not. 114: 38 (1961)

= Dactylorchis traunsteineri subsp. curvifolia (F.Nyl.) Verm.; Stud. Dactylorch. 66 (1947)

= Dactylorchis traunsteineri subsp. russowii (Klinge) Verm.; Stud. Dactylorch. 158 (1947)

= Dactylorhiza curvifolia (F.Nyl.) Czerep.; Sosud. Rast. S.S.S.R. 307 (1981)

= Dactylorhiza lapponica subsp. pseudocordigera (Neuman) Landwehr; Orchideeën 37(3): 80 (1975), isonym

= Dactylorhiza lapponica subsp. pseudocordigera (Neuman) Verm.; Stud. Dactylorch. 67 (1947)

= Dactylorhiza lapponica subsp. russowii (Klinge) H.Baumann & R.Lorenz; J. Eur. Orch. 37(3): 706 (2005)

= Dactylorhiza lapponica var. pseudocordigera (Neuman) Kreutz; Ber. Arbeitskreis. Heimische Orchid. 24(1): 152 (2007)

= Dactylorhiza majalis subsp. russowii (Klinge) H.Sund.; Europ. Medit. Orchid., ed. 3: 40 (1980)

= Dactylorhiza pseudocordigera (Neuman) Soó; Nom. Nov. Gen. Dactylorhiza 4 (1962)

= Dactylorhiza russowii (Klinge) Holub; Preslia 36: 253 (1964)

= Dactylorhiza traunsteineri subsp. curvifolia (F.Nyl.) Soó; Nom. Nov. Gen. Dactylorhiza 6 (1962)

= Dactylorhiza traunsteineri var. curvifolia (F.Nyl.) Aver.; Turczaninowia 3(1): 50 (2000)

= Dactylorhiza traunsteineri var. patula (Klinge) Soó; Nom. Nov. Gen. Dactylorhiza 6 (1962)

= Dactylorhiza traunsteineri var. russowii (Klinge) Aver.; Turczaninowia 3(1): 50 (2000)

= Orchis angustifolia f. arcuata Klinge; Revis. Orchis cordigera & O. angustifolia 69 (1893)

= Orchis angustifolia f. curvata Klinge; Revis. Orchis cordigera & O. angustifolia 69 (1893)

= Orchis angustifolia f. elongata Klinge; Revis. Orchis cordigera & O. angustifolia 69 (1893)

= Orchis angustifolia f. gruneri Klinge; Revis. Orchis cordigera & O. angustifolia 69 (1893)

= Orchis angustifolia f. intermedia Klinge; Revis. Orchis cordigera & O. angustifolia 69 (1893)

= Orchis angustifolia f. patens Klinge; Revis. Orchis cordigera & O. angustifolia 69 (1893)

= Orchis angustifolia f. stricta Klinge; Revis. Orchis cordigera & O. angustifolia 69 (1893)

= Orchis angustifolia subsp. curvifolia (F.Nyl.) E.G.Camus & A.Camus; Iconogr. Orchid. Europe Text(1): 229 (1928)

= Orchis angustifolia subsp. russowii (Klinge) Klinge; Trudy Imp. St.-Peterburgsk. Bot. Sada 17(1): 8, 31 (1898)

= Orchis angustifolia subf. concolor Klinge; Revis. Orchis cordigera & O. angustifolia 69 (1893)

= Orchis angustifolia subf. immaculata Klinge; Revis. Orchis cordigera & O. angustifolia 69 (1893), nom. illeg.

= Orchis angustifolia var. arcuata (Klinge) Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 85 (1899)

= Orchis angustifolia var. curvata (Klinge) Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 84 (1899)

= Orchis angustifolia var. curvifolia (F.Nyl.) Nyman; Consp. Fl. Eur. Suppl. 2: 291 (1890)

= Orchis angustifolia var. patula Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 77 (1899)

= Orchis angustifolia var. russowii Klinge; Arch. Naturk. Liv.- Ehst- Kurlands, Ser. 2, Biol. Naturk. 10: 84 (1893)

= Orchis curvifolia F.Nyl.; Spic. Pl. Fenn. 2: 12. n. 25 (1844)

= Orchis incarnata f. curvifolia (F.Nyl.) Cajander; Melan. Suom. Kasvio [Cajander], ed. 5: 172 (1906)

= Orchis maculata subsp. curvifolia (F.Nyl.) E.G.Camus; Monogr. Orchid. 194 (1908)

= Orchis pseudocordigera Neuman; Bot. Not. 236 (1909)

= Orchis recurva F.Nyl. ex Sw.; Summa Veg. Scand. [O.P.Swartz] 1: 61 (1814)

= Orchis russowii (Klinge) Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17 1: 154, in clavi, 177 (1899)

= Orchis russowii (Klinge) Kuppf.; Exsicc. (Herb. Norm.) 44: 107 (1902), isonym

= Orchis russowii (Klinge) Schltr.; Repert. Spec. Nov. Regni Veg. Sonderbeih. A: 1 (G. Keller & Schltr., Monogr. & Iconogr. Orch. Eur.) 170 (1927), isonym

= Orchis traunsteineri subsp. russowii (Klinge) Soó; Bot. Arch. 23: 83 (1929)

= Orchis traunsteineri var. russowii (Klinge) M.Schulze; Mitth. Thüring. Bot. Vereins, N. F. 17: 50 (1902)

= Orchis traunsteineri var. russowii (Klinge) Vollm.; Fl. Bayern [Vollmann] 164 (1914), isonym

subspecies :

subsp. turfosa (F.Proch.) Kreutz; Kompend. Eur. Orchid. 56 (-57) (2004)

Distribution: Czech Republic (SW-Bohemia)

11 CZE

Synonyms: hidden

= Dactylorhiza comosa subsp. turfosa (F.Proch.) Holub; Preslia 70(2): 103 (1998)

= Dactylorhiza majalis subsp. turfosa F.Proch.; Preslia 54(4): 289 (1982)

= Dactylorhiza turfosa (F: Proch.) Kreutz; Guide Orchids Europe N. Afr. Middle E. 423 (2024)

species :

Dactylorhiza traunsteinerioides (Pugsley) R.M.Bateman & Denholm; List Vasc. Pl. Brit. Isles [D.H.Kent] Suppl. 3: 20 (2006)

Name: Narrow-leaved Marsh-orchid (EN)

Distribution: England (N-England, NW-Wales, W-Scotland, Outer Hebrides); Ireland (Ireland, Northern Ireland); Norway (Stavanger)

10 GRB IRE NOR

Remarks: for detailed discussion see Bateman & Denholm 2012

Synonyms: hidden

= Dactylorchis traunsteinerioides (Pugsley) Verm.; Stud. Dactylorch. 66 (1947)

= Dactylorhiza hibernica (Landwehr) Kreutz; Guide Orchids Europe N. Afr. Middle E. 422 (2024)

= Dactylorhiza majalis subsp. traunsteinerioides (Pugsley) R.M.Bateman & Denholm; Watsonia 14(4): 372 (1983)

= Dactylorhiza majalis var. traunsteinerioides (Pugsley) R.M.Bateman & Denholm; Watsonia 14(4): 373 (1983)

= Dactylorhiza traunsteineri subsp. hibernica Landwehr; Orchideeën 37(3): 79 (1975), et Wilde Orchid. Eur. 2: 557 (1977)

= Dactylorhiza traunsteineri subsp. traunsteinerioides (Pugsley) Soó; Nom. Nov. Gen. Dactylorhiza 6 (1962)

= Dactylorhiza traunsteinerioides (Pugsley) Landwehr; Orchideeën 37(3): 80 (1975), nom. inval.

= Orchis majalis subsp. traunsteinerioides Pugsley; Proc. Linn. Soc. London 148: 124 (1936)

= Orchis traunsteinerioides (Pugsley) Pugsley; J. Bot. 78: 179 (1940)

variant :

var. ebudensis (Wief. ex R.M.Bateman & Denholm) R.M.Bateman & Denholm; New J. Bot. 2(1): 45 (2012)

Distribution: England (Outer Hebrides: North Uist Isl., S-Berneray Isl.)

10 GRB

Synonyms: hidden

= Dactylorhiza comosa subsp. scotica (E.Nelson) P.D.Sell; Fl. Gr. Brit. Ireland 5: 364 (1996)

= Dactylorhiza ebudensis (Wief. ex R.M.Bateman & Denholm) P.Delforge; Naturalistes Belges 81: 397 (2000)

= Dactylorhiza majalis subsp. ebudensis (Wief. ex R.M.Bateman & Denholm) M.R.Lowe; Eurorchis 15: 81 (2003)

= Dactylorhiza majalis subsp. scotica E.Nelson; Monogr. & Ikonogr. Orchid.-Gattung Dactylorhiza 90 (1976), without type, et Taxon 28 (5-6): 593 (1979), with type

= Dactylorhiza majalis var. bowmanii M.N.Jenk.; Watsonia 20(3): 272 (1995)

= Dactylorhiza majalis var. ebudensis Wief.; Wilde Orchid. Eur. 2: 557 (1977)

= Dactylorhiza majalis var. ebudensis Wief. ex R.M.Bateman & Denholm; Edinburgh J. Bot. 52(1): 57 (1995)

= Dactylorhiza majalis var. scotica (E.Nelson) R.M.Bateman & Denholm; Watsonia 44: 368 (1983)

variant :

var. francis-drucei (Wilmott) F.M.Vázquez; J. Eur. Orch. 40(1): 30 (2008)

Distribution: England (Outer Hebrides)

10 GRB

Synonyms: hidden

= Dactylorhiza francis-drucei (Wilmott) Aver.; Bot. Zhurn. (Moscow & Leningrad) 69(6): 875 (1984)

= Dactylorhiza majalis var. francis-drucei (Wilmott) R.M.Bateman & Denholm; Watsonia 14(4): 373 (1983)

= Dactylorhiza traunsteineri subsp. francis-drucei (Wilmott) Soó; Nom. Nov. Gen. Dactylorhiza 6 (1962)

= Dactylorhiza traunsteinerioides subsp. francis-drucei (Wilmott) R.M.Bateman & Denholm; New J. Bot. 2(1): 44 (2012)

= Orchis francis-drucei Wilmott; Proc. Linn. Soc. London Sess. 148, 128 (1936)

species :

Dactylorhiza umbrosa (Kar. & Kir.) Nevski; Trudy Bot. Inst. Akad. Nauk S.S.S.R., Ser. 1, Fl. Sist. Vyssh. Rast. 4: 332 (1937)

Name: Persisches Knabenkraut

Distribution: Kazakhstan; Turkmenistan; Uzbekistan; Kyrgyzstan; Tajikistan; Armenia; Azerbaijan; E-Turkey; N-Iraq; Iran; Afghanistan (Badakshan, Wakhan, Baghlan, Bamyan, Faryab, Herat, Kabul, Kunar / Nuristan, Logar, Wardak, Orozgan / Daykundi, Paktia / Khost, Parwan, Takhar); China (Xinjiang); N-Pakistan (Kurram); India (Jammu & Kashmir)

32 KAZ KGZ TKM TZK UZB 33 ARM AZE 34 AFG IRN IRQ TUR 36 CHI 40 IND PAK

Remarks: incarnata agg. Often included in D. incarnata ssp. cilicica

Synonyms: hidden

= Dactylorchis umbrosa (Kar. & Kir.) Wendelbo; Nytt Mag. Bot. 1: 24 (1952), isonym

= Dactylorhiza chuhensis Renz & Taubenheim; Fl. Turkey 8: 564 (1984), nom. nov.

= Dactylorhiza incarnata subsp. turcestanica (Klinge) H.Sund.; Europ. Medit. Orchid., ed. 3: 40 (1980)

= Dactylorhiza incarnata var. kotschyi (Rchb.f.) H.A.Pedersen, P.J.Cribb & Rolf Kühn; Field Guide Orchids Europe Medit. 139 (2019)

= Dactylorhiza kotschyi (Rchb.f.) P.F.Hunt & Summerh.; Watsonia 6: 130 (1965)

= Dactylorhiza merovensis (Grossh.) Aver.; Bot. Zhurn. (Moscow & Leningrad) 68(7): 894 (1983)

= Dactylorhiza persica (Schltr.) Soó; Nom. Nov. Gen. Dactylorhiza 4 (1962)

= Dactylorhiza renzii Aver.; Bot. Zhurn. (Moscow & Leningrad) 68(7): 893 (1983) [non H. Baumann & Künkele (1981)]

= Dactylorhiza sanasunitensis (H.Fleischm.) Soó; Nom. Nov. Gen. Dactylorhiza 4 (1962)

= Dactylorhiza umbrosa subsp. persica (Schltr.) Kreutz; Ber. Arbeitskreis. Heimische Orchid. 24(1): 156 (2007)

= Dactylorhiza umbrosa var. chuhensis (Renz & Taubenheim) Kreutz; Orchid. Türkei [Kreutz] 181 (1998)

= Dactylorhiza umbrosa var. longibracteata Renz; Fl. Iranica [Rechinger] 26: 131 (1978)

= Dactylorhiza umbrosa var. ochroleuca (Bornm.) Renz; Fl. Iranica [Rechinger] 26: 131 (1978)

= Dactylorhiza vanensis E.Nelson; Monogr. & Ikonogr. Orchid.-Gattung Dactylorhiza 59 (1976), without type, et Taxon 28 (5-6): 593 (1979), with type

= Orchis incarnata f. ochroleuca Bornm.; Repert. Spec. Nov. Regni Veg. 47: 72 (1939), nom. illeg.

