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Ver. 24.11
November 5th, 2024

034.0000 Family Dryopteridaceae Herter; Revista Sudamer. Bot. 9: 15 (1949), nom. cons.
26 genera, ca. 2100 spp. Phylogeny: Zuo et al. (2024)
Species listed in database: 2174; hybrids: 201
= Aspidiaceae Burnett; Outlines Bot. 324, 1156 (1835), nom. illeg.
= Aspidiaceae Mett. ex A.B.Frank; Syn. Pflanzenk. [Leunis] ed. 2, 3: 1469 (1877), nom. illeg.
= Bolbitidaceae (Pic.Serm.) Ching; Acta Phytotax. Sin. 16: 15 (1978)
= Elaphoglossaceae Pic.Serm.; Webbia 23: 209 (1968)
= Filicaceae Juss.; Gen. Pl. [Jussieu] 14 (1789), as ´Filices´, nom. illeg., rej.
= Peranemataceae (C.Presl) Ching; Sunyatsenia 5: 246 (1940), nom. rej.
034.4001 Genus Ctenitis C.Chr. ex C.Chr.; Man. Pteridol. 543 (1938)
Distribution: Tropical America (over 60), trop. Africa (1), Indian Ocean (39), trop. Asia, but not in Australia. Revision for South America: Viveros et al. (2017)
Species listed in database: 144
= Ataxipteris Holttum; Blumea 30: 10 (1984)
= Dryopteris subgen.Ctenitis C.Chr.; Biol. Arbejder Tilegnede Eug. Warming 77 (1911)
= Pseudotectaria Tardieu; Notul. Syst. (Paris) 15: 87 (1955)
species :
Ctenitis abyssi (Sehnem) Salino & P.O.Morais; Amer. Fern J. 93(1): 32 (2003)
Distribution: Brazil (Rio Grande do Sul)
84 BRA
Synonyms: hidden
= Dryopteris abyssi Sehnem; Fl. Ilustr. Catarin. (Aspidiaceas): 156 (1979)
species :
Ctenitis aciculata (Baker) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 292 (1938)
Distribution: Sumatra; Borneo (Sabah, Sarawak, Kalimantan); Sulawesi; Philippines (Mindanao); Java; New Guinea (S-Papua New Guinea); Solomon Isl. (Kolombangara, Ranongga, Santa Ysabel)
42 BOR JAW NWG PHI SUL SUM 43 SOL
Synonyms: hidden
= Dryopteris aciculata (Baker) C.Chr.; Index Filic. 250 (1905)
= Nephrodium aciculatum Baker; J. Linn. Soc., Bot. 22: 226 (1886)
species :
Ctenitis alteroblumei Holttum; Blumea 31(1): 27 (1985)
Distribution: S-Sumatra; Java; Lesser Sunda Isl. (Bali)
42 JAW LSI SUM
species :
Ctenitis analamazaotrensis (Rakotondr.) Hennequin & Rouhan; Ann. Missouri Bot. Gard. 102(1): 3-86 (2017)
Distribution: Madagascar
29 MDG
Synonyms: hidden
= Pseudotectaria analamazaotrensis Rakotondr.; Adansonia, sér. 3, 32(1): 32 (2010)
species :
Ctenitis angusta Holttum; Blumea 31(1): 30 (1985)
Distribution: New Guinea (Papua New Guinea)
42 NWG
Conservation Status: Known only from type
species :
Ctenitis anniesii (Rosenst.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 124 (1947)
Distribution: Brazil (Espírito Santo, Paraná, Rio de Janeiro, Rio Grande do Sul, Santa Catarina, São Paulo)
84 BRA
Synonyms: hidden
= Dryopteris anniesii Rosenst.; Hedwigia 46: 118 (1906)
= Dryopteris anniesii var. ottonis Rosenst.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 7, 10, pt. 2: 100 (1913)
species :
Ctenitis apicalis (Baker) F.Ballard; Kew Bull. 1954: 559 (1955)
Distribution: Lord Howe Isl.
50 LHI
Synonyms: hidden
= Aspidium apicale Benth.; Fl. Austral. 7: 758 (1878)
= Dryopteris apicalis (Baker) Kuntze; Revis. Gen. Pl. 2: 812 (1891)
= Nephrodium apicale Baker; Syn. Fil. [W. J. Hooker & Baker], ed. 2: 499 (1874)
species :
Ctenitis arthrothrix (Hook.) Tardieu; Amer. Fern J. 48: 32 (1958)
Distribution: Madagascar; +Mauritius
29 MAU MDG
Synonyms: hidden
= Dryopteris arthrothrix (Hook.) C.Chr.; Index Filic. 253 (1905)
= Phegopteris tomentosa (Bory) Mett.; Fil. Afr. [Kuhn] 123 (1868)
= Polypodium arthrothrix Hook.; Sp. Fil. [W. J. Hooker] 5: 14 (1863)
= Polypodium tomentosum Bory; Voy. Indes Or. [Bélanger] 2: 31 (1833) (non Thouars (1804))
= Pseudotectaria arthrothrix (Hook.) Holttum; Kew Bull. 45(2): 262 (1990)
= Tectaria arthrothrix (Hook.) J.P.Roux; Strelitzia 23: 194 (2009)
= Thelypteris arthrothrix (Hook.) C.F.Reed; Phytologia 17: 261 (1968)
species :
Ctenitis aspidioides (C.Presl) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 124 (1947)
Distribution: Brazil (Bahia, Espírito Santo, Minas Gerais, Paraná, Rio de Janeiro, São Paulo)
84 BRA
Synonyms: hidden
= Aspidium alsophilaceum Kunze; Mett., Pheg. u. Asp. 90, no. 218 (1858)
= Aspidium tijucense Fée; Crypt. Vasc. Brésil 2: 72, t. 102 (1873)
= Dryopteris alsophilacea (Kunze) Kuntze; Revis. Gen. Pl. 2: 812 (1891)
= Dryopteris itatiaiensis C.Chr.; Index Filic. 272 (1905)
= Dryopteris tenuifolia C.Chr.; Index Filic. 297 (1905)
= Lastrea aspidioides T.Moore; Index Fil. [T. Moore] 85 (1858), non (Raddi) C. Presl
= Nephrodium alsophilaceum (Kunze) Baker; Fl. Bras. [Martius] 1(2): 474 (1870)
= Phegopteris aspidioides (C.Presl) Fée; Mém. Foug. 5 (Gen. Filic.): 243 (1852)
= Polypodium aspidioides C.Presl; Delic. Prag. 1: 170 (1822)
species :
Ctenitis atrorubens Holttum; Blumea 31(1): 29 (1985)
Distribution: Borneo (Sabah); Philippines
42 BOR PHI
Synonyms: hidden
= Dryopteris intermedia var. microloba Christ; (1907)
species :
Ctenitis baulensis A.R.Sm.; Proc. Calif. Acad. Sci. ser. 4, 40: 213 (1975)
Distribution: Mexico (Chiapas)
79 MEX
Conservation Status: Known only from type
species :
Ctenitis biformis (Mett.) Tardieu; Humbert, Fl. Madagasc. Fam. 5, 1: 331 (1958)
Distribution: Madagascar; Comores (Mayotte, Mohéli)
29 COM MDG
Synonyms: hidden
= Dryopteris biformis (Boivin) C.Chr.; Index Filic. 254 (1905)
= Lastrea biformis Boivin; ms.
= Nephrodium biforme (Mett.) Diels; Nat. Pflanzenfam. [Engler & Prantl] I, 4: 178 (1899)
= Phegopteris biformis Mett.; Fil. Afr. [Kuhn] 121 (1868)
= Polypodium biforme (Mett.) Baker; Syn. Fil. [W. J. Hooker & Baker], ed. 2: 505 (1874)
= Polypodium polyxiphion Baker; Bull. Mens. Soc. Linn. Paris 1: 534 (1885)
= Pseudotectaria biformis (Mett.) Holttum; Kew Bull. 45(2): 262 (1990)
= Tectaria biformis (Mett.) J.P.Roux; Strelitzia 23: 194 (2009)
species :
Ctenitis bigarellae Schwartsb., Labiak & Salino; Brittonia 59(1): 29 (2007), nom. nov.
Distribution: Brazil (Paraná, Minas Gerais, Rio de Janeiro)
84 BRA
Synonyms: hidden
= Dryopteris falciculata var. elongata Rosenst.; Hedwigia 56: 368 (1915)
species :
Ctenitis bivestita (Mett.) Tardieu; Notul. Syst. (Paris) 15: 85 (1955)
Distribution: Mauritius
29 MAU
Synonyms: hidden
= Dryopteris bivestita (Tausch) C.Chr.; Index Filic. 254 (1905)
= Lastrea sieberiana C.Presl; Tent. Pterid. 75 (1836)
= Phegopteris bivestita (Tausch) Mett.; Abh. Senckenberg. Naturf. Ges. 2(2): [Pheg. u. Asp. 27, no. 57] (1858)
= Polypodium bivestitum Tausch; C. Presl, Tent. Pterid. 131 (1836), nomen
species :
Ctenitis blepharochlamys (C.Chr.) Tardieu; Humbert, Fl. Madagasc. Fam. 5, 1: 336 (1958)
Distribution: Madagascar
29 MDG
Synonyms: hidden
= Dryopteris blepharochlamys C.Chr.; Cat. Pl. Madag., Pterid. 28 (1932) (nomen), Dansk Bot. Ark. 7: 60, t. 15, f. 1-5 (1932)
species :
Ctenitis boholensis Holttum; Blumea 31(1): 18 (1985)
Distribution: Philippines (Bohol)
42 PHI
species :
Ctenitis borbonica (Baker) Tardieu; Notul. Syst. (Paris) 15: 84, t. 5(7-9) (1955)
Distribution: La Réunion
29 REU
Synonyms: hidden
= Dryopteris borbonica (Baker) C.Chr.; Index Filic. 255 (1905)
= Nephrodium borbonicum Baker; Ann. Bot. (Oxford) 6: 317 (1891)
species :
Ctenitis brevipetiolulata Li Bing Zhang & Yi F.Duan; Ann. Missouri Bot. Gard. 102(1): 3-86 (2017)
Distribution: Madagascar (Toamasina, Antsiranana)
29 MDG
species :
Ctenitis bullata A.R.Sm.; Proc. Calif. Acad. Sci. ser. 4, 40: 213 (1975)
Distribution: Mexico (Chiapas)
79 MEX
species :
Ctenitis bullescens Rouhan, Li Bing Zhang & Yi F.Duan; Ann. Missouri Bot. Gard. 102(1): 3-86 (2017)
Distribution: Madagascar (Antananarivo, Antsiranana, Mahajanga, Toamasina)
29 MDG
species :
Ctenitis bulusanica Holttum; Blumea 31(1): 19 (1985)
Distribution: Philippines (Luzon)
42 PHI
species :
Ctenitis calcicola M.Kato; Acta Phytotax. Geobot. 47(1): 77 (1996)
Distribution: Moluccas (Seram)
42 MOL
species :
Ctenitis cerea Li Bing Zhang & Yi F.Duan; Ann. Missouri Bot. Gard. 102(1): 3-86 (2017)
Distribution: Madagascar (Andovoranto, Antananarivo, Antsiranana, Fianarantsoa, Mahajanga, Toamasina, Toliara)
29 MDG
species :
Ctenitis cheilanthina (C.Chr.) C.V.Morton; Contr. U. S. Natl. Herb. 38: 238 (1973)
Distribution: Hispaniola (Haiti, Dominican Republic)
81 HIS
Synonyms: hidden
= Dryopteris cheilanthina C.Chr.; Kongl. Svenska Vetensk. Acad. Handl., ser. 3, 16(2). 34, t. 7(6-9) (1937)
species :
Ctenitis chiapasensis (Christ) A.R.Sm.; Proc. Calif. Acad. Sci. ser. 4, 40: 229 (1975)
Distribution: Mexico (Chiapas)
79 MEX
Conservation Status: Known only from type
Synonyms: hidden
= Aspidium chiapasense Christ; Bull. Herb. Boissier, sér. 2, 5: 727 (1905)
= Dryopteris chiapasensis (Christ) C.Chr.; Index Filic. 257 (1905)
species :
Ctenitis chiriquiana (C.Chr.) Lellinger; Amer. Fern J. 67: 58 (1977)
Distribution: Costa Rica; Panama
80 COS PAN
Synonyms: hidden
= Dryopteris chiriquiana C.Chr.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 8, 6: 36 (1920)
species :
Ctenitis christensenii R.S.Viveros & Salino; Phytotaxa 239(1): 2 (2015)
Distribution: Brazil (Espírito Santo, Minas Gerais, Bahia, Rio de Janeiro)
84 BRA
species :
Ctenitis cirrhosa (Schumach.) Ching; Sunyatsenia 5: 250 (1940)
Distribution: Guinea; Liberia; Ivory Coast; Ghana; Togo; Cameroon; Bioko Isl. [Fernando Poo]; São Tomé; Central African Republic; D.R.Congo [Zaire]; Burundi; Sudan; Ethiopia; Uganda; Kenya; Tanzania; Angola; Zambia; Malawi; Zimbabwe; Mozambique
22 GHA GUI IVO LBR TOG 23 BIO BUR CAF CMN SAO ZAI 24 ETH SUD 25 KEN TAN UGA 26 ANG MLW MOZ ZAM ZIM
Synonyms: hidden
= Aspidium cirrhosum Schumach.; Beskr. Guin. Pl. (Vid. Selsk. Afh. 4): 231 (seors. 457) (1827)
= Aspidium comorense Kuhn; Filic. Afr. 128 (1868), nom. nud.
