450.0000
Family
Asteraceae Dumort.; Comment. Bot. 55 (1822)
1620 genera, ca. 29.000 spp. (35.000 spp. incl. microspecies). Systematics taken mostly from Funk et al. (2009, 2020) and other recent publications. Barnadesioideae might be its own family, related to Calyceraceae.
Species listed in database: 35141; hybrids: 686
= Acarnaceae Link; (1829)
= Ambrosiaceae Bercht. & J.Presl; Prir. Rostlin: 254 (1820), nom. cons.
= Anthemidaceae Bercht. & J.Presl; Prir. Rostlin: 254 (1820)
= Aposeridaceae Raf.; New Fl. N. Amer. 4: 106 (1838)
= Arctotaceae Bessey; Nebraska Univ. Stud. 14: 107 (1914)
= Arctotidaceae Bercht. & J.Presl; Prir. Rostlin: 255 (1820)
= Artemisiaceae Martinov; Tekhno-Bot. Slovar: 48 (1820)
= Athanasiaceae Martinov; Tekhno-Bot. Slovar: 56 (1820)
= Calendulaceae Bercht. & J.Presl; Prir. Rostlin: 254 (1820)
= Carduaceae Bercht. & J.Presl; Prir. Rostlin: 255 (1820)
= Carlinaceae Bercht. & J.Presl; Prir. Rostlin: 255 (1820)
= Centaureaceae Bercht. & J.Presl; Prir. Rostlin: 255 (1820)
= Cichoriaceae Juss.; Gen. Pl.: 168 (1789), nom. cons.
= Cnicaceae Vest; Anleit. Stud. Bot.: 273, 297 (1818)
= Compositae Giseke; Prael. Ord. Nat. Pl.: 538 (1792), nom. cons.
= Coreopsidaceae Link; Handbuch 1: 768 (1829)
= Cynaraceae Burnett; Outlines Bot.: 926, 931, 1094, 1111 (1835)
= Echinopaceae Bercht. & J.Presl; Prir. Rostlin: 255 (1820)
= Eupatoriaceae Bercht. & J.Presl; Prir. Rostlin: 253 (1820)
= Gnaphaliaceae Link ex F.Rudolphi; Syst. Orb. Veg.: 46 (1830)
= Grindeliaceae Rchb. ex A.Heller; Muhlenbergia 2: 330 (1907)
= Heleniaceae Raf.; Cincinnati Lit. Gaz. 2: 28 (1824)
= Helianthaceae Bercht. & J.Presl; Prir. Rostlin: 254 (1820)
= Helichrysaceae Link; Handbuch 1: 712 (1829)
= Inulaceae Bercht. & J.Presl; Prir. Rostlin: 254 (1820)
= Ivaceae Raf.; Fl. Tellur. 1: 31 (1837)
= Lactucaceae Drude; J. A. Schenk, Handb. Bot. 3(2): 369 (1886)
= Lapsanaceae Martinov; (1820)
= Madiaceae A.Heller; Muhlenbergia 2: 332 (1907)
= Matricariaceae Voigt; Hort. Suburb. Calcutt.: 400 (1845)
= Mutisiaceae Burnett; Outlines Bot.: 934, 935, 1094, 1111 (1835)
= Nassauviaceae Burmeist.; Handb. Naturgesch.: 290 (1836)
= Partheniaceae Link; Handbuch 1: 816 (1829)
= Perdiciaceae Link; Handbuch 1: 728 (1829)
= Picridaceae Martinov; Tekhno-Bot. Slovar: 482 (1820)
= Santolinaceae Martinov; Tekhno-Bot. Slovar: 560 (1820)
= Senecionaceae Bercht. & J.Presl; Prir. Rostlin: 254 (1820)
= Serratulaceae Martinov; Tekhno-Bot. Slovar: 577 (1820)
= Tanacetaceae Vest; Anleit. Stud. Bot.: 273, 298 (1818)
= Tussilagaceae Bercht. & J.Presl; Prir. Rostlin: 254 (1820)
= Vernoniaceae Burmeist.; Handb. Naturgesch.: 296 (1836)
= Xanthiaceae Vest; Anleit. Stud. Bot.: 273, 298 (1818)
= Xeranthemaceae Döll; Rhein. Fl.: 498 (1843)
450.3201
Genus
Cousinia Cass.; Diot. Sci. Nat. 47: 503 (1827)
Distribution: mostly Iran (200), Afghanistan (154), C-Asia, the rest from the E-Mediterranean to Mongolia, China (only 11) and W-Himalayas, north to Caucasus and Middle Asia. The borders between Cousinia and Arctium have been redefined by Lopez-Vinyallonga et al. (2011).
Species listed in database: 675; hybrids: 12
= Auchera DC.; Coll. Mem. 9: t. 11, Prod. 6: 557 (1837)
= Lipskyella Juz.; Acta Inst. Bot. Acad. Sci. U.R.S.S., ser. I. Fasc. 3, 323 (1937)
= Tiarocarpus Rech.f.; Anz. Österr. Akad. Wiss., Mat.-Naturwiss., 108: 5 (1971 publ. (1972)
Search for: Cousinia komarowii
Nothing found!
species :
Cousinia abbreviata
Tscherneva; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Uzb. SSR 17: 87 (1962)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia acanthoblephara
Rech.f. & Gilli; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 70 (1955)
Distribution: Afghanistan (Badakshan, Kabul, Wardak, Parwan)
34 AFG
species :
Cousinia acanthodendron
Rech.f. & Podlech; Pl. Syst. Evol., 142(1-2): 2 (1983)
Distribution: Afghanistan (Samangan)
34 AFG
species :
Cousinia acanthostephana
Rech.f.; Fl. Iranica 90: 186 (1972)
Distribution: Afghanistan (Ghazni)
34 AFG
species :
Cousinia acrodroma
Tscherneva; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Uzb. SSR 17: 89 (1962)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia actinacantha
Rech.f.; Fl. Iranica 90: 302 (1972)
Distribution: Afghanistan (Faryab)
34 AFG
species :
Cousinia actinia
Boiss.; Diagn. Ser. II. 3: 55 (1856)
subspecies :
subsp. actinia
Distribution: Afghanistan (Baghlan, Kabul)
34 AFG
Synonyms: hidden
= Arctium actinia (Boiss.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
subspecies :
subsp. brachyphylla
Rech.f.; Fl. Iranica 90: 40 (1972)
Distribution: Afghanistan (Kabul)
34 AFG
species :
Cousinia adenophora
Juz.; Acta Inst. Bot. Acad. Sci. U.R.S.S., ser. I. Fasc. 3, 320 (1937)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia adenosticta
Bornm.; Bull. Herb. Boiss., ser. 2, 7: 212, t. 3, f. 2 (1907)
Distribution: Iran (EC-Iran)
34 IRN
species :
Cousinia adnata
Bunge; Mem. Acad. Sci. Pétersb. 7 sir., 9 2: 50 (1865)
Distribution: Iran (Iranian Aserbaijan, W-Iran)
34 IRN
Synonyms: hidden
= Arctium adnatum (Bunge) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia affinis
Schrenk; Enum. Pl. Nov. 1: 41 (1841)
Distribution: Kazakhstan; Uzbekistan; China (Xinjiang); Mongolia
32 KAZ UZB 36 CHI 37 MON
Synonyms: hidden
= Arctium affine (Schrenk) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Cousinia wolgensis var. affinis (Schrenk) Regel; Trudy Imp. S.-Peterburgsk. Bot. Sada 6: 317 (1879)
species :
Cousinia afghanica
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 12: 273 (1893)
Distribution: Iran (EC-Iran, E-Iran, NE-Iran: Mts.); Afghanistan (Farah, Herat, Kandahar)
34 AFG IRN
Synonyms: hidden
= Cousinia cynaroides C.Winkl. ex Aitch. & Hemsl.; Trans. Linn. Soc. Ser. II. 3: 79 (1886)
species :
Cousinia agelocephala
Tscherneva; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Uzb. SSR 17: 89 (1962)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia aggregata
DC.; Prodr. [A. P. de Candolle] 6: 553 (1838)
Distribution: Iran (EC-Iran, N-Iran, W-Iran)
34 IRN
Synonyms: hidden
= Cousinia aggregata var. glabra Assadi; Fl. Iran [ed. M. Assadi et al.] 125: 135 (2017)
= Cousinia pauciflora Bunge; Mém. Acad. Imp. Sci. Saint Pétersbourg, Sér. 7, 9(2) [Gatt. Cousinia]: 12 (1865)
species :
Cousinia agridaghensis
Tugay, Ertugrul & Ulukus; Phytotaxa 427(4): 260 (2019)
Distribution: Turkey
34 TUR
species :
Cousinia aintabensis
Boiss. & Hausskn.; Boiss., Fl. Orient. 3: 503 (1875)
Distribution: Turkey (S-Anatolia, SE-Anatolia); Iraq (NE-Iraq, NW-Iraq); Syria (C-Syrian Desert)
34 IRQ SYR TUR
Synonyms: hidden
= Arctium aintabense (Boiss. & Hausskn.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Cousinia arbelensis C.Winkl. & Bornm.; Bull. Herb. Boiss. 3 11: 567, t. 14 (1895)
= Cousinia arbelensis var. pinnata C.Winkl. & Bornm.; Bull. Herb. Boissier 3: 568 (1895)
= Cousinia birecikensis Hub.-Mor.; Notes Roy. Bot. Gard. Edinburgh 32: 53 (1972)
= Cousinia handelii Bornm.; Oesterr. Bot. Z. 62: 187 (1912)
= Cousinia sinjarensis Rech.f.; Publ. Cairo Univ. Herb.: 289 (1977)
species :
Cousinia aitchisonii
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 12: 275 (1893)
Distribution: Turkmenistan; Iran (NE-Iran: Mts.); Afghanistan (Herat)
32 TKM 34 AFG IRN
Synonyms: hidden
= Cousinia lasiosiphon Juz.; Acta Inst. Bot. Acad. Sci. U.R.S.S., ser. I. Fasc. 3, 311 (1937)
species :
Cousinia alaica
Juz. ex Tscherneva; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Uzb. SSR 17: 90 (1962)
Distribution: Uzbekistan; Kyrgyzstan; Tajikistan
32 KGZ TZK UZB
species :
Cousinia alata
Schrenk ex Fisch. & C.A.Mey.; Enum. Pl. Nov. 1: 40 (1841)
Distribution: Kazakhstan; Turkmenistan; Uzbekistan; China (Xinjiang)
32 KAZ TKM UZB 36 CHI
Synonyms: hidden
= Arctium alatum (Schrenk) Kuntze; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 197 (1887)
species :
Cousinia albertoregelia
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 91 (1887)
Distribution: Tajikistan
32 TZK
species :
Cousinia albescens
C.Winkl. & Strauss; Trudy Imp. S.-Peterburgsk. Bot. Sada 14: 223 (1896)
Distribution: Iran (SW-Iran)
34 IRN
Synonyms: hidden
= Cousinia albescens var. subsphaerica Bornm.; (1906)
species :
Cousinia albida
DC.; Prodr. [A. P. de Candolle] 6: 552 (1838)
Distribution: Iran
34 IRN
Synonyms: hidden
= Arctium albidum (DC.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia albiflora
(Bornm. & Sint.) Bornm.; Beih. Bot. Centralbl. 34 2: 285 (1916)
Distribution: Turkmenistan
32 TKM
Synonyms: hidden
= Cousinia hypopolia var. albiflora Bornm. & Sint.; (1907)
species :
Cousinia alepidea
Boiss.; Diagn. Ser. II. 3: 53 (1856)
Distribution: Afghanistan (Ghazni, Kabul, Wardak, Paktia / Khost, Zabul); Pakistan (Baluchistan, Waziristan)
34 AFG 40 PAK
Synonyms: hidden
= Arctium alepidea (Boiss.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia aleppica
Boiss.; Diagn. Pl. Orient., ser. 1, 10: 101 (1849)
Distribution: Turkey (S-Anatolia); Lebanon (C-Lebanon); Syria (C-Syrian Desert, NW-Syria)
34 LEB SYR TUR
species :
Cousinia alexeenkoana
Bornm.; Oesterr. Bot. Z. 62: 107, t. 2, f. 2 (1912)
Distribution: Iran (EC-Iran, W-Iran)
34 IRN
variant :
var. nekarmanica
(Rech.f.) Parsa; Flore de l´Iran (Publ. Univ. Teheran 1777) 10: 101 (1980)
Distribution: Iran (EC-Iran)
34 IRN
Synonyms: hidden
= Cousinia nekarmanica Rech.f.; Ann. Naturhist. Mus. Wien 57: 78 (1950)
species :
Cousinia alfredii
Bornm. & Gauba; Repert. Spec. Nov. Regni Veg. 49: 264 (1940)
Distribution: Iran (EC-Iran, N-Iran)
34 IRN
Synonyms: hidden
= Cousinia kelardashtiana Parsa; (1948)
species :
Cousinia algurdina
Rech.f.; Anz. Österr. Akad. Wiss. Math.-Naturwiss. Kl. 101: 344 (1964)
Distribution: Iraq (NE-Iraq)
34 IRQ
Synonyms: hidden
= Cousinia cynaroides var. arlgirdensis Blakelock; (1949)
species :
Cousinia aligudarzensis
Attar & Ghahr.; Novon 17(2): 145 (2007)
Distribution: Iran
34 IRN
species :
Cousinia allolepis
Tscherneva & Vved.; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Uzb. SSR 17: 91 (1962)
Distribution: Uzbekistan
32 UZB
species :
Cousinia alpina
Bunge; Del. Sem. Hort. Dorp. (1845) 3
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
Synonyms: hidden
= Arctium aciculare (Franch.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Arctium alpicola (Bunge) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Arctium alpinum (Bunge) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Cousinia acicularis Franch.; Ann. Sci. Nat., Bot., sér. 6, 16: 319 (1883)
= Cousinia alpicola I.G.Borshch. ex C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 12: 282 (1892)
= Cousinia alpicola I.G.Borshch. ex Trautv.; Trudy Imp. S.-Peterburgsk. Bot. Sada 8: 484 (1883)
species :
Cousinia ambigens
Juz.; Trans. Rubb. & Guttap. Inst., Fasc. 4, 17 (1932), et Acta Inst. Bot. Acad. Sci. U.R.S.S., ser. I. Fasc. 3, 306, 307 (1937)
Distribution: Uzbekistan; Kyrgyzstan; Tajikistan
32 KGZ TZK UZB
species :
Cousinia amicorum
Tscherneva, Joharchi & F.Ghahrem.; Bot. Zhurn. (Moscow & Leningrad) 90(3): 411 (2005)
Distribution: Iran
34 IRN
species :
Cousinia ammophila
Rech.f.; Fl. Iranica 90: 67 (1972)
Distribution: Afghanistan (Ghazni)
34 AFG
species :
Cousinia amoena
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 13: 237 (1894)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia andarabensis
Rech.f.; Anz. Österr. Akad. Wiss., Math.-Nat., ciii. 284 (1966)
Distribution: Afghanistan (Baghlan)
34 AFG
species :
Cousinia angrenii
Juz.; Trans. Rubb. & Guttap. Inst., Fasc. 4, 17 (1932)
Distribution: Uzbekistan; Kyrgyzstan; Tajikistan
32 KGZ TZK UZB
species :
Cousinia angusticeps
Juz.; Acta Inst. Bot. Acad. Sci. U.R.S.S., ser. I. Fasc. 3, 323 (1937)
Distribution: Turkmenistan
32 TKM
species :
Cousinia annua
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 88 (1887)
Distribution: Turkmenistan; Uzbekistan
32 TKM UZB
Synonyms: hidden
= Cousinia dichacantha Lipsky; Exc. bot. au de la Mer Casp. 12 (1889)
= Lipskyella annua (C.Winkl.) Juz.; Acta Inst. Bot. Acad. Sci. U.R.S.S., ser. I. Fasc. 3, 324 (1937)
species :
Cousinia anoplophylla
Rech.f.; Fl. Iranica 139a: 140, t. 134 (1979)
Distribution: Iran (SW-Iran)
34 IRN
species :
Cousinia antonowi
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 11: 135, t. 2, f. 2 (1889)
Distribution: Turkmenistan; Iran (E-Iran)
32 TKM 34 IRN
Synonyms: hidden
= Arctium antonowi (C.Winkl.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia apiculata
Tscherneva; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Uzb. SSR 17: 91 (1962)
Distribution: Turkmenistan; Uzbekistan
32 TKM UZB
species :
Cousinia aptera
Aitch. & Hemsl.; J. Linn. Soc., Bot. 19: 170 (1882)
Distribution: Afghanistan (Bamyan, Ghazni, Ghorat, Kabul, Kandahar, Logar, Wardak, Orozgan / Daykundi, Paktia / Khost, Parwan); Pakistan (Kurram)
34 AFG 40 PAK
Synonyms: hidden
= Cousinia modesta Rech.f. & Köie; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 120 (1955)
species :
Cousinia arachnoidea
Fisch. & C.A.Mey. ex DC.; Prodr. [A. P. de Candolle] 6: 553 (1838)
Distribution: Kazakhstan; Kyrgyzstan
32 KAZ KGZ
Synonyms: hidden
= Carduus pterocaulos Stephan ex DC.; Prodr. [A. P. de Candolle] 6: 553 (1838)
= Carduus pycnocephalus-alatus Steven ex DC.; Prodr. [A. P. de Candolle] 6: 553 (1838)
species :
Cousinia arakensis
Attar & Djavadi; Iranian J. Bot. 16(1): 166 (2010)
Distribution: Iran
34 IRN
species :
Cousinia araneosa
DC.; Prodr. [A. P. de Candolle] 6: 556 (1838)
Distribution: Iran (SW-Iran)
34 IRN
Synonyms: hidden
= Arctium araneosum (DC.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia araneotexta
Rech.f.; Fl. Iranica 90: 105 (1972)
Distribution: Afghanistan (Ghorat)
34 AFG
species :
Cousinia araxena
Takht.; Trans. Armen. Branch Acad. Sci. USSR, Biol. Ser. 2: 184 (1937)
Distribution: Armenia; Azerbaijan (Nachichevan)
33 ARM AZE
species :
Cousinia archibaldii
Rech.f.; Fl. Iranica 90: 147, t. 73 (1972)
Distribution: Iran (SW-Iran)
34 IRN
Synonyms: hidden
= Cousinia albescens var. adpressa Bornm.; (1914)
species :
Cousinia arctotidifolia
Bunge; Mém. Acad. Sci. Petersb. 7, ser., 9 2: 42 (1865)
Distribution: Turkmenistan; Iran (NE-Iran: Mts., N-Iran)
32 TKM 34 IRN
Synonyms: hidden
= Arctium arctotidifolium (Bunge) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Cousinia arctotidifolia var. gracilescens Bornm. & Rchb.f.; (1944)
= Cousinia arctotidifolia var. taraxacifolia Bornm. & Gauba; (1944)
= Cousinia praetermissa Bornm.; (1941)
species :
Cousinia ardalensis
Attar & Djavadi; Iranian J. Bot. 16(1): 147 (2010)
Distribution: Iran
34 IRN
species :
Cousinia arenaria
Bunge; Mém. Acad. Sci. Pétersb. 7, sér., 9 2: 29 (1865)
Distribution: Iran (E-Iran)
34 IRN
Synonyms: hidden
= Arctium arenarium (Bunge) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia argentea
Mehregan & Assadi; Willdenowia 39(2): 268 (2010), nom. nov.