= Orchis incarnata var. kotschyi Rchb.f.; Icon. Fl. Germ. Helv. [H.G.L.Reichenbach] 13-14: 53 (1851)

= Orchis kotschyi (Rchb.f.) Schltr.; Repert. Spec. Nov. Regni Veg. 19: 48 (1923)

= Orchis merovensis Grossh.; Beih. Bot. Centralbl. 44(2): 207 (1927)

= Orchis olocheilos var. sanasunitensis (H.Fleischm.) Soó; Repert. Spec. Nov. Regni Veg. 24: 30 (1927)

= Orchis orientalis subsp. turcestanica Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(1): 183 (1898)

= Orchis persica Schltr.; Repert. Spec. Nov. Regni Veg. 15: 290 (1918)

= Orchis sanasunitensis H.Fleischm.; Ann. K.K. Naturhist. Hofmus. 28: 35 (1914)

= Orchis turcestanica (Klinge) O.Fedtsch.; Trudy Imp. S.-Peterburgsk. Bot. Sada 17 1: 155, in clavi, 183 (1898), et ex B. Fedtsch., Russk. Bot. Zurn. 191 (1908)

= Orchis umbrosa Kar. & Kir.; Bull. Soc. Imp. Naturalistes Moscou 15: 504 (1842)

species :

Dactylorhiza urvilleana (Steud.) H.Baumann & Künkele; Mitt. Arbeitskreises Heimische Orchideen 13(2): 240 (1981)

subspecies :

subsp. urvilleana

Name: D´Urville-Knabenkraut (DE)

Distribution: Northern Caucasus; Georgia [Caucasus] (Abkhazia, Racha-Lechkhumi, Samegrelo, Imereti, Adzharia, Shiga Kartli, Kartli, Mtiuleti, Trialeti, Javakheti); Armenia; Azerbaijan (incl. Talysh); NE-Turkey; Iran

33 ARM AZE GEO NCS 34 IRN TUR

Remarks: praetermissa agg.

Synonyms: hidden

= Dactylorchis amblyoloba (Nevski) Soó; [nom. inval.]

= Dactylorchis lancibracteata (K.Koch) Soó; [nom. inval.]

= Dactylorhiza affinis (K.Koch) Aver.; Bot. Zhurn. (Moscow & Leningrad) 68(7): 895 (1983)

= Dactylorhiza amblyoloba (Nevski) Aver.; Novosti Sist. Vyssh. Rast. 26: 47 (1989)

= Dactylorhiza lancibracteata (K.Koch) Renz; Fl. Iranica [Rechinger] 26: 135 (1978)

= Dactylorhiza maculata subsp. triphylla (K.Koch) H.Sund.; Europ. Medit. Orchid., ed. 3: 40 (1980)

= Dactylorhiza pontica (H.Fleischm. & Hand.-Mazz.) P.Delforge; Naturalistes Belges 81(4): 397 (2000)

= Dactylorhiza saccifera subsp. amblyoloba (Nevski) Eccarius; Orchideengattung Dactylorhiza 166 (2016)

= Dactylorhiza saccifera subsp. cartaliniae (Klinge) Soó; Nom. Nov. Gen. Dactylorhiza 7 (1962) [p. p., nom. inval.]

= Dactylorhiza saccifera subsp. lancibracteata (K.Koch) Soó; Nom. Nov. Gen. Dactylorhiza 8 (1962)

= Dactylorhiza saccifera var. pontica (H.Fleischm. & Hand.-Mazz.) Soó; Nom. Nov. Gen. Dactylorhiza 8 (1962)

= Dactylorhiza triphylla (H.Sund.) Czerep.; Sosud. Rast. S.S.S.R. 309 (1981)

= Dactylorhiza urvilleana var. pontica (H.Fleischm. & Hand.-Mazz.) P.Delforge; Naturalistes Belges 89(Orchid. 21): 250 (2008)

= Orchis affinis K.Koch; Linnaea 22: 284 (1849)

= Orchis amblyoloba Nevski; Fl. URSS [Komarov et al., eds.] 4: 707, 753 (1935)

= Orchis basilica subsp. cartaliniae Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(1): 158, 196 (1898)

= Orchis cartaliniae (Klinge) Lipsky; Trudy Imp. S.-Peterburgsk. Bot. Sada 17 1: 158, in clavi, et 196 (1898)

= Orchis cartaliniae (Klinge) Medw.; Trudy Tiflissk. Bot. Sada 18: 475 (1919) [in clavi]

= Orchis lancibractea Grossh.; Fl. Kavkaza [Grossheim] 1: 269 (1928)

= Orchis lancibracteata K.Koch; Linnaea 22: 282 (1850)

= Orchis maculata subsp. lancibracteata (K.Koch) Soó; Repert. Spec. Nov. Regni Veg. 24: 32 (1927) [nom. inval.]

= Orchis maculata var. broteri Sommier & Levier; Trudy Imp. S.-Peterburgsk. Bot. Sada 16: 419 (1900)

= Orchis maculata var. cartaliniae (Klinge) Schltr.; Monogr. Iconogr. Orchid. Eur. 1: 179 (1927)

= Orchis maculata var. pontica (H.Fleischm. & Hand.-Mazz.) Soó; Repert. Spec. Nov. Regni Veg. 24: 32 (1927)

= Orchis pontica H.Fleischm. & Hand.-Mazz.; Ann. K.K. Naturhist. Hofmus. 23: 208 (1909)

= Orchis saccata d´Urv.; Mém. Soc. Linn. Paris 1: 375 (1822), nom. illeg.

= Orchis triphylla K.Koch; Linnaea 22: 283 (1850) [non Spreng. (1826)]

= Orchis urvilleana Steud.; Nomencl. Bot. [Steudel], ed. 2, 2: 225 (1841)

= xDactyloglossum ×ponticum Kohlmüller; Ber. Arbeitskreis. Heimische Orchid. 16(1): 74 (1999)

subspecies :

subsp. bithynica (H.Baumann) H.Baumann & R.Lorenz; J. Eur. Orch. 37(3): 707 (2005)

Distribution: Turkey (W-Anatolia)

34 TUR

Synonyms: hidden

= Dactylorhiza bithynica H.Baumann; Mitt. Arbeitskreises Heimische Orchideen 15(1): 68 (1983)

= Dactylorhiza saccifera subsp. bithynica (H.Baumann) Kreutz; Kompend. Eur. Orchid. 54 (2004)

subspecies :

subsp. ilgazica (Kreutz) Kreutz; Kompend. Eur. Orchid. 58 (2004)

Distribution: NE-Turkey

34 TUR

Synonyms: hidden

= Dactylorhiza ilgazica Kreutz; Eurorchis 9: 49 (1997)

subspecies :

subsp. phoenissa B.Baumann & H.Baumann; J. Eur. Orch. 37(2): 251, 254 (2005)

Distribution: Lebanon

34 LEB

Synonyms: hidden

= Dactylorhiza phoenissa (B.Baumann & H.Baumann) P.Delforge; Naturalistes Belges 96(Orchid. 28): 15 (2015)

species :

Dactylorhiza viridis (L.) R.M.Bateman, Pridgeon & M.W.Chase; Lindleyana 12(3): 113-141 (1997)

Name: Grüne Hohlzunge (DE), Frog Orchid (EN)

Distribution: Iceland; England (England, Wales, +Isle of Man, Scotland, Outer Hebrides, Orkney Isl., Shetland Isl., +St. Kilda Isl.); Ireland (Ireland, Northern Ireland); Denmark; Norway; Sweden; Finland; Netherlands; Belgium; Luxembourg; Germany (Brandenburg, Baden-Württemberg, Bayern, Hessen, +Niedersachsen, Nordrhein-Westfalen, Rheinland-Pfalz, Saarland, Sachsen, Sachsen-Anhalt, Thüringen); Switzerland; Liechtenstein; Austria; Poland; Czech Republic; Slovakia; Hungary; Spain; Andorra; France; Italy; ?Sicily; Slovenia; Croatia; Bosnia & Hercegovina; Montenegro; Serbia; Kosovo; North Macedonia; Albania; Romania; Bulgaria; Greece (mountains of N-Greece); Estonia; Latvia; Lithuania; Belarus; Ukraine; Crimea; European Russia (N-European Russia, W-European Russia, C-European Russia); Siberia (Altai, Buryatia, Chita, Irkutsk, Krasnoyarsk, Tuva, W-Siberia, Yakutia); Russian Far East (Amur, Kamchatka, Khabarovsk, Magadan, Primorye, Commander Isl.); Kazakhstan; Kyrgyzstan; Tajikistan; Northern Caucasus; Georgia [Caucasus] (Abkhazia, Samegrelo, Imereti, Adzharia, Shiga Kartli, Kartli, Mtiuleti); Armenia; Azerbaijan (incl. Talysh); N-Turkey; Iran; China (Gansu, Hebei, Heilongjiang, Henan, Hubei, Jilin, Liaoning, Nei Mongol, Ningxia, Qinghai, Shaanxi, Shanxi, Sichuan, Xinjiang, NW-Yunnan); NE-Tibet; Mongolia; North Korea; South Korea; Japan (Hokkaido, Honshu); Taiwan; Pakistan (Baltistan, Karakorum); Nepal; Bhutan; India (Himachal Pradesh, Jammu & Kashmir); Aleutes; C-Alaska; Canada (Alberta, British Columbia, Manitoba, New Brunswick, Newfoundland, N. W. T., Nova Scotia, Ontario, Prince Edward Isl., Québec, Saskatchewan, Yukon, Anticosti Isl.); USA (Arizona, Colorado, Connecticut, Illinois, Indiana, Iowa, Maine, Maryland, Massachusetts, Michigan, Minnesota, Missouri, Montana, Nebraska, New Hampshire, New Jersey, New Mexico, New York, North Carolina, North Dakota, Ohio, Pennsylvania, Rhode Island, South Dakota, Tennessee, Utah, Vermont, Virginia, Washington, West Virginia, Wisconsin, Wyoming)

10 DEN FIN GRB ICE IRE NOR SWE 11 AUT BGM CZE GER HUN LIE LUX NET POL SLO SWI 12 ADO FRA SPA 13 ALB BIH BUL CRO GRC ITA KOS MNT NMA ROM SLV SRB 14 BLR EST KRY LAT LIT RUS UKR 30 SIB 31 RFE 32 KAZ KGZ TZK 33 ARM AZE GEO NCS 34 IRN TUR 36 CHI TIB 37 MON 38 JAP KOR NKO TAI 40 BHU IND NEP PAK 70 ASK CDN 71 CDN 72 CDN 73 USA 74 USA 75 USA 76 USA 77 USA 78 USA

Synonyms: hidden

= Chamorchis viridis (L.) Dumort.; Fl. Belg. [Dumortier] 133 (1827)

= Coeloglossum bracteatum (Muhl. ex Willd.) Parl.; Fl. Ital. [Parlatore] 3: 409 (1858)

= Coeloglossum bracteatum (Muhl. ex Willd.) Schltr.; Repert. Spec. Nov. Regni Veg. 16 (463-467): 374 (1920), isonym

= Coeloglossum bracteatum var. kashmiricum Soó; Ann. Hist.-Nat. Mus. Natl. Hung. 26: 356 (1929)

= Coeloglossum bracteatum var. majus (Maxim.) Nakai; J. Jap. Bot. 18: 282 (1942)

= Coeloglossum coreanum (Nakai) Schltr.; Repert. Spec. Nov. Regni Veg. 16: 374 (1920)

= Coeloglossum kaschmirianum Schltr.; Repert. Spec. Nov. Regni Veg. 16: 374 (1920)

= Coeloglossum nankotaizanense (Masam.) Masam.; J. Geobot. 16: 55 (1968)

= Coeloglossum nankotaizanensis (Masam.) S.S.Ying; Quart. J. Chin. Forest. 8(4): 149 (1975), isonym

= Coeloglossum taiwanianum S.S.Ying; Alp. Pl. Taiwan Colour 1: 74 (1975)

= Coeloglossum vaillantii (Ten.) Guss.; Icon. Fl. Germ. Helv. [H.G.L.Reichenbach] 13-14: 130 (1851)

= Coeloglossum viride (L.) Hartm.; Handb. Skand. Fl. 329 (1820)

= Coeloglossum viride f. bracteatum (Muhl. ex Willd.) Bolzon; Bull. Soc. Bot. Ital. 1909: 70 (1909)

= Coeloglossum viride f. brevibracteatum Asch. & Graebn.; Syn. Mitteleur. Fl. [Ascherson & Graebner] 3: 807 (1907)

= Coeloglossum viride f. collinum (Rupp ex W.Zimm.) Höppner; Verh. Naturhist. Vereines Preuss. Rheinl. Westphalens 81: 266 (1924 publ. 1925)

= Coeloglossum viride f. dentatum Zapal.; Consp. Fl. Galic. Crit. 1: 214 (1906)

= Coeloglossum viride f. gracillimum Schur; Enum. Pl. Transsilv. 645 (1866)

= Coeloglossum viride f. grandiflorum Zapal.; Consp. Fl. Galic. Crit. 1: 214 (1906)

= Coeloglossum viride f. integrum Soó; Acta Bot. Acad. Sci. Hung. 16: 367 (1970 publ. 1971), no type indicated.