= Aspidium pinnatifidoserratum Mett.; Fil. Afr. [Kuhn]: 139 (1868)
= Aspidium welwitschii (Baker) Kuhn; Filic. Afr. 209 (1868)
= Ctenitis crinibulbon (Hook.) Ching; Sunyatsenia 5: 250 (1940)
= Dryopteris cirrhosa (Schumach.) Kuntze; Revis. Gen. Pl. 2: 812 (1891)
= Dryopteris crinibulbon (Hook.) C.Chr.; Index Filic., Suppl. Tert.: 84 (1934)
= Dryopteris nimbaensis Tardieu; Notul. Syst. (Paris) 13: 370, t. 1(9) (1949)
= Dryopteris trachyrachis Christ ex Bonap.; Notes Ptéridol. 15: 17 (1924), nom. nud.
= Dryopteris welwitschii (Baker) Kuntze; Revis. Gen. Pl. 2: 814 (1891)
= Lastrea crinibulbon (Hook.) J.Sm.; Hist. Fil. 214 (1875)
= Nephrodium cirrhosum (Schumach.) Baker; Syn. Fil. [W. J. Hooker & Baker], ed. 2: 263 (1874)
= Nephrodium crinibulbon Hook.; Sp. Fil. [W. J. Hooker] 4: 92, t. 244 (1862)
= Nephrodium welwitschii Baker; Syn. Fil. [W. J. Hooker & Baker] 274 (1867)
species :
Ctenitis clathrata M.Kato; Acta Phytotax. Geobot. 47(1): 75 (1996)
Distribution: Moluccas (Seram)
42 MOL
species :
Ctenitis comorensis Li Bing Zhang & Yi F.Duan; Ann. Missouri Bot. Gard. 102(1): 3-86 (2017)
Distribution: Comores (Grande Comore, Mohéli)
29 COM
species :
Ctenitis coriacea M.Kato; Acta Phytotax. Geobot. 47(1): 75 (1996)
Distribution: Moluccas (Seram)
42 MOL
species :
Ctenitis crinigera (C.Chr.) Hennequin & Rouhan; Ann. Missouri Bot. Gard. 102(1): 3-86 (2017)
Distribution: Madagascar (Toliara)
29 MDG
Synonyms: hidden
= Aspidium crinigerum C.Chr.; Notes Ptéridol. [Bonaparte] 16: 34 (1925)
= Pseudotectaria crinigera (C.Chr.) Tardieu; Notul. Syst. (Paris) 15: 87 (1955), et Amer. Fern J. 48: 33 (1958)
= Tectaria crinigera (C.Chr.) C.Chr.; Dansk Bot. Ark. 7: 66, t. 18 (1932)
species :
Ctenitis crinita (Poir.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 290 (1939)
Distribution: Tanzania; Madagascar; Mauritius
25 TAN 29 MAU MDG
Synonyms: hidden
= Alsophila erubescens Kunze; Bot. Zeitung (Berlin) 344 (1844)
= Aspidium crinitum (Poir.) Wall.; Numer. List. [Wallich] no. 357 (1829), nomen
= Aspidium mascarenhense Fée; Mém. Foug. 5 (Gen. Filic.): 296 (1852)
= Aspidium nitidum Bory ex Willd.; Sp. Pl., ed. 4 [Willdenow] 5(1): 251 (1810)
= Aspidium sieberianum Mett.; Salomon, Nomencl. Gefäßkrypt. 70 (1883)
= Aspidium strigosum Mett.; Abh. Senckenberg. Naturf. Ges. 2(2): [Pheg. u. Asp. 89, no. 217] (1858) (non Willd. (1810))
= Aspidium sulcatum Kaulf. ex Bory; Voy. Indes Or. [Bélanger] 2: 58 (1833)
= Ctenitis crinita var. atrata Holttum; Kew Bull. 38(1): 122 (1983)
= Ctenitis crinita var. setacea Holttum; Kew Bull. 38(1): 122 (1983)
= Ctenitis nitida (Bory ex Willd.) Tardieu; Notul. Syst. (Paris) 15: 81, t. 4(4-6) (1955)
= Dryopteris crinita (Poir.) Kuntze; Revis. Gen. Pl. 2: 812 (1891)
= Dryopteris strigosa (Mett.) C.Chr.; Index Filic. 295 (1905)
= Lastrea crinita (Poir.) T.Moore; Index Fil. [T. Moore] 89 (1857)
= Lastrea mascarenensis (Fée) T.Moore; Index Fil. [T. Moore] 96 (1858)
= Nephrodium crinitum (Poir.) Desv.; Mem. Soc. Linn. Paris 6(2): 257 (1827)
= Nephrodium maritimum Cordem.; Bull. Soc. Sci. Île Réunion 1890-1891: 186 (1890)
= Phegopteris crinita (Poir.) Fée; Mém. Foug. 5 (Gen. Filic.): 243 (1852)
= Polypodium crinitum Poir.; Encycl. [J. Lamarck et al.] 5: 528 (1804)
= Polypodium phegopteroides Desv.; Encycl. [J. Lamarck et al.], Suppl. 3: 264 (1813)
= Polypodium thelypteroides Desv.; Mag. Neuesten Entdeck. Gesammten Naturk. Ges. Naturf. Freunde Berlin 5: 317 (1811)
species :
Ctenitis croftii Holttum; Blumea 31(1): 26 (1985)
Distribution: New Guinea (Papua New Guinea: Morobe Prov.)
42 NWG
species :
Ctenitis crystallina (Kunze) Proctor; Bull. Inst. Jamaica, Sci. Ser. 5: 18 (1953)
Distribution: Cuba
81 CUB
Synonyms: hidden
= Cystopteris brevinervis Fée; Mém. Foug. 5 (Gen. Filic.): 300 (1852), et Mém. Foug. 7 (Icon. Esp. Nouv.): 66, t. 26, f. 2
= Dryopteris crystallina (Kunze) Maxon; C. Chr., Ind. 259 (1905)
= Lastrea crystallina (Kunze) T.Moore; Index Fil. [T. Moore] 89 (1858)
= Polypodium crystallinum Kunze; Farnkräuter 2: 85, t. 135 (1851)
species :
Ctenitis cumingii Holttum; Blumea 31(1): 34 (1985)
Distribution: Pitcairn Isl.
61 PIT
Conservation Status: IUCN E
species :
Ctenitis cyclochlamys (Fée) Holttum; Kew Bull. 38(1): 120 (1983)
Distribution: La Réunion
29 REU
Synonyms: hidden
= Aspidium cyclochlamys Fée; Mém. Foug. 5 (Gen. Filic.): 296 (1852)
= Aspidium dasychlamys Fée; Mém. Soc. Sci. Nat. Strasbourg 6(1): 38, t. 41, fig. 4 (1865) [doubtful synonymy, material possibly collected in Madagascar]
= Lastrea cyclochlamys (Fée) T.Moore; Index Fil. [T. Moore] 89 (1858)
species :
Ctenitis decaryana (C.Chr.) Hennequin & Rouhan; Ann. Missouri Bot. Gard. 102(1): 3-86 (2017)
Distribution: Madagascar
29 MDG
Synonyms: hidden
= Aspidium decaryanum (C.Chr.) C.Chr.; Cat. Pl. Madag., Pterid.31 (1932)
= Pseudotectaria decaryana (C.Chr.) Tardieu; Amer. Fern J. 48: 33 (1958)
= Tectaria decaryana C.Chr.; Dansk Bot. Ark. 7: 66 (1932)
species :
Ctenitis decurrentipinnata (Ching) Tardieu & C.Chr.; Notul. Syst. (Paris) 7: 86 (1938)
Distribution: China (Hainan); Vietnam; Philippines (Luzon)
36 CHI 41 VIE 42 PHI
Synonyms: hidden
= Dryopteris decurrentipinnata Ching; Bull. Fan Mem. Inst. Biol. Bot. 2: 195, t. 9 (1931)
species :
Ctenitis deflexa (Kaulf.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 124 (1947)
Distribution: Brazil (Alagoas, Bahia, Espírito Santo, Minas Gerais, Paraná, Rio de Janeiro, Santa Catarina, São Paulo)
84 BRA
Synonyms: hidden
= Aspidium basilare Fée; Crypt. Vasc. Brésil 1: 135 t. 43 f. 2 (1869)
= Aspidium raddianum Mett.; Abh. Senckenberg. Naturf. Ges. 2(2) [Pheg. u. Asp.]: 375 (1858)
= Aspidium vestitum (Raddi) Fée; Crypt. Vasc. Brésil 1: 134 (1869)
= Dryopteris deflexa (Kaulf.) C.Chr.; Index Filic. 261 (1905)
= Dryopteris deflexa var. aschersonii Mett.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 7, 10, pt. 2: 99 (1913)
= Dryopteris vestita (Raddi) Kuntze; Revis. Gen. Pl. 2: 814 (1891)
= Lastrea vestita (Raddi) J.Sm.; Compan. Bot. Mag. 33 (1846)
= Nephrodium raddianum (Mett.) Hook.; Sp. Fil. [W. J. Hooker] 4: 98, t. 245 (1862)
= Nephrodium vestitum (Raddi) Baker; Syn. Fil. [W. J. Hooker & Baker] 266 (1867)
= Nephrodium vestitum var. squamigerum Baker; Fl. Bras. [Martius] 1(2): 474 (1870)
= Phegopteris fluminensis Fée; Crypt. Vasc. Brésil 1: 97 (1869)
= Polypodium deflexum Kaulf.; Enum. Filic. [Kaulfuss] 114 (1824)
= Polypodium lepigerum Schrad.; Gött. Gel. Anz. 868 (1824)
= Polypodium vestitum Raddi; Opusc. Sci. 3: 288 (1819) (non G. Forst. (1786))
species :
Ctenitis dentata Li Bing Zhang & Yi F.Duan; Ann. Missouri Bot. Gard. 102(1): 3-86 (2017)
Distribution: Madagascar (Fianarantsoa, Alaotra-Mangoro)
29 MDG
species :
Ctenitis desvauxii (Mett.) Tardieu; Notul. Syst. (Paris) 16: 82, t. 5(4-6) (1955)
Distribution: Mauritius
29 MAU
Synonyms: hidden
= Aspidium desvauxii Mett.; Fil. Afr. [Kuhn] 131 (1868), nom. nov. for Polypodium obtusilobum Desv.
= Dryopteris obtusiloba (Desv.) C.Chr.; Index Filic. 280 (1905), non (Baker) Kuntze
= Phegopteris obtusiloba (Desv.) Christ; Farnkr. Erde 273 (1897)
= Polypodium obtusilobum Desv.; Mag. Neuesten Entdeck. Gesammten Naturk. Ges. Naturf. Freunde Berlin 5: 317 (1811)
species :
Ctenitis dianguiensis (W.M.Chu & H.G.Zhou) S.Y.Dong; Novon 18(1): 35 (2008)
Distribution: China (Yunnan, Guangxi, Hainan); Vietnam
36 CHI 41 VIE
Synonyms: hidden
= Ataxipteris dianguiensis W.M.Chu & H.G.Zhou; H. G. Zhou & W. M. Chu, Acta Bot. Yunnan. 17(4): 423-424, f. 1 (1995), et Acta Phytotax. Sin. 42(4): 376 (2004)
species :
Ctenitis dingnanensis Ching; Acta Phytotax. Sin. 19(1): 122 (1981)
Distribution: China (Guangdong, Jiangxi)
36 CHI
species :
Ctenitis distans (Brack.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 277 (1938)
Distribution: Brazil (Alagoas, Bahia, Ceará, Espírito Santo, Minas Gerais, Pernambuco, Paraná, Rio de Janeiro, Santa Catarina, São Paulo)
84 BRA
Synonyms: hidden
= Aspidium amaurolepis Fée; Crypt. Vasc. Brésil 1: 137, t. 44, f. 2 (1869)
= Aspidium ctenitis Link; Hort. Berol. [Link] 2: 122 (1833)
= Dryopteris ctenitis (Link) Kuntze; Revis. Gen. Pl. 2: 812 (1891)
= Dryopteris ctenitis f. amaurolepis (Fée) C.Chr.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 7, 10, pt. 2: 94 (1913)
= Lastrea ctenitis (Link) T.Moore; Index Fil. [T. Moore] 89 (1858)
= Lastrea distans Brack.; U.S. Expl. Exped., Filic. 16: 192 (1854)
= Nephrodium ctenitis (Link) Baker; Syn. Fil. [W. J. Hooker & Baker] 265 (1867)
variant :
var. isabellina (Fée) R.S.Viveros & Salino; Phytotaxa 335(1): 41 (2018)
Distribution: Brazil (Espírito Santo, Minas Gerais, Paraná, Rio de Janeiro, São Paulo)
84 BRA
Synonyms: hidden
= Aspidium isabellinum Fée; Crypt. Vasc. Brésil 1: 137, t. 45, f. 2 (1869)
= Dryopteris ctenitis f. isabellina (Fée) C.Chr.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 7, 10, pt. 2: 94 (1913)
= Dryopteris isabellina (Fée) C.Chr.; Index Filic. 272 (1905)
species :
Ctenitis dolphinensis Proctor; Amer. Fern J. 48: 108 (1958)
Distribution: Jamaica
81 JAM
species :
Ctenitis dumrongii Tagawa & K.Iwats.; Acta Phytotax. Geobot. 23: 115, f. 11 (1968)
Distribution: Myanmar (Kachin); N-Thailand (Chiang Mai, Chiang Rai)
41 MYA THA
Conservation Status: DD in Thailand
species :
Ctenitis eatonii (Baker) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 291 (1938)
Distribution: China (Guangdong, Guangxi, Guizhou, Hubei, Hunan, Jiangxi, Sichuan); Ryukyu Isl.; Taiwan; Vietnam
36 CHI 38 NNS TAI 41 VIE
Synonyms: hidden
= Aspidium eatonii (Baker) Christ; Monsunia 1: 81 (1900)
= Aspidium intermedium Christ; (1902) (non Willd. 1810, nec Blume 1828)
= Aspidium leucostipes (Baker) Christ; Bull. Herb. Boissier, sér. 2, 4: 615 (1904)
= Ctenitis changanensis Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 297 (1938)
= Ctenitis confusa Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 290 (1938)
= Ctenitis leucostipes (Baker) Tardieu; Bull. Soc. Bot. France 87: 366 (1941)
= Dryopteris eatonii (Baker) Kuntze; Revis. Gen. Pl. 2: 812 (1891)
= Dryopteris eatonii var. iriomotensis H.Itô; (1971)
= Dryopteris leucostipes (Baker) C.Chr.; Index Filic. 274 (1905)
= Dryopteris subtripinnata var. leucostipes (Baker) H.Itô; (1938)
= Nephrodium eatonii Baker; Syn. Fil. [W. J. Hooker & Baker] 276 (1867)
= Nephrodium eatonii var. formosana Harr.; J. Linn. Soc., Bot. 16: 29 (1877)
= Nephrodium leucostipes Baker; J. Bot. 23: 105 (1885)
species :
Ctenitis elata Holttum; Blumea 31(1): 24 (1985), nom. nov.
