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Ver. 24.11
November 5th, 2024

044.0000 Family Thelypteridaceae Ching ex Pic.Serm.; Webbia 24: 711 (1970)
5-34 genera (depending on genus concept), 1100-1200 spp. New genus concept adopted here from Fawcett & Smith (2021), based on molecular results.
Species listed in database: 1180; hybrids: 38
044.2012 Genus Cyclosorus Link; Hort. Berol. [Link] 2: 128 (1833) sensu stricto
Distribution: 1 pan- and subtropical, 1 Africa
Species listed in database: 3
= Dryopteris subgen.Cyclosorus (Link) C.Chr.; Index Filic. 21 (1906)
= Thelypteris subgen.Cyclosoriopsis K.Iwats.; Mem. Coll. Sci. Univ. Kyoto, Ser. B, Biol. 31: 28 (1964)
= Cyclosorus subgen.Cyclosoriopsis (K.Iwats.) Panigrahi; Abstr. & Souv. Nation. Symp. Curr. Trends Pterid. 4-6 October 1991: 13 (1991)
= Thelypteris subgen.Cyclosorus (Link) C.V.Morton; Amer. Fern J. 53: 153 (1963)
= Nephrodium Schott; Gen. Fil. (Vindob.) t. 10 (1834)
species :
Cyclosorus interruptus (Willd.) H.Itô; Bot. Mag. (Tokyo) 51: 714 (1937)
Name: Hottentot Fern, Willdenows maiden fern
Distribution: Morocco [I]; Algeria [I]; Senegal; Gambia; Guinea; Liberia; Ivory Coast; Niger; Bioko Isl. [Fernando Poo]; Bioko Isl. [Fernando Poo]; Congo [Brazzaville]; Central African Republic; D.R.Congo [Zaire]; Rwanda; Burundi; Chad; Sudan; Sudan; Ethiopia; Ethiopia; Uganda; Kenya; Pemba Isl.; Tanzania; Zanzibar; Angola; Zambia; Malawi; Zimbabwe; Mozambique; Madagascar; Comores (Mayotte); Mauritius; La Réunion; Chagos Arch. (Diego Garcia); China (Fujian, Guangdong, Guangxi, Hainan, Jiangxi, S-Yunnan, Macao); South Korea (Cheju Isl.); Japan; Ryukyu Isl.; Taiwan; Nepal; India (Andhra Pradesh, Arunachal Pradesh, Assam State, Goa, Karnataka, Kerala, Madhya Pradesh, Manipur, Maharashtra, Meghalaya, Nagaland, Odisha, Tamil Nadu, Tripura, West Bengal); Maldives; Sri Lanka; Bangladesh; Andaman Isl.; Myanmar; Thailand; Laos; Cambodia; Vietnam; peninsular Malaysia; Singapore; Sumatra; Borneo; Brunei; Sulawesi; Palawan; Philippines; Java; Lesser Sunda Isl. (Bali, Lombok, Alor, Sumbawa, Flores, Timor); Moluccas; New Guinea (Irian Jaya, Papua New Guinea); Bismarck Arch. (Admiralty Isl.); Solomon Isl. (Rennell, Tulagi); Australia (Western Australia, Northern Territory, E-Queensland, NE-New South Wales, Victoria); New Zealand (North Isl.); New Caledonia; Vanuatu; Rotuma Isl. [I]; Tonga (Tongatapu); Fiji (Viti Levu); American Samoa (Tutuila, Aunu´u, Ta´u); [Western] Samoa (Savai´i, ´Upolu); Cook Isl. (Mangaia, Aitutaki, Atiu, Mauke); Society Isl. (Tahiti, Hitiaa); Austral Isl. (Rimatara, Rurutu); Easter Isl.; Easter Isl.; Palau (Angaur, Namonuito); Guam; Fed. States of Micronesia (Truk, Piis, Lukunor, Satawan, Pohnpei, Kosrae); Marshall Isl.. (Arno, Jaluit); Hawaii (Kauai, Oahu, Molokai, Lanai, Maui, Hawaii Isl.); USA (Florida, SE-Louisiana); Mexico (Campeche, Chiapas, Hidalgo, Jalisco, Nayarit, Quintana Roo, Tabasco, Veracruz); Guatemala; El Salvador; Honduras; Nicaragua; Costa