Distribution: Iran
34 IRN
Synonyms: hidden
= Jurinea pungens Boiss.; Fl. Orient. 3: 569 (1875) emend. Rech.f., Repert. Spec. Nov. Regni Veg. 48: 148, t. 306, f. 1 & 2 (1940)
species :
Cousinia ariana
Bornm.; Beih. Bot. Centralbl. 34: 11. 278 (1916)
Distribution: Afghanistan (Baghlan, Bamyan, Wardak, Parwan); Pakistan (Swat, Quetta)
34 AFG 40 PAK
species :
Cousinia arida
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 12: 190 (1893)
Distribution: Iran (E-Iran); Afghanistan (Herat)
34 AFG IRN
Synonyms: hidden
= Cousinia deserti Aitch. & Hemsl.; Trans. Linn. Soc. Ser. II. 3: (1886) 77 (non Bunge)
species :
Cousinia armena
Takht.; Trans. Armen. Branch Acad. Sci. USSR, Biol. Ser. 2: 182 (1937)
Distribution: Armenia
33 ARM
species :
Cousinia aspera
(Kult.) Karmysch.; Fl. Kazakhst. 9: 243 (1966)
Distribution: Kazakhstan
32 KAZ
Synonyms: hidden
= Cousinia chrysantha var. aspera Kult.; Trudy Sredne-Aziatsk. Gosud. Univ., ser. 8b, Bot. 6: 25 (1929)
species :
Cousinia assadii
Attar; Iranian J. Bot. 17(2): 153 (2011)
Distribution: Iran
34 IRN
species :
Cousinia assyriaca
Jaub. & Spach; Ill. Pl. Or. 2: 87, t. 169 (1846)
Distribution: Iran (EC-Iran, W-Iran)
34 IRN
Synonyms: hidden
= Arctium assyriacum (Jaub. & Spach) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Cousinia leucochlora Bunge; Mém. Acad. Sci. Pétersb. 7 sér., 9 2: 24 (1865)
= Cousinia turkmenorum var. flaviflora Parsa; (1948)
species :
Cousinia astracanica
(Spreng.) Tamamsch.; Not. Syst. (Leningrad) 16: 468 (1954)
Distribution: S-European Russia; Kazakhstan; Turkmenistan
14 RUS 32 KAZ TKM
Synonyms: hidden
= Arctium wolgense (M.Bieb. ex Willd.) Kuntze; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 198 (1887)
= Carduus astracanicus Biehler; Pl. Nov. Herb. Spreng.: 34 (1807)
= Carduus wolgensis M.Bieb. ex Willd.; Enum. Pl. 2: 839 (1809)
= Cousinia wolgensis (M.Bieb. ex Willd.) C.A.Mey. ex DC.; Prodr. [A. P. de Candolle] 6: 555 (1838)
species :
Cousinia atrobracteata
Attar; Iranian J. Bot. 17(2): 152 (2011)
Distribution: Iran
34 IRN
species :
Cousinia atropatana
Bunge; Mém. Acad. Sci. Pétersb. 7 sér., 9 2: 17 (1865)
Distribution: Iran (Iranian Aserbaijan)
34 IRN
Synonyms: hidden
= Arctium atropatanum (Bunge) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia attariae
Assadi & Joharchi; Iranian J. Bot. 16(2): 192 (194-195, fig. 2) (2010)
Distribution: Iran
34 IRN
species :
Cousinia aucheri
DC.; Prodr. [A. P. de Candolle] 6: 555 (1838)
Distribution: Turkey (E-Anatolia, SE-Anatolia)
34 TUR
Synonyms: hidden
= Arctium aucheri (DC.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Cousinia montbretiana Bornm.; Repert. Spec. Nov. Regni Veg. 46: 275 (1939)
species :
Cousinia auriculata
Boiss.; Diagn. Ser. II. 3: 58 (1856)
Distribution: Tajikistan (Pamir-Alai); Afghanistan (Wakhan, Bamyan, Ghorat, Kabul, Wardak); Pakistan (Waziristan, Kurram); NW-India (Jammu & Kashmir)
32 TZK 34 AFG 40 IND PAK
Synonyms: hidden
= Arctium auriculatum (Boiss.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Cousinia kotandarica Rech.f. & Gilli; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 112 (1955)
= Cousinia pamirica Bornm.; Beih. Bot. Centralbl. 34(2): 182 (1916) ex O. Fedtsch., Trudy Imp. S.-Peterburgsk. Bot. Sada 31: 463 (1915)
= Cousinia semidecurrens C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 574 (1837)
species :
Cousinia austrojordanica
F.G.Davies & Boulos; Publ. Cairo Univ. Herb. 7 & 8: 287 (1977)
Distribution: Jordania (S-Jordania)
34 JOR
species :
Cousinia autranii
C, Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 12: 193 (1892)
Distribution: Afghanistan (Ghazni, Ghorat, Kabul, Wardak, Zabul)
34 AFG
Synonyms: hidden
= Cousinia ghazniensis Rech.f. & Gilli; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 97 (1955)
species :
Cousinia azerbaidjanica
Djavadi, Attar & Najafi; Iranian J. Bot. 13(1): 44 (2007)
Distribution: Iran
34 IRN
species :
Cousinia azmarensis
S.A.Ahmad, Rastegar & Attar; Harvard Pap. Bot. 22(1): 71 (2017)
Distribution: Iraq
34 IRQ
species :
Cousinia baberi
Boiss.; Diagn. Ser. II. 3: 53 (1856)
Distribution: Afghanistan (Kabul, Logar, Wardak)
34 AFG
Synonyms: hidden
= Arctium baberi (Boiss.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia bachtiarica
Boiss. & Hausskn.; Boiss., Fl. Orient. 3: 475 (1875)
Distribution: Iran (SW-Iran)
34 IRN
Synonyms: hidden
= Arctium bachtiaricum (Boiss. & Hausskn.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia badghysi
Kult.; Korovin, Kult. & Popov, Descr. Pl. Nov. Turkmenistan. (Bodenk. u. Pflanzengeogr. Beob. u. Unters. 1: Flussgeb. d. Amu Darya u. Syr Darya, II.), 76 (1916)
Distribution: Turkmenistan
32 TKM
species :
Cousinia balchanica
Tscherneva; Novosti Sist. Vyssh. Rast. 30: 153 (1996)
Distribution: Kazakhstan
32 KAZ
species :
Cousinia baluchistanica
Rech.f.; Anz. Österr. Akad. Wiss., Mat.-Naturwiss., 125: 53 (1988 publ. 1989)
Distribution: Pakistan (Baluchistan)
40 PAK
species :
Cousinia bamianica
Rech.f. & Köie; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 77 (1955)
Distribution: Afghanistan (Bamyan, Wardak)
34 AFG
Synonyms: hidden
= Cousinia actinostegia Rech.f.; Anz. Österr. Akad. Wiss., Math.-Nat., ciii. 282 (1966)
species :
Cousinia baranovii
Tscherneva; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Uzb. SSR. 17: 92 (1962)
Distribution: Kazakhstan; Kyrgyzstan
32 KAZ KGZ
species :
Cousinia barbeyi
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 12: 274 (1893)
Distribution: Iran (SW-Iran)
34 IRN
species :
Cousinia barezica
Assadi; Iranian J. Bot. 15(1): 37 (2009)
Distribution: Iran
34 IRN
species :
Cousinia batalinii
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 12: 248 (1892)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia bazoftensis
Attar; Iranian J. Bot. 17(2): 154 (2011)
Distribution: Iran
34 IRN
species :
Cousinia beauverdiana
Bornm.; Oesterr. Bot. Z. 63: 55 (1913)
Distribution: Iran (SW-Iran)
34 IRN
Synonyms: hidden
= Cousinia arctotidifolia var. laeviseta Winkl. & Bornm.; (1897)
species :
Cousinia beckeri
Trautv.; Trudy Imp. S.-Peterburgsk. Bot. Sada 9: 457 (1886)
Distribution: Turkmenistan
32 TKM
species :
Cousinia belangeri
DC.; Prodr. [A. P. de Candolle] 6: 553 (1838)
Distribution: Iran (EC-Iran, Iranian Aserbaijan, W-Iran)
34 IRN
Synonyms: hidden
= Arctium belangeri (DC.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Arctium verutum (Bunge) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Cousinia carlinoides DC.; Prodr. [A. P. de Candolle] 6: 554 (1838)
= Cousinia pugionifera var. decurrens Parsa; (1948)
= Cousinia teheranica Bornm.; Beih. Bot. Centralbl. 34(2): 274 (1916)
= Cousinia verutum Bunge; Mém. Acad. Sci. Pétersb. 7 sér., 9 2: 31 (1865)
= Cousinia verutum subsp. teheranica Bornm.; (1917)
species :
Cousinia bicolor
Freyn & Sint.; Oesterr. Bot. Z. 42 6: 204 (1892)
Distribution: Turkey (E-Anatolia, NE-Anatolia)
34 TUR
species :
Cousinia bienerti
Bunge; Mém. Acad. Sci. Pétersb. 7 sér., 9 2: 27 (1865)
Distribution: Iran (NE-Iran: Mts.)
34 IRN
Synonyms: hidden
= Arctium bienerti (Bunge) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia bijarensis
Rech.f.; Fl. Iranica 90: 152 (1972)
Distribution: Iran (SW-Iran)
34 IRN
species :
Cousinia bipinnata
Boiss.; Diagn. Ser. II. 3: 52 (1856)
Distribution: Afghanistan (Kandahar); Pakistan (Quetta, Baluchistan)
34 AFG 40 PAK
Synonyms: hidden
= Arctium bipinnatum (Boiss.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia birandiana
Hub.-Mor.; Notes Roy. Bot. Gard. Edinburgh 32: 49 (1972)
Distribution: Turkey (Inner Anatolia, C-Anatolia)
34 TUR
species :
Cousinia blepharobasis
Rech.f. & Gilli; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 78 (1955)
Distribution: Afghanistan (Bamyan, Kabul, Wardak)
34 AFG
species :
Cousinia bobekii
Rech.f.; Fl. Iranica 90: 238, t. 120 (1972)
Distribution: Iran (Iranian Aserbaijan)
34 IRN
species :
Cousinia bobrovii
Juz.; Acta Inst. Bot. Acad. Sci. U.R.S.S., ser. I. Fasc. 3, 308 (1937)
Distribution: Turkmenistan; Uzbekistan
32 TKM UZB
species :
Cousinia boissieri
Buhse; Boiss. & Buhse, Nouv. Mém. Soc. Nat. Moscou 12 18: 127 (1860)
Distribution: Turkey (E-Anatolia, SE-Anatolia); Iraq (SE-Iraq: Mesopotamia); Iran (Iranian Aserbaijan, W-Iran)
34 IRN IRQ TUR
Synonyms: hidden
= Arctium boissieri (Buhse) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Cousinia layardii Ball & Barbey; Cousinia layardi 2 (1890)
species :
Cousinia bonvalotii
Franch.; Ann. Sci. Nat. Ser. VI. 16: 320 (1883)
Distribution: Kazakhstan; Uzbekistan; Kyrgyzstan
32 KAZ KGZ UZB
Synonyms: hidden
= Arctium bonvalotii (Franch.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia bornmulleriana
Attar & Maroofi; Nova Biol. Reperta 9(2): 145 (2022), non C. bornmuelleri C. Winkl.