Distribution: Sumatra (Korinchi Peak)
42 SUM
Synonyms: hidden
= Dryopteris subarborea var. glabrior Alderw.; (1914)
species :
Ctenitis equestris (Kunze) Ching; Sunyatsenia 5: 250 (1940)
Distribution: Mexico (Chiapas, Colima, Guerrero, Hidalgo, Jalisco, Mexico State, Michoacan, Morelos, Nayarit, Oaxaca, Sinaloa, Tamaulipas, Veracruz); Revillagigedos Isl.; Guatemala; Belize; El Salvador; Honduras; Nicaragua; Costa Rica; Panama; Colombia (Antioquia, Santander); Venezuela (Venezuela Distrito Federal, Lara); Galapagos
79 MEX RGI 80 BLZ COS ELS GUA HON NIC PAN 82 CLM VEN 83 GAL
Synonyms: hidden
= Aspidium bourgaei E.Fourn.; Mexic. Pl. 1: 98 (1872)
= Aspidium ciliatum (Liebm.) Mett.; Salomon, Nomencl. Gefäßkrypt. 46 (1883), nom. illeg.
= Aspidium equestre Kunze; Linnaea 18: 347 (1844)
= Aspidium scabriusculum Davenp.; Bot. Gaz. 21: 256 (1896) (non Mett. ex Salom (1883))
= Dryopteris bourgaei (E.Fourn.) C.Chr.; Index Filic. 255 (1905)
= Dryopteris davenportii C.Chr.; Index Filic. 260 (1905), nom. nov. for Aspidium scabriusculum Davenp. 1896
= Dryopteris equestris (Kunze) C.Chr.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 8, 6: 54 (1920)
= Dryopteris equestris var. mentiens C.Chr.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 8, 6: 56 (1920)
= Lastrea ciliata Liebm.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 5, 1: 273 (seors. 121) (1849)
= Lastrea equestris (Kunze) Liebm.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 5, 1: 273 (seors. 121) (1849)
= Nephrodium ciliatum (Liebm.) Kiaersk.; Cat. Fil. hb. hort. Haun. 63 (1874) (non Desv. (1827))
= Polypodium alsophiloides Liebm.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 5, 1: 208 (seors. 56) (1849)
variant :
var. erosa Stolze; Amer. Fern J. 67(2): 41 (1977)
Distribution: Mexico (Chiapas, Oaxaca, Guerrero, Hidalgo); El Salvador; Honduras
79 MEX 80 ELS HON
species :
Ctenitis erinacea A.R.Sm.; Mem. New York Bot. Gard. 88 [Pterid. Mexico]: 223 (2004)
Distribution: Mexico (Chiapas, Hidalgo, Oaxaca, Puebla, Veracruz); Guatemala; El Salvador; Honduras
79 MEX 80 ELS GUA HON
species :
Ctenitis eriocaulis (Fée) Alston; Lilloa 30: 112 (1960)
Distribution: Brazil (Alagoas, Bahia, Espírito Santo, Minas Gerais, Paraná, Rio de Janeiro, São Paulo); Paraguay (Canindeyu)
84 BRA 85 PAR
Synonyms: hidden
= Aspidium eriocaulon Fée; Crypt. Vasc. Brésil 1: 136, t. 44, f. 1 (1869)
= Dryopteris cirrhosa var. eriocaulis (Fée) C.Chr.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 7, 10, pt. 2: 102 (1913)
= Dryopteris eriocaulis (Fée) Kuntze; Revis. Gen. Pl. 2: 812 (1891)
= Nephrodium eriocaulon (Fée) Baker; Syn. Fil. [W. J. Hooker & Baker], ed. 2: 495 (1874)
= Nephrodium ramentaceum Baker; Fl. Bras. [Martius] 1(2): 473 (1870)
species :
Ctenitis erythradenia Holttum; Blumea 31(1): 29 (1985)
Distribution: Philippines (Luzon)
42 PHI
species :
Ctenitis exaggerata (Baker) Ching; Sunyatsenia 5: 250 (1940)
Distribution: Madagascar (Antananarivo, Toamasina)
29 MDG
Synonyms: hidden
= Dryopteris exaggerata (Baker) C.Chr.; Dansk Bot. Ark. 7: 60, t. 16, f. 4-7 (1932)
= Megalastrum exaggeratum (Baker) Holttum; Gard. Bull. Singapore 39(2): 161 (1986 publ. 1987)
= Nephrodium crinitum var. exaggeratum Baker; Ann. Bot. (Oxford) 5: 319 (1892)
species :
Ctenitis excelsa (Desv.) Proctor; Rhodora 63: 34 (1961)
Distribution: Mexico (Guerrero, Chiapas, Veracruz, Oaxaca); Guatemala; Belize; El Salvador; Honduras; Nicaragua; Costa Rica; Panama; Guadeloupe; Dominica; Martinique
79 MEX 80 BLZ COS ELS GUA HON NIC PAN 81 DOM GDL MAR
Synonyms: hidden
= Aspidium effusum var. muticum Christ; Prim. Fl. Costaric. 3: 32 (1901)
= Ctenitis equestris var. mutica (Christ) Lellinger; Proc. Biol. Soc. Wash. 89(61): 709 (1977)
= Ctenitis gomezii R.C.Moran; Brenesia 21: 449 (1983)
= Dryopteris equestris var. heterolepis C.Chr.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 8, 6: 56 (1920)
= Dryopteris excelsa (Desv.) C.Chr.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 8, 6: 54 (1920)
= Phegopteris inaequalis Fée; Mém. Foug. 8 (Icon. Esp. Nouv.): 91 (1857)
= Polypodium excelsum Desv.; Mem. Soc. Linn. Paris 6(2): 243 (1827) (non Baker (1874))
species :
Ctenitis falciculata (Raddi) Ching; Sunyatsenia 5: 250 (1940)
Distribution: Guyana; Brazil (Bahia, Espírito Santo, Minas Gerais, Pernambuco, Paraná, Rio de Janeiro, Rio Grande do Sul, Santa Catarina, São Paulo); Paraguay (Canindeyu)
82 GUY 84 BRA 85 PAR
Synonyms: hidden
= Aspidium chrysolobum Kaulf.; Hort. Berol. [Link] 2: 117 (1833)
= Aspidium falciculatum Raddi; Opusc. Sci. 8: 289 (1819)
= Aspidium mucronatum (C.Presl) Beyr. ex Mett.; Abh. Senckenberg. Naturf. Ges. 2(2) [Pheg. u. Asp.]: 90 (1857), nom. illeg.
= Aspidium schomburgkii Klotzsch; Linnaea 20: 369 (1847)
= Aspidium schottianum Kunze ex Baker; Syn. Fil. [W. J. Hooker & Baker] 262 (1867)
= Aspidium sericeum Fée; Crypt. Vasc. Brésil 1: 144, t. 42, f. 1 (1869)
= Dryopteris chrysoloba (Kaulf. ex Link) Kuntze; Revis. Gen. Pl. 2: 812 (1891)
= Dryopteris falciculata (Raddi) Kuntze; Revis. Gen. Pl. 3(2): 378 (1898)
= Dryopteris falciculata f. sericea (Fée) C.Chr.; (1913)
= Lastrea chrysoloba (Kaulf. ex Link) C.Presl; Tent. Pterid. 75 (1836)
= Lastrea falciculata (Raddi) C.Presl; Tent. Pterid. 75 (1836)
= Lastrea mucronata C.Presl; Tent. Pterid. 75 (1849)
= Nephrodium chrysolobum (Kaulf. ex Link) Fée; Mém. Foug. 5 (Gen. Filic.): 305 (1852)
= Nephrodium falciculatum (Raddi) Desv.; Mem. Soc. Linn. Paris 6(2): 259 (1827)
= Phegopteris falciculata (Raddi) Fée; Crypt. Vasc. Brésil 1: 99 (1869)
= Polypodium ciliatum Spreng.; Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 10: 229 (1821)
= Polypodium distans Kaulf.; Enum. Filic. [Kaulfuss] 113 (1824)
= Polypodium lanuginosum Vell.; Fl. Flumin. 11: t. 70 (1831)
= Polystichum falciculatum (Raddi) Gaudich.; Voy. Uranie [Freycinet], Bot. 328 (1827)
= Thelypteris chrysoloba (Kaulf. ex Link) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 251 (1941)
species :
Ctenitis fenestralis (C.Chr.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 124 (1947)
Distribution: Brazil (Ceará, Rio de Janeiro, Santa Catarina, São Paulo, Paraná, Rio Grande do Sul)
84 BRA
Synonyms: hidden
= Dryopteris fenestralis C.Chr.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 7, 10: 100, f. 8 (1913)
= Dryopteris fenestralis var. spannagelii Rosenst. ex C.Chr.; (1913)
species :
Ctenitis fibrillosa (Baker) Li Bing Zhang & Yi F.Duan; Ann. Missouri Bot. Gard. 102(1): 3-86 (2017)
Distribution: Madagascar
29 MDG
Synonyms: hidden
= Aspidium fibrillosum (Baker) Kuhn; Reisen Ost-Afr. [Decken] 38. Bot. 65 (1879)
= Ctenitis poolii (C.Chr.) Tardieu; Humbert, Fl. Madagasc. Fam. 5, 1: 330 (1958)
= Dryopteris blepharorachis C.Chr.; Ark. Bot. 14(19) 3, t. 1, f. 1-2 (1916)
= Dryopteris poolii C.Chr.; Index Filic. 285 (1905), nom. nov., non D. fibrillosa (Baker) C. Chr. (1905)
= Nephrodium fibrillosum Baker; J. Linn. Soc., Bot. 16: 418 (1876)
= Pseudotectaria fibrillosa (Baker) Holttum; Kew Bull. 45(2): 261 (1990)
= Tectaria blepharorachis (C.Chr.) Li Bing Zhang & Yi F.Duan; Phytotaxa 162(2): 108 (2014)
= Tectaria fibrillosa (Baker) J.P.Roux; Strelitzia 23: 194 (2009)
species :
Ctenitis fijiensis (Hook.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 124 (1947)
Distribution: Fiji (Viti Levu, Vanua Levu, Ovalau); ?[Western] Samoa
60 FIJ
Synonyms: hidden
= Aspidium amoenum Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 236 (1941)
= Aspidium chrysotrichum Christ; Bot. Jahrb. Syst. 28: 352 (1896)
= Ctenitis tenuifrons (C.Chr.) Ching; Sunyatsenia 5: 250 (1940) (non (Hayata) Ching)
= Dryopteris chrysotrichoides Rosenst.; Meded. Rijks-Herb. no. 31: 5 (1917)
= Dryopteris fijiensis (Hook.) C.Chr.; Index Filic. 264 (1905)
= Dryopteris tenuifolia (Brack.) Kuntze; Revis. Gen. Pl. 2: 813 (1891)
= Dryopteris tenuifrons C.Chr.; Index Filic. 297 (1905)
= Lastrea fijiensis (Hook.) Carruth.; Fl. Vit. [Seemann] 360 (1873)
= Lastrea tenuifolia Brack.; U.S. Expl. Exped., Filic. 16: 199 (1854) (non C. Presl (1849))
= Nephrodium fijiense Hook.; Sec. Cent. Ferns t. 67 (1861)
= Nephrodium tenuifolium (Brack.) Hook.; Sp. Fil. [W. J. Hooker] 4: 144 (1862)
species :
Ctenitis flexuosa (Fée) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 124 (1947)