Rica; Panama; Cuba; Cayman Isl. (Grand Cayman); Jamaica; Hispaniola (Haiti, Dominican Republic); Puerto Rico; St. Kitts; Montserrat; Guadeloupe; Marie-Galante; Martinique; Colombia (Antioquia, Bolívar, Cundinamarca, Valle); Venezuela (Anzoategui, Apure, Aragua, Barinas, Bolívar, Carabobo, Cojedes, Venezuela Distrito Federal, Falcon, Guarico, Lara, Merida, Miranda, Monagas, Portuguesa, Sucre, Tachira, Trujillo, Zulia); Guyana; Surinam; French Guiana; Ecuador; Galapagos; Peru; Bolivia (Beni, Chuquisaca, Cochabamba, Pando, Santa Cruz); Brazil (Alagoas, Amazonas, Amapá, Bahia, Ceará, Brasilia Distrito Federal, Espírito Santo, Goiás, Maranhão, Minas Gerais, Mato Grosso do Sul, Mato Grosso, Paraíba, Pernambuco, Paraná, Rio de Janeiro, Rio Grande do Sul, Santa Catarina, São Paulo); Paraguay (Alto Paraná, Central, Cordillera, Guaira, Itapua, Misiones, Neembucu, Paraguari, Pres. Hayes, San Pedro); Uruguay (Rio Negro); Argentina (Buenos Aires, Chaco, Corrientes, Entre Rios, Jujuy, Misiones, Santa Fe)
20 alg mor 22 GAM GUI IVO LBR NGR SEN 23 BIO BUR CAF CON RWA ZAI 24 CHA ETH SUD 25 KEN TAN UGA 26 ANG MLW MOZ ZAM ZIM 29 CGS COM MAU MDG REU 36 CHI 38 JAP KOR NNS TAI 40 BAN IND MDV NEP SRL 41 AND CBD LAO MYA THA VIE 42 BOR BRN JAW LSI MLY MOL NWG PHI PLW SNG SUL SUM 43 BIS SOL 50 AUS 51 NZL 60 ASA FIJ NWC rot SAM TON VAN 61 AUI COO EAS SCI 62 GUM MIC PLU 63 HAW 78 USA 79 MEX 80 COS ELS GUA HON NIC PAN 81 CAY CUB GDL HIS JAM MAR MTS PUE STK 82 CLM FRG GUY SUR VEN 83 BOL ECU GAL PER 84 BRA 85 ARG PAR URU
Synonyms: hidden
= Aspidium aneitense E.Fourn.; Ann. Sci. Nat., Bot., sér. 5, 18: 297 (1873)
= Aspidium continuum Desv.; Mag. Neuesten Entdeck. Gesammten Naturk. Ges. Naturf. Freunde Berlin 5: 320 (1811)
= Aspidium dissectum Mett.; Ann. Mus. Bot. Lugduno-Batavi 1: 232 (1863) (non Polypodium dissectum G. Forst. (1786))
= Aspidium ecklonii Kunze; Linnaea 10: 546 (1836)
= Aspidium gongylodes Schkuhr; Deutschl. Krypt. Gew. 1: 193, t. 33 C (1809) (goggilodus)
= Aspidium lucens Bojer ex Kunze; Linnaea 28: 228 (1850)
= Aspidium obtusatum Sw.; J. Bot. (Schrader) 1801(2): 33 (1800 publ. 1801)
= Aspidium paludosum (Liebm.) Mett.; Abh. Senckenberg. Naturf. Ges. 2(2) [Pheg. u. Asp.]: 385 (1858), nom. illeg.
= Aspidium plantianum (Pappe & Rawson) Kuhn; Filic. Afr. 139 (1868)
= Aspidium pohlianum C.Presl; Delic. Prag. 1: 173 (1822)
= Aspidium propinquoides Mett. ex Salomon; Nomencl. Gefäßkrypt. 66 (1883)
= Aspidium propinquum (R.Br.) Sw.; Adnot. Bot. 67 (1829)
= Aspidium resiniferum Kaulf.; Enum. Filic. [Kaulfuss] 237 (1824)
= Aspidium serra Schkuhr; 24. Kl. Linn. Pfl.-Syst. 1: 35 (1806), nom. illeg.
= Aspidium serratum Sw.; Syn. Fil. [Swartz] 47, 246 (1806)
= Aspidium unitum var. hirsutum Mett.; Ann. Mus. Bot. Lugduno-Batavi 1: 230 (1863)
= Aspidium unitum var. propinquum (R.Br.) F.M.Bailey; Syn. Queensland Fl. p. ? (1883)
= Aspidium venulosum Blume; Enum. Pl. Javae 2: 151 (1828)
= Cyclosorus goggilodus f. paucipilosus H.Itô; Bot. Mag. (Tokyo) 51(608): 714 (1937)