= Coeloglossum viride f. lancifolium Rohlena; Sitzungsber. Königl. Böhm. Ges. Wiss. Prag, Math.-Naturwiss. Cl. 1912(Beitr.): 118 (1913)

= Coeloglossum viride f. longibracteatum Asch. & Graebn.; Syn. Mitteleur. Fl. [Ascherson & Graebner] 3: 807 (1907)

= Coeloglossum viride f. microbracteatum Schur; Oesterr. Bot. Z. 20: 294 (1870)

= Coeloglossum viride f. parviflorum Höppner; Verh. Naturhist. Vereines Preuss. Rheinl. Westphalens 81: 266 (1924 publ. 1925)

= Coeloglossum viride f. purpureum Schur; Enum. Pl. Transsilv. 646 (1866)

= Coeloglossum viride f. roseum Bolzon; Bull. Soc. Bot. Ital. 1910: 71 (1910)

= Coeloglossum viride f. rubens Norm.; Forh. Vidensk.-Selsk. Kristiania 1893 (16): 39 (1893)

= Coeloglossum viride f. subalpinum Neuman; Sver. Fl. 633 (1901)

= Coeloglossum viride f. thuringiacum (Rupp ex W.Zimm.) Höppner; Verh. Naturhist. Vereines Preuss. Rheinl. Westphalens 81: 266 (1924 publ. 1925)

= Coeloglossum viride f. virescens (Muhl. ex Willd.) O.Gruss & M.Wolff; Orchid. Atlas 77 (2007)

= Coeloglossum viride subsp. bracteatum (Muhl. ex Willd.) Hultén; Acta Univ. Lund. 2, 39, no. 1: 472 (1943)

= Coeloglossum viride subsp. coreanum (Nakai) Satomi; J. Jap. Bot. 44: 127 (1969)

= Coeloglossum viride subsp. eurasiaticum Selander; Acta Phytogeogr. Suec. 28: 10 (1950)

= Coeloglossum viride var. akaishimontanum Satomi; J. Jap. Bot. 44: 127 (1969)

= Coeloglossum viride var. bracteatum (Muhl. ex Willd.) A.Gray; Manual [A.Gray] ed. 5: 500 (1867)

= Coeloglossum viride var. bracteatum (Muhl. ex Willd.) K.Richt.; Pl. Eur. 1: 278 (1890)

= Coeloglossum viride var. brevibracteatum Asch. & Graebn.; Syn. Mitteleur. Fl. [Ascherson & Graebner] 3: 807 (1907)

= Coeloglossum viride var. collinum Rupp ex W.Zimm.; Form. Orchid. Deutschl. 61 (1912)

= Coeloglossum viride var. dentatum Zapal.; Consp. Fl. Galic. Crit. 1: 219 (1906)

= Coeloglossum viride var. grandiflorum Zapal.; Consp. Fl. Galic. Crit. 1: 219 (1906)

= Coeloglossum viride var. interjecta (Fernald) Miyabe & Kudô; J. Fac. Agric. Hokkaido Imp. Univ. 26: 360 (1932)

= Coeloglossum viride var. labellifidum (Costa ex Vayr.) E.G.Camus; Monogr. Orchid. 320 (1908)

= Coeloglossum viride var. longibracteatum Asch. & Graebn.; Syn. Mitteleur. Fl. [Ascherson & Graebner] 3: 807 (1907), nom. illeg.

= Coeloglossum viride var. macrobracteatum Schur; 294 (1870)

= Coeloglossum viride var. microbracteatum Schur; 294 (1870)

= Coeloglossum viride var. rhenanum Höppner; Verh. Naturhist. Vereines Preuss. Rheinl. Westphalens 81: 266 (1925)

= Coeloglossum viride var. subalpinum Rupp ex W.Zimm.; Form. Orchid. Deutschl. 61 (1912)

= Coeloglossum viride var. thuringiacum Rupp ex W.Zimm.; Form. Orchid. Deutschl. 61 (1912)

= Coeloglossum viride var. vaillantii (Ten.) Thell.; Fl. Schweiz [Schinz] ed. 3: 73 (1914)

= Coeloglossum viride var. virescens (Muhl. ex Willd.) Luer; Native Orch. U.S. & Canada 172 (1975)

= Dactylorhiza viridis var. coreana (Nakai) N.S.Lee; Ill. Fl. Korean Orchids 100 (2011)

= Dactylorhiza viridis var. virescens (Muhl. ex Willd.) Baumbach; Orchidee (Hamburg) 64: 137 (2013)

= Entaticus viridis (L.) Gray; Nat. Arr. Brit. Pl. 2: 206 (1821 publ. 1822)

= Gymnadenia bracteata (Muhl. ex Willd.) C.Presl; Reliq. Haenk. 1: 92 (1827)

= Gymnadenia viridis (L.) Rich.; Mém. Mus. Hist. Nat. 4: 57 (1818)

= Gymnadenia viridis var. brevibracteata (Bréb.) Bréb.; 259 (1849)

= Habenaria bracteata (Muhl. ex Willd.) R.Br.; Hort. Kew., ed. 2 [W.T.Aiton] 5: 192 (1813)

= Habenaria flava var. virescens (Muhl. ex Willd.) Fernald; Rhodora 23: 148 (1921)

= Habenaria virescens (Willd.) Spreng.; Syst. Veg., ed. 16 [Sprengel] 3: 688 (1826)

= Habenaria viridis (L.) R.Br.; Hort. Kew., ed. 2 [W.T.Aiton] 5: 192 (1813)

= Habenaria viridis subsp. bracteata (Muhl. ex Willd.) R.T.Clausen; Cornell Univ. Agric. Exp. Sta. Mem. 291: 11 (1949)

= Habenaria viridis var. bracteata (Muhl. ex Willd.) A.Gray; Manual [A.Gray] ed. 5: 500 (1867)

= Habenaria viridis var. interjecta Fernald; Rhodora 28: 173 (1926)

= Herminium nankotaizanense Masam.; J. Soc. Trop. Agric. 4: 194 (1932)

= Himantoglossum viride (L.) Rchb.; Fl. Germ. Excurs. 120 (1830)

= Orchis batrachites Schrank; Baier. Reise 86 (1786)

= Orchis bractealis Salisb.; Parad. Lond. 2: t. 110 (1808), nom. superfl.

= Orchis bracteata Muhl. ex Willd.; Sp. Pl., ed. 4 [Willdenow] 4(1): 34 (1805)

= Orchis coreana Nakai; Bot. Mag. (Tokyo) 28: 302 (1914)

= Orchis ferruginea (F.W.Schmidt) F.W.Schmidt; Samml. Phys.-Oekon. Aufsätze 1: 238 (1795)

= Orchis flava var. virescens (Muhl. ex Willd.) Green; Trans. Soc. Promot. Useful Arts 3: 120 (1814)

= Orchis obsoleta Muhl. ex Willd.; Sp. Pl., ed. 4 [Willdenow] 4: 34 (1805)

= Orchis virens Scop.; Fl. Carniol., ed. 2, 2: 199 (1771)

= Orchis virescens (L.) Muhl. ex Willd.; Sp. Pl., ed. 4 [Willdenow] 4: 37 (1805)

= Orchis viridis (L.) Crantz; Stirp. Austr. Fasc., ed. 2, 6: 491 (1769)

= Orchis viridis var. major Tinant; Fl. Luxemb. 442 (1836)

= Orchis viridis var. vaillantii Ten.; Syll. Pl. Neapol. 629 (1831)

= Peristylus bracteatus (Muhl. ex Willd.) Lindl.; Gen. Sp. Orchid. Pl. 298 (1835)

= Peristylus bracteatus f. major Franch. & Sav.; Enum. Pl. Jap. 2: 31 (1879)

= Peristylus montanus Schur; Enum. Pl. Transsilv. 645 (1866)

= Peristylus virescens (Muhl. ex Willd.) Lindl.; Gen. Sp. Orchid. Pl. 298 (1835)

= Peristylus viridis (L.) Lindl.; Syn. Brit. Fl., ed. 2: 261 (1835)

= Peristylus viridis var. bracteata (Muhl. ex Willd.) Rchb.f. ex Kurtz; Bot. Jahrb. Syst. 19: 408 (1894)

= Peristylus viridis var. macrobracteatus Schur; Enum. Pl. Transsilv. 645 (1866)

= Perularia virescens (Muhl. ex Willd.) A.Gray; Bot. Gaz. 5: 63 (1880)

= Platanthera bracteata (Muhl. ex Willd.) Torr.; Fl. New York 2: 279 (1843)

= Platanthera coreana (Nakai) Nakai; Fl. M´t. Paik-Tu-San 57 (1918)

= Platanthera nankotaizanensis (Masam.) Masam.; Bot. Mag. (Tokyo) 46: 773 (1932)

= Platanthera viridis (L.) Lindl.; Syn. Brit. Fl., ed. 1: 261 (1829)

= Platanthera viridis var. bracteata (Muhl. ex Willd.) Rchb.f.; Icon. Fl. Germ. Helv. [H.G.L.Reichenbach] 13/14(2): 130, t. 83 (- t. 435) (1851)

= Platanthera viridis var. densiflora Regel; Bull. Soc. Imp. Naturalistes Moscou 41(1): 430 (1868)

= Satyrium alpinum F.W.Schmidt; Fl. Boëm. Cent. 1: 63 (1793)

= Satyrium bracteale Salisb.; Trans. Hort. Soc. London 1: 290 (1812)

= Satyrium bracteatum (Muhl. ex Willd.) Pers.; Syn. Pl. [Persoon] 2: 507 (1807) [non Lindl.]

= Satyrium ferrugineum F.W.Schmidt; Fl. Boëm. Cent. 1: 62 (1793)

= Satyrium fuscum Huds.; Fl. Angl. [Hudson] ed. 1: 337 (1762)

= Satyrium lingulatum Vill.; Hist. Pl. Dauphiné [Villars] 2: 42 (err. typ. 24) (1787)

= Satyrium virescens (Muhl. ex Willd.) Pers.; Syn. Pl. [Persoon] 2: 507 (1807)

= Satyrium viride L.; Sp. Pl. [Linnaeus] 2: 944 (1753)

= Sieberia viridis (L.) Spreng.; Anleit. Kenntn. Gew., ed. 2, 2(1): 282 (1817)

variant :

var. islandicum (Lindl.) M.Schultze; Oesterr. Bot. Z. 48: 113 (1898)

Distribution: Iceland; Faeroe Isl.