Distribution: Iran
34 IRN
species :
Cousinia botschantzevii
Juz. ex Tscherneva; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Uzb. SSR 17: 93 (1962)
Distribution: Uzbekistan
32 UZB
species :
Cousinia boyerahmadica
Rastegar, Attar & Mirtadz.; Phytotaxa 343(3): 228 (2018)
Distribution: Iran
34 IRN
species :
Cousinia brachyptera
DC.; Prodr. [A. P. de Candolle] 6: 554 (1838)
Distribution: Georgia [Caucasus]; Armenia; Turkey (E-Anatolia, Inner Anatolia, NE-Anatolia)
33 ARM GEO 34 TUR
Synonyms: hidden
= Arctium brachypterum (DC.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Cousinia gundelia K.Koch; Linnaea 24: 387 (1851)
= Cousinia heterolepis K.Koch; Linnaea 24: 386 (1851)
= Cousinia squarrosa K.Koch; Linnaea 24: 387 (1851)
species :
Cousinia brevicaulis
Attar, Mozaff. & Mirtadz.; Nordic J. Bot. 34(2): 176 (2016)
Distribution: Iran
34 IRN
species :
Cousinia bucharica
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: (1887) 473 et 477
Distribution: Uzbekistan (Buchara)
32 UZB
Synonyms: hidden
= Arctium bucharicum (C.Winkl.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia bungeana
Regel & Schmalh.; Trudy Imp. S.-Peterburgsk. Bot. Sada 6: 318 (1879)
Distribution: Kazakhstan; Uzbekistan; Tajikistan
32 KAZ TZK UZB
Synonyms: hidden
= Arctium bungeanum (Regel & Schmalh.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia buphthalmoides
Regel; Bull. Soc. Imp. Naturalistes Moscou 40: (1867) II. 155
Distribution: Tajikistan; Afghanistan (Badakshan, Wakhan, Baghlan, Bamyan, Ghorat, Kabul, Laghman, Logar, Wardak, Paktia / Khost, Parwan, Samangan); Pakistan (Kurram, Chitral, Drosh, Swat, Gilgit)
32 TZK 34 AFG 40 PAK
Synonyms: hidden
= Arctium buphthalmoides (Regel) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Cousinia auriculata Hook.f.; Fl. Brit. India [J. D. Hooker] 3: 360 (1881), nom. illeg.
species :
Cousinia butkovii
Tschern. & Vved.; Not. Syst. Herb. Inst. Bot. Acad. Sci. . 16: 54 (1961)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia caesarea
Boiss. & Balansa; Boiss., Diagn. Pl. Orient., ser. 2, 6: 116 (1859)
Distribution: Turkey (Central Anatolia, Inner Anatolia)
34 TUR
Synonyms: hidden
= Arctium caesareum (Boiss. & Balansa) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia caespitosa
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 93 (1887)
Distribution: Kazakhstan; Kyrgyzstan; China (W-Xinjiang)
32 KAZ KGZ 36 CHI
species :
Cousinia calcitrapa
Boiss.; Fl. Orient. 3: 490 (1875)
Distribution: Iran (Iranian Aserbaijan, S-Iran, W-Iran)
34 IRN
Synonyms: hidden
= Arctium calcitrapa (Boiss.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Cousinia larvea C.Winkl. & Bornm.; Bull. Herb. Boissier 5: 167 (1897)
variant :
var. interrupta
Heimerl; Denkschr. Kaiserl. Akad. Wiss., Wien. Math.-Naturwiss. Kl. 50: 62 (1885)
Distribution: Iran
34 IRN
Synonyms: hidden
= Cousinia xiphacantha C.Winkl. & Strauss ex C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 14: 225 (1897)
= Cousinia xiphacantha var. lachnopoda Bornm.; (1912)
species :
Cousinia calcitrapella
Bornm.; Notizbl. Bot. Gart. Berlin-Dahlem 15: 118 (1940)
Distribution: India (Punjab)
40 IND
species :
Cousinia calocephala
Jaub. & Spach; Ill. Pl. Or. 2: 96 (1846)
subspecies :
subsp. calocephala
Distribution: Turkey (E-Anatolia, SE-Anatolia); Iran (EC-Iran, N-Iran, Iranian Aserbaijan, W-Iran)
34 IRN TUR
Synonyms: hidden
= Arctium calocephalum (Jaub. & Spach) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Arctium squarrosum (Boiss.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Cousinia squarrosa Boiss.; Diagn. Pl. Orient., ser. 1, 10: 102 (1849)
= Cousinia vanensis Hub.-Mor.; Notes Roy. Bot. Gard. Edinburgh 32: 55 (1972).
subspecies :
subsp. asterocephala
(Hausskn. & Bornm.) Mehregan; Willdenowia 38(2): 348 (2008)
Distribution: Iran
34 IRN
Synonyms: hidden
= Cousinia asterocephala Hausskn. & Bornm.; Beih. Bot. Centralbl. 20(2): 160 (1906)
subspecies :
subsp. behboudiana
(Rech.f. & Esfand.) Mehregan; Willdenowia 38(2): 347 (2008)
Distribution: Iran (EC-Iran)
34 IRN
Synonyms: hidden
= Cousinia behboudiana Rech.f. & Esfand.; Ann. Naturhist. Mus. Wien 57: 81 (1950)
= Cousinia squarrosa var. behboudiana (Rech.f. & Esfand.) Parsa; Flore de l´Iran (Publ. Univ. Teheran 1777) 10: 128 (1980)
species :
Cousinia calolepis
Boiss.; Diagn. Pl. Orient., ser. 1, 6: 116 (1846)
Distribution: Iran (EC-Iran, N-Iran)
34 IRN
Synonyms: hidden
= Arctium anisopterum (Jaub. & Spach) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Cousinia anisoptera Jaub. & Spach; Illustr. 2: 88, t. 170 (1846)
species :
Cousinia calva
Juz.; Acta Inst. Bot. Acad. Sci. U.R.S.S., ser. I. Fasc. 3, 317 (1937)
Distribution: Tajikistan
32 TZK
species :
Cousinia campyloraphis
Tscherneva; Not. Syst. Herb. Inst. Bot. Acad. Sci. . 15: 37 (1959)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia candicans
Juz.; Acta Inst. Bot. Acad. Sci. U.R.S.S., ser. I. Fasc. 3, 300 (1937)
Distribution: Tajikistan
32 TZK
species :
Cousinia candolleana
Jaub. & Spach; Ill. Pl. Or. 2: 74, t. 156 (1845)
Distribution: Iran (SW-Iran)
34 IRN
Synonyms: hidden
= Arctium orientale (DC.) Kuntze; Revis. Gen. Pl. 1: 367 (1891)
= Auchera orientalis DC.; Coll. Mem. 9: t. 11 (1838)
species :
Cousinia canescens
DC.; Prodr. [A. P. de Candolle] 6: 556 (1838)
Distribution: Armenia; Turkey (E-Anatolia, Inner Anatolia, NE-Anatolia, S-Anatolia, N-Anatolia, SE-Anatolia); Iran (Iranian Aserbaijan, W-Iran)
33 ARM 34 IRN TUR
Synonyms: hidden
= Arctium eriocephalum (Boiss. & Hausskn.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Cousinia eriocephala Boiss. & Hausskn.; Boiss., Fl. Orient. 3: 504 (1875)
= Cousinia satdagensis Hub.-Mor.; Notes Roy. Bot. Gard. Edinburgh 32: 55 (1972)
= Cousinia wettsteiniana Bornm.; Verh. K. K. Zool.-Bot. Ges. Wien 60: 137 (1910)
species :
Cousinia carduncelloidea
Regel & Schmalh.; Pl. Nov. Fedsch. 50 (1882)
Distribution: Kyrgyzstan; Tajikistan
32 KGZ TZK
Synonyms: hidden
= Arctium carduncelloideum (Regel & Schmalh.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Cousinia carduncelloidea var. sonchifolia Bornm.
= Cousinia carduncellus Regel & Schmalh. ex O.Fedtsch. & B.Fedtsch.; Consp. Fl. Turkest. 4: 252 (1902)
= Cousinia sonchifolia (Bornm.) Bornm.; Repert. Spec. Nov. Regni Veg., Beih. 108: 42 (1938)
species :
Cousinia caroli-henrici
Attar & Ghahr.; Rostaniha 7(Suppl. 2): 317 (2007)
Distribution: Iran
34 IRN
species :
Cousinia carthamoides
Aitch. & Hemsl.; J. Linn. Soc., Bot. 19: 171 (1882)
Distribution: Afghanistan (Kunar / Nuristan, Paktia / Khost); Pakistan (Kurram)
34 AFG 40 PAK
Synonyms: hidden
= Cousinia carthamoides var. stenorhachis Bornm.; (1938)
species :
Cousinia cataonica
Boiss. & Hausskn.; Boiss., Fl. Orient. 3: 483 (1875)
Distribution: Turkey (Inner Anatolia, SE-Anatolia)
34 TUR
Synonyms: hidden
= Arctium cataonicum (Boiss. & Hausskn.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia catenata
Rech.f.; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 82 (1955)
Distribution: Afghanistan (Balkh, Jawzjan / Sar-e-Pol)
34 AFG
species :
Cousinia cavarae
Bornm.; Mitt. Thür. Bot. Ver. 38: 2, 49 (1929)
Distribution: Iran (NE-Iran: Mts.)
34 IRN
species :
Cousinia centauroides
Fisch. & C.A.Mey.; Bull. Soc. Imp. Naturalistes Moscou (1839) 159 ex Bunge
Distribution: Turkmenistan
32 TKM
Synonyms: hidden
= Arctium centauroides (Fisch. & C.A.Mey.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia ceratophora
Kult.; Acta Univ. As. Med. Ser. VIII. Bot. Fasc. VI. 24 (1929)
Distribution: Tajikistan
32 TZK
species :
Cousinia chaetocephala
Kult.; Acta Univ. As. Med., ser. 8b 7: 3 (1929)
Distribution: Turkmenistan; Iran (NE-Iran: Mts.)
32 TKM 34 IRN
species :
Cousinia chaetolepis
Rech.f.; Fl. Iranica 90: 207 (1972)
Distribution: C-Afghanistan
34 AFG
species :
Cousinia chamaepeuce
Boiss.; Diagn. Pl. Orient., ser. 1, 6: 120 (1846)
Distribution: Iran (EC-Iran, N-Iran)
34 IRN
Synonyms: hidden
= Arctium chamaepeuce (Boiss.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Cousinia chamaepeucides Bornm.; Bull. Herb. Boiss. Ser. 2(7): 212 (1907)
species :
Cousinia chejrabadensis
Kult.; Fl. Turkm. 7: 380 (1960)
Distribution: Turkmenistan
32 TKM
species :
Cousinia chionophila
Rech.f. & Köie; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 84 (1955)
Distribution: Afghanistan (Badakshan, Wakhan, Baghlan, Bamyan Ghazni, Kunar / Nuristan, Parwan); Pakistan (Chitral)
34 AFG 40 PAK
species :
Cousinia chitralensis
Rech.f.; Fl. Iranica 90: 306 (1972)
Distribution: Pakistan (Chitral)
40 PAK
species :
Cousinia chlorocephala
C.A.Mey.; Prodr. [A. P. de Candolle] 6: 554 (1838)
Distribution: Armenia; Azerbaijan (incl. Nachichevan); Iran (N-Iran)
33 ARM AZE 34 IRN
Synonyms: hidden
= Arctium chlorocephalum (C.A.Mey.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Cousinia intermedia C.A.Mey.; Prodr. [A. P. de Candolle] 6: 553 (1838)
species :
Cousinia chlorosphaera
Bornm.; Beih. Bot. Centralbl. 28 2: 253 (1911)
Distribution: Iran (SW-Iran)
34 IRN
Synonyms: hidden
= Cousinia chlorosphaera var. straminea Bornm.; (1914)
= Cousinia chlorosphaera var. producta Bornm.; (1914)
species :
Cousinia chlorothyrsa
Rech.f. & Köie; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 86 (1955)
Distribution: Afghanistan (Bamyan, Ghorat, Orozgan / Daykundi)
34 AFG
species :
Cousinia chrysacantha
Jaub. & Spach; Ill. Pl. Or. 2: 78, t. 160 (1845)
Distribution: Iran (EC-Iran, Iranian Aserbaijan, W-Iran)
34 IRN
Synonyms: hidden
= Arctium chrysacanthum (Jaub. & Spach) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia chrysandra
Bornm. & Gauba; Mitt. Thür. Bot. Ver. 47: 115, t. 11, f. 1 (1941)
Distribution: Iran (NE-Iran: Mts.)
34 IRN
species :
Cousinia chrysantha
Kult.; Acta Univ. As. Med. Ser. VIII. Bot. Fasc. VI. 25 (1929)
Distribution: Kazakhstan; Uzbekistan
32 KAZ UZB
species :
Cousinia chrysochlora
Rech.f. & Köie; Dansk Biol. Skr. 8, 2: 88 (1955)
Distribution: Iran (E-Iran); Afghanistan (Farah, Herat)
34 AFG IRN
species :
Cousinia cirsioides
Boiss. & Balansa; Boiss., Diagn. Pl. Orient., ser. 2, 6: 117 (1859)
Distribution: Turkey (Inner Anatolia)
34 TUR
Synonyms: hidden
= Arctium cirsioides (Boiss. & Balansa) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia cisdarvasica
V.V.Nikitin; Not. Syst. Herb. Inst. Bot. Acad. Sci. U.R.S.S. 19: 385 (1959)
Distribution: Tajikistan
32 TZK
species :
Cousinia coerulea
Kult. ex Tscherneva; Not. Syst. Herb. Inst. Bot. Acad. Sci. . 15: 51 (1959)
Distribution: Tajikistan
32 TZK
Synonyms: hidden
= Cousinia subinermis Tschern.; Bot. Mater. Gerb. Inst. Bot. Zool. Akad. Nauk Uzbeksk. S.S.R. 15: 40 (1959)
species :
Cousinia commutata
Bunge; Mém. Acad. Sci. Pétersb. 7 sér., 9 2: 24 (1865)
Distribution: Iran (EC-Iran, N-Iran)
34 IRN
Synonyms: hidden
= Arctium commutatum (Bunge) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Cousinia campylacme Bornm. & Gauba; Repert. Spec. Nov. Regni Veg. 40: 340 (1936)
= Cousinia dichroantha Bornm. & Gauba; Repert. Spec. Nov. Regni Veg. 40: 344 (1936)
= Cousinia larica Parsa; Kew Bull. 3: 205 (1948)
species :
Cousinia concinna
Boiss. & Hausskn.; Boiss., Fl. Orient. 3: 513 (1875)
Distribution: Iran (SW-Iran)
34 IRN
Synonyms: hidden
= Arctium concinnum (Boiss. & Hausskn.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia concolor
Bunge; Mém. Acad. Sci. Pétersb. 7 sér., 9 2: 33 (1865)
Distribution: Iran (EC-Iran, E-Iran)
34 IRN
Synonyms: hidden
= Arctium concolor (Bunge) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia congesta
Bunge; Mém. Acad. Sci. Pétersb. 7 sér., 9 2: 12 (1865)
Distribution: Turkmenistan; Tajikistan; Iran (EC-Iran, E-Iran, NE-Iran: Mts., N-Iran, W-Iran); Afghanistan (Balkh, Faryab, Ghorat, Herat, Kabul, Qunduz, Samangan); Pakistan (Baluchistan)
32 TKM TZK 34 AFG IRN 40 PAK
Synonyms: hidden
= Arctium congestum (Bunge) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia consanguinea
(Bornm.) Rech.f.; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 89 (1955)
Distribution: Afghanistan (locality unknown)
34 AFG
Synonyms: hidden
= Cousinia scariosa subsp. consanguinea Bornm.; Beih. Bot. Centralbl. 34: 11. 282 (1916)
species :
Cousinia contumax
C.Winkl. & Bornm.; Bull. Herb. Boiss. 5: 168, t. 4 (1897)
Distribution: Iran (EC-Iran, S-Iran)
34 IRN
species :
Cousinia cordifolia
Djavadi & Attar; Feddes Repert. 117(7-8): 453 (2006)
Distribution: Iran
34 IRN
species :
Cousinia coronata
Franch.; Ann. Sci. Nat. Ser. VI. 16: 321 (1883)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
Synonyms: hidden
= Arctium coronatum (Franch.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Cousinia campylacantha Juz.; Trudy Bot. Inst. Akad. Nauk S.S.S.R., ser. 1, Fl. Sist. Vyssh. Rast. 3: 312 (1937)
species :
Cousinia corymbosa
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 89 (1887)
Distribution: Tajikistan
32 TZK
species :
Cousinia crassipes
Kult.; Acta Univ. As. Med., ser. 8b 7: 2 (1929)
Distribution: Iran (NE-Iran: Mts.)