= Cyclosorus goggilodus var. hirsutus (Mett.) Farw.; Amer. Midl. Naturalist 12: 259 (1931)
= Cyclosorus gongylodes (Schkuhr) Link; Hort. Berol. [Link] 2: 128 (1833) (goggilodus)
= Cyclosorus gongylodes var. glaber (Mett.) H.Itô; Bot. Mag. (Tokyo) 51: 714 (1917)
= Cyclosorus gongylodes var. hirsutus (Mett.) Farw.; Amer. Midl. Naturalist 12(8): 259 (1931)
= Cyclosorus interruptus (Willd.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 184 (1938), nom. superfl.
= Cyclosorus lenormandii (C.Chr.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 246 (1941)
= Dryopteris gongylodes (Schkuhr) Kuntze; Revis. Gen. Pl. 2: 811 (1891), as ´goggilodus´, emend. Fosberg, Occas. Pap. Bernice P. Bishop Mus. 16: 337 (1942)
= Dryopteris gongylodes subsp. propinqua (R.Br.) C.Chr.; Index Filic. 268 (1905)
= Dryopteris gongylodes var. glabra (Mett.) C.Chr.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 7, 10(2): 193 (1913)
= Dryopteris gongylodes var. glabra (Mett.) H.Itô; J. Jap. Bot. 11: 786 (1935), isonym
= Dryopteris gongylodes var. hirsuta (Mett.) Bonap.; Notes Ptéridol. 1: 49, 110 (1915), isonym.
= Dryopteris gongylodes var. hirsuta (Mett.) C.Chr.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 7, 10, pt. 2: 193 (1913)
= Dryopteris gongylodes var. longipinna C.Chr.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 7, 10(2): 194 (1913)
= Dryopteris gongylodes var. longipinnulata Bonap.; Notes Ptéridol. 15: 13 (1924) nom. nud.
= Dryopteris gongylodes var. propinqua (R.Br.) Bonap.; Bull. Jard. Bot. État. Bruxelles 4: 31 (1914)
= Dryopteris interrupta (Willd.) Ching; Lingnan Sci. J. 12: 566 (1933)
= Dryopteris lenormandii C.Chr.; Index Filic. 274 (1906)
= Dryopteris obtusata (Sw.) F.Ballard; Bull. Misc. Inform. Kew 1939: 75 (1939)
= Dryopteris propinqua (R.Br.) Gilib.; Bull. Torrey Bot. Club 4: 659 (1896)
= Dryopteris unita (Sw.) Maxon; Proc. U.S. Natl. Mus. 23: 639 (1901)
= Filix goggilodes (Schkuhr) Farw.; Rep. (Annual) Michigan Acad. Sci. 18: 81-83 (1916)
= Goniopteris cheilocarpa Fée; Mém. Foug. 5 (Gen. Filic.): 251 (1852)
= Goniopteris pennigera C.Presl; Tent. Pterid. 183 (1836)
= Hypopeltis marginifera Bory; Voy. Indes Or. [Bélanger] 2: 69 (1833)
= Hypopeltis propinquoides Bory; Voy. Indes Or. [Bélanger] 2: 69 (1833)
= Nephrodium aneitense (E.Fourn.) Baker; Ann. Bot. (Oxford) 6: 326 (1891)
= Nephrodium consanguineum T.Moore; Index Fil. [T. Moore] 101 (1858)
= Nephrodium contiguum Schott; Gen. Fil. (Vindob.) ad t. 10 (1834)
= Nephrodium continuum (Desv.) Desv.; Mem. Soc. Linn. Paris 6(2): 254 (1827)
= Nephrodium dissectum (Mett.) Baker; Syn. Fil. [W. J. Hooker & Baker], ed. 1: 290 (1867)
= Nephrodium ecklonianum (Kunze) Pappe & Rawson; Syn. Filic. Afr. Austr. 14 (1858)
= Nephrodium ecklonii (Kunze) C.Presl; Abh. Königl. Böhm. Ges. Wiss., ser. 5, 6 [Epim. Bot.]: 49 (1851)
= Nephrodium gongylodes (Schkuhr) Schott; Gen. Fil. (Vindob.) ad t. 10 (1834), as ´gongyloides´
= Nephrodium gongylodes var. ecklonii (Kunze) Kümmerle; Magyar Bot. Lapok 32: 60 (1933)