10 FOR ICE

Synonyms: hidden

= Platanthera islandica (Lindl.) Kraenzl.; Repert. Spec. Nov. Regni Veg., Beih. 65: 53 (1931)

= Peristylus islandicus Lindl.; Gen. Sp. Orchid. Pl. 297 (1835)

= Coeloglossum viride subsp. islandicum (Lindl.) Kreutz; Ber. Arbeitskreis. Heimische Orchid. 24(1): 150 (2007), isonym

= Coeloglossum viride f. islandicum (Lindl.) O.Gruss & M.Wolff; Orchid. Atlas 77 (2007)

= Coeloglossum islandicum (Lindl.) Kom.; Fl. Kamtchatka 1: 313 (1927), isonym

= Coeloglossum islandicum (Lindl.) Nyman; Syll. Fl. Eur. 359 (1855)

= Coeloglossum viride subsp. islandicum (Lindl.) Selander; Acta Phytogeogr. Suec. 28: 10 (1950)

species :

Dactylorhiza ×abantiana H.Baumann & Künkele; Mitt. Arbeitskreises Heimische Orchideen 13(2): 261 (1981)

Distribution: Turkey

34 TUR

Remarks: Dactylorhiza iberica x nieschalkiorum

species :

Dactylorhiza ×abeliana (Klotzsch) Peitz; Jahresber. Naturwiss. Vereins Wuppertal 25: 173 (1972)

Distribution: Europe

Error

Synonyms: hidden

= Orchis traunsteineri f. abeliana Klotzsch

= Orchis angustifolia var. abeliana Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 87 (1899)

species :

Dactylorhiza ×aldenii H.Baumann; Mitt. Arbeitskreises Heimische Orchideen 15(1): 74 (1983)

Distribution: Greece

13 GRC

Remarks: Dactylorhiza iberica x (majalis ssp.) kalopissii

species :

Dactylorhiza ×altobracensis (Coste & Soulié) Soó; Nom. Nov. Gen. Dactylorhiza 10 (1962)

Distribution: C-Europe; Austria

11 AUT

Remarks: Dactylorhiza fuchsii x sambucina

Synonyms: hidden

= Dactylorhiza ×influenza (Sennholz) Soó; Nom. Nov. Gen. Dactylorhiza 9 (1962)

= Orchis ×altobracensis Coste & Soulié; Bull. Soc. Bot. France 44(Sess. Extraord.): 117 (1898)

= Orchis ×influenza Sennholz; Verh. K.K. Zool.-Bot. Ges. Wien 12, Sitzb. 40 (1891), et Oesterr. Bot. Z. 41: 381 (1891)

species :

Dactylorhiza ×alutiiqorum (A.Baum & H.Baum) J.M.H.Shaw; Orchid Rev., Suppl., 122 (1306): 37 (2014)

Distribution: Aleutes

70 ASK

Remarks: Dactylorhiza viridis x D. aristata

Synonyms: hidden

= Dactylorhiza ×alutiiqorum (A.Baum & H.Baum) Eccarius; Orchideengattung Dactylorhiza 580 (2016), isonym

= xDactyloglossum ×alutiiqorum A.Baum & H.Baum; Native Orchid Conf. J. 6(2): 3 (2009)

species :

Dactylorhiza ×aschersoniana (Hausskn.) Borsos & Soó; Ann. Univ. Sci. Budapest. Rolando Eötvös, Sect. Biol., 3 (Geobot. Monogr. Orchid. Pannon. & Karpat. Fl. IV.) 113 (1960) [nothosubsp.]

subspecies :

subsp. aschersoniana

Distribution: Europe (widespread); European Russia

14 RUS

Remarks: Dactylorhiza incarnata x Dactylorhiza majalis ssp. majalis

Synonyms: hidden

= Dactylorchis ×pseudotraunsteineri (A.Fuchs) Verm.; Stud. Dactylorch. 66 (1947)

= Dactylorhiza ×bavarica (A.Fuchs) Soó; Nom. Nov. Gen. Dactylorhiza 6 (1962)

= Dactylorhiza ×eifliaca (A.Fuchs) Soó; Nom. Nov. Gen. Dactylorhiza 6 (1962)

= Dactylorhiza ×gennachensis (A.Fuchs) Soó; Nom. Nov. Gen. Dactylorhiza 6 (1962)

= Dactylorhiza ×koningweeniana (A.Fuchs) Soó; Nom. Nov. Gen. Dactylorhiza 6 (1962)

= Dactylorhiza ×pseudojunialis R.Doll; Feddes Repert. 89: 346 (1978)

= Orchis ×aschersoniana Hausskn.; Mitt. Geogr. Ges. Thüringen (Jena) 2: 223 (1884 publ. 1885)

= Orchis ×pseudotraunsteineri A.Fuchs; Ber. Naturwiss. Vereins Schwaben Neuburg 42: 24 (1919)

= Orchis ×pseudotraunsteineri subsp. bavarica A.Fuchs; Ber. Naturwiss. Vereins Schwaben Neuburg 42: 100 (1919)

= Orchis ×pseudotraunsteineri subsp. eifliaca A.Fuchs; Ber. Naturwiss. Vereins Schwaben Neuburg 42: 154 (1919)

= Orchis ×pseudotraunsteineri var. gennachensis A.Fuchs; Ber. Naturwiss. Vereins Schwaben Neuburg 42: 141 (1919)

= Orchis ×pseudotraunsteineri var. koningweeniana A.Fuchs; Ber. Naturwiss. Vereins Schwaben Neuburg 42: 158 (1919)

variant :

var. mulignensis (Gsell) Kümpel; Ber. Arbeitskreis. Heimische Orchid. 18(2): 23 (2001 publ. 2002) [nothovar.]

Distribution: Switzerland

11 SWI

Remarks: Dactylorhiza incarnata var. serotina x Dactylorhiza majalis

Synonyms: hidden

= Dactylorhiza ×mulignensis (Gsell) Soó; Nom. Nov. Gen. Dactylorhiza 9 (1962)

= Orchis ×mulignensis Gsell; Jahresber. Naturf. Ges. Graubündens, N. F. 74: 17 (1936)

variant :

var. uliginosa Kümpel; Ber. Arbeitskreis. Heimische Orchid. 18(2): 24 (2001 publ. 2002) [nothovar.]

Distribution: Germany

11 GER

Remarks: Dactylorhiza incarnata var. serotina x Dactylorhiza majalis ssp. brevifolia

variant :

var. wisniewskii (Hemke) Gathoye & D.Tyteca; Lejeunia 143: 61 (1994) [nothovar.]

Distribution: Czech Republic; Slovakia

11 CZE SLO

Remarks: Dactylorhiza incarnata ssp. haematodes x Dactylorhiza majalis ssp. majalis

Synonyms: hidden

= Dactylorhiza ×aschersoniana nothosubsp. wiesniewskii (Hemke) Potucek; Kl´úc na Urcovanie Vstavacovit ch Ceskoslovenska (Rosalia, Mimor. Vyd.): 135 (1990)

= Dactylorhiza ×wisniewskii Hemke; Mitteilungsbl. Arbeitskr. Heim. Orch. DDR 8: 51 (1978), contrary to Art. 37.3 ICBN (1994)

= Dactylorhiza ×aschersoniana nothovar. wisniewskii Kümpel; Ber. Arbeitskreis. Heimische Orchid. 18(2): 24 (2001 publ. 2002)

subspecies :

subsp. hochreutineriana (K.Hellm.) Gathoye & D.Tyteca; Lejeunia 143: 61 (1994) [nothosubsp.]

Distribution: Switzerland

11 SWI

Synonyms: hidden

= Dactylorhiza ×hochreutinerana (K.Hellm.) Soó; Feddes Repert. 83: 159 (1972)

= Dactylorhiza ×hochreutineriana (K.Hellm.) Aver.; Bot. Zhurn. (Moscow & Leningrad) 71(1): 92 (1986), isonym

= Orchis ×hochreutineriana K.Hellm.; Boissiera 7: 387 (1943)

subspecies :

subsp. ishorica (Aver.) Eccarius; Orchideengattung Dactylorhiza 585 (2016) [nothosubsp.]

Distribution: European Russia

14 RUS

Remarks: Dactylorhiza incarnata x baltica ssp. longifolia

Synonyms: hidden

= Dactylorhiza ×ishorica Aver.; Bot. Zhurn. (Moscow & Leningrad) 71(1): 93 (1986)

subspecies :

subsp. templinensis (Potucek) H.Kretzschmar; Orchideen Deutschlands [Baumann et al.] 696 (2005) [nothosubsp.]

Distribution: Germany

11 GER

Remarks: Dactylorhiza incarnata ssp. ochroleuca x majalis

Synonyms: hidden

= Dactylorhiza ×aschersoniana nothovar. templinensis (Potucek) Kümpel; Ber. Arbeitskreis. Heimische Orchid. 18(2): 24 (2001 publ. 2002)

= Dactylorhiza ×templinensis Potucek; Gleditschia 3: 31 (1975)

species :

Dactylorhiza ×baicalica Aver.; Bot. Zhurn. (Moscow & Leningrad) 71(1): 92 (1986)

Distribution: Siberia (Buryatia)

30 SIB

Remarks: Dactylorhiza (incarnata ssp.) cruenta x Dactylorhiza salina

species :

Dactylorhiza ×balabaniana H.Baumann; Mitt. Arbeitskreises Heimische Orchideen 15(1): 75 (1983)

Distribution: Turkey

34 TUR

Remarks: Dactylorhiza iberica x urvilleana

Synonyms: hidden

= Dactylorhiza ×balabaniana nothosubsp. mizabensis B.Baumann & H.Baumann; J. Eur. Orch. 37: 254 (2005)

species :

Dactylorhiza ×bayburtiana H.Baumann; Mitt. Arbeitskreises Heimische Orchideen 15(1): 76 (1983)

Distribution: Turkey

34 TUR

Remarks: Dactylorhiza euxina x umbrosa (incarnata ssp. cilicica)

species :

Dactylorhiza ×beckeriana (Höppner) Soó; Nom. Nov. Gen. Dactylorhiza 6 (1962)

Distribution: Germany; Switzerland

11 GER SWI

Remarks: Dactylorhiza incarnata x D. maculata x D. majalis

Synonyms: hidden

= Dactylorchis ×beckeriana (Höppner) Verm.; Stud. Dactylorch. 68 (1947)

= Dactylorhiza ×genevensis (Klinge ex M.Schultze) Soó; Nom. Nov. Gen. Dactylorhiza 10 (1962)

= Orchis ×beckeriana Höppner; Sitzungsber. Naturhist. Vereins Preuss. Rheinl. 3 (1926 publ. 1927)

= Orchis incarnata var. genevensis Klinge ex Schulze; Mitth. Thüring. Bot. Vereins, N. F. 17: 49 (1902)

= Orchis ×beckeriana Höppner ex G.Keller & Soó; Repert. Spec. Nov. Regni Veg. Sonderbeih. A: 2: 251, 258 (1933)

species :

Dactylorhiza ×boluiana H.Baumann; Mitt. Arbeitskreises Heimische Orchideen 15(1): 76 (1983)

Distribution: Turkey

34 TUR

Remarks: Dactylorhiza maculata ssp. saccifera x D. majalis ssp. nieschalkiorum

species :

Dactylorhiza ×braunii (Halácsy) Borsos & Soó; Ann. Univ. Sci. Budapest. Rolando Eötvös, Sect. Biol., 4 (Geobot. Monogr. Orchid. Pannon. & Karpat. Fl. 5): 82 (1961) [nothosubsp.]

subspecies :

subsp. braunii

Distribution: Europe; European Russia; Germany (?Bayern, ?Mecklenburg-Vorpommern, Rheinland-Pfalz, ?Schleswig-Holstein, ?Sachsen); Czech Republic

11 CZE GER 14 RUS

Remarks: Dactylorhiza fuchsii ssp. fuchsii x Dactylorhiza majalis ssp. majalis

Synonyms: hidden

= Dactylorhiza ×braunii nothovar. alpina (E.G.Camus) Soó; Nom. Nov. Gen. Dactylorhiza 9 (1962)

= Dactylorhiza ×dinglensis nothosubsp. braunii (Halácsy) M.H.J.van der Meer; Cact. Phantast. 2019(5)-1: 1 (2019)

= Orchis ×braunii nothovar. alpina E.G.Camus; Iconogr. Orchid. Europe 1: 279 (1927)

= Orchis ×braunii Halácsy; Oesterr. Bot. Z. 31: 137 (1881)

subspecies :

subsp. lilacina F.Proch.; Preslia 54(4): 291 (1982) [nothosubsp.]

Distribution: Czech Republic

11 CZE

Remarks: Dactylorhiza fuchsii ssp. fuchsii x Dactylorhiza majalis ssp. turfosa

Synonyms: hidden

= Dactylorhiza ×braunii nothosubsp. lilacina (F.Proch.) Holub; Preslia 70(2): 103 (1998)

= Dactylorhiza ×braunii nothovar. lilacina F.Proch.; Preslia 54(4): 291 (1982)

subspecies :

subsp. monticola Potucek; Preslia 48(2): 134 (1976) [nothosubsp.]

Distribution: Slovakia

11 SLO

Remarks: Dactylorhiza fuchsii ssp. psychrophila x Dactylorhiza majalis ssp. majalis

Synonyms: hidden

= Dactylorhiza ×braunii nothosubsp. monticola (Potucek) Oddone; GIROS Orch. Spont. Eur. 59: 355 (2016)

= Dactylorhiza ×braunii nothovar. monticola Potucek; Preslia 48: 134 (1976)

= Dactylorhiza ×monticola (Potucek) Gathoye & D.Tyteca; Lejeunia 143: 65 (1994)

= Dactylorhiza ×townsendiana nothosubsp. monticola (Potucek) Eccarius; Orchideengattung Dactylorhiza 587 (2016)

subspecies :

subsp. smitakii Batousek; J. Eur. Orch. 29(4): 643 (1997) [nothosubsp.]