Distribution: Brazil (Rio de Janeiro, ?Santa Catarina)
84 BRA
Synonyms: hidden
= Aspidium flexuosum Fée; Crypt. Vasc. Brésil 1: 138, t. 46, f. 2 (1869)
= Aspidium flexuosum var. moelleri Christ; Bull. Herb. Boissier, sér. 2, 2: 633 (1902)
= Dryopteris camptocaulis (Fée) C.Chr.; Index Filic. 256 (1905)
= Dryopteris flexuosa (Fée) Kuntze; Revis. Gen. Pl. 2: 812 (1891)
= Nephrodium flexuosum (Fée) Baker; Fl. Bras. [Martius] 1(2): 483 (1870)
= Phegopteris camptocaulon Fée; Crypt. Vasc. Brésil 2: 60, t. 98, f. 1 (1873)
= Polypodium camptocaulon (Fée) Baker; Ann. Bot. (Oxford) 5: 459 (1891)
species :
Ctenitis glandulosa R.S.Viveros & Salino; Phytotaxa 239(1): 6 (2015)
Distribution: Guyana; Brazil (Espírito Santo, Pernambuco, Bahia, Minas Gerais, São Paulo)
82 GUY 84 BRA
species :
Ctenitis grisebachii (Baker) Ching; Sunyatsenia 5: 250 (1940)
Distribution: Mexico (Chiapas, Guerrero, Nayarit, Veracruz); Guatemala; Honduras; Nicaragua; Costa Rica; Cuba; Jamaica; Hispaniola (Haiti); Galapagos
79 MEX 80 COS GUA HON NIC 81 CUB HIS JAM 83 GAL
Synonyms: hidden
= Ctenitis molinae Stolze; Amer. Fern J. 67(2): 40 (1977)
= Ctenitis sotoana A.Rojas; Revista Biol. Trop. 49: 468 (2001) [fide Viveros et al. 2017]
= Dryopteris grisebachii (Baker) Kuntze; Revis. Gen. Pl. 2: 812 (1891)
= Lastrea ameristoneura T.Moore; Index Fil. [T. Moore] 85 (1858)
= Nephrodium amplum Hook.; Sp. Fil. [W. J. Hooker] 4: 264 (1862)
= Nephrodium grisebachii Baker; Syn. Fil. [W. J. Hooker & Baker] 285 (1867)
species :
Ctenitis guidianensis H.G.Zhou & W.M.Chu; Acta Bot. Yunnan. 17(4): 424-426, f. 2 (1995)
Distribution: China (Yunnan, Guangxi)
36 CHI
species :
Ctenitis harrisii Proctor; Brit. Fern Gaz. 9: 216 (1965)
Distribution: Jamaica; Hispaniola (Haiti, Dominican Republic)
81 HIS JAM
species :
Ctenitis hemsleyana (Baker ex Hemsl.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 124 (1947)
Distribution: Mexico (Chiapas, Hidalgo, Oaxaca, Puebla, Veracruz); Guatemala; El Salvador; Honduras; Nicaragua; Costa Rica; Panama
79 MEX 80 COS ELS GUA HON NIC PAN
Synonyms: hidden
= Aspidium hemsleyanum (Baker ex Hemsl.) Christ; Bull. Herb. Boissier, sér. 2, 6: 57 (1906)
= Ctenitis costaricensis R.C.Moran; Brenesia 21: 450 (1983)
= Dryopteris hemsleyana (Baker ex Hemsl.) C.Chr.; Index Filic. 270 (1905)
= Polypodium hemsleyanum Baker ex Hemsl.; Biol. Centr.-Amer., Bot. 3: 660 (1885)
species :
Ctenitis hirta (Sw.) Ching; Sunyatsenia 5: 250 (1940)
Distribution: Bahamas; Cuba; Jamaica; Hispaniola (Dominican Republic, Haiti)
81 BAH CUB HIS JAM
Synonyms: hidden
= Aspidium crystallinum Mett.; Fil. Hort. Bot. Lips. 90 (1856)
= Aspidium hirtum (Sw.) Sw.; J. Bot. (Schrader) 1800(2): 39 (1801)
= Ctenitis hirta f. cubana (C.Chr.) Duek; Adansonia, sér. 2, 11: 723 (1971)
= Cystopteris hirta (Sw.) Klotzsch; Linnaea 20: 361 (1847)
= Cystopteris rufescens Fée; Mém. Foug. 5 (Gen. Filic.): 300 (1852)
= Dryopteris hirta (Sw.) Kuntze; Revis. Gen. Pl. 2: 813 (1891)
= Dryopteris hirta f. cubana C.Chr.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 8: 6: 38 (1920)
= Dryopteris hirta f. domingana C.Chr.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 8, 6: 38 (1920)
= Lastrea hirta (Sw.) C.Presl; Tent. Pterid. 76 (1836)
= Nephrodium hirtum (Sw.) Hook.; Sp. Fil. [W. J. Hooker] 4: 128 (1862)
= Polypodium barbatum Kunze; Syst. Veg., ed. 16 [Sprengel] 4(1): 57 (1827)
= Polypodium hirtum Sw.; Prodr. Veg. Ind. Occ. 133 (1788)
= Thelypteris hirta (Sw.) Copel.
variant :
var. portoricana (C.Chr.) Proctor; Mem. New York Bot. Gard. 53: 248 (1989)
Distribution: Puerto Rico
81 PUE
Synonyms: hidden
= Dryopteris hirta f. portoricana C.Chr.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 8, 6: 39 (1920)
species :
Ctenitis hispida (Kuhn) Li Bing Zhang & Yi F.Duan; Ann. Missouri Bot. Gard. 102(1): 3-86 (2017)
Distribution: Mauritius
29 MAU
Synonyms: hidden
= Aspidium crinitum var. hispidum Willd. ex Kuhn; Filic. Afr. 130 (1868), excl. syn. Alsophila erubescens Kunze
= Ctenitis crinita var. hispida (Kuhn) Tardieu; Fl. Madagasc. Fam. 5e: 337 (1958)
species :
Ctenitis humida (Cordem.) Holttum; Kew Bull. 38(1): 126 (1983)
Distribution: La Réunion
29 REU
Synonyms: hidden
= Dryopteris humida (Cordem.) C.Chr.; Index Filic. 271 (1905)
= Nephrodium crinitum var. nudatum Baker; Fl. Mauritius [J.G.Baker] 495 (1877)
= Nephrodium humidum Cordem.; Bull. Soc. Sci. Île Réunion 1890-1891: 187 (1890), et Fl. Réunion (E. J. de Cordemoy): 74 (1895)
species :
Ctenitis humilis Holttum; Blumea 31(1): 22 (1985)
Distribution: Philippines (Bohol, Mindoro)
42 PHI
species :
Ctenitis interjecta (C.Chr.) Ching; Sunyatsenia 5: 250 (1940)
Distribution: Mexico (Chiapas, Veracruz, Oaxaca); Guatemala; Belize; El Salvador; Honduras; Nicaragua; Costa Rica
79 MEX 80 BLZ COS ELS GUA HON NIC
Synonyms: hidden
= Dryopteris interjecta C.Chr.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 8, 6: 43 (1920)
species :
Ctenitis iriomotensis (H.Itô) Nakaike; Enum. Pterid. Jap., Filic. 195 (1975)
Distribution: Ryukyu Isl. (Iriomote Isl.)
38 NNS
Conservation Status: IUCN R. VU in Japan
Synonyms: hidden
= Ctenitis eatonii var. iriomotensis H.Itô; J. Jap. Bot. 46: 305, f. 1 (1971)
species :
Ctenitis jinfoshanensis Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(4): 14-15, f. 43 (1984), et Phytotaxa 205(4): 299 (2015)
Distribution: China (Sichuan)
36 CHI
species :
Ctenitis jouyana (Rakotondr.) Hennequin & Rouhan; Ann. Missouri Bot. Gard. 102(1): 3-86 (2017)
Distribution: Madagascar
29 MDG
Synonyms: hidden
= Pseudotectaria jouyana Rakotondr.; Adansonia, sér. 3, 32(1): 34 (2010)
species :
Ctenitis kinabaluensis Holttum; Blumea 31(1): 16 (1985)
Distribution: Borneo (Sabah, Sarawak, E-Kalimantan); Brunei
42 BOR BRN
variant :
var. crassisquama Holttum; Blumea 31(1): 16 (1985)
Distribution: Borneo (Kalimantan)
42 BOR
Conservation Status: Known only from type
species :
Ctenitis kjellbergii (C.Chr. ex Kjellb. & C.Chr.) Ching; Sunyatsenia 5: 250 (1940)
Distribution: C-Sulawesi; New Guinea (Yapen Isl.)
42 NWG SUL
Synonyms: hidden
= Dryopteris kjellbergii C.Chr.; Bot. Jahrb. Syst. 66: 44 (1933)
species :
Ctenitis koordersii Holttum; Blumea 31(1): 25 (1985)
Distribution: Sulawesi
42 SUL
species :
Ctenitis laetevirens (Rosenst.) Salino & P.O.Morais; Amer. Fern J. 93(1): 34 (2003)
Distribution: Brazil (Santa Catarina, São Paulo)
84 BRA
Synonyms: hidden
= Dryopteris laetevirens Rosenst.; Hedwigia 66: 368 (1915)
species :
Ctenitis lanceolata (Baker) A.R.Sm.; Proc. Calif. Acad. Sci. ser. 4, 40: 229 (1975)
Distribution: Mexico (Chiapas); Guatemala
79 MEX 80 GUA
Synonyms: hidden
= Aspidium tricholepis (Baker) Christ; Bull. Herb. Boissier, sér. 2, 6: 259 (1906)
= Dryopteris lanceolata (Baker) C.Chr.; Index Filic. 298 (1905)
= Dryopteris lanceolata (Baker) Kuntze; Revis. Gen. Pl. 2: 813 (1891)
= Dryopteris lanceolata var. deltoideo-lanceolata C.Chr.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 7, 10, pt. 2: 107 (1913)
= Dryopteris lanceolata var. tricholepis (Baker) C.Chr.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 7, 10, pt. 2: 107, fig. 10d (1913)
= Dryopteris tricholepis (Baker) C.Chr.; Index Filic. 298 (1905)
= Filix-mas lanceolata (Baker) Farw.; Amer. Midl. Naturalist 12: 234-251 (1931)
= Nephrodium lanceolatum Baker; Syn. Fil. [W. J. Hooker & Baker], ed. 2: 498 (1874)
= Nephrodium tricholepis Baker ex Hemsl.; Biol. Centr.-Amer., Bot. 3: 651 (1885)
species :
Ctenitis latifrons (Brack.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 124 (1947)
Distribution: Hawaii (Kauai, Oahu, Molokai, Lanai, Maui, ?Hawaii Isl.)
63 HAW
Synonyms: hidden
= Aspidium latifrons (Brack.) Mann; Proc. Amer. Acad. Arts 7: 217 (1868)
= Ctenitis honolulensis (Hook.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 124 (1947)
= Dryopteris honolulensis (Hook.) C.Chr.; Index Filic. 271 (1905)
= Dryopteris latifrons (Brack.) Kuntze; Revis. Gen. Pl. 2: 813 (1891)
= Filix-mas latifrons (Brack.) Farw.; Amer. Midl. Naturalist 12: 234-251 (1931)
= Lastrea latifrons Brack.; U.S. Expl. Exped., Filic. 16: 196 (1854)
= Nephrodium latifrons (Brack.) Hook.; Sp. Fil. [W. J. Hooker] 4: 138 (1862)
= Phegopteris hillebrandii Luerss.; Flora 58: 436 (1875), nom. nov. superfl. for Polypodium honolulense Hook.