34 IRN
species :
Cousinia crispa
Jaub. & Spach; Ill. Pl. Or. 2: 81 (1846)
Distribution: Iran (EC-Iran, N-Iran)
34 IRN
Synonyms: hidden
= Arctium crispum (Jaub. & Spach) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Cousinia barbarae Gauba; Mitth. Thüring. Bot. Vereins n.f. 47: 116 (1941)
= Cousinia kandewanensis Parsa; Kew Bull. 3(2): 205 (1948)
species :
Cousinia cryptadena
Juz.; Not. Syst. Herb. Hort. Petrop. 5: 111 (1924)
Distribution: Turkmenistan
32 TKM
Synonyms: hidden
= Cousinia leucocitrina Kult.; Bot. Mater. Gerb. Glavn. Bot. Sada S. S. S. R. 6: 30 (1926)
species :
Cousinia curvibracteata
Mehregan; Iranian J. Bot. 17(2): 144 (2011)
Distribution: Iran
34 IRN
species :
Cousinia cylindracea
Boiss.; Diagn. Pl. Orient., ser. 1, 6: 115 (1846)
Distribution: Iran (EC-Iran)
34 IRN
Synonyms: hidden
= Arctium cylindraceum (Boiss.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Cousinia cylindracea var. patula Heimerl; (1885)
species :
Cousinia cylindrocephala
Jaub. & Spach; Ill. Pl. Or. 2: 84, t. 166 (1846)
Distribution: Iran (EC-Iran)
34 IRN
Synonyms: hidden
= Arctium cylindrocephalum (Jaub. & Spach) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia cynaroides
(M.Bieb.) C.A.Mey.; Verz. Pfl. Caucasus 230 (1831)
Distribution: Turkmenistan; Northern Caucasus; Armenia; Azerbaijan; Iran (EC-Iran, Iranian Aserbaijan, W-Iran)
32 TKM 33 ARM AZE NCS 34 IRN
Synonyms: hidden
= Arctium cynaroides (M.Bieb.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Carthamus cynaroides M.Bieb.; Beschr. Länder Terek Kasp. 195 (1800)
= Onobroma cynaroides (M.Bieb.) Spreng.; Syst. Veg. ed. 16, 3: 392 (1826)
= Onopordon cynaroides (M.Bieb.) Steven ex Link; Enum. Hort. Berol. Alt. 2: 303 (1822)
species :
Cousinia czatkalica
Tscherneva; Novosti Sist. Vyssh. Rast. 29: 122 (1993)
Distribution: Kyrgyzstan
32 KGZ
species :
Cousinia czerniakowskae
Kult.; Acta Univ. As. Med., ser. 8b 7: 5 (1929)
Distribution: Iran (NE-Iran: Mts.)
34 IRN
Synonyms: hidden
= Cousinia tscherniakowskiana Kult.; (1930)
variant :
var. adenostegia
(Rech.f.) Assadi; Fl. Iran [ed. M. Assadi et al.] 125: 64 (2017)
Distribution: Iran (NE-Iran: Mts.)
34 IRN
Synonyms: hidden
= Cousinia adenostegia Rech.f.; Fl. Iranica 139a: 151, t. 147 (1979)
species :
Cousinia dalahuensis
Attar & Ghahr.; Iranian J. Bot. 8(2): 264 (2000)
Distribution: Iran
34 IRN
species :
Cousinia daralaghezica
Takht.; Trans. Armen. Branck Acad. Sci. USSR, Biol. Ser. 2: 181 (1937)
Distribution: Armenia; Azerbaijan (incl. Nachichevan)
33 ARM AZE
species :
Cousinia darwasica
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 12: 249 (1892)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia dasylepis
Kult.; Acta Univ. As. Med., ser. 8b 7: 4 (1929)
Distribution: Iran (NE-Iran: Mts.)
34 IRN
species :
Cousinia davisiana
Hub.-Mor.; Notes Roy. Bot. Gard. Edinburgh 32: 56 (1972)
Distribution: Turkey (SSW-Anatolia)
34 TUR
species :
Cousinia decipiens
Boiss. & Buhse; Nouv. Mém. Soc. Nat. Moscou 12 18: 125 (1860)
Distribution: Iran (EC-Iran, N-Iran)
34 IRN
Synonyms: hidden
= Arctium decipiens (Boiss. & Buhse) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Cousinia daryoushiana Parsa; Kew Bull. 3(2): 203 (1948)
= Cousinia hyrcanica Bornm.; Oesterr. Bot. Z. 75: 233 (1926)
species :
Cousinia decolorans
Freyn & Sint.; Oesterr. Bot. Z. 42 6: 206 (1892)
Distribution: Turkey (Inner Anatolia, E-Anatolia)
34 TUR
species :
Cousinia decumbens
Rech.f.; Ann. Naturhist. Mus. Wien 57: 77 (1950)
Distribution: Iran (EC-Iran)
34 IRN
Synonyms: hidden
= Cousinia meyamensis var. decumbens (Rech.f.) Parsa; Flore de l´Iran (Publ. Univ. Teheran 1777) 10: 99 (1980)
species :
Cousinia decurrens
Regel; Bull. Soc. Imp. Naturalistes Moscou 40: (1867) II. 154
Distribution: Uzbekistan; Tajikistan; Afghanistan (Badakshan, Badghys, Baghlan, Balkh, Bamyan, Faryab, Ghazni, Kabul, Wardak, Qunduz, Takhar); Pakistan (Baluchistan, Quetta)
32 TZK UZB 34 AFG 40 PAK
Synonyms: hidden
= Arctium decurrens (Regel) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia decurrentifolia
Juz. ex Tscherneva; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Uzb. SSR 17: 93 (1962)
Distribution: Uzbekistan
32 UZB
species :
Cousinia deluensis
Attar, Mozaff. & Mirtadz.; Phytotaxa 343(3): 231 (2018)
Distribution: Iran
34 IRN
species :
Cousinia denaensis
Attar & Djavadi; Iranian J. Bot. 16(1): 140 (2010)
Distribution: Iran
34 IRN
species :
Cousinia deserti
Bunge; Mém. Acad. Sci. Pétersb. 7 sér., 9 2: 29 (1865)
Distribution: Iran (EC-Iran, NE-Iran: Mts.); Afghanistan (Farah, Herat, Zabul)
34 AFG IRN
Synonyms: hidden
= Arctium deserti (Bunge) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Cousinia leuconeura Bornm. & Gauba; Repert. Spec. Nov. Regni Veg. 36: 324 (1934)
variant :
var. longispinosa
Djavadi & Attar; Feddes Repert. 120(1-2): 33 (2009)
Distribution: Iran
34 IRN
species :
Cousinia dichotoma
Bunge; Del. Sem. Hort. Dorpat. (1845) 3
Distribution: Turkmenistan; Uzbekistan
32 TKM UZB
Synonyms: hidden
= Arctium dichotomum (Bunge) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia dichromata
Kult.; Acta Univ. As. Med. Ser. VIII. Bot. Fasc. VI. 14 (1929)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia diezii
Rech.f.; Fl. Iranica 90: 275, t. 149 (1972)
Distribution: Iran (E-Iran, NE-Iran: Mts.)
34 IRN
species :
Cousinia dimoana
Kult.; Korovin, Kult. & Popov, Descr. Pl. Nov. Turkmenistan. (Bodenk. u. Pflanzengeogr. Beob. u. Unters. 1: Flussgeb. d. Amu Darya u. Syr Darya, II.), 79 (1916)
Distribution: Turkmenistan; Uzbekistan
32 TKM UZB
species :
Cousinia dipterocarpa
Bornm. & Rech.f.; Repert. Spec. Nov. Regni Veg. 48: 147, t. 313, f. 2 (1940)
Distribution: Iran (NE-Iran: Mts.)
34 IRN
species :
Cousinia discolor
Bunge; Mém. Acad. Sci. Pétersb. 7 sér., 9 2: 34 (1865)
Distribution: Iran (NE-Iran: Mts.)
34 IRN
Synonyms: hidden
= Arctium discolor (Bunge) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Cousinia discolor var. glabra Assadi; Fl. Iran [ed. M. Assadi et al.] 125: 66 (2017)
species :
Cousinia dissecta
Kar. & Kir.; Bull. Soc. Imp. Naturalistes Moscou 15: 391 (1842)
Distribution: Kazakhstan; China (NW-Xinjiang)
32 KAZ 36 CHI
Synonyms: hidden
= Arctium dissectum (Kar. & Kir.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia dissectifolia
Kult.; Trans. Sci. Soc. Turk. 1: 111 (1923)
Distribution: Turkmenistan; Uzbekistan
32 TKM UZB
species :
Cousinia divaricata
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 11: 334 (1890)
Distribution: Tajikistan
32 TZK
species :
Cousinia dolichoclada
Juz.; Not. Syst. Herb. Hort. Petrop. 2: 57 (1921)
Distribution: Uzbekistan
32 UZB
species :
Cousinia dolicholepis
Schrenk; Bull. Phys.-Math. Acad. Petersb. 3: 107 (1845)
Distribution: Kazakhstan; Kyrgyzstan
32 KAZ KGZ
Synonyms: hidden
= Arctium dolicholepis (Schrenk) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia dshisakensis
Kult.; Bull. Inst. Pedol. & Geobot. Univ. As. Centr. 1: 115 (1925)
Distribution: Uzbekistan
32 UZB
species :
Cousinia dubia
Popov; Popov & Androsov, Rast. Zapovedn. Guralash i Zaaminsk. Lesnich. 37 (1936)
Distribution: Uzbekistan; Kyrgyzstan; Tajikistan
32 KGZ TZK UZB
species :
Cousinia eburnea
Bornm.; Oesterr. Bot. Z. 62: 105, t. 2, f. 1 (1912)
Distribution: Iran (SW-Iran)
34 IRN
Synonyms: hidden
= Cousinia ebrahim Parsa; Flore de l´Iran (Publ. Univ. Teheran 1777) 10: 121 (1980)
species :
Cousinia ecbatanensis
Bornm.; Oesterr. Bot. Z. 62: 184, t. 3, f. 3-3b (1912)
Distribution: Iran (SW-Iran)
34 IRN
species :
Cousinia echinocephala
Sennikov; Phytotaxa 5: 24 (2010)
Distribution: Kyrgyzstan
32 KGZ
species :
Cousinia edelbergii
Rech.f.; Fl. Iranica 90: 182 (1955)
Distribution: C-Afghanistan
34 AFG
Remarks: provisionally accepted name
species :
Cousinia edmondsonii
Rech.f.; Fl. Iranica 90: 321 (1972)
Distribution: Iran (NE-Iran: Mts., N-Iran)
34 IRN
species :
Cousinia egens
Juz.; Acta Inst. Bot. Acad. Sci. U.R.S.S., ser. I. Fasc. 3, 304 (1937)
Distribution: Tajikistan
32 TZK
species :
Cousinia elburzensis
Attar, Mahdigholi & Ghahr.; Novon 17(1): 3 (2007)
Distribution: Iran (Mazandaran)
34 IRN
species :
Cousinia eleonorae
Hub.-Mor.; Notes Roy. Bot. Gard. Edinburgh 32: 50 (1972)
Distribution: Turkey (Inner Anatolia)
34 TUR
species :
Cousinia elephantina
Rech.f.; Fl. Iranica 90: 182 (1972)
Distribution: Afghanistan (Herat)
34 AFG
species :
Cousinia elwendensis
Bornm.; Beih. Bot. Centralbl. 28 2: 255 (1911)
Distribution: Iran (SW-Iran)
34 IRN
species :
Cousinia elymaitica
Attar, Mozaff. & Mirtadz.; Nova Biol. Reperta 5(3): 306, fig. 1 (2018)
Distribution: Iran
34 IRN
species :
Cousinia erectispina
Tscherneva; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Uzb. SSR 17: 94 (1962)
Distribution: Turkmenistan; Uzbekistan
32 TKM UZB
species :
Cousinia erinacea
Jaub. & Spach; Ill. Pl. Or. 2: 83, t. 165 (1846)
Distribution: Iran (N-Iran)
34 IRN
Synonyms: hidden
= Arctium erinaceum (Jaub. & Spach) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Cousinia lindsayana Rech.f.; Oesterr. Bot. Z. 100: 465 (1953)
species :
Cousinia eriobasis
Bunge; Mem. Acad. Sci. Pétersb. 7 sér., 9 2: 37 (1865)
Distribution: Iran (EC-Iran, S-Iran, SW-Iran)
34 IRN
Synonyms: hidden
= Arctium eriobasis (Bunge) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Cousinia deickii Bornm.; Notizbl. Bot. Gart. Berlin 14: 278 (1938)
species :
Cousinia eriophylla
(Kult.) Bornm.; Rech.f., Repert. Spec. Nov. Regni Veg. 48: 147 (1940)
Distribution: Iran (NE-Iran: Mts.)
34 IRN
Synonyms: hidden
= Cousinia freynii var. eriophylla Kult.; Acta Univ. As. Med. Ser. 8b 7: 7 (1929)
species :
Cousinia eriorrhiza
Bornm.; Beih. Bot. Centralbl. 28 2: 251 (1911)
Distribution: Iran (SW-Iran)
34 IRN
species :
Cousinia eriotricha
Juz.; Acta Inst. Bot. Acad. Sci. U.R.S.S., ser. I. Fasc. 3, 321 (1937)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia erivanensis
Bornm.; Vestn. Tiflissk. Bot. Sada 30: 15 (1913)
Distribution: Armenia; Azerbaijan (Nachichevan)
33 ARM AZE
species :
Cousinia ermenekensis
Hub.-Mor.; Notes Roy. Bot. Gard. Edinburgh 32: 50 (1972)
Distribution: Turkey (SSW-Anatolia)
34 TUR
species :
Cousinia eryngioides
Boiss.; Diagn. Pl. Orient., ser. 1, 10: 101 (1849)
Distribution: Turkmenistan; Iran (EC-Iran, E-Iran, NE-Iran: Mts., N-Iran)
32 TKM 34 IRN
Synonyms: hidden
= Arctium eryngioides (Boiss.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia esfandiarii
Rech.f. & Aellen; Ann. Naturhist. Mus. Wien 57: 75 (1950)
Distribution: Iran (EC-Iran, NE-Iran: Mts., N-Iran)
34 IRN
species :
Cousinia euchlora
Bornm. & Rech.f.; Repert. Spec. Nov. Regni Veg. 48: 13 1, t. 309, f. 2 (1940)
Distribution: Iran (E-Iran)
34 IRN
Synonyms: hidden
= Cousinia euchlora var. arachnoidea Assadi; Fl. Iran [ed. M. Assadi et al.] 125: 69 (2017)
species :
Cousinia eugenii
Kult.; Not. Syst. Herb. Hort. Petrop. 5: 141 (1924), et Not. Syst. Herb. Hort. Bot. Russia 6: 28 (1926)
Distribution: Turkmenistan; Uzbekistan
32 TKM UZB
species :
Cousinia euphratica
Hub.-Mor.; Notes Roy. Bot. Gard. Edinburgh 32: 51 (1972)
Distribution: Turkey (SE-Anatolia)
34 TUR
species :
Cousinia eurylepis
Rech.f.; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 95 (1955)
Distribution: Afghanistan (Bamyan, etc.)