= Nephrodium gongylodus (Schkuhr) Watts; J. Bot. 153 (1920), isonym
= Nephrodium inaequilaterum Colenso; Trans. & Proc. New Zealand Inst. 20: 229 (1888)
= Nephrodium lucens Bojer; Fée, Mém. Foug. 5 (Gen. Filic.): 305 (1850)
= Nephrodium microcarpum C.Presl; Tent. Pterid. 81 (1836)
= Nephrodium obtusatum (Sw.) C.Presl; Reliq. Haenk. 1: 35 (1825)
= Nephrodium obtusatum (Sw.) Desv.; Mem. Soc. Linn. Paris 6(2): 254 (1827), isonym
= Nephrodium paludosum Liebm.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 5, 1: 275 (1849)
= Nephrodium plantianum Pappe & Rawson; Syn. Filic. Afr. Austr. 14 (1858)
= Nephrodium pohlianum (C.Presl) C.Presl; Tent. Pterid. 81 (1836)
= Nephrodium propinquum R.Br.; Prodr. Fl. Nov. Holland. 148 (1810)
= Nephrodium pteroides (Retz.) Hook. & Baker; Syn. Fil. [W. J. Hooker & Baker] 289 (1867)
= Nephrodium resiniferum (Kaulf.) Hook. & Arn.; Bot. Beechey Voy. 105 (1832)
= Nephrodium serratum Desv.; Mem. Soc. Linn. Paris 6(2): 254 (1827)
= Nephrodium unitum var. glabrum Davenp.; List N. Amer. Pterid.: 17 (1901)
= Nephrodium unitum var. propinquum (R.Br.) Baker; Fl. Bras. [Martius] 1(2): 487 (1870)
= Nephrodium venulosum Desv.; Mem. Soc. Linn. Paris 6(2): 255 (1827)
= Phymatodes secunda C.Presl; Tent. Pterid. 197 (1836)
= Polypodium pennigerum C.Presl; Reliq. Haenk. 1: 25 (1825), nom. illeg.
= Polypodium secundum Wall.; Numer. List. [Wallich] no. 301 (1829), nomen
= Polystichum gongylodes (Schkuhr) Gaudich.; Voy. Uranie [Freycinet], Bot. 326 (1828), as ´goggilodus´
= Polystichum propinquum (R.Br.) Gaudich.; Voy. Uranie [Freycinet], Bot. 330 (1827)
= Polystichum serratum (Sw.) Gaudich.; Voy. Uranie [Freycinet], Bot. 32 (1827)
= Pteris crenata Vell.; Fl. Flumin. 11: t. 86 (1831)
= Pteris interrupta Willd.; Phytographia 1: 13, t. 10, f. 1 (1794)
= Tectaria marginalis Cav. ex Sw.; Syn. Fil. [Swartz] 47 (1806), pro syn.
= Thelypteris aneitensis (E.Fourn.) C.F.Reed; Phytologia 17: 259 (1968)
= Thelypteris goggilodus (Schkuhr) Small; Ferns S. E. States [Small] 248 cum tab., 475 (1938)
= Thelypteris gongylodes (Schkuhr) K.Iwats.; Mem. Coll. Sci. Univ. Kyoto, Ser. B, Biol. 31(3): 184 (1965), nom. superfl.
= Thelypteris interrupta (Willd.) K.Iwats.; J. Jap. Bot. 38: 314 (1963)
= Thelypteris lenormandii (C.Chr) Christenh.; Global Fl. 1(4): 33 (2018)
= Thelypteris totta f. paucipilosa (H.Itô) C.F.Reed; Phytologia 17(4): 320 (1968)
= Thelypteris totta var. glabra (Mett. ex H.Itô) C.F.Reed; Phytologia 17(4): 320 (1968)
= Thelypteris totta var. hirsuta (Mett.) C.V.Morton; Contr. U. S. Natl. Herb. 38: 73 (1967)
= Thelypteris totta var. longipinna (C.Chr.) C.V.Morton; Contr. U. S. Natl. Herb. 38: 74 (1967)
species :
Cyclosorus striatus (Schumach.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 249 (1941)
Distribution: Senegal; Gambia; Guinea-Bissau; Guinea; Sierra Leone; Liberia; Mali; Burkina Faso; Ivory Coast; Ghana; Togo; Benin; Niger; Nigeria; Bioko Isl. [Fernando Poo]; São Tomé; Annobon Isl.; Equatorial Guinea (Rio Muni); Gabon; Congo [Brazzaville]; Central African Republic; D.R.Congo [Zaire]; Burundi; Chad; Sudan; Uganda; Tanzania; Angola; Malawi