Distribution: Czech Republic

11 CZE

Remarks: Dactylorhiza fuchsii ssp. sooana x majalis ssp. majalis

Synonyms: hidden

= Dactylorhiza ×dinglensis nothosubsp. smitakii (Batousek) Batousek, Taraska & Trávn.; Pl. Syst. Evol. 307(4)-51: 18 (2021)

species :

Dactylorhiza ×breviceras Renz & Taubenheim; Notes Roy. Bot. Gard. Edinburgh 41(2): 269 (1983)

Distribution: Turkey

34 TUR

Remarks: Dactylorhiza osmanica x urvilleana

species :

Dactylorhiza ×carnea Soó; Nom. Nov. Gen. Dactylorhiza 9 (1962)

Distribution: Germany (Niedersachsen); Czech Republic; France; European Russia

11 CZE GER 12 FRA 14 RUS

Remarks: Dactylorhiza elodes x Dactylorhiza incarnata

Synonyms: hidden

= Dactylorhiza ×claudiopolitana nothosubsp. carnea (Soó) Oddone; GIROS Orch. Spont. Eur. 59(2): 355 (2016)

= Orchis ×carnea E.G.Camus; J. Bot. [Morot] 6: 419 (1892), nom. illeg.

species :

Dactylorhiza ×carpathiana Soó, Anghelescu, L.Balogh, Mih.Balogh & Kigyossy; Sci. Papers, Ser. B Hort. 68(2): 634 (2024), nom. inval.

Distribution: Romania

13 ROM

Remarks: Dactylorhiza cordigera x majalis

species :

Dactylorhiza ×conigera (Norman) B.Bock; Bull. Soc. Bot. Centre-Ouest 42: 267 (2011 publ. 2012)

Distribution: Austria; etc.

11 AUT

Remarks: Dactylorhiza viridis x Dactylorhiza maculata

Synonyms: hidden

= Coeloglossum ×conigerum Norman; Norges Arktiske Fl. 1: Sp. Pl. 1050 (1900)

= Dactylorhiza ×conigera nothosubsp. evae (R.Breiner) Oddone; GIROS Orch. Spont. Eur. 59(2): 354 (2016)

= Orchis ×dominiana E.G.Camus; Monogr. Orchid. 322 (1908)

= Orchis ×ullmanii P.M.Hall; Rep. Bot. Soc. Exch. Club Brit. Isles 8: 790 (1928 publ. 1929), nom. provis.

= xCoeloglossorchis ×dominiana (E.G.Camus) P.Fourn.; Quatre Fl. France 201 (1935)

= xDactyloglossum ×dominianum (E.G.Camus) Soó; Ann. Univ. Sci. Budapest. Rolando Eötvös, Sect. Biol. 8: 317 (1966)

= xDactyloglossum ×conigerum (Norman) Rauschert; Feddes Repert. 83 (9-10): 658 (1972 publ. 1973)

= xDactyloglossum ×evae R.Breiner; Ber. Arbeitskreis. Heimische Orchid. 18(2): 90, 89 (2001 publ. 2002)

= xDactyloglossum ×ullmanii (Hall) Peitz; Jahresber. Naturwiss. Vereins Wuppertal 25: 189 (1972)

= xOrchicoeloglossum ×conigerum (Norman) Nordh.; Norsk Fl. [Nordhagen]: 65 (1940)

= xOrchicoeloglossum ×dominianum E.G.Camus; Monogr. Orchid. 322 (1908)

species :

Dactylorhiza ×csatoi (Soó) Soó; Ann. Univ. Sci. Budapest. Rolando Eötvös, Sect. Biol., 4 (Geobot. Monogr. Orchid. Pannon. & Karpat. Fl. 5) 81 (1961)

Distribution: Romania

13 ROM

Remarks: Dactylorhiza cordigera x maculata

Synonyms: hidden

= Orchis ×csatoi Soó; Repert. Spec. Nov. Regni Veg. 24: 36 (1927)

species :

Dactylorhiza ×daunia W.Rossi, Ard., Cianchi, Piemont. & Bullini; Webbia 50(1): 11 (1995)

Distribution: Italy

13 ITA

Remarks: Dactylorhiza maculata ssp. saccifera x D. majalis ssp. pindica

Synonyms: hidden

= Dactylorhiza ×kerasovinensis B.Willing & E.Willing; Ber. Arbeitskreis. Heimische Orchid. 3(2): 217 (1986 publ. 1987)

species :

Dactylorhiza ×delamainii (G.Keller ex T.Stephenson) Soó; Nom. Nov. Gen. Dactylorhiza 9 (1962)

Distribution: France

12 FRA

Remarks: Dactylorhiza elata x D. maculata

Synonyms: hidden

= Orchis ×delamainii G.Keller ex T.Stephenson; Rep. Bot. Soc. Exch. Club Brit. Isles 8: 494 (1927 publ. 1928)

species :

Dactylorhiza ×dinglensis (Wilmott) Soó; Nom. Nov. Gen. Dactylorhiza 10 (1962) [nothosubsp.]

subspecies :

subsp. dinglensis

Distribution: Ireland

10 IRE

Remarks: Dactylorhiza maculata ssp. ericetorum x majalis ssp. occidentalis

Synonyms: hidden

= Orchis ×dinglensis Wilmott; Proc. Linn. Soc. London Sess. 148. 128 (1936)

subspecies :

subsp. robertsii F.Horsman; Watsonia 18(4): 397 (1991) [nothosubsp.]

Distribution: England

10 GRB

Remarks: Dactylorhiza maculata ssp. ericetorum x majalis ssp. cambrensis

Synonyms: hidden

= Dactylorhiza ×formosa nothovar. robertsii (F.Horsman) Oddone; GIROS Orch. Spont. Eur. 59(2): 356 (2016)

species :

Dactylorhiza ×drucei (A.Camus) Eccarius; Orchideengattung Dactylorhiza 589 (2016)

Distribution: England; Czech Republic; etc.

10 GRB 11 CZE

Remarks: Dactylorhiza viridis x Dactylorhiza majalis

Synonyms: hidden

= Dactylorhiza drucei (A.Camus) Oddone; GIROS Orch. Spont. Eur. 59(2): 354 (2016)

= Orchis tschiebozi G.Keller; Repert. Spec. Nov. Regni Veg. Sonderbeih. A: 2: 196 (1932)

= xDactyloglossum ×drucei (A.Camus) Soó; Ann. Univ. Sci. Budapest. Rolando Eötvös, Sect. Biol. 8: 317 (1966)

= xOrchicoeloglossum ×drucei A.Camus; Iconogr. Orchid. Europe 377 (1928)

species :

Dactylorhiza ×dubreuilhii (G.Keller & Jeanj.) Soó; Nom. Nov. Gen. Dactylorhiza 9 (1962)

Distribution: France

12 FRA

Remarks: Dactylorhiza elata (sesquipedalis) x incarnata

Synonyms: hidden

= Dactylorhiza ×bourdonii D.Tyteca & Gathoye; Orchidophile (Paris) 81: 59 (1988)

= Orchis ×dubreuilhii G.Keller & Jeanj.; Repert. Spec. Nov. Regni Veg. Sonderbeih. A: 2: 253 (1933)

species :

Dactylorhiza ×dufftii (Hausskn.) Peitz; Jahresber. Naturwiss. Vereins Wuppertal 25: 176 (1972)

subspecies :

subsp. dufftii

Distribution: Austria; Czech Republic; France; European Russia

11 AUT CZE 12 FRA 14 RUS

Remarks: Dactylorhiza incarnata x D. lapponica

Synonyms: hidden

= Dactylorhiza angustata (Arv.-Touv.) D.Tyteca & Gathoye; Orchidophile (Paris) 97: 115 (1991)

= Dactylorhiza delphinensis D.Tyteca & Gathoye; Naturalistes Belges 69(2): 52 (1988)

= Dactylorhiza incarnata nothosubsp. dufftii (Hausskn.) B.Bock; Bull. Soc. Bot. Centre-Ouest 42: 269 (2011 publ. 2012) [nothosubsp.]

= Dactylorhiza lapponica subsp. angustata (Arv.-Touv.) Kreutz; Kompend. Eur. Orchid. 44 (2004)

= Dactylorhiza ×approximata (Rouy) P.Delforge; Naturalistes Belges 89(Orchid. 21): 250 (2008)

= Dactylorhiza ×dufftiana (M.Schultze) Soó; Nom. Nov. Gen. Dactylorhiza 10 (1962)

= Orchis angustata Arv.-Touv.; Essai Pl. Dauphiné 62 (1872)

= Orchis divaricata Boreau; Fl. Centre France [Boreau], éd. 2: 522 (1849), et éd. 3, 2: 646 (1857)

= Orchis strictiflora Opiz; Naturalientausch 10: 217 (1825)

= Orchis ×approximata Rouy; Fl. France [Rouy & Foucaud] 13: 172, hybr. (1912)

= Orchis ×dufftiana M.Schultze; Orch. Deutschl. Österr. Schweiz sub No. 21 [p. 9] (1894)

= Orchis ×dufftii Hausskn.; Mitt. Geogr. Ges. Thüringen (Jena) 2: 225 (1884 publ. 1885)

subspecies :

subsp. gotlandica (Kreutz) Oddone; GIROS Orch. Spont. Eur. 59(2): 355 (2016) [nothosubsp.]

Distribution: Sweden

10 SWE

Remarks: Dactylorhiza incarnata ssp. ochroleuca x D. lapponica

Synonyms: hidden

= Dactylorhiza ×gotlandica Kreutz; Mitt. Arbeitskreises Heimische Orchideen 25(4): 436 (1993)

species :

Dactylorhiza ×elcitoi Benigni, Barigelli & Petroselli; Micol. Marche, n. s. 5(2): 41 (2021)

Distribution: Italy

13 ITA

species :

Dactylorhiza ×erdingeri (A.Kern.) B.Bock; Bull. Soc. Bot. Centre-Ouest 42: 267 (2011 publ. 2012)

Distribution: Austria; Czech Republic; France; etc.

11 AUT CZE 12 FRA

Remarks: Dactylorhiza viridis x Dactylorhiza sambucina

Synonyms: hidden

= Coeloglossum ×erdingeri A.Kern.; Oesterr. Bot. Z. 14: 140 (1864)

= Dactylorhiza ×erdingeri f. elongata (E.G.Camus) B.Bock; Bull. Soc. Bot. Centre-Ouest 42: 267 (2011 publ. 2012)

= Orchis ×erdingeri (A.Kern.) Sennholz; Verh. K.K. Zool.-Bot. Ges. Wien 41(Sitzb.): 41 (1891)

= Platanthera ×erdingeri (A.Kern.) A.Kern.; Verh. K.K. Zool.-Bot. Ges. Wien 15: 229 (1865)

= xCoeloglossorchis ×erdingeri (A.Kern.) Guétrot; Pl. Hybr. France [Guétrot] 2: 57 (1926 publ. 1927)

= xDactyloglossum ×sennholzii Peitz; Jahresber. Naturwiss. Vereins Wuppertal 25: 189 (1972), nom. nov.

= xDactyloglossum ×erdingeri (A.Kern.) Janch. ex Soó; Ann. Univ. Sci. Budapest. Rolando Eötvös, Sect. Biol. 8: 317 (1966)

= xDactyloglossum ×erdingeri nothof. elongatum (E.G.Camus) Gathoye & D.Tyteca; Lejeunia 143: 72 (1994)

= xOrchicoeloglossum ×erdingeri (A.Kern.) Asch. & Graebn.; Syn. Mitteleur. Fl. [Ascherson & Graebner] 3: 849 (1907)

= xOrchicoeloglossum ×erdingeri var. elongatum E.G.Camus; Iconogr. Orchid. Europe 2: 375 (1929)

species :

Dactylorhiza ×estonica (Klinge) Jagiello & Kuusk; Fragm. Florist. Geobot. 34 (34): 328 (1989)

Distribution: Czech Republic; France; Estonia; European Russia; etc.

11 CZE 12 FRA 14 EST RUS

Remarks: Dactylorhiza fuchsii x D. (majalis ssp.) lapponica

Synonyms: hidden

= Dactylorhiza traunsteineri var. estonica (Klinge) Soó; Nom. Nov. Gen. Dactylorhiza 6 (1962)

= Dactylorhiza ×silvae-gabretae F.Proch.; Preslia 71: 236 (1999)

= Orchis angustifolia var. estonica Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 77 (1899)

species :

Dactylorhiza ×flixensis (Gsell) Soó; Nom. Nov. Gen. Dactylorhiza 9 (1962)

Distribution: Switzerland

11 SWI

Remarks: Dactylorhiza incarnata ssp. pulchella x traunsteineri

Synonyms: hidden

= Orchis ×flixensis Gsell; Jahresber. Naturf. Ges. Graubündens, N. F. 74: 17 (1936)

species :

Dactylorhiza ×formosa (T.Stephenson & T.A.Stephenson) Soó; Nom. Nov. Gen. Dactylorhiza 10 (1962)

Distribution: England (N-England, Wales, Scotland, Outer Hebrides, Orkney Isl., Shetland Isl.); Ireland (N-Ireland, Northern Ireland); Norway

10 GRB IRE NOR

Remarks: Dactylorhiza maculata subsp. ericetorum x purpurella

Synonyms: hidden

= Orchis ×formosa T.Stephenson & T.A.Stephenson; J. Bot. 60: 33 (1922), nom. illeg.

species :

Dactylorhiza ×fourkensis B.Willing & E.Willing; Ber. Arbeitskreis. Heimische Orchid. 4(1): 61 (1987)

Distribution: Greece

13 GRC

Remarks: Dactylorhiza majalis ssp. pindica x D. sambucina

species :

Dactylorhiza ×gabretana (A.Fuchs) Soó; Nom. Nov. Gen. Dactylorhiza 10 (1962)

Distribution: Germany; etc.