34 AFG
species :
Cousinia fabrorum
Rech.f.; Fl. Iranica 139a: 113 (1979)
Distribution: Iran (E-Iran)
34 IRN
species :
Cousinia falcinella
Bornm.; Repert. Spec. Nov. Regni Veg. 26: 179 (1929)
Distribution: Iran (EC-Iran)
34 IRN
species :
Cousinia falconeri
Hook.f.; Fl. Brit. India [J. D. Hooker] 3(8): 360 (1881)
Distribution: W-Tibet; NW-India (Zanskar, Jammu & Kashmir)
36 TIB 40 IND
Synonyms: hidden
= Arctium falconeri (Hook.f.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia fallax
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 471 (1887)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia farimanensis
Assadi; Iranian J. Bot. 17(1): 6 (2011), nom. inval., et Phytotaxa 227(1): 99 (2015)
Distribution: Iran
34 IRN
species :
Cousinia farsistanica
Bornm.; Oesterr. Bot. Z. 62: 185, t. 3, f. 2 (1912)
Distribution: Iran (SW-Iran)
34 IRN
species :
Cousinia fascicularis
Juz.; Acta Inst. Bot. Acad. Sci. U.R.S.S., ser. I. Fasc. 3, 299 (1937)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia fedorovii
Takht.; Trans. Armen. Branch Acad. Sci. USSR, Biol. Ser. 2: 183 (1937)
Distribution: Armenia
33 ARM
species :
Cousinia ferghanensis
Bornm.; Beih. Bot. Centralbl. 34: 11. 195 (1916)
Distribution: Uzbekistan; Kyrgyzstan; Tajikistan
32 KGZ TZK UZB
Synonyms: hidden
= Cousinia popovii Kult.; Trudy Turkestansk. Gosud. Univ. 4: 66 (1922)
species :
Cousinia ferruginea
Kult.; Not. Syst. Herb. Hort. Petrop. 5: 142 (1924)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
Synonyms: hidden
= Cousinia truncata Juz.; Trudy Bot. Inst. Akad. Nauk S.S.S.R., ser. 1, Fl. Sist. Vyssh. Rast. 3: 318 (1937)
species :
Cousinia fetissowii
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 12: 247 (1892)
Distribution: Kazakhstan; Kyrgyzstan
32 KAZ KGZ
species :
Cousinia finitima
Juz.; Trans. Rubb. & Guttop. Inst., Fasc. 4, 24 (1932)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia firuzkuhensis
Rech.f.; Fl. Iranica 139a: 120, t. 111 (1979)
Distribution: Iran (EC-Iran)
34 IRN
species :
Cousinia foliosa
Boiss. & Balansa; Boiss., Diagn. Pl. Orient., ser. 2, 6: 116 (1859)
Distribution: Turkey (Inner Anatolia, S-Anatolia, SE-Anatolia)
34 TUR
Synonyms: hidden
= Arctium foliosum (Boiss. & Balansa) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia fragilis
C.Winkl. & Bornm.; Bull. Herb. Boiss. 5: 169, t. 6 (1897)
Distribution: Iran (SW-Iran)
34 IRN
species :
Cousinia fragillima
Rech.f.; Oesterr. Bot. Z. 100: 458, f. 2 (1953)
Distribution: Iran (E-Iran)
34 IRN
species :
Cousinia franchetii
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 578 (1887), nom. nov.
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
Synonyms: hidden
= Arctium franchetianum (C.Winkl.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Cousinia canescens Franch.; Ann. Sci. Nat. Ser. VI. 16: 323 (1883)
species :
Cousinia fraternella
Bornm.; Beih. Bot. Centralbl. 34: 11. 273 (1916)
Distribution: Afghanistan (Bamyan, Ghorat, Wardak, Parwan)
34 AFG
species :
Cousinia freitagii
Rech.f.; Fl. Iranica 90: 86 (1972)
Distribution: Afghanistan (Baghlan)
34 AFG
species :
Cousinia freynii
Bornm. & Sint.; Russk. Bot. Zhurn. 1911: 4 (1911)
Distribution: Turkmenistan; Iran (NE-Iran: Mts., N-Iran)
32 TKM 34 IRN
species :
Cousinia fursei
Rech.f.; Fl. Iranica 90: 247, t. 128 (1972)
Distribution: Iran (SW-Iran)
34 IRN
species :
Cousinia gabrielae
Bornm.; Beih. Bot. Centralbl. 59 2: 301 (1939)
Distribution: Iran (S-Iran); Afghanistan (Farah)
34 AFG IRN
species :
Cousinia gaharensis
Attar & Djavadi; Iranian J. Bot. 16(1): 149 (2010)
Distribution: Iran
34 IRN
species :
Cousinia gatchsaranica
Mehregan, Assadi & Attar; Willdenowia 33(1): 108 (2003)
Distribution: Iran
34 IRN
species :
Cousinia gaubae
Bornm.; Repert. Spec. Nov. Regni Veg. 36: 326, t. 181B (1934)
Distribution: Iran (EC-Iran, N-Iran)
34 IRN
species :
Cousinia gedrosiaca
Bornm. & Gauba; Mitt. Thür. Bot. Ver. 47: 121, t. 15 (1941)
Distribution: Iran (S-Iran, SW-Iran); Pakistan (Baluchistan)
34 IRN 40 PAK
species :
Cousinia ghahremanii
Mirtadz. & Attar; Iranian J. Bot. 15(2): 147 (2009)
Distribution: Iran
34 IRN
species :
Cousinia ghorana
Rech.f.; Anz. Österr. Akad. Wiss., Math.-Nat., 101: 346 (1964)
Distribution: Afghanistan (Ghorat)
34 AFG
species :
Cousinia gigantolepis
Rech.f.; Fl. Iranica 90: 257, t. 136 (1972)
Distribution: Armenia; Iran (Iranian Aserbaijan)
33 ARM 34 IRN
species :
Cousinia gigantoptera
Rech.f. & Podlech; Pl. Syst. Evol., 142(1-2): 2 (1983)
Distribution: Afghanistan (Takhar)
34 AFG
species :
Cousinia gilanica
Bornm.; Oesterr. Bot. Z. 62: 108, t. 2, f. 3 (1912)
Distribution: Iran (EC-Iran, N-Iran)
34 IRN
species :
Cousinia gilliatii
Rech.f.; Fl. Iranica 90: 244, t. 124 (1972)
Distribution: Iran (Iranian Aserbaijan)
34 IRN
species :
Cousinia ginuae
Attar; Taxon. Biosyst. 3(6): 60 (2011), et Ann. Bot. Fenn. 57(4-6): 238 (2020)
Distribution: Iran
34 IRN
species :
Cousinia glabriseta
Kult.; Acta Univ. As. Med. Ser. VIII. Bot. Fasc. VI. 7 (1929)
Distribution: Turkmenistan; Uzbekistan
32 TKM UZB
species :
Cousinia glandulosa
Kult.; Acta Univ. As. Med. Ser. VIII. Bot. Fasc. VI. 13 (1929)
Distribution: Turkmenistan
32 TKM
species :
Cousinia glaphyra
Rech.f. & Köie; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 99 (1955)
Distribution: Afghanistan (Bamyan)
34 AFG
species :
Cousinia glaphyrocephala
Juz. ex Tscherneva; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Uzb. SSR 17: 94 (1962)
Distribution: Uzbekistan
32 UZB
species :
Cousinia glaucopsis
Bornm. & Rech.f.; Repert. Spec. Nov. Regni Veg. 48: 134, t. 310, f. 2 (1940)
Distribution: Iran (EC-Iran, N-Iran)
34 IRN
variant :
var. spinellosissima
Bornm. & Rech.f.; Repert. Spec. Nov. Regni Veg. 48: 134, t. 310, f. 2 (1940)
Distribution: Iran
34 IRN
species :
Cousinia glochidiata
Kult.; Acta Univ. As. Med. Ser. VIII. Bot. Fasc. VI. 3 (1929)
Distribution: Turkmenistan
32 TKM
species :
Cousinia gmelinii
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 12: 241 (1893)
Distribution: Iran (N-Iran)
34 IRN
species :
Cousinia gnezdilloi
Tscherneva; Not. Syst. Herb. Inst. Bot. Acad. Sci. . 15: 47 (1959)
Distribution: Turkmenistan; Uzbekistan; Tajikistan
32 TKM TZK UZB
species :
Cousinia golestanica
Attar; Taxon. Biosyst. 3(6): 63 (2011)
Distribution: Iran
34 IRN
species :
Cousinia gomolitzkii
Juz. ex Tscherneva; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Uzb. SSR 17: 95 (1962)
Distribution: Kazakhstan
32 KAZ
species :
Cousinia gontscharowii
Juz.; Acta Inst. Bot. Acad. Sci. U.R.S.S., ser. I. Fasc. 3, 311 (1937)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia gracilis
Boiss.; Diap., ser. 1, 6: 117 (1846)
Distribution: Iran (SW-Iran)
34 IRN
Synonyms: hidden
= Arctium gracile (Boiss.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Cousinia elymaea Hausskn. & Winkl. ex C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 14: 225 (1897)
species :
Cousinia graminifolia
Rech.f.; Fl. Iranica 90: 178 (1972)
Distribution: Afghanistan (Baghlan)
34 AFG
species :
Cousinia grandis
C.A.Mey.; Prodr. [A. P. de Candolle] 6: 557 (1838)
Distribution: Armenia; Turkey (SE-Anatolia); Iran (Iranian Aserbaijan)
33 ARM 34 IRN TUR
Synonyms: hidden
= Arctium grande (C.A.Mey.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia grantii
Rech.f.; Fl. Iranica 90, 245, t. 125 (1972)
Distribution: Iran (Iranian Aserbaijan)
34 IRN
species :
Cousinia greuteri
Rech.f.; Fl. Iranica 90: 191 (1972)
Distribution: Afghanistan (Baghlan, Samangan)
34 AFG
species :
Cousinia griffithiana
Boiss.; Diagn. Ser. II. 3: 55 (1856)
Distribution: Afghanistan (Ghazni, Zabul)
34 AFG
Synonyms: hidden
= Arctium griffithianum (Boiss.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia grigoriewii
Juz.; Acta Inst. Bot. Acad. Sci. U.R.S.S., ser. I. Fasc. 3, 313 (1937)
Distribution: Tajikistan
32 TZK
species :
Cousinia grisea
Kult.; Acta Univ. As. Med. Ser. VIII. Bot. Fasc. VI. 5 (1929)
Distribution: Uzbekistan
32 UZB
species :
Cousinia gulczensis
Kult.; Acta Univ. As. Med. Ser. VIII b, Bot. Fasc. 6, 16 (1929)
Distribution: Kyrgyzstan
32 KGZ
species :
Cousinia gymnoclada
Rech.f. & Köie; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 102 (1955)
Distribution: Afghanistan (Herat)
34 AFG
species :
Cousinia hablitzii
C.A.Mey.; Prodr. [A. P. de Candolle] 6: 554 (1838)
Distribution: Iran (N-Iran)
34 IRN
Synonyms: hidden
= Arctium hablitzii (C.A.Mey.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Carduus macranthus (Schrank) Hablitz; Boiss., Fl. Orient. 3: 496 (1875)
= Chamaepeuce macrantha Schrank; Bull. Sci. Acad. Petersb. 10: 354 (1842)
= Cirsium macranthum (Schrenk) B.Fedtsch.; Rastitel´n. Turkestana: 755 (1915)
species :
Cousinia haeckeliae
Bornm.; Repert. Spec. Nov. Regni Veg., Beih. 108: 40 (1938)
Distribution: Afghanistan (Bamyan, Kunar / Nuristan, Parwan); Pakistan (Chitral)
34 AFG 40 PAK
species :
Cousinia hakkarica
Hub.-Mor.; Notes Roy. Bot. Gard. Edinburgh 32: 54 (1972)
Distribution: Turkey (SE-Anatolia)
34 TUR
species :
Cousinia halysensis
Hub.-Mor.; Notes Roy. Bot. Gard. Edinburgh 32: 51 (1972)
Distribution: Turkey (Inner Anatolia)
34 TUR
species :
Cousinia hamadae
Juz.; Trudy Sk. Bazy 8: 523 (1940)
Distribution: Turkmenistan; Uzbekistan; Kyrgyzstan; Tajikistan
32 KGZ TKM TZK UZB
Synonyms: hidden
= Cousinia pseudaffinis var. bucharica Kult.; Trudy Sredne-Aziatsk. Gosud. Univ., ser. 8b, Bot 6: 11 (1929)
species :
Cousinia hamadanensis
Rech.f.; Fl. Iranica 139a: 142, t. 136 (1979)
Distribution: Iran (SW-Iran)
34 IRN
species :
Cousinia hamosa
C.A.Mey.; Prodr. [A. P. de Candolle] 6: 555 (1838)
Distribution: Armenia; Iran (Iranian Aserbaijan)
33 ARM 34 IRN
Synonyms: hidden
= Arctium hamosum (C.A.Mey.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Cousinia buhsei Bunge; Mém. Acad. Imp. Sci. Saint Pétersbourg, Sér. 7, 9(2): 22 (1865)
= Cousinia hamulosa C.A.Mey.; Boiss., Fl. Orient. [Boiss. ] 3: 483 (1875)
species :
Cousinia harazensis
Rech.f.; Fl. Iranica 90: 259, t. 137 (1972)
Distribution: Iran (EC-Iran, N-Iran)
34 IRN
species :
Cousinia hastifolia
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 571 (1887)
Distribution: Kyrgyzstan; Tajikistan
32 KGZ TZK
Synonyms: hidden
= Cousinia taldykensis Juz.; Trudy Bot. Inst. Akad. Nauk S.S.S.R., ser. 1, Fl. Sist. Vyssh. Rast. 3: 299 (1937)
species :
Cousinia haussknechtii
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 12: 238 (1893)
Distribution: Iraq (NE-Iraq); Iran (W-Iran)
34 IRN IRQ
Synonyms: hidden
= Cousinia serotina Rech.f.; Anz. Österr. Akad. Wiss. Math.-Naturwiss. Kl. 101: 349 (1964)
species :
Cousinia hazarensis
Mirtadz. & Attar; Willdenowia 34(1): 191 (2004)
Distribution: Iran
34 IRN
species :
Cousinia hedgei
Rech.f.; Fl. Iranica 90: 43 (1972)
Distribution: Afghanistan (Samangan)
34 AFG
species :
Cousinia heliantha
Bunge; Mém. Acad. Sci. Petersb. 7 ser., 9 2: 39 (1865)
Distribution: Iran (EC-Iran, NE-Iran: Mts.)
34 IRN
Synonyms: hidden
= Arctium helianthum (Bunge) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia hemsleyana
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 12: 228 (1892)
Distribution: Afghanistan (Herat)
34 AFG
species :
Cousinia hergtiana
Bornm.; Beih. Bot. Centralbl. 32 2: 400 (1914)
Distribution: Iran (SW-Iran)
34 IRN
species :
Cousinia heteroloba
Rech.f.; Anz. Österr. Akad. Wiss., Math.-Nat., 101: 347 (1964)
Distribution: Afghanistan (Baghlan, Bamyan)
34 AFG
species :
Cousinia heterophylla
Boiss.; Diagn. Ser. II. 3: 54 (1856)
Distribution: Afghanistan (Ghazni, Ghorat, Herat, Orozgan / Daykundi, Paktia / Khost, Zabul); Pakistan (Quetta, Baluchistan)
34 AFG 40 PAK
Synonyms: hidden
= Arctium heterophyllum (Boiss.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia hilariae
Kult.; Acta Univ. As. Med. Ser. VIII. Bot. Fasc. VI. 22 (1929)
Distribution: Tajikistan
32 TZK
species :
Cousinia hohenackeri
Fisch. & C.A.Mey.; Ind. Sem. Hort. Petrop. 2: 34 (1835)
Distribution: Azerbaijan (Talysh); Iran
33 AZE 34 IRN
Synonyms: hidden
= Arctium hohenackeri (Fisch. & C.A.Mey.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia hololeuca
Bunge; Mém. Acad. Sci. Petersb. 7 sér., 9 2: 27 (1865)
Distribution: Iran (EC-Iran)
34 IRN
Synonyms: hidden
= Arctium hololeucum (Bunge) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia hoplites
Bornm.; Beih. Bot. Centralbl. 34: 11. 277 (1916)
Distribution: Afghanistan (Ghazni, Wardak)
34 AFG
species :
Cousinia hoplophylla
Tscherneva; Not. Syst. Herb. Inst. Bot. Acad. Sci. . 15: 49 (1959)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia horrida
Kult.; Acta Univ. As. Med. Ser. 8b 7: 1, f. 1 (1929)
Distribution: Iran (NE-Iran: Mts.)