= Phegopteris honolulensis (Hook.) Mann; Proc. Amer. Acad. Arts 7: 218 (1868)
= Polypodium honolulense Hook.; Sp. Fil. [W. J. Hooker] 4: 288 (nota) (1862)
species :
Ctenitis leonii A.Rojas; Amer. J. Pl. Sci. 2017(9): 1331 (2017)
Distribution: Honduras
80 HON
species :
Ctenitis longivillosa Li Bing Zhang & Yi F.Duan; Ann. Missouri Bot. Gard. 102(1): 3-86 (2017)
Distribution: Madagascar (Toliara, Toamasina)
29 MDG
species :
Ctenitis lorencei Holttum; Kew Bull. 38(1): 123 (1983)
Distribution: Rodrigues
29 ROD
Conservation Status: IUCN R, probably in danger of extinction
species :
Ctenitis madagascariensis Tardieu; Naturaliste Malgache 7(1): 91, t. 1(1-5) (1955)
Distribution: Madagascar (Toliara)
29 MDG
species :
Ctenitis magna (Baker) Tardieu; Humbert, Fl. Madagasc. Fam. 5, 1: 344 (1958)
Distribution: Madagascar
29 MDG
Synonyms: hidden
= Dryopteris baronii (Baker) C.Chr.; Index Filic. 254 (1905)
= Dryopteris magna (Baker) C.Chr.; Index Filic. 276 (1905)
= Megalastrum magnum (Baker) Holttum; Gard. Bull. Singapore 39(2): 161 (1986 publ. 1987)
= Nephrodium magnum Baker; J. Bot. 22: 142 (1884)
= Polypodium baronii Baker; J. Linn. Soc., Bot. 22: 535 (1887)
species :
Ctenitis mannii (C.Hope) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 289 (1938)
Distribution: China (Yunnan); India (Assam, Meghalaya); Myanmar; peninsular Malaysia; Sumatra; Borneo (E-Kalimantan); Java
36 CHI 40 IND 41 MYA 42 BOR JAW MLY SUM
Remarks: C. mannii in strict sense is endemic to NE-India
Conservation Status: IUCN EN for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Ctenitis fulgens Ching & C.H.Wang; Acta Phytotax. Sin. 19(1): 120 (1981)
= Dryopteris intermedia var. mannii (C.Hope) Christ; Philipp. J. Sci. 7: 215 (1907)
= Dryopteris mannii (C.Hope) Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 3, 2: 145 (1920)
= Lastrea intermedia Bedd.; Ferns Brit. India Correct. 11 (1870), non Bedd. (1836)
= Nephrodium mannii C.Hope; J. Bot. 28: 145 (1810) (non Dryopteris manniana C. Chr.)
species :
Ctenitis marginalisora Li Bing Zhang & Yi F.Duan; Ann. Missouri Bot. Gard. 102(1): 3-86 (2017)
Distribution: Madagascar (Antananarivo)
29 MDG
species :
Ctenitis maritima (Cordem.) Tardieu; Notul. Syst. (Paris) 15: 84 (1955)
Distribution: Mauritius; La Réunion
29 MAU REU
Synonyms: hidden
= Nephrodium crinitum var. maritimum Cordem.; Fl. Réunion (E. J. de Cordemoy): 74 (1895)
species :
Ctenitis mayottensis Li Bing Zhang & Yi F.Duan; Ann. Missouri Bot. Gard. 102(1): 3-86 (2017)
Distribution: Comores (Mayotte)
29 COM
species :
Ctenitis megalastriformis R.S.Viveros & Salino; Brittonia 69(3): 317 (2017)
Distribution: Peru
83 PER
species :
Ctenitis melanochlamys (Fée) Ching; Sunyatsenia 5: 250 (1940)
Distribution: Cuba
81 CUB
Synonyms: hidden
= Aspidium lomatopelta Kunze ex Mett.; Abh. Senckenberg. Naturf. Ges. 2(2) [Pheg. u. Asp.]: 356 (1858)
= Aspidium melanochlamys Fée; Mém. Foug. 5 (Gen. Filic.): 294 (1852)
= Dryopteris lomatopelta (Kunze ex Mett.) C.Chr.; Index Filic. 275 (1905)
= Dryopteris melanochlamys (Fée) Kuntze; Revis. Gen. Pl. 2: 813 (1891)
= Dryopteris nemorosa var. melanochlamys (Fée) C.Chr.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 8, 6: 41 (1920)
= Lastrea lomatopelta (Kunze ex Mett.) T.Moore; Index Fil. [T. Moore] 95 (1858)
= Lastrea melanochlamys (Fée) T.Moore; Index Fil. [T. Moore] 96 (1858)
= Polystichum melanochlamys (Fée) Diels; Nat. Pflanzenfam. [Engler & Prantl] I, 4: 193 (1899)
species :
Ctenitis melanosticta (Kunze) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 124 (1947)
Distribution: Mexico (Chiapas, Hidalgo, Morelos, Oaxaca, Puebla, Queretaro, San Luis Potosi, Tabasco, Tamaulipas, Veracruz); Guatemala; Belize; Honduras; Nicaragua; Costa Rica
79 MEX 80 BLZ COS GUA HON NIC
Synonyms: hidden
= Aspidium bullatum Christ; Bull. Herb. Boissier, sér. 2, 6: 53 (1906)
= Aspidium expansum Fée; Mém. Foug. 10 (Icon. Esp. Nouv.): 42 (1865)
= Aspidium extensum Fée; Mém. Foug. 5 (Gen. Filic.): 294 (1852) (non Blume (1828))
= Aspidium melanostictum Kunze; Linnaea 13: 148 (1839)
= Ctenitis melanosticta var. bullata (Christ) Lellinger; Amer. Fern J. 67(2): 58 (1977)
= Dryopteris bullata (Christ) C.Chr.; Index Filic., Suppl. 2 (1906-1912): 80 (1913)
= Dryopteris melanosticta (Kunze) Kuntze; Revis. Gen. Pl. 2: 813 (1891)
= Dryopteris melanosticta var. bullata (Christ) C.Chr.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 8, 6: 43 (1920)
= Lastrea melanosticta (Kunze) T.Moore; Index Fil. [T. Moore] 91 (1858)
= Polystichum melanostictum (Kunze) Liebm.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 5, 1: 276 (1849)
species :
Ctenitis meridionalis (Poir.) Ching; Sunyatsenia 5: 250 (1940)
Distribution: St. Eustatius; Nevis; St. Kitts; Montserrat; Guadeloupe; Martinique; ?Grenada
81 GDL MAR MTS STE STK
Synonyms: hidden
= Aspidium consobrinum Fée; Mém. Foug. 11 (Hist. Foug. Antil.): 85 (1866)
= Aspidium frondosum Wikstr.; Kongl. Vetensk. Acad. Handl. 1825: 437 (1825)
= Aspidium nemorosum Jenman; Gard. Chron. 1894: ? (1894)
= Dryopteris meridionalis (Poir.) C.Chr.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 8, 6: 46, f. 8 (1920)
= Nephrodium nemorosum (Jenman) Jenman; Ferns Brit. W. Ind. 223 (1908) (non Desv. (1827))
= Polypodium meridionale Poir.; Encycl. [J. Lamarck et al.] 5: 553 (1804)
variant :
var. speluncae (C.Chr.) K.U.Kramer; Fl. Netherl. Antilles 1: 59 (1962)
Distribution: Bermuda
81 BER
Synonyms: hidden
= Ctenitis meridionalis var. speluncae (C.Chr.) C.V.Morton; Contr. U. S. Natl. Herb. 38(6): 238 (1973)
= Dryopteris meridionalis var. speluncae C.Chr.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 8, 6: 47 (1920)
species :
Ctenitis mexicana A.R.Sm.; Mem. New York Bot. Gard. 88 [Pterid. Mexico]: 228 (2004)
Distribution: Mexico (Hidalgo, Puebla, Queretaro, Veracruz)
79 MEX
species :
Ctenitis microchlaena (Fée) Stolze; Fieldiana, Bot., n. s. 27: 9 (1991)
Distribution: Mexico (Chiapas, Veracruz); Honduras; Nicaragua; Costa Rica; Colombia (Santander, Valle); Venezuela (Barinas, Venezuela Distrito Federal, Falcon, Lara, Miranda, Portuguesa, Zulia); Ecuador; Peru; Bolivia (La Paz, Santa Cruz)
79 MEX 80 COS HON NIC 82 CLM VEN 83 BOL ECU PER
Synonyms: hidden
= Aspidium karstenii A.Braun; Index Sem. (B, Berolinensis) app. 3 (1867)
= Aspidium microcarpon Fée; Mém. Foug. 8 (Icon. Esp. Nouv.): 105 (1857) (non Blume (1828))
= Aspidium microchlaena Fée; Mém. Foug. 8 (Icon. Esp. Nouv.): 102 (1857)
= Aspidium obtusilobum Fée; Mém. Foug. 8 (Icon. Esp. Nouv.): 105 (1857)
= Dryopteris huatuscensis C.Chr.; Index Filic. 271 (1905), nom. nov. for Aspidium obtusilobum Fée 1857
= Dryopteris karstenii (A.Braun) C.Chr.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 7, 10(2): 98 (1913)
= Dryopteris microchlaena (Fée) C.Chr.; Index Filic. 278 (1905)
= Lastrea microchlaena (Fée) T.Moore; 97 (1858)
= Lastrea obtusiloba (Fée) T.Moore; Index Fil. [T. Moore] 98 (1858)
species :
Ctenitis microlepigera (Nakai) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 280 (1938)
Distribution: Bonin Isl. (Hahajima)
38 OGA
Conservation Status: IUCN R. CR in Japan
Synonyms: hidden
= Dryopteris microlepigera Nakai; Bot. Mag. (Tokyo) 43: 5 (1929)
species :
Ctenitis minutiloba Holttum; Blumea 31(1): 23 (1985)
Distribution: Borneo (Mt. Kinabalu, Sabah)
42 BOR
species :
Ctenitis multilobata Li Bing Zhang, Yi F.Duan & Rouhan; Ann. Missouri Bot. Gard. 102(1): 3-86 (2017)
Distribution: Madagascar
29 MDG
species :
Ctenitis muluensis Holttum; Fern Gaz. 12(6): 320 (1984)
Distribution: Borneo (Sarawak)
42 BOR
species :
Ctenitis nemorosa (Willd.) Ching; Sunyatsenia 5: 250 (1940)
Distribution: Hispaniola (Haiti, Dominican Republic); Puerto Rico
81 HIS PUE
Synonyms: hidden
= Aspidium lepidotrichum Desv.; Mag. Neuesten Entdeck. Gesammten Naturk. Ges. Naturf. Freunde Berlin 5: 321 (1811)
= Aspidium nemorosum Willd.; Sp. Pl., ed. 4 [Willdenow] 5(1): 255 (1810)
= Dryopteris myriolepis (Baker) C.Chr.; Index Filic. 279 (1905)
= Dryopteris nemorosa (Willd.) Urb.; Symb. Antill. [Urban] 4: 15 (1903)
= Lastrea nemorosa (Willd.) T.Moore; Index Fil. [T. Moore] 95 (1858)
= Nephrodium myriolepis Baker; J. Bot. 26: 34 (1888)
= Nephrodium nemorosum (Willd.) Desv.; Mem. Soc. Linn. Paris 6(2): 261 (1827)
species :
Ctenitis nervata (Fée) R.S.Viveros & Salino; Brittonia 69(3): 320 (2017)
Distribution: Colombia (Antioquia, Huila, Meta, Risaralda); Bolivia (La Paz, Santa Cruz); Brazil (Paraná, Rio de Janeiro, Santa Catarina, São Paulo)
82 CLM 83 BOL 84 BRA
Synonyms: hidden
= Aspidium nervatum Fée; Crypt. Vasc. Brésil 1: 136 (1869)
= Aspidium pedicellatum Christ; Denkschr. Kaiserl. Akad. Wiss., Wien. Math.-Naturwiss. Kl. 79(1): 20 (1908)
= Ctenitis pedicellata (Christ) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 124 (1947)
= Dryopteris indecora Rosenst.; Hedwigia 46: 117 (1906) (non C. Chr. (1905))
= Dryopteris pedicellata (Christ) C.Chr.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 7, 10(2): 88 (1913)
species :
Ctenitis nigrovenia (Christ) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 124 (1947)
Distribution: Mexico (Chiapas, Hidalgo, Oaxaca, Puebla, Queretaro, San Luis Potosi, Veracruz); Guatemala; Belize; El Salvador; Honduras; Nicaragua; Costa Rica; Panama; Trinidad; Colombia (Antioquia, Risaralda); Venezuela (Aragua, Barinas, Lara, Zulia); Ecuador; Peru; Bolivia (Cochabamba, La Paz, Santa Cruz); Brazil (Bahia, Mato Grosso, Pará)
79 MEX 80 BLZ COS ELS GUA HON NIC PAN 81 TRT 82 CLM VEN 83 BOL ECU PER 84 BRA
Synonyms: hidden
= Aspidium nigrovenium Christ; Bull. Herb. Boissier, sér. 2, 6: 162 (1906)
= Aspidium setosum Klotzsch; Linnaea 20: 371 (1847) (non Sw. (1801))
= Aspidium tonduzii Christ; Prim. Fl. Costaric. 3: 34 (1901)
= Ctenitis thelypteroides A.R.Sm.; Proc. Calif. Acad. Sci. ser. 4, 40: 215 (1975) [fide Viveros et al. 2017]
= Ctenitis tonduzii (Christ) L.D.Gómez; Brenesia 12-13: 64 (1977)
= Ctenitis tonduzii (Christ) R.M.Tryon & A.F.Tryon; Rhodora 84(837): 127 (1982), isonym
= Dryopteris nigrovenia (Christ) C.Chr.; Index Filic. 279 (1905)
= Dryopteris tonduzii (Christ) C.Chr.; Index Filic. 664 (1906)
= Nephrodium nigrovenium Christ; , Bot. Gaz. 20: 545 (1895)
= Nephrodium setosum (Klotzsch) Keyserl.; Polyp. Herb. Bunge 48 (1873)
species :
Ctenitis oophylla (C.Chr.) Ching; Sunyatsenia 5: 250 (1940)
Distribution: Jamaica; Hispaniola (Haiti, Dominican Republic)
81 HIS JAM
Synonyms: hidden
= Dryopteris oophylla C.Chr.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 8, 6: 39 (1920)
species :
Ctenitis palatangana (Hook.) Ching; Sunyatsenia 5: 250 (1940)
Distribution: Ecuador
83 ECU
Synonyms: hidden
= Dryopteris palatangana (Hook.) C.Chr.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 8, 6: 56 (1920)
= Nephrodium palatanganum Hook.; Sp. Fil. [W. J. Hooker] 4: 260 (1862)
species :
Ctenitis paleolata Copel.; Philipp. J. Sci. 81: 24 (1952)
Distribution: Philippines (Luzon: Benguet)
42 PHI
Conservation Status: IUCN Ex/E. Known only from type
species :
Ctenitis pallens (Brack.) M.G.Price; Kalikasan 12: 155 (1983)
Distribution: Sulawesi; Palawan; Philippines (Luzon, Samar, Negros, Mindanao); New Guinea (Papua New Guinea); Bismarck Arch. (New Ireland)
42 NWG PHI PLW SUL 43 BIS
Synonyms: hidden
= Lastrea pallens Brack.; U.S. Expl. Exped., Filic. 16: 197 (1854)
= Ctenitis rizalensis (Christ) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 125 (1947)
= Dryopteris rizalensis Christ; Bull. Herb. Boissier, sér. 2, 6: 1001 (1906)
species :
Ctenitis paranaensis (C.Chr.) Lellinger; Amer. Fern J. 74(2): 56 (1984)
Distribution: Brazil (Paraná, Rio Grande do Sul, Santa Catarina)
84 BRA
Synonyms: hidden
= Dryopteris falciculata f. glabrata Hieron.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 7, 10, pt. 2: 92 (1913)
= Dryopteris falciculata var. paranaensis C.Chr.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd. VII. 10(2): 92 (1913)
= Dryopteris paranaensis (C.Chr.) Sehnem; Fl. Ilustr. Catarin. (Aspidiaceas): 158 (1979)
species :
Ctenitis parvula Proctor; Brit. Fern Gaz. 9: 217 (1965)
Distribution: Jamaica
81 JAM
Synonyms: hidden
= Nephrodium pubescens Desv.; Mem. Soc. Linn. Paris 6(2): 261 (1827), nom. nud.