11 GER

Remarks: Dactylorhiza incarnata x maculata x sambucina

Synonyms: hidden

= Orchis ×gabretana A.Fuchs; Mitt. Bayer. Bot. Ges. 3: 381 (1917)

species :

Dactylorhiza ×godferyana (Soó) Peitz; Jahresber. Naturwiss. Vereins Wuppertal 25: 177 (1972)

Distribution: England

10 GRB

Remarks: Dactylorhiza majalis x praetermissa

Synonyms: hidden

= Orchis ×godferyana Soó; Repert. Spec. Nov. Regni Veg. Sonderbeih. A: 2: 257 (1933)

= Dactylorhiza majalis nothosubsp. godferyana (Soó) M.H.J.van der Meer; Cact. Phantast. 2019(5)-1: 3 (2019)

species :

Dactylorhiza ×grandis (Druce) P.F.Hunt; Orchid Rev. 79 (935): 139 (1971) [nothosubsp.]

Distribution: England (England, Wales); France; Channel Isl. (Jersey); etc.

10 GRB 12 CIS FRA

Remarks: Dactylorhiza fuchsii x D. praetermissa

Synonyms: hidden

= Dactylorhiza ×dinglensis nothosubsp. grandis (Druce) M.H.J.van der Meer; Cact. Phantast. 2019(5)-1: 1 (2019)

variant :

var. montis-mosae (Druce) comb.ined.

Distribution: Netherlands; Belgium; Luxembourg

11 BGM LUX NET

Remarks: Dactylorhiza fuchsii x praetermissa var. integrata

Synonyms: hidden

= Dactylorhiza ×montis-mosae (Kreutz) Kreutz; Orchid. Benelux (Field Guide) 79 (2021)

= Dactylorhiza ×grandis nothosubsp. montis-mosae Kreutz; Natuurhist. Maandblad 106(5): 97-98 (2017)

species :

Dactylorhiza ×guilhotii (E.G.Camus) B.Bock; Bull. Soc. Bot. Centre-Ouest 42: 268 (2011 publ. 2012)

Distribution: France

12 FRA

Remarks: Dactylorhiza viridis x Dactylorhiza incarnata

Synonyms: hidden

= xCoeloglossorchis ×guilhotii (E.G.Camus) P.Fourn.; Quatre Fl. France 201 (1935)

= xDactyloglossum ×guilhotii (E.G.Camus) Soó; Ann. Univ. Sci. Budapest. Rolando Eötvös, Sect. Biol. 8: 317 (1966)

= xOrchicoeloglossum ×guilhotii E.G.Camus; Monogr. Orchid. 321 (1908)

species :

Dactylorhiza ×guillaumeae C.Bernard; Bull. Soc. Bot. France, Lett. Bot. 130(2): 154 (1983)

Distribution: France

12 FRA

Remarks: Dactylorhiza incarnata x sambucina

species :

Dactylorhiza ×gustavssonii H.Baumann; Mitt. Arbeitskreises Heimische Orchideen 15(1): 77 (1983)

Distribution: Greece; Turkey

13 GRC 34 TUR

Remarks: Dactylorhiza iberica x saccifera

Synonyms: hidden

= Dactylorhiza ×sultandagi Renz & Taubenheim; Notes Roy. Bot. Gard. Edinburgh 41(2): 270 (1983)

species :

Dactylorhiza ×hallii (Druce) Soó; Nom. Nov. Gen. Dactylorhiza 10 (1962) [nothosubsp.]

subspecies :

subsp. hallii

Distribution: England (S-England, Wales); Netherlands; France; Channel Isl. (Guernsey, Jersey); etc.

10 GRB 11 NET 12 CIS FRA

Remarks: Dactylorhiza maculata ssp. ericetorum x praetermissa

Synonyms: hidden

= Dactylorhiza ×dinglensis nothosubsp. hallii (Druce) M.H.J.van der Meer; Cact. Phantast. 2019(5)-1: 2 (2019)

= Orchis ×hallii Druce; Rep. Bot. Soc. Exch. Club Brit. Isles 4: 24 (1914 publ. 1915)

subspecies :

subsp. nummiana (P.Fourn.) Gathoye & D.Tyteca; Lejeunia 143: 67 (1994) [nothosubsp.]

Distribution: France

12 FRA

Remarks: Dactylorhiza maculata ssp. elodes x praetermissa

Synonyms: hidden

= Dactylorhiza ×nummiana (P.Fourn.) Soó; Nom. Nov. Gen. Dactylorhiza 10 (1962)

= Orchis ×nummiana P.Fourn.; Bull. Soc. Bot. France 78: 433 (1931)

species :

Dactylorhiza ×hepburnii (Druce) Soó; Nom. Nov. Gen. Dactylorhiza 9 (1962)

Distribution: Scotland

10 GRB

Remarks: Dactylorhiza foliosa x D. incarnata x D. fuchsii

Synonyms: hidden

= Orchis ×hepburnii Druce; Rep. Bot. Soc. Exch. Club Brit. Isles 4: 211 (1915 publ. 1916)

species :

Dactylorhiza ×insignis (T.Stephenson & T.A.Stephenson) Soó; Nom. Nov. Gen. Dactylorhiza 10 (1962)

Distribution: England

10 GRB

Remarks: Dactylorhiza praetermissa x purpurella

Synonyms: hidden

= Dactylorhiza ×salteri (T.Stephenson) Soó; Jahresber. Naturwiss. Vereins Wuppertal 21-22: 18 (1968)

= Orchis ×insignis T.Stephenson & T.A.Stephenson; J. Bot. 60: 33 (1922)

= Orchis ×salteri T.Stephenson; J. Bot. 80: 104 (1942)

species :

Dactylorhiza ×isculana Seiser; Ber. Arbeitskreis. Heimische Orchid. 19(1): 23 (2002), pro sp.

Distribution: Austria (Salzkammergut)

11 AUT

Remarks: probably Dactylorhiza fuchsii x D. viridis

species :

Dactylorhiza ×jenensis (Brand) Soó; Nom. Nov. Gen. Dactylorhiza 10 (1962)

Distribution: Europe; Germany; France

11 GER 12 FRA

Remarks: Dactylorhiza maculata ssp. ericetorum x traunsteineri (not lapponica)

Synonyms: hidden

= Orchis ×jenensis Brand; Syn. Deut. Schweiz. Fl., ed. 3 [Hallier & Brand] 3: 2430 (1904)

= Orchis ×schulzei C.Richt.; Pl. Eur. 1: 274 (1890), et J. Bot. [Morot] 6: 418 (1892)

species :

Dactylorhiza ×jestrebiensis Businsky; Preslia 61(4): 306 (1989)

Distribution: Czech Republic

11 CZE

Remarks: Dactylorhiza traunsteineri ssp. bohemica x Dactylorhiza majalis ssp. majalis

species :

Dactylorhiza ×juennensis Perko; Carinthia II, 187/107(1): 91 (1997)

Distribution: Austria

11 AUT

Remarks: Dactylorhiza traunsteineri ssp. rhaetica (not lapponica) x fuchsii

species :

Dactylorhiza ×katarana H.Baumann; Mitt. Arbeitskreises Heimische Orchideen 15(1): 78 (1983)

Distribution: Greece

13 GRC

Remarks: Dactylorhiza maculata ssp. saccifera x D. majalis ssp. kalopissii

species :

Dactylorhiza ×kelleriana (Cif. & Giacom.) Soó; Nom. Nov. Gen. Dactylorhiza 9 (1962)

Distribution: England; Ireland; etc.

10 GRB IRE

Remarks: Dactylorhiza fuchsii x Dactylorhiza traunsteinerioides

Synonyms: hidden

= xOrchicoeloglossum ×kellerianum Duffort ex G.Keller & Soó; Repert. Spec. Nov. Regni Veg. Sonderbeih. A: 2: 299 (1935), et adnot.

species :

Dactylorhiza ×kernerorum (Soó) Soó; Nom. Nov. Gen. Dactylorhiza 9 (1962) [nothosubsp.]

subspecies :

subsp. kernerorum

Distribution: England (England, Wales); Ireland (Ireland, Northern Ireland); Germany; Czech Republic; Slovakia; European Russia; etc.

10 GRB IRE 11 CZE GER SLO 14 RUS

Remarks: Dactylorhiza fuchsii x incarnata

Synonyms: hidden

= Dactylorhiza ×claudiopolitana (Simonk.) Borsos & Soó; Ann. Univ. Sci. Budapest. Rolando Eötvös, Sect. Biol., 4 (Geobot. Monogr. Orchid. Pannon. & Karpat. Fl. 5) 81 (1961)

= Dactylorhiza ×pomeranica Wucherpf.; J. Eur. Orch. 44(4): 700 (2012)

= Orchis ×ambigua var. claudiopolitana Simonk.; Enum. Fl. Transsilv. 500 (1887)

= Orchis ×curtisii Druce; Rep. Bot. Soc. Exch. Club Brit. Isles 9: 669 (1931 publ. 1932)

= Orchis ×kernerorum Soó; Notizbl. Bot. Gart. Berlin-Dahlem 9: 904 (1926)

= Orchis ×maculatoformis Rouy; Fl. France [Rouy & Foucaud] 13: 174 (1912)

= Orchis ×maculotoformis nothomorph. claudiopolitana (Simonk.) Pauca; Fl. Reipubl. Popul. Roman. (Fl. Republ. Socialist. Romania) 12: 707 (1972)

= Orchis ×rigida Höppner; Verh. Naturhist. Vereines Preuss. Rheinl. Westphalens 1924, 270-273

subspecies :

subsp. ampolai (Hautz.) Eccarius; Orchideengattung Dactylorhiza 581 (2016) [nothosubsp.]

Distribution: Italy

13 ITA

Remarks: Dactylorhiza cruenta x maculata

Synonyms: hidden

= Dactylorhiza ×ampolai Hautz.; Verh. Zool.-Bot. Ges. Wien 112: 115 (1972)

= Dactylorhiza ×kerneri nothosubsp. ampolai (Hautz.) Oddone; GIROS Orch. Spont. Eur. 59(2): 356 (2016)

= Dactylorhiza ×kernerorum nothosubsp. ampolai (Hautz.) Eccarius; Orchideengattung Dactylorhiza 581 (2016)

subspecies :

subsp. lillsundica (Kreutz) Oddone; GIROS Orch. Spont. Eur. 59(2): 356 (2016) [nothosubsp.]

Distribution: Sweden

10 SWE

Remarks: Dactylorhiza fuchsii x incarnata ssp. ochroleuca

Synonyms: hidden

= Dactylorhiza ×lillsundica Kreutz; Mitt. Arbeitskreises Heimische Orchideen 25(4): 435 (1993)

subspecies :

subsp. variabilis (Hesl.-Harr.) P.D.Sell; Fl. Gr. Brit. Ireland 5: 364 (1996) [nothosubsp.]

Distribution: England (Scotland)

10 GRB

Synonyms: hidden

= Dactylorhiza ×variabilis (Hesl.-Harr.) Soó; Nom. Nov. Gen. Dactylorhiza 9 (1962)

= Orchis ×variabilis Hesl.-Harr.; J. Bot. 79: 168 (1941), et Proc. Univ. Durham Phil. Soc. 10: 308 (1941), descr.

species :

Dactylorhiza ×komiensis Aver.; Bot. Zhurn. (Moscow & Leningrad) 67(3): 310 (1982)

Distribution: European Russia

14 RUS

Remarks: Dactylorhiza hebridensis x maculata

species :

Dactylorhiza ×kopdagiana H.Baumann; Mitt. Arbeitskreises Heimische Orchideen 15(1): 78 (1983)

Distribution: Turkey

34 TUR

Remarks: Dactylorhiza iberica x umbrosa

species :

Dactylorhiza ×koutsourana B.Willing & E.Willing; Ber. Arbeitskreis. Heimische Orchid. 6(1): 36 (1989)

Distribution: Greece

13 GRC

Remarks: Dactylorhiza baumanniana x smolikana

species :

Dactylorhiza ×krylovii (Soó) Soó; Nom. Nov. Gen. Dactylorhiza 8 (1962)

Distribution: E-European Russia; Siberia (W-Siberia)

14 RUS 30 SIB

Remarks: Dactylorhiza incarnata ssp. cruenta x D. incarnata ssp. incarnata

Synonyms: hidden

= Orchis ×krylovii Soó; Repert. Spec. Nov. Regni Veg. Sonderbeih. A: 2: 211 (1933)

species :

Dactylorhiza ×kuehnensis Presser & Riech.; J. Eur. Orch. 32(1): 109 (2000)

Distribution: Germany

11 GER

Remarks: Dactylorhiza majalis x ruthei

species :

Dactylorhiza ×kuuskiae E.Breiner & R.Breiner; Ber. Arbeitskreis. Heimische Orchid. 15(2): 34 (1998)

Distribution: Estonia

14 EST

Remarks: Dactylorhiza (majalis ssp.) baltica x vironii

Synonyms: hidden

= Dactylorhiza ×kuehnensis nothosubsp. kuuskiae (E.Breiner & R.Breiner) Eccarius; Orchideengattung Dactylorhiza 588 (2016)

species :

Dactylorhiza ×latirella (P.M.Hall) Soó; Nom. Nov. Gen. Dactylorhiza 9 (1962)

Distribution: England

10 GRB

Remarks: Dactylorhiza incarnata x (majalis ssp.) purpurella

Synonyms: hidden

= Orchis ×latirella P.M.Hall; J. Bot. 74: 329 (1936)

species :

Dactylorhiza ×maculatiformis (Rouy) Borsos & Soó; Ann. Univ. Sci. Budapest. Rolando Eötvös, Sect. Biol., 4 (Geobot. Monogr. Orchid. Pannon. & Karpat. Fl. 5) 81 (1961)

Distribution: Germany (?Baden-Württemberg, ?Bayern, Rheinland-Pfalz, Schleswig-Holstein); European Russia; etc.