34 IRN
species :
Cousinia humilis
Boiss.; Ann. Sci. Nat., ser. 4, 2: 248 (1854)
Distribution: Turkey (Inner Anatolia)
34 TUR
Synonyms: hidden
= Arctium humile (Boiss.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia hymenostephanus
(Rech.f.) Podlech; Vasc. Pl. Afghanistan 162 (2013)
Distribution: Afghanistan (Ghorat)
34 AFG
Synonyms: hidden
= Tiarocarpus hymenostephanus Rech.f.; Anz. Österr. Akad. Wiss., Mat.-Naturwiss., 108: 7 (1971 publ. 1972)
species :
Cousinia hypochionea
Bornm.; Bull. Herb. Boiss., ser. 2, 7: 217, t. 7, f. 1 (1907)
Distribution: Iran (N-Iran)
34 IRN
species :
Cousinia hypoleuca
Boiss.; Fl. Orient. 3: 464 (1875)
Distribution: Iran (EC-Iran, E-Iran, NE-Iran: Mts., N-Iran, W-Iran)
34 IRN
Synonyms: hidden
= Arctium hypoleucum (Boiss.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Cousinia assasinorum Bornm.; Bull. Herb. Boiss. Ser. 2(7): 210 (1907)
= Cousinia hypoleuca var. congesta Bornm.; (1917)
= Cousinia hypoleuca var. teheranica Parsa; (1948)
= Cousinia tafreshensis Rech.f.; Ann. Naturhist. Mus. Wien 42: 73 (1950)
species :
Cousinia hypopolia
Bornm. & Sint.; Bull. Herb. Boiss., ser. 2, 7: 214, t. 7, f. 4 (1907)
Distribution: Turkmenistan; Iran (EC-Iran, NE-Iran: Mts., N-Iran)
32 TKM 34 IRN
species :
Cousinia hystricocephala
Sennikov; Phytotaxa 25: 25 (2011)
Distribution: Uzbekistan; Kyrgyzstan; Tajikistan
32 KGZ TZK UZB
species :
Cousinia iconica
Hub.-Mor.; Notes Roy. Bot. Gard. Edinburgh 32: 52 (1972)
Distribution: Turkey (Central Anatolia, Inner Anatolia)
34 TUR
species :
Cousinia ilicifolia
Jaub. & Spach; Ill. Pl. Or. 2: 82 (1846)
Distribution: Iran (S-Iran, SW-Iran)
34 IRN
Synonyms: hidden
= Arctium ilicifolium (Jaub. & Spach) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia iljinii
Takht.; Trans. Armen. Branch Acad. Sci. USSR, Biol. Ser. 2: 187 (1937)
Distribution: Armenia; Azerbaijan (Nachichevan)
33 ARM AZE
species :
Cousinia immitans
Rech.f.; Anz. Österr. Akad. Wiss., Math.-Nat. 103: 285 (1966)
Distribution: Afghanistan (Baghlan, Parwan)
34 AFG
species :
Cousinia immitantiformis
Rech.f.; Fl. Iranica 90: 125 (1972)
Distribution: Afghanistan (Herat)
34 AFG
species :
Cousinia incompta
DC.; Prodr. [A. P. de Candolle] 6: 553 (1838)
Distribution: Iran
34 IRN
Synonyms: hidden
= Arctium incomptum (DC.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Cousinia incompta var. continua Assadi; Fl. Iran [ed. M. Assadi et al.] 125: 331 (2017)
species :
Cousinia inflata
Boiss. & Hausskn.; Boiss., Fl. Orient. 3: 512 (1875)
Distribution: Iraq (NE-Iraq, W-Iraq); Iran (W-Iran)
34 IRN IRQ
Synonyms: hidden
= Arctium inflatum (Boiss. & Hausskn.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Cousinia straussii Hausskn. & C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 14: 235 (1897)
species :
Cousinia infundibularis
Rech.f.; Fl. Iranica 90: 60 (1972)
Distribution: Afghanistan (Ghorat)
34 AFG
species :
Cousinia insignis
Rech.f.; Fl. Iranica 90: 106 (1972)
Distribution: Afghanistan (Orozgan / Daykundi)
34 AFG
species :
Cousinia integrifolia
Franch.; Ann. Sci. Nat. Ser. VI. 16: 322 (1883)
Distribution: Turkmenistan; Uzbekistan; Tajikistan
32 TKM TZK UZB
Synonyms: hidden
= Arctium integrifolium (Franch.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia intermedia
DC.; Prodr. [A. P. de Candolle] 6: 553 (1838)
Distribution: Armenia; Azerbaijan (Nachichevan)
33 ARM AZE
Synonyms: hidden
= Arctium intermedium (C.A.Mey. ex DC.) Kuntze; Revis. Gen. Pl. 1: 308 (1891), nom. illeg.
species :
Cousinia intertexta
Freyn & Sint.; Oesterr. Bot. Z. 42 5: 169 (1892)
Distribution: Turkey (E-Anatolia, Inner Anatolia, N-Anatolia, SE-Anatolia)
34 TUR
species :
Cousinia iranshahriana
Attar & Maroofi; Iranian J. Bot. 16(2): 197 (-199, fig. 1) (2010)
Distribution: Iran
34 IRN
species :
Cousinia iranshahrii
Rech.f.; Fl. Iranica 139a: 152, t. 148 (1979)
Distribution: Iran (NE-Iran: Mts.)
34 IRN
species :
Cousinia irritans
Rech.f.; Anz. Österr. Akad. Wiss. Math.-Naturwiss. Kl. 57: 72 (1950)
Distribution: Iran (EC-Iran, NE-Iran: Mts.)
34 IRN
species :
Cousinia isfahanica
Assadi; Iranian J. Bot. 15(1): 37 (2009)
Distribution: Iran
34 IRN
species :
Cousinia jacobsii
Rech.f.; Fl. Iranica 90: 237, t. 119 (1972)
Distribution: Iran (SW-Iran)
34 IRN
species :
Cousinia jassyensis
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 472 (1887)
Distribution: Kyrgyzstan
32 KGZ
Synonyms: hidden
= Arctium jassyense (C.Winkl.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia joharchii
Assadi & Mehregan; Phytotaxa 257(3): 276 (2016)
Distribution: Iran
34 IRN
species :
Cousinia juzepczukii
Tscherneva; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Uzb. SSR 17: 95 (1962)
Distribution: Uzbekistan
32 UZB
species :
Cousinia kadereitii
Mehregan & Assadi; Willdenowia 39(2): 268 (2010)
Distribution: Iran
34 IRN
species :
Cousinia kaluensis
Rech.f.; Fl. Iranica 90: 84 (1972)
Distribution: Afghanistan (Bamyan)
34 AFG
species :
Cousinia kandavanensis
Attar; Taxon. Biosyst. 3(6): 64 (2011)
Distribution: Iran
34 IRN
variant :
var. golydaghi
Parsa; Kew Bull. 3(2): 205 (1948)
Distribution: Iran
34 IRN
species :
Cousinia karkasensis
Mehregan & Djavadi; Iranian J. Bot. 16(2): 200 (-203, fig. 1, map) (2010)
Distribution: Iran
34 IRN
species :
Cousinia kasachstanica
Sennikov; Phytotaxa 5: 21 (2010)
Distribution: Kazakhstan
32 KAZ
species :
Cousinia kataghanica
Rech.f.; Anz. Österr. Akad. Wiss., Math.-Naturwiss. Kl. 105: 6 (1969)
Distribution: Afghanistan (Baghlan, Parwan)
34 AFG
species :
Cousinia kazachorum
Juz. ex Tscherneva; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Uzb. SSR 17: 96 (1962)
Distribution: Kazakhstan; Kyrgyzstan
32 KAZ KGZ
species :
Cousinia keredjensis
Bornm. & Gauba; Repert. Spec. Nov. Regni Veg. 36: 328, t. 227 (1934)
Distribution: Iran (EC-Iran)
34 IRN
species :
Cousinia kermanensis
Mirtadz. & Attar; Nordic J. Bot. 32(3): 279 (2014)
Distribution: Iran
34 IRN
species :
Cousinia kermanshahensis
Attar, Ghahr. & Assadi; Iranian J. Bot. 9(1): 56 (2001)
Distribution: Iran
34 IRN
species :
Cousinia kerstanii
Bornm.; Repert. Spec. Nov. Regni Veg., Beih. 108: 49 (1938)
Distribution: Afghanistan (Kunar / Nuristan); Pakistan (Chitral); NW-India (Jammu & Kashmir)
34 AFG 40 IND PAK
species :
Cousinia khashensis
Rech.f.; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 105 (1955)
Distribution: Afghanistan (Badakshan, Wakhan)
34 AFG
species :
Cousinia khorasanica
Djavadi & Attar; Rostaniha 7(Suppl. 2): 170 (2007)
Distribution: Iran
34 IRN
species :
Cousinia khorramabadensis
Bornm.; Köie, Danish Scient. Invest. Iran 4, Beitr. Sw. Iran 1: 25, f. 4 (1945)
Distribution: Iran (SW-Iran)
34 IRN
variant :
var. purpurea
Attar & Ghahr.; Iranian J. Bot. 9(2): 166 (2002)
Distribution: Iran
34 IRN
species :
Cousinia kilouyensis
Djavadi & Attar; Willdenowia 39(1): 89 (2009)
Distribution: Iran
34 IRN
species :
Cousinia kirrindica
Bornm. & Rech.f.; Repert. Spec. Nov. Regni Veg. 48: 141 (1940)
Distribution: Iran (SW-Iran)
34 IRN
Synonyms: hidden
= Cousinia lurorum var. recurvata Bornm.; (1936)
= Cousinia lurorum var. lancigera Bornm.; (1936)
species :
Cousinia knorringiae
Bornm.; Beih. Bot. Centralbl. 34: 11. 201 (1916)
Distribution: Kyrgyzstan
32 KGZ
species :
Cousinia koelzii
Rech.f.; Biol. Skr. 8(2): 109 (1955)
Distribution: Afghanistan (Badakshan)
34 AFG
species :
Cousinia kokanica
Regel & Schmalh. ex Regel; Pl. Nov. Fedsch. 49 (1882)
Distribution: Kyrgyzstan; Tajikistan
32 KGZ TZK
Synonyms: hidden
= Arctium kokanicum (Regel & Schmalh.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia komaroffii
(Kuntze) C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 470 (1889)
Distribution: Turkmenistan; Iran (NE-Iran: Mts.)
32 TKM 34 IRN
Synonyms: hidden
= Arctium komaroffii Kuntze; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 197 (1887)
species :
Cousinia komidjanensis
Mehregan; Iranian J. Bot. 17(2): 140 (2011)
Distribution: Iran
34 IRN
species :
Cousinia kopi-karadaghensis
Rech.f.; Fl. Iranica 90: 249, t. 130 (1972)
Distribution: Iraq (NE-Iraq); Iran (W-Iran)
34 IRN IRQ
species :
Cousinia kornhuberi
Heimerl; Denkschr. Akad. Wiss. Math.-Naturwiss. Kl. 50: 62 (1885)
Distribution: Iran (SW-Iran)
34 IRN
Synonyms: hidden
= Cousinia medorum Bornm. & Gauba; Repert. Spec. Nov. Regni Veg. 40: 345 (1936)
species :
Cousinia korowiakowii
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 14: 226 (1897)
Distribution: Iran (EC-Iran)
34 IRN
species :
Cousinia kotschyi
Boiss.; Diagn. Pl. Orient., ser. 1, 6: 120 (1846)
subspecies :
subsp. kotschyi
Distribution: Iran (EC-Iran, SW-Iran)
34 IRN
Synonyms: hidden
= Lappa kotschyi (Boiss.) Boiss.; Fl. Orient. 3: 458 (1875)
subspecies :
subsp. khansarica
(Attar & Ghahr.) Mehregan; Willdenowia 38(2): 340 (2008)
Distribution: Iran
34 IRN
Synonyms: hidden
= Cousinia khansarica Attar & Ghahr.; Nordic J. Bot. 23(5): 592 (2005)
species :
Cousinia kovalevskiana
Tscherneva; Novosti Sist. Vyssh. Rast. 26: 162 (1989)
Distribution: Kyrgyzstan
32 KGZ
species :
Cousinia krauseana
Regel & Schmalh. ex Regel; Pl. Nov. Fedsch. 48 (1882)
Distribution: Uzbekistan; Kyrgyzstan
32 KGZ UZB
Synonyms: hidden
= Arctium krauseanum (Regel & Schmalh.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia kuekenthalii
Bornm.; Beih. Bot. Centralbl. 34: 11. 146 (1916)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia kuramensis
Bornm.; Beih. Bot. Centralbl. 34: 11. 150 (1916)
Distribution: Afghanistan (Paktia / Khost); Pakistan (Kurram)
34 AFG 40 PAK
species :
Cousinia kurdistanica
Attar; Iranian J. Bot. 16(1): 169 (2010)
Distribution: Iran
34 IRN
species :
Cousinia lachnobasis
Rech.f. & Köie; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 113 (1955)
Distribution: Afghanistan (Farah)
34 AFG
species :
Cousinia lachnopoda
Juz.; Trudy sk. Bazy 8: 525 (1940)
Distribution: Tajikistan
32 TZK
species :
Cousinia lachnosphaera
Bunge; Mém. Acad. Sci. Pétersb. 7 sér., 9 2: 36 (1865)
Distribution: Iran (EC-Iran, E-Iran, NE-Iran: Mts.); Afghanistan (Bamyan, Ghorat, Herat, Wardak, Orozgan / Daykundi)
34 AFG IRN
Synonyms: hidden
= Arctium lachnosphaerum (Bunge) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Cousinia sven-hediniana Bornm.; Beih. Bot. Centralbl. 34(2): 268 (1916)
variant :
var. kashanensis
(Rech.f. & Esfand.) Parsa; Flore de l´Iran (Publ. Univ. Teheran 1777) 10: 108 (1980)
Distribution: Iran (EC-Iran)
34 IRN
Synonyms: hidden
= Cousinia kashanensis Rech.f. & Esfand.; Ann. Naturhist. Mus. Wien 57: 76 (1950)
species :
Cousinia lactiflora
Rech.f.; Fl. Iranica 139a: 143, t. 138 (1979)
Distribution: Iran (SW-Iran)
34 IRN
species :
Cousinia laetevirens
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 576 (1887)
Distribution: Turkmenistan; Uzbekistan; Tajikistan
32 TKM TZK UZB
species :
Cousinia lanata
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 13: 238 (1894)
Distribution: Turkmenistan; Uzbekistan; Tajikistan
32 TKM TZK UZB
species :
Cousinia lancifolia
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 572 (1887)
Distribution: Tajikistan
32 TZK
species :
Cousinia laniceps
(Bornm.) Juz.; Acta Inst. Bot. Acad. Sci. U.R.S.S., ser. I. Fasc. 3, 316 (1937)
Distribution: Kyrgyzstan; Tajikistan
32 KGZ TZK
Synonyms: hidden
= Cousinia schtschurowskiana var. laniceps Bornm.; Beih. Bot. Centralbl. 34(2): 187 (1916)
species :
Cousinia lasiandra
Bunge; Mém. Acad. Sci. Pétersb. 7 sér., 9 2: 40 (1865)
Distribution: Iran (EC-Iran, NE-Iran: Mts.)
34 IRN
Synonyms: hidden
= Arctium lasiandrum (Bunge) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia lasiolepis
Boiss.; Diagn. Pl. Orient., ser. 1, 6: 118 (1846)
Distribution: Iran (EC-Iran, E-Iran, S-Iran, SW-Iran); Afghanistan (Bamyan, Ghorat)
34 AFG IRN
Synonyms: hidden
= Arctium lasiolepis (Boiss.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Cousinia machaerophora C.Winkl. & Bornm.; (1897)
species :
Cousinia lasiophylla
C.Shih; Bull. Bot. Res., Harbin 4(2): 59-60 (1984)
Distribution: China (W-Xinjiang)
36 CHI
species :
Cousinia leiocephala
(Regel) Juz.; Not. Inst. Bot. Acad. Sci. U.R.S.S., ser. 1, 3: 314 (1936)
Distribution: Kazakhstan; Uzbekistan; Kyrgyzstan; China (Xinjiang)
32 KAZ KGZ UZB 36 CHI
Synonyms: hidden
= Cousinia sewertzowii var. leiocephala Regel; Trudy Imp. S.-Peterburgsk. Bot. Sada 6(2): 314 (1880)
species :
Cousinia lepida
Bunge; Boiss., Fl. Orient. 3: 476 (1875)
Distribution: Iran (EC-Iran, E-Iran, NE-Iran: Mts.)