species :
Ctenitis pauciflora (Kaulf. ex Spreng.) Holttum; Kew Bull. 38(1): 123 (1983)
Distribution: Mauritius
29 MAU
Synonyms: hidden
= Aspidium pauciflorum Kaulf. ex Spreng.; Syst. Veg., ed. 16 [Sprengel] 4(1): 101 (1827)
= Dryopteris crinita var. pauciflora (Kaulf. ex Spreng.) Bonap.; Notes Ptéridol. 4: 17 (1917)
= Lastrea pauciflora (Kaulf. ex Spreng.) C.Presl; Tent. Pterid. 75 (1836)
= Nephrodium crinitum var. pauciflorum (Kaulf. ex Spreng.) Baker; Fl. Mauritius [J.G.Baker] 495 (1877)
= Nephrodium pauciflorum (Kaulf. ex Spreng.) Kiaersk.; Cat. Fil. hb. hort. Haun. 63 (1874)
species :
Ctenitis propinqua (J.Sm. ex C.Presl) Copel.; Fern Fl. Philipp. 2: 289 (1960)
Distribution: Philippines (Luzon)
42 PHI
Synonyms: hidden
= Aspidium propinquum (J.Sm. ex C.Presl) Fée; Mém. Foug. 5 (Gen. Filic.): 292 (1852)
= Ctenitis preslii (Baker) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 124 (1947)
= Dryopteris preslii (Baker) Kuntze; Revis. Gen. Pl. 2: 813 (1891)
= Lastrea propinqua J.Sm.; J. Bot. (Hooker) 3: 412 (1841)
= Lastrea propinqua J.Sm. ex C.Presl; Abh. Königl. Böhm. Ges. Wiss., ser. 5, 6 [Epim. Bot.]: 38 (1851) (non Nephrodium propinquum R. Br. (1810))
= Nephrodium preslii Baker; Syn. Fil. [W. J. Hooker & Baker] 272 (1867), nom. nov.
species :
Ctenitis pseudorhodolepis Ching & Chu H.Wang; Acta Phytotax. Sin. 19(1): 121 (1981)
Distribution: China (Sichuan, Hunan, Guizhou)
36 CHI
species :
Ctenitis pulvinata (C.Chr.) Ching; Sunyatsenia 5: 250 (1940)
Distribution: Madagascar (Antananarivo)
29 MDG
Remarks: possibly to C. cirrhosa from Africa
Synonyms: hidden
= Dryopteris pulvinata C.Chr.; Notes Ptéridol. [Bonaparte] 16: 177, t. 6 (1925)
species :
Ctenitis rapensis (E.D.Br.) Holttum; Blumea 31(1): 34 (1985)
Distribution: Austral Isl. (Rapa Iti)
61 AUI
Synonyms: hidden
= Dryopteris sciaphila var. rapensis E.D.Br.; Bull. Bernice P. Bishop Mus. 89 [Fl. S. E. Polynesia II]: 33 (1931)
species :
Ctenitis refulgens (Klotzsch ex Mett.) Vareschi; Fl. Venez. 1: 404 (1969)
Distribution: Mexico (Chiapas); Guatemala; Belize; Honduras; Nicaragua; Panama; Trinidad; Colombia (Chocó, Meta); Venezuela (Amazonas, Apure, Barinas, Bolívar, Zulia); Guyana; Surinam; French Guiana; Ecuador; Peru; Bolivia (Beni, La Paz, Pando); Brazil (Acre, Amazonas, Amapá, Maranhão, Mato Grosso, Pará, Roraima)
79 MEX 80 BLZ GUA HON NIC PAN 81 TRT 82 CLM FRG GUY SUR VEN 83 BOL ECU PER 84 BRA
Synonyms: hidden
= Dryopteris refulgens (Klotzsch) C.Chr.; Index Filic. 288 (1905)
= Nephrodium refulgens (Klotzsch) Diels; Nat. Pflanzenfam. [Engler & Prantl] I, 4: 170 (1891)
= Phegopteris refulgens (Klotzsch ex Mett.) Mett.; Ann. Sci. Nat., Bot., sér. 5, 2: 240 (1864)
= Phegopteris tricholepis Fée; Crypt. Vasc. Brésil 1: 98, t. 32, f. 1 (1869)
= Polypodium refulgens Klotzsch ex Mett.; Syn. Fil. [W. J. Hooker & Baker] 307 (1857)
variant :
var. peruviana (C.Chr.) R.S.Viveros & Salino; Phytotaxa 335(1): 68 (2018)
Distribution: Peru; Bolivia (La Paz)
83 BOL PER
Synonyms: hidden
= Dryopteris refulgens var. peruviana C.Chr.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 7, 10, pt. 2: 90, fig. 5 (1913)
species :
Ctenitis reunionensis Hennequin, Li Bing Zhang & Yi F.Duan; Ann. Missouri Bot. Gard. 102(1): 3-86 (2017)
Distribution: La Réunion
29 REU
species :
Ctenitis salvinii (Baker) Stolze; Amer. Fern J. 67(2): 43 (1977)
Distribution: Mexico (Tabasco, Chiapas, Veracruz); Guatemala; ?Belize
79 MEX 80 GUA
Synonyms: hidden
= Aspidium lindenii (Kuhn) E.Fourn.; Mexic. Pl. 1: 97 (1872) (non Kuhn 1869)
= Aspidium lindenii Kuhn; Linnaea 36: 116 (1869) (non Fourn (1872))
= Ctenitis lindenii (Kuhn) A.R.Sm.; Proc. Calif. Acad. Sci. ser. 4, 40: 229 (1975)
= Dryopteris lindenii (Kuhn) Kuntze; Revis. Gen. Pl. 2: 813 (1891)
= Dryopteris moreletii C.Chr.; Index Filic. 278 (1905)
= Dryopteris salvinii (Baker) Kuntze; Revis. Gen. Pl. 2: 813 (1891)
= Nephrodium lindenii (Kuhn) Baker; Syn. Fil. [W. J. Hooker & Baker], ed. 2: 493 (1874)
= Nephrodium salvinii Baker; Syn. Fil. [W. J. Hooker & Baker] 274 (1867)
species :
Ctenitis samoensis (C.Chr.) Holttum; Blumea 31(1): 32 (1985)
Distribution: American Samoa (Tutuila); [Western] Samoa (´Upolu)
60 ASA SAM
Synonyms: hidden
= Dryopteris samoensis C.Chr.; Index Filic. 290 (1905)
= Nephrodium intermedium Baker; Syn. Fil. [W. J. Hooker & Baker] 283 (1867)
= Polypodium paleaceum Powell; Syn. Fil. [W. J. Hooker & Baker] 505, 187 (1867) (non Hook.f. (1847)
species :
Ctenitis santae-clarae (C.Chr.) Ching; Sunyatsenia 5: 250 (1940)
Distribution: Cuba
81 CUB
Synonyms: hidden
= Dryopteris santae-clarae C.Chr.; J. Wash. Acad. Sci. 22: 166 (1932)
species :
Ctenitis sciaphila (Maxon) Ching; Sunyatsenia 5: 250 (1940)
Distribution: Cook Isl. (Rarotonga); Society Isl. (Tahiti, Bora Bora, Huahine, Moorea)
61 COO SCI
Synonyms: hidden
= Dryopteris sciaphila Maxon; Univ. Calif. Publ. Bot. 12: 24 (1924)
variant :
var. raivavensis (E.D.Br.) Holttum; Blumea 31(1): 33 (1985)
Distribution: Austral Isl. (Raivavae, Tubuai, Rurutu)
61 AUI
Synonyms: hidden
= Dryopteris sciaphila var. raivavensis E.D.Br.; Bull. Bernice P. Bishop Mus. 89 [Fl. S. E. Polynesia II]: 32-33 (1931)
species :
Ctenitis seramensis Holttum; Blumea 31(1): 19 (1985)
Distribution: Moluccas (Seram, Ambon)
42 MOL
species :
Ctenitis setosa (C.Presl) Holttum; Novit. Bot. Delect. Seminum Horti Bot. Univ. Carol. Prag. 1968: 20 (1969)
Distribution: Philippines
42 PHI
Conservation Status: very rare
Synonyms: hidden
= Dryopteris setosa (C.Presl) C.Chr.; Index Filic. 292 (1905), nom. illeg.