11 GER 14 RUS

Remarks: Dactylorhiza incarnata x Dactylorhiza maculata

Synonyms: hidden

= Dactylorhiza ×ambigua (A.Kern.) H.Sund.

= Dactylorhiza ×carnea nothosubsp. maculatiformis (Rouy) Gathoye & D.Tyteca; Lejeunia 143: 63 (1994)

= Dactylorhiza ×claudiopolitana nothosubsp. maculatiformis (Rouy) Oddone; GIROS Orch. Spont. Eur. 59(2): 355 (2016)

= Orchis ×ambigua A.Kern.; Verh. K.K. Zool.-Bot. Ges. Wien 15: 205 (1865), nom. illeg.

= Orchis ×maculatiformis Rouy; Fl. France [Rouy & Foucaud] 13: 174 (1912)

species :

Dactylorhiza ×megapolitana (Bisse) Soó; Ann. Univ. Sci. Budapest. Rolando Eötvös, Sect. Biol. 11: 71 (1969)

Distribution: Germany (Mecklenburg-Vorpommern); European Russia

11 GER 14 RUS

Remarks: Dactylorhiza fuchsii x Dactylorhiza russowii

Synonyms: hidden

= Dactylorchis ×megapolitana Bisse; Feddes Repert. Spec. Nov. Regni Veg. 67: 186 (1963)

= Dactylorhiza ×silvae-gabretae nothosubsp. megapolitana (Bisse) Eccarius; Orchideengattung Dactylorhiza 582 (2016)

species :

Dactylorhiza ×metsowonensis B.Baumann & H.Baumann; Mitt. Arbeitskreises Heimische Orchideen 16(1): 128 (1984)

Distribution: Greece

13 GRC

Remarks: Dactylorhiza latifolia x kalopissii

species :

Dactylorhiza ×mixta (Asch. & Graebn.) B.Bock; Bull. Soc. Bot. Centre-Ouest 42: 268 (2011 publ. 2012)

Distribution: England; Slovakia; France; etc.

10 GRB 11 SLO 12 FRA

Remarks: Dactylorhiza viridis x Dactylorhiza fuchsii

Synonyms: hidden

= Habenaria ×mixta (Asch. & Graebn.) Druce; Rep. Bot. Soc. Exch. Club Brit. Isles 8: 876 (1928 publ. 1929)

= Orchis ×kaspari Domin & Podp.; Klíc Kvét. Ceskoslov. 796 (1928)

= Orchis ×mixta Domin; Sitzungsber. Königl. Böhm. Ges. Wiss. Prag, Math.-Naturwiss. Cl. 22: 7 (1902 publ. 1903), nom. illeg.

= xDactyloglossum ×mixtum (Asch. & Graebn.) P.F.Hunt; Orchid Rev. 79 (935): 140 (1971), isonym

= xDactyloglossum ×mixtum (Asch. & Graebn.) Rauschert; Feddes Repert. 79: 413 (1969)

= xOrchicoeloglossum ×mixtum Asch. & Graebn.; Syn. Mitteleur. Fl. [Ascherson & Graebner] 3: 847 (1907)

species :

Dactylorhiza ×mortonii (Druce) Soó; Nom. Nov. Gen. Dactylorhiza 9 (1962)

Distribution: England; etc.

10 GRB

Remarks: Dactylorhiza fuchsii x D. majalis ssp. elatior

Synonyms: hidden

= Dactylorhiza ×grandis nothosubsp. robustior (Hartm.) J.M.H.Shaw; Orchid Rev. 125 (1319, Suppl.): 57 (2017)

= Orchis angustifolia f. robustior (Hartm.) Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 83 (1899)

= Orchis latifolia var. robustior Hartm.; Handb. Skand. Fl., ed. 3: 204 (1838)

= Orchis ×grandis Druce; Fl. Buckinghamshire 328 (1926)

= Orchis ×mortonensis E.G.Camus & A.Camus; Iconogr. Orchid. Europe 280 (1928)

= Orchis ×mortonii Druce; Rep. Bot. Soc. Exch. Club Brit. Isles 8: 419 (1927 publ. 1928)

species :

Dactylorhiza ×nevskii H.Baumann & Künkele; Mitt. Arbeitskreises Heimische Orchideen 13(2): 262 (1981)

Distribution: Turkey

34 TUR

Remarks: Dactylorhiza osmanica x umbrosa

species :

Dactylorhiza ×ornonensis (G.Keller & Jeanj.) Soó; Nom. Nov. Gen. Dactylorhiza 9 (1962)

Distribution: France

12 FRA

Remarks: Dactylorhiza elata ssp. sesquipedalis x incarnata x maculata

Synonyms: hidden

= Orchis ×ornonensis G.Keller & Jeanj.; Actes Soc. Linn. Bordeaux 89: Proc.-Verb. 34 (1937)

species :

Dactylorhiza ×panoramica Willing; Mitt. Arbeitskreises Heimische Orchideen 17(4): 516 (1985)

Distribution: Greece

13 GRC

Remarks: Dactylorhiza baumanniana x saccifera

species :

Dactylorhiza ×paridaeniana Kreutz; Eurorchis 2: 65 (1990)

Distribution: Netherlands

11 NET

species :

Dactylorhiza ×perez-chiscanoi F.M.Vázquez; J. Eur. Orch. 40(1): 36 (2008)

Distribution: Spain

12 SPA

Remarks: Dactylorhiza elata x D. lapponica

species :

Dactylorhiza ×predaensis (Gsell) Soó ex Rauschert; Feddes Repert. 85 (9-10): 656 (1974)

Distribution: Switzerland; European Russia

11 SWI 14 RUS

Remarks: Dactylorhiza (incarnata ssp.) cruenta x Dactylorhiza majalis

Synonyms: hidden

= Dactylorchis ×predaensis Gsell ex Soó in Rothm.; 88 (1963), nom. inval.

= Dactylorhiza ×aschersoniana nothosubsp. predaensis (Gsell) Oddone; GIROS Orch. Spont. Eur. 59(2): 355 (2016)

= Dactylorhiza ×predaensis Gsell ex Soó ex Soó; 656 (1974), nom. inval.

= Dactylorhiza ×scampstonii (Druce) Soó; Nom. Nov. Gen. Dactylorhiza 9 (1962)

= Orchis ×predaensis Gsell; Jahresber. Naturf. Ges. Graubündens, N. F. 74: 17 (1936), nom. inval.

= Orchis ×scampstonii Druce; Rep. Bot. Soc. Exch. Club Brit. Isles 5: 53 (1917 publ. 1918)

species :

Dactylorhiza ×prochazkana Businsky; Preslia 61(4): 305 (1989)

Distribution: Czech Republic

11 CZE

Remarks: Dactylorhiza bohemica x Dactylorhiza maculata ssp. maculata

species :

Dactylorhiza ×reitaluae Hennecke, E.Breiner & R.Breiner; Ber. Arbeitskreis. Heimische Orchid. 15: 36 (1998)

Distribution: Estonia

14 EST

Remarks: Dactylorhiza incarnata x vironii [described as incarnata x ruthei]

species :

Dactylorhiza ×renzii H.Baumann & Künkele; Mitt. Arbeitskreises Heimische Orchideen 13(2): 261 (1981)

Distribution: Turkey

34 TUR

Remarks: Dactylorhiza incarnata x nieschalkiorum

species :

Dactylorhiza ×rizeana Renz & Taubenheim; Notes Roy. Bot. Gard. Edinburgh 41(2): 269 (1983)

Distribution: Turkey

34 TUR

Remarks: Dactylorhiza euxina x urvilleana

species :

Dactylorhiza ×rombucina (Cif. & Giacom.) Soó; Nom. Nov. Gen. Dactylorhiza 10 (1962)

Distribution: C-Europe

Error

Remarks: Dactylorhiza romana x sambucina

Synonyms: hidden

= Orchis ×rombucina Cif. & Giacom.; Nomenclator Fl. Ital. 1: 165 (1950)

species :

Dactylorhiza ×ruppertii (M.Schultze) Borsos & Soó; Ann. Univ. Sci. Budapest. Rolando Eötvös, Sect. Biol., 3 (Geobot. Monogr. Orchid. Pannon. & Karpat. Fl. 4) 122 (1960)

Distribution: Germany; Czech Republic; Slovakia; etc.

11 CZE GER SLO

Remarks: Dactylorhiza majalis ssp. majalis x Dactylorhiza sambucina

Synonyms: hidden

= Orchis ×ruppertii M.Schultze; Oesterr. Bot. Z. 49: 264 (1899)

species :

Dactylorhiza ×salictina B.Willing & E.Willing; Ber. Arbeitskreis. Heimische Orchid. 6(1): 35 (1989)

Distribution: Greece

13 GRC

Remarks: Dactylorhiza pindica x smolikana

Synonyms: hidden

= Dactylorhiza ×souflikensis nothosubsp. salictina (B.Willing & E.Willing) Eccarius; Orchideengattung Dactylorhiza 580 (2016)

species :

Dactylorhiza ×samnaunensis (Gsell) Soó; Nom. Nov. Gen. Dactylorhiza 8 (1962)

Distribution: Switzerland; European Russia

11 SWI 14 RUS

Remarks: Dactylorhiza cruenta x Dactylorhiza maculata

Synonyms: hidden

= Orchis ×samnaunensis Gsell; Jahresber. Naturf. Ges. Graubündens, N. F. 74: 17 (1936)

species :

Dactylorhiza ×serbica (H.Fleischm.) Soó; Nom. Nov. Gen. Dactylorhiza 9 (1962)

Distribution: Serbia

13 SRB

Remarks: Dactylorhiza incarnata x saccifera

Synonyms: hidden

= Dactylorhiza ×naris W.Rossi & Mazzola; Atti Soc. Tosc. Sci. Nat. Pisa, Mem., Ser. B, 89: 271 (1982 publ. 1983)

= Orchis ×serbica H.Fleischm.; Mitt. Naturwiss. Vereines Steiermark 45: 179 (1908 publ. 1909)

species :

Dactylorhiza ×serreana H.Baumann; Mitt. Arbeitskreises Heimische Orchideen 15(1): 79 (1983)

Distribution: Greece

13 GRC

Remarks: Dactylorhiza majalis ssp. cordigera x D. cordigera ssp. bosniaca

species :

Dactylorhiza ×sivasiana Baumann & Künkele ex Renz & Taubenheim; Notes Roy. Bot. Gard. Edinburgh 41(2): 270 (1983)

Distribution: Turkey

34 TUR

Remarks: Dactylorhiza umbrosa x urvilleana

species :

Dactylorhiza ×sooi (Ruppert ex Soó) Soó; Nom. Nov. Gen. Dactylorhiza 10 (1962)

Distribution: C-Europe

Error

Remarks: Dactylorhiza majalis var. alpestris x fuchsii

Synonyms: hidden

= Orchis ×sooi Ruppert ex Soó; Monogr. Iconogr. Orchid. Eur. 2: 398 (1936)

species :

Dactylorhiza ×souflikensis B.Willing & E.Willing; Ber. Arbeitskreis. Heimische Orchid. 4(1): 60 (1987)

Distribution: Greece

13 GRC

Remarks: Dactylorhiza baumanniana x pindica

species :

Dactylorhiza ×stagni-novi D.Tyteca & Gathoye; Orchidophile (Paris) 81: 58 (1988)

Distribution: Spain; France

12 FRA SPA

Remarks: Dactylorhiza brennensis x fuchsii

Synonyms: hidden

= Dactylorhiza ×hjertsonii P.P.Ferrer, J.L.Lozano, R.Roselló, Feliu & Peña-Riv.; Fl. Montiber. 71: 100 (2018)

species :

Dactylorhiza ×stenostachys (Murr) Rauschert; Feddes Repert. 85 (9- 101: 656 (1974)

subspecies :

subsp. stenostachys

Distribution: Switzerland; European Russia

11 SWI 14 RUS

Remarks: Dactylorhiza incarnata ssp. incarnata x D. (majalis ssp.) lapponica

Synonyms: hidden

= Dactylorhiza ×thellungiana (Braun-Blanq.) Soó; Nom. Nov. Gen. Dactylorhiza 9 (1962)

= Orchis instabilis G.Keller ex G.Keller & Soó; Repert. Spec. Nov. Regni Veg. Sonderbeih. A: 2: 252 (1933)

= Orchis ×stenostachys Murr; Neue Übers. Bl.-Pfl. Vorarlberg 281 (1923)

= Orchis ×thellungiana Braun-Blanq.; Veröff. Geobot. Inst. Rübel Zürich 7: 349 (1932)

subspecies :

subsp. lehmannii (Klinge) Eccarius; Orchideengattung Dactylorhiza 586 (2016) [nothosubsp.]