22 BEN BKN GAM GHA GNB GUI IVO LBR MLI NGA NGR SEN SIE TOG 23 ANN BIO BUR CAF CON EQG GAB SAO ZAI 24 CHA SUD 25 TAN UGA 26 ANG MLW
Synonyms: hidden
= Aspidium grunowii Bolle; Bonplandia (Hannover) 3: 123 (1855)
= Aspidium pallidivenium Hieron.; Pflanzenw. Ost-Afrikas [Engler] C: 85 (1895)
= Aspidium striatum Schumach.; Beskr. Guin. Pl. (Vid. Selsk. Afh. 4): 230 (seors. 456) (1827)
= Cyclosorus grunowii (Bolle) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 245 (1941)
= Cyclosorus interruptus var. striatus (Schumach.) Verdc.; Fl. Trop. E. Africa, Thelypterid. 27 (2006)
= Dryopteris grunowii (Bolle) C.Chr.; Index Filic. 269 (1905)
= Dryopteris hemitelioides Christ; Ann. Mus. Congo Belge, Bot., sér. 5, 8: 26 (1909)
= Dryopteris pallidivenia Kuntze; Revis. Gen. Pl. 2: 813 (1891)
= Dryopteris striata (Schumach.) C.Chr.; Index Filic. 294 (1905)
= Nephrodium grunowii T.Moore; Index Fil. [T. Moore] 93 (1858)
= Nephrodium pallidivenium (Hook.) Baker; Syn. Fil. [W. J. Hooker & Baker] 290 (1867)
= Polypodium pallidivenium Hook.; Sp. Fil. [W. J. Hooker] 5: 8 (1863)
= Thelypteris grunowii (Bolle) C.F.Reed; Phytologia 17: 280 (1968)
= Thelypteris striata (Schumach.) Schelpe; J. S. African Bot. 31: 268 (1965)
variant :
var. molundensis (Brause) Tardieu; Notul. Syst. (Paris) 14(4): 345 (1953)
Distribution: Cameroon; Central African Republic; D.R.Congo [Zaire]; Burundi
23 BUR CAF CMN ZAI
Synonyms: hidden
= Cyclosorus molundensis (Brause) Pic.Serm.; Webbia 32(1): 77 (1977)
= Dryopteris molundensis Brause; Bot. Jahrb. Syst. 68: 378 (1916)
species :
Cyclosorus tottus (Thunb.) Pic.Serm.; Webbia 23: 173 (1968)
Distribution: Burundi; Namibia; Botswana; South Africa (Limpopo, Mpulamanga, KwaZulu-Natal, W-Cape Prov., E-Cape Prov.); Eswatini [Swaziland]; etc.
23 BUR 27 BOT NAM SAF SWZ
Remarks: frequently included in C. interruptus s. lat.
Synonyms: hidden
= Aspidium unitum var. glabrum Mett.; Ann. Mus. Bot. Lugduno-Batavi 1(8): 230 (1863), as ´glabra´
= Polypodium tottum Thunb.; Prodr. Pl. Cap. 2: 172 (1800)
= Thelypteris totta (Thunb.) Schelpe; J. S. African Bot. 29: 91 (1963)
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A complete, synonymic checklist of the Higher Plants of the World

Version 24.11 (November 5th, 2024)

Covering 355,576 accepted species,
8,715 hybrids, 47,863 accepted infraspecific taxa and 1,000,581 synonyms,
in total 1,412,785* names

[*about 25,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus" are deliberately omitted]

With complete, detailed distribution data (countries and regions) and literature references

Arranged in linear phylogenetic sequence reflecting true relationships

Fully searchable and continously updated

With 38,209 pictures of 15,787 species

Assembled through 40 years independently from other databases

In cooperation with Catalogue of Life