= Lastrea setosa C.Presl; Abh. Königl. Böhm. Ges. Wiss., ser. 5, 6 [Epim. Bot.]: 40 (1851)
= Nephrodium hirtum (C.Presl) Copel.; Polypodiaceae Philipp. 25 (1905) (non Hook. (1862))
= Phegopteris hirta (C.Presl) Christ; Bull. Herb. Boissier, sér. 1, 6: 195 (1898)
= Polypodium hirtum C.Presl; Reliq. Haenk. 1: 27 (1825) (non Sw. (1788))
species :
Ctenitis sieberiana (Kaulf. ex Spreng.) Hennequin & Rouhan; Ann. Missouri Bot. Gard. 102(1): 3-86 (2017)
Distribution: Mauritius
29 MAU
Synonyms: hidden
= Dryopteris cyatheifolia (Desv.) C.Chr.; Index Filic. 260 (1905)
= Heterogonium cyatheifolium (Desv.) Tardieu; Amer. Fern J. 48: 33 (1958)
= Heterogonium sieberianum (Kaulf. ex Spreng.) Holttum; Kew Bull. 38(1): 115 (1983)
= Phegopteris cyatheifolia (Desv.) Mett.; Fil. Afr. [Kuhn] 122 (1868)
= Phegopteris sieberiana (Kaulf. ex Spreng.) Fée; Mém. Foug. 5 (Gen. Filic.): 243 (1852)
= Polypodium cyatheifolium Desv.; Mem. Soc. Linn. Paris 6(2): 239 (1827)
= Polypodium sieberianum Kaulf. ex Spreng.; Syst. Veg., ed. 16 [Sprengel] 4(1): 56 (1827)
species :
Ctenitis silvatica Holttum; Blumea 31(1): 20 (1985)
Distribution: Philippines (many localities); New Guinea (Papua New Guinea)
42 NWG PHI
species :
Ctenitis sinii (Ching) Ohwi; Fl. Jap. [Ohwi] 92 (1957)
Distribution: China (Fujian, Guangdong, Guangxi, Hunan, Jiangxi, Zhejiang); Japan; Vietnam
36 CHI 38 JAP 41 VIE
Conservation Status: IUCN R
Synonyms: hidden
= Ataxipteris sinii (Ching) Holttum; Blumea 30(1): 10 (1984)
= Ctenitopsis sinii (Ching) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 319 (1938)
= Tectaria sinii Ching; Bull. Biol. Dept. Sci. Coll. Sun Yatsen Univ. 6 22 (1933)
species :
Ctenitis sloanei (Poepp. ex Spreng.) C.V.Morton; Amer. Fern J. 59: 66 (1969)
Name: Florida lacefern, Red-hair Comb Fern
Distribution: USA (S-Florida); Bermuda; Mexico (Chiapas); El Salvador; Honduras; Nicaragua; Costa Rica; Panama; Cuba; Jamaica; Hispaniola (Dominican Republic); St. Vincent; Grenada; Trinidad; Colombia (Antioquia, Chocó, Cundinamarca, Magdalena, Risaralda, Valle); Venezuela (Anzoategui, Aragua, Nueva Esparta, Sucre, Venezuela Distrito Federal, Lara, Monagas); Ecuador; Galapagos; Peru; Bolivia (Beni, Chuquisaca, Cochabamba, La Paz, Santa Cruz); Brazil (Minas Gerais, Ceará, Pernambuco); Argentina (Jujuy)
78 USA 79 MEX 80 COS ELS HON NIC PAN 81 BER CUB GRE HIS JAM STV TRT 82 CLM VEN 83 BOL ECU GAL PER 84 BRA 85 ARG
Remarks: unification with C. ampla fide Viveros et al. 2017
Synonyms: hidden
= Aspidium amplum (Humb. & Bonpl. ex Willd.) Mett.; Abh. Senckenberg. Naturf. Ges. 2(2) [Pheg. u. Asp.]: 358 (1858)
= Aspidium catocarpum Kunze; Linnaea 9: 95 (1834)
= Aspidium culcita Christ; Bull. Herb. Boissier, sér. 2, 6: 54 (1906)
= Aspidium furcatum Klotzsch; Linnaea 20: 371 (1847)
= Aspidium nemophilum Kunze; Linnaea 9: 95 (1834)
= Ctenitis ampla (Humb. & Bonpl. ex Willd.) Ching; Sunyatsenia 5: 250 (1940)
= Ctenitis catocarpa (Kunze) C.V.Morton; Steyerm. et al., Fieldiana, Bot. 28: 12 (1951)
= Ctenitis nemophila (Kunze) Ching; Sunyatsenia 5: 250 (1940)
= Dryopteris ampla (Humb. & Bonpl. ex Willd.) Kuntze; Revis. Gen. Pl. 2: 812 (1891)
= Dryopteris catocarpa (Kunze) Kuntze; Revis. Gen. Pl. 2: 812 (1891)
= Dryopteris culcita (Christ) C.Chr.; Index Filic., Suppl. 2 (1906-1912): 31 (1913)
= Dryopteris furcata (Klotzsch) Kuntze; Revis. Gen. Pl. 2: 812 (1891)
= Dryopteris nemophila (Kunze) C.Chr.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 8, 6: 57, f. 11 (1920)
= Filix ampla (Humb. & Bonpl. ex Willd.) Farw.; Rep. (Annual) Michigan Acad. Sci. 18: 81 (1916)
= Lastrea ampla (Humb. & Bonpl. ex Willd.) T.Moore; Index Fil. [T. Moore] 85 (1858)
= Lastrea furcata (Klotzsch) T.Moore; Index Fil. [T. Moore] 92 (1858)
= Lastrea nemophila (Kunze) T.Moore; Index Fil. [T. Moore] 87 (1858)
= Nephrodium amplum (Humb. & Bonpl. ex Willd.) Baker; Syn. Fil. [W. J. Hooker & Baker] 285 (1867) (non Hook. (1862))
= Nephrodium catocarpum (Kunze) Hook.; Sp. Fil. [W. J. Hooker] 4: 259 (1862)
= Nephrodium furcatum (Klotzsch) Hook.; Sp. Fil. [W. J. Hooker] 4: 136 (1862)
= Nephrodium paleaceum (Hook.f.) Diels; Nat. Pflanzenfam. [Engler & Prantl] I, 4: 177 (1891)
= Phegopteris ampla (Humb. & Bonpl. ex Willd.) Fée; Mém. Foug. 5 (Gen. Filic.): 243 (1852)
= Phegopteris lachnopodia J.Sm.; Cat. Cult. Ferns: 16 (1857)
= Phegopteris sloanei (Poepp. ex Spreng.) Fée; Mém. Foug. 5 (Gen. Filic.): 243 (1852)
= Polypodium amplum Humb. & Bonpl. ex Willd.; Sp. Pl., ed. 4 [Willdenow] 5(1): 207 (1810)
= Polypodium lachnopodium J.Sm.; Compan. Bot. Mag. 8 (1846)
= Polypodium paleaceum Hook.f.; Trans. Linn. Soc. London 20: 166 (1847) (non Borkh. (1798))
= Polypodium sloanei Poepp. ex Spreng.; Syst. Veg., ed. 16 [Sprengel] 4(1): 59 (1827)
species :
Ctenitis sparsa Li Bing Zhang & Yi F.Duan; Ann. Missouri Bot. Gard. 102(1): 3-86 (2017)
Distribution: Madagascar
29 MDG
species :
Ctenitis spekei (Baker) Li Bing Zhang & Yi F.Duan; Ann. Missouri Bot. Gard. 102(1): 3-86 (2017)
Distribution: Comores (Grande Comore, Mayotte, Anjouan)
29 COM
Synonyms: hidden
= Aspidium spekei (Baker) Kuhn; Filic. Afr. 209 (1868)
= Dryopteris spekei (Baker) Kuntze; Revis. Gen. Pl. 2: 813 (1891)
= Nephrodium spekei Baker; Syn. Fil. [W. J. Hooker & Baker] 263 (1867)
= Thelypteris spekei (Baker) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 254 (1941)
species :
Ctenitis squamigera (Hook. & Arn.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 125 (1947)
Distribution: Hawaii (Kauai, Oahu, Molokai, Lanai, Maui, Hawaii Isl.)
63 HAW
Conservation Status: IUCN E
Synonyms: hidden
= Aspidium squamigerum (Hook. & Arn.) Mann; Proc. Amer. Acad. Arts 7: 217 (1868)
= Dryopteris squamigera (Hook. & Arn.) Kuntze; Revis. Gen. Pl. 2: 813 (1891)
= Filix-mas squamigera (Hook. & Arn.) Farw.; Amer. Midl. Naturalist 12: 234-251 (1931)
= Lastrea squamigera (Hook. & Arn.) Brack.; U.S. Expl. Exped., Filic. 16: 198 (1854)
= Nephrodium squamigerum Hook. & Arn.; Bot. Beechey Voy.: 106 (1832)
species :
Ctenitis strigilosa (Davenp.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 125 (1947)
Distribution: Mexico (Veracruz, Oaxaca, Chiapas); Guatemala
79 MEX 80 GUA
Synonyms: hidden
= Aspidium strigilosum Davenp.; Bot. Gaz. 21: 256 (1896)
= Dryopteris strigilosa Davenp.; Bot. Gaz. 21: 257 (1896), nom. altern.
= Dryopteris strigilosa var. cookii Maxon; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 7, 10(2): 105 (1913)
= Nephrodium strigilosum Davenp.; Bot. Gaz. 21: 257 (1896), nom. altern.
= Lastrea strigilosa (Davenp.) Davenp.; Bot. Gaz. 21: 257 (1896)
species :
Ctenitis subconnexa (Christ) Holttum; Blumea 31(1): 17 (1985)
Distribution: Moluccas (Bacan)
42 MOL
Synonyms: hidden
= Dryopteris subconnexa (Christ) C.Chr.; Index Filic. 295 (1905)
= Phegopteris subconnexa Christ; Warb., Monsunia 1: 83 (1900)
variant :
var. alstonii Holttum; Blumea 31(1): 17 (1985)
Distribution: Moluccas (Bacan)
42 MOL
Conservation Status: Known only from type
species :
Ctenitis subcrenulata (Baker) Li Bing Zhang, Rouhan & Yi F.Duan; Ann. Missouri Bot. Gard. 102(1): 3-86 (2017)
Distribution: Madagascar
29 MDG
Synonyms: hidden
= Aspidium subcrenulatum (Baker) Kuhn; Reisen Ost-Afr. [Decken] 3(3): Bot. 65 (1879)
= Ctenitis modesta (C.Chr.) Ching; Sunyatsenia 5: 250 (1940)
= Ctenitis ochrorachis (Baker) Tardieu; Humbert, Fl. Madagasc. Fam. 5, 1: 332 (1958)
= Ctenitis ochrorachis var. violacea Rakotondr.; Bull. Mus. Nat. Hist. Nat., B, Adansonia, ser. 4, 10(4): 375 (1988 publ. 1989)
= Dryopteris modesta C.Chr.; Notes Ptéridol. [Bonaparte] 16: 171, t. 5 (1925)
= Dryopteris ochrorachis (Baker) C.Chr.; Index Filic. 280 (1905)
= Dryopteris subcrenulata (Baker) C.Chr.; Index Filic. 295 (1905)
= Nephrodium ochrorachis Baker; J. Bot. 22: 142 (1884)
= Nephrodium subcrenulatum Baker; J. Linn. Soc., Bot. 16: 202 (1877)
species :
Ctenitis subdryopteris (Christ) Lellinger; Proc. Biol. Soc. Wash. 89: 710 (1977)
Distribution: Costa Rica; Panama
80 COS PAN
Synonyms: hidden
= Dryopteris ampla var. subdryopteris (Christ) C.Chr.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 8, 6: 53 (1920)
= Dryopteris subdryopteris (Christ) C.Chr.; Index Filic. 295 (1905)
= Phegopteris subdryopteris Christ; Prim. Fl. Costaric. 3: 36 (1901)
species :
Ctenitis subglandulosa (Hance) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 302 (1938)
Distribution: China (Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hubei, Hunan, Jiangxi, Sichuan, Yunnan, Zhejiang); Tibet; Japan; Ryukyu Isl.; Bonin Isl. (Chichijima, Anijima, Otohtojima, Hahajima); Volcano Isl. (Kita-Iwojima, Minami-Iwojima); Taiwan; India (Arunachal Pradesh, Assam State, Meghalaya, Nagaland, West Bengal); Bangladesh; Myanmar; N-Thailand; Vietnam; peninsular Malaysia; Philippines (Luzon, Panay, Mindoro); E-Java; Lesser Sunda Isl. (Bali); Moluccas (Seram); New Guinea (Papua New Guinea); Vanuatu; Fiji; Northern Marianas (Rota Isl.); Guam
36 CHI TIB 38 JAP KZN NNS OGA TAI 40 BAN IND 41 MYA THA VIE 42 JAW LSI MLY MOL NWG PHI 60 FIJ VAN 62 GUM MRN
Synonyms: hidden
= Alsophila subglandulosa Hance; Ann. Sci. Nat., Bot., sér. 5, 6: 253 (1866)
= Aspidium blumei Thwaites; Enum. Pl. Zeyl. [Thwaites]: 392 (1864) (non Kunze ex Mett. (1856))
= Aspidium intermedium var. rhodolepis (C.B.Clarke) Christ; Bull. Acad. Int. Géogr. Bot. 1902: 256 (1902)
= Aspidium lepigerum (Baker) Christ; Monsunia 1: 82 (1900)
= Aspidium subtripinnatum Miq.; Ann. Mus. Bot. Lugduno-Batavi 8: 179 (1867)
= Ctenitis anyuanensis Ching & C.H.Wang; Acta Phytotax. Sin. 19(1): 118 (1981)
= Ctenitis calcarea Ching & Chu H.Wang; Acta Phytotax. Sin. 19(1): 118 (1981)
= Ctenitis chungyiensis Ching & Chu H.Wang; Acta Phytotax. Sin. 19(1): 118 (1981)
= Ctenitis costulisora Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 299 (1938)
= Ctenitis holttumii Sarn.Singh & Panigrahi; Ferns Fern Allies Arunachal Pradesh 2: 624 (2005) (fide Fraser-Jenkins 2008)
= Ctenitis lepigera (Baker) Tagawa; Acta Phytotax. Geobot. 8: 95 (1939)
= Ctenitis maolanensis P.S.Wang; Acta Bot. Yunnan. 9(4): 399, f. 2: 1-4 (1987)
= Ctenitis membranifolia Ching & Chu H.Wang; Acta Phytotax. Sin. 19(1): 121 (1981)
= Ctenitis rhodolepis (C.B.Clarke) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 300 (1938)
= Ctenitis subtripinnata (Miq.) H.Itô; Nov. Fl. Jap. [Nakai & Honda] 4: 74 (1939)
= Ctenitis yunnanensis Ching & Chu H.Wang; Acta Phytotax. Sin. 19(1): 124 (1981)
= Cyathea subglandulosa (Hance) Copel.; Philipp. J. Sci. 4: 46 (1909)
= Dryopteris lepigera (Baker) Kuntze; Revis. Gen. Pl. 2: 813 (1891)
= Dryopteris maximowicziana var. rhodolepis (C.B.Clarke) Koidz.; Fl. Symb. Orient.-Asiat. 22 (1930)
= Dryopteris oldhamii (Baker) C.Chr.; Index Filic. 280 (1905)
= Dryopteris rhodolepis (C.B.Clarke) C.Chr.; Index Filic. 288 (1905)
= Dryopteris subglandulosa (Hance) Hayata; Icon. Pl. Formosan. 6: 101 (1917), nom. illeg., non Kuntze
= Dryopteris subtripinnata (Miq.) Kuntze; Revis. Gen. Pl. 2: 811 (1891)
= Lastrea blumei (Thwaites) Bedd.; Ferns S. India t. 249 (1864)
= Lastrea intermedia var. blumei Bedd.; Suppl. Ferns Brit. Ind. 61 (1892)
= Lastrea intermedia var. rhodolepis (C.B.Clarke) Bedd.; Ferns Brit. India Suppl. 62 (1892)
= Nephrodium blumei Hook.; Sp. Fil. [W. J. Hooker] 4: 135 (1862)
= Nephrodium lepigerum Baker; Syn. Fil. [W. J. Hooker & Baker] 284 (1867)
= Nephrodium rhodolepis C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 526, t. 72 (1880)
= Nephrodium subtripinnatum (Miq.) Baker; Syn. Fil. [W. J. Hooker & Baker], ed. 1: 455 (1868)
= Polypodium oldhamii Baker; Syn. Fil. [W. J. Hooker & Baker] 311 (1867)
species :
Ctenitis submarginalis (Langsd. & Fisch.) Ching; Sunyatsenia 5: 250 (1940)
Name: Brownhair lacefern
Distribution: USA (Florida, S-Louisiana); Mexico (Chiapas, Hidalgo, Oaxaca, Puebla, Queretaro, San Luis Potosi, Veracruz); Guatemala; Honduras; Nicaragua; Hispaniola (Haiti, Dominican Republic); Colombia (Antioquia, Boyacá, Caldas, Cundinamarca, Risaralda, Santander, Valle); Venezuela (Aragua, Carabobo, Venezuela Distrito Federal, Lara, Merida, Miranda, Monagas, Yaracuy, Zulia); Ecuador; Peru; Bolivia (Chuquisaca, Cochabamba, La Paz, Santa Cruz, Tarija); Brazil (Alagoas, Bahia, Ceará, Espírito Santo, Goiás, Minas Gerais, Mato Grosso do Sul, Mato Grosso, Pernambuco, Paraná, Rio de Janeiro, Rio Grande do Norte, Rio Grande do Sul, Santa Catarina, São Paulo); Paraguay (Alto Paraná, Caazapa, Canindeyu); Uruguay (Artigas, Cerro Largo, Rivera, Rocha, San Jose, Soriano, Tacuarembo, Trenta y Tres); Argentina (Buenos Aires, Chaco, Corrientes, Entre Rios, Formosa, Jujuy, Misiones, Salta, Santa Fe, Tucuman)
78 USA 79 MEX 80 GUA HON NIC 81 HIS 82 CLM VEN 83 BOL ECU PER 84 BRA 85 ARG PAR URU
Synonyms: hidden
= Aspidium caripense (Humb. & Bonpl. ex Willd.) Mett.; Fil. Hort. Bot. Lips. 90, t. 18, f. 9 (1856)
= Ctenitis submarginalis f. caripensis (Humb. & Bonpl. ex Willd.) Lellinger; Proc. Biol. Soc. Wash. 89(61): 710 (1977)
= Dryopteris caripensis (Humb. & Bonpl. ex Willd.) Kuntze; Revis. Gen. Pl. 2: 812 (1891)
= Dryopteris collina Christ; Bull. Herb. Boissier, sér. 2, 7: 922 (1907)
= Dryopteris lagerheimii (Sodiro) C.Chr.; Index Filic. 273 (1905)
= Dryopteris sellowii Hieron.; Hedwigia 46: 324, t. 3, f. 1 (1906)
= Dryopteris soriloba Christ; Repert. Spec. Nov. Regni Veg. 6: 350 (1908)
= Dryopteris submarginalis (Langsd. & Fisch.) C.Chr. & Cogn.; Index Filic. 296 (1905)
= Dryopteris submarginalis f. caripensis (Humb. & Bonpl. ex Willd.) C.Chr.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 7, 10, pt. 2: 97 (1913)
= Dryopteris submarginalis f. glaucescens (Sodiro) C.Chr.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 7, 10, pt. 2: 97 (1913)
= Dryopteris submarginalis var. exigua Dutra; Anais Reunião Sul-Amer. Bot. 2: 47 (1938 publ. 1939)
= Dryopteris submarginalis var. lagerheimii (Sodiro) C.Chr.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 7, 10, pt. 2: 98 (1913)
= Dryopteris submarginalis var. tarapotensis (Hook.) C.Chr.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 7, 10, pt. 2: 98 (1913)
= Lastrea caripensis (Humb. & Bonpl. ex Willd.) T.Moore; Index Fil. [T. Moore] 87 (1858)
= Lastrea microcarpa T.Moore; Index Fil. [T. Moore] 96 (1858)
= Nephrodium caripense (Humb. & Bonpl. ex Willd.) Hook.; Sp. Fil. [W. J. Hooker] 4: 99 (1862)
= Nephrodium caripense var. squamigerum Baker; (1870)
= Nephrodium crinitum Sodiro; Crypt. Vasc. Quit.: 250, 642 (1893) (non Desv. (1827))
= Nephrodium crinitum var. glaucescens Sodiro; Crypt. Vasc. Quit.: 250, 642 (1893)
= Nephrodium lagerheimii Sodiro; Crypt. Vasc. Quit.: 252 (1893)
= Nephrodium squamigerum (Mett.) Rosenst.; Hedwigia 43: 224 (1904), nom. illeg.