34 IRN
Synonyms: hidden
= Arctium lepidum (Bunge) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia leptacma
Tscherneva; Not. Syst. Herb. Inst. Bot. Acad. Sci. . 15: 35 (1959)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia leptocampyla
Bornm.; Beih. Bot. Centralbl. 34: 11. 179 (1916)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia leptocephala
Fisch. & C.A.Mey.; Bull. Soc. Imp. Naturalistes Moscou (1839) 159 ex Bunge
Distribution: Turkmenistan
32 TKM
Synonyms: hidden
= Arctium leptocephalum (Fisch. & C.A.Mey.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia leptoclada
Kult.; Acta Univ. As. Med. Ser. VIII. Bot. Fasc. VI. 15 (1929)
Distribution: Turkmenistan; Uzbekistan
32 TKM UZB
species :
Cousinia leptocladoides
Tscherneva; Not. Syst. Herb. Inst. Bot. Acad. Sci. . 15: 45 (1959)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia leptolepis
(Bornm. & Gauba) Rech.f.; Fl. Iranica 90: 1 1 1, t. 49 (1972)
Distribution: Iraq (NE-Iraq); Iran (EC-Iran)
34 IRN IRQ
Synonyms: hidden
= Cousinia adenosticta var. leptolepis Bornm. & Gauba; Repert. Spec. Nov. Regni Veg. 36: 33 (1934)
species :
Cousinia leptomera
Rech.f.; Anz. Österr. Akad. Wiss., Math.-Nat., 101: 348 (1964)
Distribution: Afghanistan (Ghorat)
34 AFG
species :
Cousinia leucantha
Bornm. & Sint; J. Russe de Bot., 1 (1911)
Distribution: Turkmenistan
32 TKM
Synonyms: hidden
= Cousinia simplex Kult.; (1960)
species :
Cousinia libanotica
DC.; Prodr. [A. P. de Candolle] 6: 556 (1838)
Distribution: Lebanon; Syria; Israel (coastal W-Israel); Jordania (S-Jordania)
34 ISR JOR LEB SYR
Synonyms: hidden
= Arctium hermonis (Boiss.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Arctium libanoticum (DC.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Cousinia dayi Post; Pl. Postianae 2: 13 (1891)
= Cousinia hermonis Boiss.; Diagn. Pl. Orient., ser. 1, 10: 102 (1849)
species :
Cousinia lignosissima
Rech.f.; Fl. Iranica 139a: 113, t. 106 (1979)
Distribution: Iran (NE-Iran: Mts.)
34 IRN
species :
Cousinia linczewskii
Juz.; Acta Inst. Bot. Acad. Sci. U.R.S.S., ser. 1, 3 (1936)
Distribution: Turkmenistan; Iran (NE-Iran: Mts.)
32 TKM 34 IRN
species :
Cousinia litwinowiana
Bornm.; Bull. Herb. Boiss. Ser. II. 7: 215 (1907)
Distribution: Uzbekistan
32 UZB
species :
Cousinia lomakinii
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 14: 220 (1897)
Distribution: Armenia
33 ARM
species :
Cousinia longibracteata
Attar & Mirtadz.; Iranian J. Bot. 15(2): 149 (2009)
Distribution: Iran
34 IRN
species :
Cousinia longifolia
C.Winkl. & Bornm.; Bull. Herb. Boiss. 5: 168, t. 8 (1897)
Distribution: Iran (EC-Iran, SW-Iran)
34 IRN
species :
Cousinia lordeganensis
Mehregan; Willdenowia 38(2): 341 (2008)
Distribution: Iran
34 IRN
species :
Cousinia lucida
DC.; Prodr. [A. P. de Candolle] 6: 553 (1838)
Distribution: Iran (EC-Iran, SW-Iran)
34 IRN
Synonyms: hidden
= Arctium lucidum (DC.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Cousinia lucida var. arachnoidea Assadi & Mehregan; Fl. Iran [ed. M. Assadi et al.] 125: 132 (2017)
species :
Cousinia lurestanica
Attar & Djavadi; Iranian J. Bot. 16(1): 153 (2010)
Distribution: Iran
34 IRN
species :
Cousinia lurorum
Bornm.; Repert. Spec. Nov. Regni Veg. 40: 346 (1936)
Distribution: Iran (SW-Iran)
34 IRN
Synonyms: hidden
= Cousinia cymbolepis var. subintegriloba Bornm.; (1914)
species :
Cousinia lyrata
Bunge; Mém. Acad. Sci. Pétersb. 7 sér., 9 2: 53 (1865)
Distribution: Turkmenistan; Iran (NE-Iran: Mts.); NW-Afghanistan
32 TKM 34 AFG IRN
Synonyms: hidden
= Arctium lyratum (Bunge) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia maassoumii
Assadi; Iranian J. Bot. 15(1): 40 (2009)
Distribution: Iran
34 IRN
species :
Cousinia macrocephala
C.A.Mey.; Verz. Pfl. Caucasus 231 (1831)
Distribution: Armenia; Azerbaijan (incl. Nachichevan); Iran
33 ARM AZE 34 IRN
Synonyms: hidden
= Arctium macrocephalum (C.A.Mey.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Cousinia gabrieljaniae Takht. & Tamanian; Bot. Zhurn. (Moscow & Leningrad) 73(11): 1609 (1988)
= Cousinia takhtajanii Tamanian; Fl. Rast. Rast. Res. Armenii (Sborn. Nauch. Trud. 13): 17 (1991)
species :
Cousinia macrolepis
Boiss. & Hausskn.; Boiss., Fl. Orient. 3: 505 (1875)
Distribution: Iraq (NE-Iraq)
34 IRQ
Synonyms: hidden
= Arctium macrolepis (Boiss. & Hausskn.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Cousinia gigantosphaera Rech.f.; Anz. österr. Akad. Wiss. Math.-Naturwiss. Kl. 101: 345 (1964)
species :
Cousinia macroptera
C.A.Mey.; Prodr. [A. P. de Candolle] 6: 555 (1838)
Distribution: Armenia; Azerbaijan (incl. Nachichevan); Turkey (E-Anatolia); Iran (Iranian Aserbaijan, W-Iran)
33 ARM AZE 34 IRN TUR
Synonyms: hidden
= Arctium macropterum (C.A.Mey.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Arctium obovatum (Boiss.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Cousinia carlinoides Boiss. & Buhse; Nouv. Mém. Soc. Imp. Naturalistes Moscou 12: 126 (1860)
= Cousinia compacta Turrill; Bull. Misc. Inform. Kew 1929(7): 229 (1929)
= Cousinia macroptera var. obovata (Boiss.) Winkl.; (1893)
= Cousinia obovata Boiss.; Fl. Orient. 3: 492 (1875)
species :
Cousinia magnifica
Juz.; Acta Inst. Bot. Acad. Sci. U.R.S.S., ser. I. Fasc. 3, 319 (1937)
Distribution: Tajikistan
32 TZK
species :
Cousinia malacophylla
Rech.f.; Fl. Iranica 90: 176 (1972)
Distribution: Afghanistan (Ghorat)
34 AFG
species :
Cousinia malurensis
Rech.f.; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 119 (1955)
Distribution: Afghanistan (Faryab)
34 AFG
species :
Cousinia manouchehrii
Rech.f. & Esfand.; Ann. Naturhist. Mus. Wien 57: 79 (1950)
Distribution: Iran (EC-Iran)
34 IRN
species :
Cousinia maracandica
Juz.; Trans. Rubb. & Guttap. Inst., Fasc. 4, 30 (1932)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia margaritae
Kult.; Acta Univ. As. Med. Ser. VIII. Bot. Fasc. VI. 5 (1929)
Distribution: Kyrgyzstan
32 KGZ
species :
Cousinia margiana
Juz.; Acta Inst. Bot. Acad. Sci. U.R.S.S., ser. I. Fasc. 3, 303 (1937)
Distribution: Turkmenistan
32 TKM
species :
Cousinia masulehensis
Attar & Rastegar; Phytotaxa 343(3): 231 (2018)
Distribution: Iran
34 IRN
species :
Cousinia mattfeldii
Bornm.; Repert. Spec. Nov. Regni Veg., Beih. 108: 38 (1938)
Distribution: Pakistan (Chitral); NW-India (Jammu & Kashmir)
40 IND PAK
species :
Cousinia megalomastix
Rech.f.; Fl. Iranica 90: 316 (1972)
Distribution: Afghanistan (Faryab)
34 AFG
species :
Cousinia meghrica
Takht.; Trans. Armen. Branch Acad. Sci. USSR, Biol. Ser. 2: 186 (1937)
Distribution: Armenia; Azerbaijan (Nachichevna)
33 ARM AZE
species :
Cousinia mehreganii
Assadi; Iranian J. Bot. 17(1): 8 (2011)
Distribution: Iran
34 IRN
species :
Cousinia meshhedensis
Bornm. & Rech.f.; Repert. Spec. Nov. Regni Veg. 48: 144, t. 312, f. 1 (1940)
Distribution: Iran (EC-Iran, NE-Iran: Mts., N-Iran)
34 IRN
Synonyms: hidden
= Cousinia strictissima Rech.f.; Ann. Naturhist. Mus. Wien 57: 80 (1950)
species :
Cousinia meyqanensis
Rastegar, Attar & Mirtadz.; Ann. Bot. Fenn. 61(1): 98 (2024)
Distribution: Iran (NE-Iran: Semnan)
34 IRN
species :
Cousinia microcarpa
Boiss.; Diagn. Pl. Orient., ser. 2, 3: 59 (1856)
Distribution: Kazakhstan; Turkmenistan; Uzbekistan; Kyrgyzstan; Tajikistan; Iran (E-Iran, NE-Iran: Mts.); Afghanistan (Badakshan, Badghys, Baghlan, Balkh, Bamyan, Farah, Faryab, Ghorat, Herat, Jawzjan / Sar-e-Pol, Kabul, Orozgan / Daykundi, Paktia / Khost, Parwan, Qunduz, Samangan, Takhar); Pakistan (Waziristan, Kurram, Pakistani Punjab); NW-India (Jammu & Kashmir)
32 KAZ KGZ TKM TZK UZB 34 AFG IRN 40 IND PAK
Synonyms: hidden
= Arctium microcarpum (Boiss.) Kuntze; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 197 (1887)
= Cousinia acanthoptera Rech.f.; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 72 (1955)
= Cousinia hystrix C.B.Clarke; Compos. Ind. 214 (1876)
= Cousinia xeranthemoides Bunge; Boiss., Fl. Orient. 3: 497 (1865)
species :
Cousinia microcephala
C.A.Mey.; Prodr. [A. P. de Candolle] 6: 552 (1838)
Distribution: Armenia; Azerbaijan (incl. Nachichevan); Iran (Iranian Aserbaijan)
33 ARM AZE 34 IRN
Synonyms: hidden
= Arctium microcephalum (C.A.Mey.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Cousinia grossheimii Bornm.; Oesterr. Bot. Z. 75: 235 (1926)
= Cousinia microcephala var. glabrescens Turrill; (1930)
species :
Cousinia millefontana
Rech.f.; Fl. Iranica 90: 248 (1972)
Distribution: Iran (Iranian Aserbaijan, W-Iran)
34 IRN
species :
Cousinia mindshelkensis
B.Fedtsch.; Repert. Spec. Nov. Regni Veg. 10: 164 (1911)
Distribution: Kazakhstan
32 KAZ
species :
Cousinia minkwitziae
Bornm.; Beih. Bot. Centralbl. 34: 11. 158 (1916)
Distribution: Kazakhstan; Kyrgyzstan
32 KAZ KGZ
species :
Cousinia minuticapitata
Attar; Taxon. Biosyst. 3(6): 66 (2011)
Distribution: Iran
34 IRN
species :
Cousinia miserabilis
Rech.f.; Fl. Iranica 90: 171 (1972)
Distribution: Afghanistan (Ghorat)
34 AFG
species :
Cousinia moabitica
Bornm. & Nábelek; Bornm., Oesterr. Bot. Z. 63: 60 (1913)
Distribution: Jordania (S-Jordania)
34 JOR
species :
Cousinia mobayenii
Ghahr. & Attar; Iranian J. Bot. 8(2): 260 (2000)
Distribution: Iran
34 IRN
species :
Cousinia mogensii
Rech.f.; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 107 (1955)
Distribution: Afghanistan (Bamyan, Ghazni)
34 AFG
species :
Cousinia mogoltavica
Tscherneva & Vved.; Not. Syst. Herb. Inst. Bot. Acad. Sci. . 16: 55 (1961)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia mollis
Schrenk; Bull. Phys-Math. Acad. Petersb. 3: 108 (1845)
Distribution: Kazakhstan; Turkmenistan; Uzbekistan; Kyrgyzstan; Tajikistan; Afghanistan (Samangan)
32 KAZ KGZ TKM TZK UZB 34 AFG
Synonyms: hidden
= Arctium molle (Schrenk) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia monocephala
Bunge; Mém. Acad. Sci. Pétersb. 7 sér., 9 2: 52 (1865)
Distribution: Iran (NE-Iran: Mts.)
34 IRN
Synonyms: hidden
= Arctium monocephalum (Bunge) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia mozaffarianii
Attar, Assadi & Ghahr.; Pakistan J. Bot. 32(2): 293 (2000)
Distribution: Iran
34 IRN
species :
Cousinia mozdouranensis
Djavadi & Attar; Feddes Repert. 116(5-6): 287 (2005), nom. inval., et Phytotaxa 227(1): 99 (2015)
Distribution: Iran
34 IRN
species :
Cousinia mucida
Kult. ex Tscherneva; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Uzb. SSR 17: 97 (1962)
Distribution: Turkmenistan; Uzbekistan
32 TKM UZB
species :
Cousinia mulgediifolia
Bornm.; Beih. Bot. Centralbl. 34: 11. 139 (1916)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia multiloba
DC.; Prodr. [A. P. de Candolle] 6: 554 (1838)
Distribution: Turkmenistan; Tajikistan; Iran (EC-Iran, NE-Iran: Mts., N-Iran, S-Iran, W-Iran); Afghanistan (Badghys, Bamyan, Ghazni, Ghorat, Jawzjan / Sar-e-Pol, Kabul, Logar, Wardak, Orozgan / Daykundi, Paktia / Khost, Parwan); Pakistan (Kurram, Baluchistan, Chitral)
32 TKM TZK 34 AFG IRN 40 PAK
Synonyms: hidden
= Arctium multilobum (DC.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Cousinia elegans Aitch. & Hemsl.; J. Linn. Soc., Bot. 19: 172 (1882)
= Cousinia gnaphalodes Bornm.; Repert. Spec. Nov. Regni Veg. 46: 281 (1939)
= Cousinia multiloba var. afghanorum Bornm.