Distribution: Germany (Mecklenburg-Vorpommern); European Russia

11 GER 14 RUS

Remarks: Dactylorhiza incarnata var. longebracteata x Dactylorhiza traunsteineri ssp. russowii

Synonyms: hidden

= Dactylorhiza ×lehmannii (Klinge) Soó; Nom. Nov. Gen. Dactylorhiza 9 (1962)

= Orchis ×lehmannii Klinge; Revis. Orchis cordigera & O. angustifolia 102 (1893)

subspecies :

subsp. stenkyrkae (Hautz.) Eccarius; Orchideengattung Dactylorhiza 586 (2016) [nothosubsp.]

Distribution: Sweden; Switzerland; etc.

10 SWE 11 SWI

Remarks: Dactylorhiza incarnata subsp. cruenta x traunsteineri

Synonyms: hidden

= Dactylorhiza ×dufftii nothosubsp. stenkyrkae (Hautz.) Oddone; GIROS Orch. Spont. Eur. 59(2): 355 (2016)

= Dactylorhiza ×engadinensis (Cif. & Giacom.) Soó; Nom. Nov. Gen. Dactylorhiza 8 (1962), nom. inval.

= Dactylorhiza ×stenkyrkae Hautz.; Verh. Zool.-Bot. Ges. Wien 115: 48 (1976)

= Orchis ×engadinensis Cif. & Giacom.; Nomenclator Fl. Ital. 1: 162 (1950)

species :

Dactylorhiza ×subalpina Anghelescu, L.Balogh, Mih.Balogh & Kigyossy; Sci. Papers, Ser. B Hort. 68(2): 634 (2024)

Distribution: Romania

13 ROM

Remarks: Dactylorhiza cordigera subsp. siculorum x incarnata

species :

Dactylorhiza ×surensis (Gsell) Soó; Nom. Nov. Gen. Dactylorhiza 9 (1962)

Distribution: Switzerland

11 SWI

Remarks: provisionally accepted name

Synonyms: hidden

= Orchis ×surensis Gsell; Jahresber. Naturf. Ges. Graubündens, N. F. 74: 17 (1936)

species :

Dactylorhiza ×szaboiana (Soó) Soó; Ann. Univ. Sci. Budapest. Rolando Eötvös, Sect. Biol., 4 (Geobot. Monogr. Orchid. Pannon. & Karpat. Fl. 5) 81 (1961)

Distribution: Hungary; Romania

11 HUN 13 ROM

Remarks: Dactylorhiza maculata ssp. maculata x majalis ssp. cordigera

Synonyms: hidden

= Dactylorhiza ×csatoi nothosubsp. szaboiana (Soó) Eccarius; Orchideengattung Dactylorhiza 580 (2016)

= Orchis ×csatoi var. szaboiana Soó; Repert. Spec. Nov. Regni Veg. 24: 36 (1927)

= Orchis ×szaboiana (Soó) Soó; Arch. Bot. (Leipzig) 23: 125 (1928)

species :

Dactylorhiza ×transiens (Druce) Soó; Nom. Nov. Gen. Dactylorhiza 9 (1962) [nothosubsp.]

subspecies :

subsp. transiens

Distribution: England (England, Wales, Isle of Man, Scotland, Outer Hebrides); Ireland (Ireland, Northern Ireland); European Russia; Slovakia; France; Channel Isl. (Jersey); etc.

10 GRB IRE 11 SLO 12 CIS FRA 14 RUS

Remarks: Dactylorhiza fuchsii x Dactylorhiza maculata. Probably widespread but difficult to recognize and therefore under-recorded

Synonyms: hidden

= Dactylorhiza maculata nothosubsp. transiens (Druce) M.H.J.van der Meer; Cact. Phantast. 2019(5)-1: 2 (2019)

= Orchis ×transiens Druce; Rep. Bot. Soc. Exch. Club Brit. Isles 4: 213 (1915 publ. 1916)

subspecies :

subsp. corylensis (Hesl.-Harr.) P.D.Sell; Fl. Gr. Brit. Ireland 5: 364 (1996) [nothosubsp.]

Distribution: England (Scotland)

10 GRB

Synonyms: hidden

= Dactylorhiza ×corylensis (Hesl.-Harr.) Soó; Nom. Nov. Gen. Dactylorhiza 9 (1962)

= Orchis ×corylensis Hesl.-Harr.; J. Bot. 79: 168 (1941), et Proc. Univ. Durham Phil. Soc. 10: 308 (1941), descr.

species :

Dactylorhiza ×turcestanica (G.Keller & Soó) Oddone; GIROS Orch. Spont. Eur. 59(2): 354 (2016)

Distribution: Tajikistan

32 TZK

Remarks: Dactylorhiza viridis x D. umbrosa

Synonyms: hidden

= xOrchicoeloglossum ×turcestanicum G.Keller & Soó; Repert. Spec. Nov. Regni Veg. Sonderbeih. A: 2: 299 (1935)

= xDactyloglossum ×turcestanicum (G.Keller & Soó) Soó; Ann. Univ. Sci. Budapest. Rolando Eötvös, Sect. Biol. 8: 317 (1966)

species :

Dactylorhiza ×vallis-peenae Kümpel; Ber. Arbeitskreis. Heimische Orchid. 18(2): 28 (2001 publ. 2002)

Distribution: Germany

11 GER

Remarks: Dactylorhiza majalis x Dactylorhiza traunsteineri ssp. russowii

species :

Dactylorhiza ×venusta (T.Stephenson & T.A.Stephenson) Soó; Nom. Nov. Gen. Dactylorhiza 9 (1962) [nothosubsp.]

subspecies :

subsp. venusta

Distribution: England (N-England, Wales, Scotland, Shetland Isl.); Ireland (Ireland, Northern Ireland)

10 GRB IRE

Remarks: Dactylorhiza fuchsii x purpurella

Synonyms: hidden

= Orchis ×venusta T.Stephenson & T.A.Stephenson; J. Bot. 60: 34 (1922)

subspecies :

subsp. hebridella (Wilmott) P.D.Sell; Fl. Gr. Brit. Ireland 5: 364 (1996) [nothosubsp.]

Distribution: England (Outer Hebrides)

10 GRB

Remarks: Dactylorhiza fuchsii ssp. hebridensis x purpurella

Synonyms: hidden

= Dactylorhiza ×hebridella (Wilmott) Soó; Nom. Nov. Gen. Dactylorhiza 9 (1962)

= Orchis ×hebridella Wilmott; J. Bot. 77: 193 (1939)

species :

Dactylorhiza ×vermeuleniana Soó; Jahresber. Naturwiss. Vereins Wuppertal 21-22: 17 (1968), et Acta Bot. Acad. Sci. Hung. 18 (1- 2): 174 (1973)

Distribution: Czech Republic; etc.

11 CZE

Remarks: Dactylorhiza incarnata x maculata x majalis

Synonyms: hidden

= Dactylorhiza ×senayi nothosubsp. vermeuleniana (Soó) Oddone; GIROS Orch. Spont. Eur. 59(2): 356 (2016)

variant :

var. senayi (Alleiz.) Kümpel; Ber. Arbeitskreis. Heimische Orchid. 18(2): 28 (2001 publ. 2002) [nothovar.]

Distribution: France

12 FRA

Remarks: Dactylorhiza maculata ssp. elodes x majalis

Synonyms: hidden

= Dactylorhiza ×senayi (Alleiz.) Soó; Nom. Nov. Gen. Dactylorhiza 10 (1962)

= Orchis ×senayi Alleiz.; Bull. Soc. Bot. France 94: 338 (1948)

variant :

var. townsendiana (Rouy) Kümpel; Ber. Arbeitskreis. Heimische Orchid. 18(2): 29 (2001 publ. 2002) [nothovar.]

Distribution: Germany

11 GER

Remarks: Dactylorhiza maculata ssp. maculata x D. majalis ssp. majalis

Synonyms: hidden

= Dactylorhiza ×senayi nothosubsp. townsendiana (Rouy) Oddone; GIROS Orch. Spont. Eur. 59(2): 356 (2016)

= Dactylorhiza ×townsendiana (Rouy) Soó; Nom. Nov. Gen. Dactylorhiza 10 (1962)

= Orchis ×braunii var. townsendiana Rouy; Fl. France [Rouy & Foucaud] 13: 173 (1912)

species :

Dactylorhiza ×viridella (Hesl.-Harr.) Oddone; GIROS Orch. Spont. Eur. 59(2): 354 (2016)

Distribution: England

10 GRB

Remarks: Dactylorhiza viridis x D. purpurella

Synonyms: hidden

= Orchis ×viridella Hesl.-Harr.; Vasculum 34: 22 (1949)

= xDactyloglossum ×viridellum (Hesl.-Harr.) Soó; Jahresber. Naturwiss. Vereins Wuppertal 21-22: 18 (1968)

species :

Dactylorhiza ×vitosana H.Baumann; Mitt. Arbeitskreises Heimische Orchideen 15(1): 79 (1983)

Distribution: Bulgaria

13 BUL

Remarks: Dactylorhiza lagotis x saccifera

species :

Dactylorhiza ×vogtiana H.Baumann; Mitt. Arbeitskreises Heimische Orchideen 15(1): 80 (1983)

Distribution: Turkey

34 TUR

Remarks: Dactylorhiza iberica x incarnata

species :

Dactylorhiza ×vorasica B.Willing & E.Willing; Ber. Arbeitskreis. Heimische Orchid. 4(1): 63 (1987)

Distribution: Greece

13 GRC

Remarks: Dactylorhiza majalis ssp. cordigera x sambucina

species :

Dactylorhiza ×weissenbachiana Perko; Carinthia II, 184/104(1): 211 (1994)

Distribution: Austria

11 AUT

Remarks: Dactylorhiza incarnata x lapponica

species :

Dactylorhiza ×wiefelspuetziana D.Tyteca; Bull. Soc. Roy. Bot. Belgique 114(1): 24 (1981)

Distribution: Belgium

11 BGM

Remarks: Dactylorhiza maculata ssp. arduennensis x sphagnicola

Synonyms: hidden

= Dactylorhiza ×senayi nothosubsp. wiefelspuetziana (D.Tyteca) Oddone; GIROS Orch. Spont. Eur. 59(2): 356 (2016)

Loading ...

Prev genus

Top

Next genus

A complete, synonymic checklist of the Higher Plants of the World

Search

+ Family

Filter

Order

Family (14)

Genus (738)

Display options

A complete, synonymic checklist of the Higher Plants of the World

Version 25.03 (March 6th, 2025)

Covering 356,112 accepted species,
8,776 hybrids, 48,085 accepted infraspecific taxa and 1,002,454 synonyms,
in total 1,415,427* names

[about 25,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus*" are deliberately omitted]

With complete, detailed distribution data (countries and regions) and literature references

Arranged in linear phylogenetic sequence reflecting true relationships

Fully searchable and continously updated

With 38,209 pictures of 15,787 species

Assembled through 40 years independently from other databases

In cooperation with Catalogue of Life

Contact

How to Cite:

Search

+ Family

Filter

Order

Family (14)

Genus (738)

Display options

A complete, synonymic checklist of the Higher Plants of the World

Version 25.03 (March 6th, 2025)

Covering 356,112 accepted species, 8,776 hybrids, 48,085 accepted infraspecific taxa and 1,002,454 synonyms, in total 1,415,427* names

[*about 25,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus" are deliberately omitted]

With complete, detailed distribution data (countries and regions) and literature references

Arranged in linear phylogenetic sequence reflecting true relationships

Fully searchable and continously updated

With 38,209 pictures of 15,787 species

Assembled through 40 years independently from other databases

In cooperation with Catalogue of Life

Covering 356,112 accepted species,
8,776 hybrids, 48,085 accepted infraspecific taxa and 1,002,454 synonyms,
in total 1,415,427* names

[about 25,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus*" are deliberately omitted]