= Nephrodium submarginale (Langsd. & Fisch.) Hicken; Anales Soc. Ci. Argent. 63: 7 (1907)
= Nephrodium tarapotense Hook.; Sp. Fil. [W. J. Hooker] 4: 107 (1862)
= Phegopteris submarginalis (Langsd. & Fisch.) J.Sm.; Cat. Cult. Ferns: 16 (1857)
= Polypodium bonplandii Desv.; Mem. Soc. Linn. Paris 6(2): 241 (1827)
= Polypodium caripense Humb. & Bonpl. ex Willd.; Sp. Pl., ed. 4 [Willdenow] 5(1): 202 (1810)
= Polystichum caripense (Humb. & Bonpl. ex Willd.) Keyserl.; Polyp. Herb. Bunge 45 (1873)
= Polypodium submarginale Langsd. & Fisch.; Pl. Voy. Russes Monde, Icon. Fil. 12 (1810)
= Thelypteris submarginalis (Langsd. & Fisch.) Small; Ferns S. E. States [Small] 258 cum tab. 476 (1938)
variant :
var. tenuifolia (C.Presl) R.S.Viveros & Salino; Phytotaxa 335(1): 73 (2018)
Distribution: Brazil (Alagoas, Ceara, Bahia, Espírito Santo, Mato Grosso do Sul, Minas Gerais, Pernambuco, Rio de Janeiro, São Paulo)
84 BRA
Synonyms: hidden
= Aspidium sancti-pauli Christ; Denkschr. Kaiserl. Akad. Wiss., Wien. Math.-Naturwiss. Kl. 79(1): 21 (1908)
= Dryopteris submarginalis f. tenuifolia (C.Presl) C.Chr.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 7, 10, p. 2: 96 (1913)
= Lastrea tenuifolia C.Presl; Abh. Königl. Böhm. Ges. Wiss., ser. 5, 6 [Epim. Bot.]: 37 (1851)
= Phegopteris blanchetiana Fée; Mém. Foug. 5 (Gen. Filic.): 245 (1852)
= Phegopteris oreopteridastrum Fée; Crypt. Vasc. Brésil 1: 97 (1869)
species :
Ctenitis subobscura (Christ) Holttum; Fern Gaz. 12(6): 320 (1984)
Distribution: peninsular Thailand; SW-Thailand; peninsular Malaysia; Sumatra; Borneo (Sabah, Kalimantan); Philippines (Mindanao); Java
41 THA 42 BOR JAW MLY PHI SUM
Synonyms: hidden
= Dryopteris squamulifera Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 16: 9 (1914)
= Dryopteris subobscura (Christ) C.Chr.; Index Filic. 296 (1905)
= Phegopteris subobscura Christ; Bull. Herb. Boissier, sér. 1, 6: 836 (1898)
species :
Ctenitis sumbawensis Holttum; Blumea 31(1): 26 (1985)
Distribution: Lesser Sunda Isl. (Sumbawa, Lombok); Moluccas (Seram)
42 LSI MOL
species :
Ctenitis tabacifera (Alderw.) Ching; Sunyatsenia 5: 250 (1940)
Distribution: Moluccas (Seram, Ambon)
42 MOL
Synonyms: hidden
= Dryopteris tabacifera Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 3, 2: 147 (1920)
species :
Ctenitis tardieu-blotiae Li Bing Zhang & Yi F.Duan; Ann. Missouri Bot. Gard. 102(1): 3-86 (2017)
Distribution: Madagascar
29 MDG
species :
Ctenitis thwaitesii Holttum; Blumea 31(1): 31 (1985)
Distribution: Sri Lanka
40 SRL
species :
Ctenitis truncicola (C.Chr.) Ching; Sunyatsenia 5: 250 (1940)
Distribution: Madagascar; ?La Réunion
29 MDG
Synonyms: hidden
= Dryopteris truncicola C.Chr.; Notes Ptéridol. [Bonaparte] 16: 181, t. 5 (1925)
species :
Ctenitis ursina A.R.Sm.; Proc. Calif. Acad. Sci. ser. 4, 40: 216 (1975)
Distribution: Mexico (Chiapas)
79 MEX
species :
Ctenitis velata (Kunze ex Mett.) R.M.Tryon & A.F.Tryon; Rhodora 84(837): 127 (1982)
Distribution: Cuba
81 CUB
Synonyms: hidden
= Aspidium velatum Kunze ex Mett.; Abh. Senckenberg. Naturf. Ges. 2(2) [Pheg. u. Asp.]: 363 [Pheg. u. Asp. 79, no. 190] (1858)
= Dryopteris velata (Kunze ex Mett.) Kuntze; Revis. Gen. Pl. 2: 814 (1891)
= Lastrea velata (Kunze ex Mett.) T.Moore; Index Fil. [T. Moore] 107 (1858)
= Nephrodium velatum (Kunze ex Mett.) Hook.; Sp. Fil. [W. J. Hooker] 4: 101, t. 247 (1862)
= Thelypteris velata (Kunze ex Mett.) C.F.Reed; Phytologia 17: 323 (1968)
species :
Ctenitis vellea (Willd.) Proctor; Amer. Fern J. 40: 227 (1950)
Distribution: Cuba; Jamaica
81 CUB JAM
Synonyms: hidden
= Aspidium aureovestitum (Hook.) Griseb.; Fl. Brit. W.I. [Grisebach] 692 (1864)
= Aspidium velleum Willd.; Sp. Pl., ed. 4 [Willdenow] 5(1): 256 (1810)
= Dryopteris vellea (Willd.) Kuntze; Revis. Gen. Pl. 2: 814 (1891)
= Lastrea vellea (Willd.) T.Moore; Index Fil. [T. Moore] 107 (1858)
= Nephrodium aureovestitum Hook.; Sp. Fil. [W. J. Hooker] 4: 101, t. 246 (1862)
= Nephrodium velleum (Willd.) Desv.; Mem. Soc. Linn. Paris 6(2): 261 (1827)
= Phegopteris aureovestita (Hook.) Fée; Mém. Foug. 11 (Hist. Foug. Antil.): 52 (1866)
species :
Ctenitis vilis (Kunze) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 290 (1938)
Distribution: peninsular Thailand; peninsular Malaysia (Perak); Sumatra; Borneo (Sarawak, Sabah, Kalimantan); Java; Lesser Sunda Isl. (Lombok, Bali, Timor); Moluccas (Ambon)
41 THA 42 BOR JAW LSI MLY MOL SUM
Synonyms: hidden
= Aspidium intermedium Blume; Enum. Pl. Javae 2: 161 (1828) (non Willd. (1810))
= Aspidium setosum Blume; (non Sw. (1801) nec Schkuhr (1809))
= Aspidium vile Kunze; Bot. Zeitung (Berlin) 4: 474 (1846)
= Dryopteris intermedia (Keyserl.) Kuntze; Revis. Gen. Pl. 2: 813 (1891), ex Alderw., Bull. Jard. Bot. Buitenzorg, sér. 3, 2: 144 (1920)
= Dryopteris millettii C.Chr.; Index Filic. 278 (1905)
= Dryopteris padangensis (Bedd.) C.Chr.; Index Filic. 282 (1905)
= Dryopteris sarawakensis (Baker) Alderw.; Malayan Ferns 200 (1909)
= Dryopteris setosa (Blume) Kuntze; Revis. Gen. Pl. 2: 813 (1891)
= Dryopteris vilis (Kunze) Kuntze; Revis. Gen. Pl. 2: 814 (1891)
= Lastrea blumei T.Moore; Index Fil. [T. Moore] 94 (1858) (non Nephrodium blumei J. Sm. (1841))
= Lastrea padangensis Bedd.; Suppl. Ferns Brit. Ind. 60 (1892)
= Lastrea setosa (Blume) J.Sm.; Ferns Brit. Foreign, ed. 2: 306 (1877)
= Lastrea vilis (Kunze) T.Moore; Index Fil. [T. Moore] 107 (1858)
= Nephrodium sarawakense Baker; J. Linn. Soc., Bot. 22: 225 (1886)
= Nephrodium vile (Kunze) Baker; Syn. Fil. [W. J. Hooker & Baker], ed. 2: 499 (1874)
= Polypodium asplenioides Bory; Voy. Indes Or. [Bélanger] 2(1): 33 (1833), nom. illeg.
= Polystichum intermedium Keyserl.; Polyp. Herb. Bunge 45 (1873)
species :
Ctenitis warburii (C.Chr.) Tardieu; Amer. Fern J. 48: 32 (1958)
Distribution: Madagascar
29 MDG
Synonyms: hidden
= Dryopteris warburii C.Chr.; Cat. Pl. Madag., Pterid. 29 (1932) (nomen), Dansk Bot. Ark. 7: 58, t. 14 (1932)
= Pseudotectaria warburii (C.Chr.) Holttum; Kew Bull. 45(2): 262 (1990)
= Tectaria warburii (C.Chr.) J.P.Roux; Strelitzia 23: 196 (2009)
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A complete, synonymic checklist of the Higher Plants of the World


Version 24.11 (November 5th, 2024)

Covering 355,576 accepted species,
8,715 hybrids, 47,863 accepted infraspecific taxa and 1,000,581 synonyms,
in total 1,412,785* names

[*about 25,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus" are deliberately omitted]


With complete, detailed distribution data (countries and regions) and literature references

Arranged in linear phylogenetic sequence reflecting true relationships

Fully searchable and continously updated

With 38,209 pictures of 15,787 species

Assembled through 40 years independently from other databases

In cooperation with Catalogue of Life