= Cousinia palmatiloba Jaub. & Spach; Illustr. 2: 80, t. 162 (1846)
species :
Cousinia murgabica
Tscherneva; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Uzb. SSR 17: 97 (1962)
Distribution: Turkmenistan
32 TKM
species :
Cousinia mutehensis
Rech.f.; Fl. Iranica 139a: 129, t. 122 (1979)
Distribution: Iran (EC-Iran)
34 IRN
species :
Cousinia myrioglochis
Rech.f.; Fl. Iranica 90: 57 (1972)
Distribution: Afghanistan (Baghlan, Parwan)
34 AFG
species :
Cousinia myriolepis
Rech.f. & Köie; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 124 (1955)
Distribution: Afghanistan (Logar, Paktia / Khost, Parwan)
34 AFG
species :
Cousinia myriotoma
Rech.f. & Köie; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 126 (1955)
Distribution: Iran (NE-Iran: Mts.); Afghanistan (Farah, Herat)
34 AFG IRN
species :
Cousinia nabelekii
Bornm.; Oesterr. Bot. Z. 63: 59, t. 1, f. 5, 6 (1913)
Distribution: Turkey (E-Anatolia, SE-Anatolia)
34 TUR
species :
Cousinia nana
Attar, Ghahr. & Assadi; Nordic J. Bot. 20(6): 698 (2000 publ. 2001)
Distribution: Iran
34 IRN
species :
Cousinia neglecta
Juz.; Not. Syst. Herb. Hort. Petrop. 4: 78 (1923)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia neubaueri
Rech.f.; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 128 (1955)
Distribution: Afghanistan (Bamyan)
34 AFG
Synonyms: hidden
= Tiarocarpus neubaueri (Rech.f.) Rech.f.; Anz. Österr. Akad. Wiss., Mat.-Naturwiss., 108: 6 (1971 publ. 1972)
species :
Cousinia neurocentra
Bunge; Mém. Acad. Sci. Pétersb. 7 ser., 9 2: 29 (1865)
Distribution: Iran (EC-Iran, E-Iran)
34 IRN
Synonyms: hidden
= Arctium neurocentrum (Bunge) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia newesskyana
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 11: 333 (1896)
Distribution: Uzbekistan
32 UZB
species :
Cousinia ninae
Juz.; Acta Inst. Bot. Acad. Sci. U.R.S.S., ser. I. Fasc. 3, 315 (1937)
Distribution: Kazakhstan; Kyrgyzstan
32 KAZ KGZ
species :
Cousinia niveocoronata
C.Jeffrey ex Grey-Wilson; Kew Bull. 29(1): 50 (1974)
Distribution: Afghanistan (Balkh)
34 AFG
species :
Cousinia noeana
Boiss.; Diagn. Pl. Orient., ser. 2, 3: 57 (1856)
Distribution: Iran (SW-Iran)
34 IRN
Synonyms: hidden
= Arctium noeanum (Boiss.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia novissima
Rech.f.; Fl. Iranica 90: 310 (1972)
Distribution: Afghanistan (Ghazni, Ghorat)
34 AFG
species :
Cousinia nujianensis
Attar, Ghahr., Saber & Zarre; Iranian J. Bot. 11(1): 66 (2005)
Distribution: Iran
34 IRN
species :
Cousinia odontolepis
DC.; Prodr. [A. P. de Candolle] 6: 556 (1838)
subspecies :
subsp. odontolepis
Distribution: Iraq (NE-Iraq, NW-Iraq); Iran (EC-Iran, Iranian Aserbaijan, W-Iran)
34 IRN IRQ
Synonyms: hidden
= Arctium cymbolepis (Boiss.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Arctium odontolepis (DC.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Cousinia acanthophysa Rech.f.; Fl. Iranica 90: 227, t. 115 (1972)
= Cousinia baueri Bornm. & Nábelek ex Bornm.; Oesterr. Bot. Z. 63: 61 (1913)
= Cousinia cymbolepis Boiss.; Diagn. Pl. Orient., ser. 1, 6: 119 (1846)
= Cousinia leatherdalei Rech.f.; Oesterr. Bot. Z. 100: 462, f. 3-4 (1953)
= Cousinia mazu-shirinensis Rech.f.; Fl. Iranica 90: 226, t. 114 (1972)
= Cousinia odontolepis var. leiolepis Bornm.; Repert. Spec. Nov. Regni Veg. 46: 282 (1939)
= Cousinia schultziana Jaub. & Spach; Illustr. 2: 95, t. 177 (1846)
subspecies :
subsp. kurdica
(C.Winkl. & Bornm.) Mehregan; Willdenowia 38(2): 315 (2008)
Distribution: Iraq (NE-Iraq)
34 IRQ
Synonyms: hidden
= Cousinia carduchorum C.Winkl. & Bornm.; Bull. Herb. Boiss., ser. 1, 3: 568, t. 15 (1895)
= Cousinia carduchorum var. major C.Winkl. & Bornm.; Bull. Herb. Boissier 3: 569 (1895)
= Cousinia carduchorum var. minor C.Winkl. & Bornm.; Bull. Herb. Boissier 3: 568 (1895), nom. inval.
= Cousinia kurdica C.Winkl. & Bornm.; Bull. Herb. Boiss. 3: 567, t. 13 (1895)
= Cousinia qandilica Rech.f.; Fl. Iranica 90: 223 (1972)
species :
Cousinia olgae
Regel & Schmalh.; Regel, Descr. Pl. Nov. Fedtsch., Izv. Obsh. Ljub. Estv. Antrop. Etnogr. 34 2: 48 (1882)
Distribution: Turkmenistan; Uzbekistan; Kyrgyzstan; Tajikistan; Iran (E-Iran); Afghanistan (Badghys, Baghlan, Balkh, Faryab, Herat, Jawzjan / Sar-e-Pol)
32 KGZ TKM TZK UZB 34 AFG IRN
species :
Cousinia oligocephala
Boiss.; Diagn. Pl. Orient., ser. 1, 6: 116 (1846)
Distribution: Iran (SW-Iran)
34 IRN
Synonyms: hidden
= Arctium oligocephalum (Boiss.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia olivieri
DC.; Prodr. [A. P. de Candolle] 6: 552 (1838)
Distribution: ?Iran
Synonyms: hidden
= Arctium olivieri (DC.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Cousinia pyrrhocephala Jaub. & Spach; Ill. Pl. Orient. 2(17-18): 85 (1846)
species :
Cousinia omissa
Tscherneva; Novosti Sist. Vyssh. Rast. 30: 155 (1996)
Distribution: Kyrgyzstan
32 KGZ
species :
Cousinia omphalodes
Tscherneva; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Uzb. SSR 17: 98 (1962)
Distribution: Kyrgyzstan
32 KGZ
species :
Cousinia onopordioides
Ledeb.; Eichw., Pl. Nov. It. Caspio-Caucasus 40 (1833)
Distribution: Kazakhstan; Turkmenistan; Northern Caucasus; Azerbaijan (incl. Talysh); Iran (EC-Iran, E-Iran, NE-Iran: Mts., N-Iran); Afghanistan (Ghazni, Logar, Paktia / Khost, Parwan); Pakistan (Baluchistan)
32 KAZ TKM 33 AZE NCS 34 AFG IRN 40 PAK
Synonyms: hidden
= Arctium albicaule (Boiss. & Buhse) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Arctium grandiceps (Bunge) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Arctium karelinii (Less. ex Kar.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Arctium polyacanthum (Eichw.) Kuntze; Revis. Gen. Pl. 1: 367 (1891)
= Cousinia albicaulis Boiss. & Buhse; Nouv. Mém. Soc. Nat. Moscou 12 18: 128, t. 37 (1860)
= Cousinia bornmuelleri C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 14: 235 (1897)
= Cousinia grandiceps Bunge; Mém. Acad. Sci. Pétersb. 7 sér., 9 2: 54 (1865)
= Cousinia karelinii Less. ex Kar.; Bull. Soc. Imp. Naturalistes Moscou 13: 159 (1839)
= Cousinia onopordioides var. glabra Assadi & Mehregan; Fl. Iran [ed. M. Assadi et al.] 125: 224 (2017)
= Cousinia subzevarensis Rech.f.; Fl. Iranica 139a: 134 (1979)
= Onopordum polyacanthum Eichw.; Pl. Casp.-Caucas.: 3 (1831), nom. nud.
species :
Cousinia onopordon
Freyn & Sint.; Oesterr. Bot. Z. 42: 207 (1892)
Distribution: ?Transcaucasus
species :
Cousinia oophora
Rech.f.; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 133 (1955)
Distribution: Afghanistan (Faryab)
34 AFG
species :
Cousinia oopoda
Juz.; Acta Inst. Bot. Acad. Sci. U.R.S.S., ser. I. Fasc. 3, 310 (1937)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia oreodoxa
Bornm. & Sint.; Mitt. Thür. Bot. Ver. 23: 99 (1908)
Distribution: Turkmenistan; Iran (NE-Iran: Mts.)
32 TKM 34 IRN
species :
Cousinia oreoxerophila
Kult.; Fl. Turkm. 7: 380 (1960)
Distribution: Turkmenistan
32 TKM
species :
Cousinia orientalis
(Adams) K.Koch; Linnaea 24: 386 (1851)
Distribution: Northern Caucasus; Georgia [Caucasus]; Azerbaijan
33 AZE GEO NCS
Synonyms: hidden
= Arctium carduiforme (Cass.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Arctium orientale (Adams) Kuntze; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 197 (1887)
= Carduus orientalis Adams; Weber & Mohr, Beitr. 1: 63 (1805)
= Cousinia carduiformis Cass.; Dict. Sci. Nat. 47: 503 (1827)
= Cousinia orientalis (Adams) Grossh.; Grossh. & Schischk., Sched. Pl. Or. Exsicc. Fasc. X. 14 (1928)
= Cousinia orientalis (Adams) O.Hoffm.; Nat. Pflanzenfam. [Engler & Prantl] IV, 5: 319 (1890)
species :
Cousinia orthacantha
Tscherneva; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Uzb. SSR 17: 98 (1962)
Distribution: Uzbekistan; Tajikistan; Afghanistan (Balkh, Qunduz, Samangan)
32 TZK UZB 34 AFG
species :
Cousinia orthoclada
Hausskn. & Bornm.; Beih. Bot. Centralbl. 20 2: 159 (1906)
Distribution: Iran (S-Iran, SW-Iran)
34 IRN
Synonyms: hidden
= Cousinia gracilis var. erecto-patula Parsa; (1948)
= Cousinia orthoclada var. subappendiculata Bornm.; (1914)
= Cousinia orthoclada var. subrecurvata Bornm.; (1917)
species :
Cousinia ortholepis
Juz. ex Tscherneva; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Uzb. SSR 17: 99 (1962)
Distribution: Uzbekistan
32 UZB
species :
Cousinia orthoneura
Rech.f.; Fl. Iranica 90: 120 (1972)
Distribution: Afghanistan (Laghman, Parwan)
34 AFG
species :
Cousinia oshtorankuhensis
Attar; Iranian J. Bot. 17(2): 155 (2011)
Distribution: Iran
34 IRN
species :
Cousinia ottonis
Bornm.; Oesterr. Bot. Z. 62: 106, t. 3, f. 484a (1912)
Distribution: Iran (SW-Iran)
34 IRN
species :
Cousinia outichaschensis
Franch.; Ann. Sci. Nat. Ser. VI. 16: 322 (1883)
Distribution: Turkmenistan; Uzbekistan; Tajikistan
32 TKM TZK UZB
Synonyms: hidden
= Arctium outichaschense (Franch.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Cousinia commixta Kult.; Trudy Sredne-Aziatsk. Gosud. Univ., ser. 8b, Bot. 6: 18 (1929)
= Cousinia outichaschensis f. commixta (Kult.) Kamelin; Fl. Ushchel. Reki Varzob 343 (2021)
= Cousinia outichaschensis f. glauciifolia (Kult.) Kamelin; Fl. Ushchel. Reki Varzob 343 (2021)
species :
Cousinia ovczinnikovii
Tscherneva; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Uzb. SSR 17: 99 (1962)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia oxiana
Tscherneva; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Uzb. SSR 17: 100 (1962)
Distribution: Turkmenistan; Uzbekistan; Tajikistan; Iran (NE-Iran: Mts.); Afghanistan (Badghys, Balkh, Farah, Faryab)
32 TKM TZK UZB 34 AFG IRN
species :
Cousinia oxytoma
Rech.f.; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 135 (1955)
Distribution: Tajikistan (Petr I Mts.); Afghanistan (Badakshan, Baghlan, Kunar / Nuristan); Pakistan (Chitral)
32 TZK 34 AFG 40 PAK
Synonyms: hidden
= Cousinia batalinii var. virescens Bornm.; Beih. Bot. Centralbl. 34(2): 168 (1916)
species :
Cousinia panjshirensis
Rech.f.; Fl. Iranica 90: 128 (1972)
Distribution: Afghanistan (Parwan)
34 AFG
species :
Cousinia pannosa
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 578 (1887)
Distribution: Tajikistan
32 TZK
Synonyms: hidden
= Cousinia resupinata Juz.; Trudy Bot. Inst. Akad. Nauk S.S.S.R., ser. 1, Fl. Sist. Vyssh. Rast. 3: 318 (1937)
species :
Cousinia pannosiformis
Tscherneva; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Uzb. SSR 17: 101 (1962)
Distribution: Kyrgyzstan; Tajikistan
32 KGZ TZK
species :
Cousinia papillosa
Djavadi & Attar; Rostaniha 8(2): 64 (2007)
Distribution: Iran
34 IRN
species :
Cousinia parjumanensis
Rech.f.; Fl. Iranica 90: 101 (1972)
Distribution: Afghanistan (Farah, Ghorat)
34 AFG
species :
Cousinia parsana
Ghahr., Iranshahr & Attar; Iranian J. Bot. 8(1): 16 (1999)
Distribution: Iran
34 IRN
species :
Cousinia parviceps
Rech.f. & Köie; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 136 (1955)
Distribution: Afghanistan (Badghys, Bamyan, Faryab, Ghazni, Ghorat, Herat, Wardak, Orozgan / Daykundi, Paktia / Khost, Parwan)
34 AFG
species :
Cousinia parviziana
Parsa; Flore de l´Iran (Publ. Univ. Teheran 1777) 10: 110 (1980)
Distribution: Iran
34 IRN
species :
Cousinia pasargardensis
Attar; Iranian J. Bot. 16(1): 139 (2010)
Distribution: Iran
34 IRN
species :
Cousinia patentispina
Tscherneva; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Uzb. SSR 17: 101 (1962)
Distribution: Kazakhstan; Turkmenistan; Uzbekistan
32 KAZ TKM UZB
species :
Cousinia pauciramosa
Kult.; Not. Syst. Herb. Hort. Petrop. 5: 139 (1924)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia pectinata
Rech.f.; Ann. Naturhist. Mus. Wien 57: 76 (1950)
Distribution: Afghanistan (Kabul)
34 AFG
Synonyms: hidden
= Cousinia gillii Rech.f.; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 99 (1955)
species :
Cousinia peduncularis
Juz. ex Tscherneva; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Uzb. SSR 17: 101 (1962)
Distribution: Tajikistan
32 TZK
species :
Cousinia pergamacea
Boiss. & Hausskn.; Boiss., Fl. Orient. 3: 513 (1875)
subspecies :
subsp. pergamacea
Distribution: Iraq (NE-Iraq); Iran (W-Iran)
34 IRN IRQ
Synonyms: hidden
= Arctium pergamaceum (Boiss. & Hausskn.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Cousinia wheeler-hainesii Rech.f.; Anz. Österr. Akad. Wiss., Math.-Naturwiss. Kl. 101: 353 (1964)
subspecies :
subsp. sardashtensis
(Rech.f.) Mehregan; Willdenowia 38(2): 331 (2008)
Distribution: Iran (Iranian Aserbaijan)
34 IRN
Synonyms: hidden
= Cousinia sardashtensis Rech.f.; Fl. Iranica 139a: 139, t. 132 (1979)
species :
Cousinia perovskiensis
(Bornm.) Juz. ex Tscherneva; Kom., Fl. U.R.S.S. 27: 161 (1962)
Distribution: Kazakhstan
32 KAZ
Synonyms: hidden
= Cousinia dissecta var. perovskiensis Bornm.
species :
Cousinia persarum
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 12: 227 (1893)
Distribution: Iran (S-Iran)
34 IRN
species :
Cousinia persica
Djavadi & Attar; Feddes Repert. 117(7-8): 455 (2006)
Distribution: Iran
34 IRN
species :
Cousinia perspolitana
Attar & Ghahr.; Nordic J. Bot. 23(5): 589 (2005)
Distribution: Iran
34 IRN
species :
Cousinia pestalozzae
Boiss.; Fl. Orient. 3: 471 (1875)
Distribution: Lebanon (Antilebanon); Syria (C-Syrian Desert)
34 LEB SYR
Synonyms: hidden
= Arctium pestalozzae (Boiss.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia phyllocephala
Bornm. & Gauba; Repert. Spec. Nov. Regni Veg. 36: 330, t. 181 (1934)
Distribution: Iran (SW-Iran)
34 IRN
Synonyms: hidden
= Cousinia koieana Bornm.; Danish Sci. Invest. Iran 4: 26 (1945)
= Cousinia koeieana var. adenoloba Bornm.; (1945)
species :
Cousinia pichleriana
Bornm.; Repert. Spec. Nov. Regni Veg. 46: 282 (1939), in adnot., et ex Rech.f., Fl. Iranica 139a: 141, t. 135 (1979)
Distribution: Iran (SW-Iran)
34 IRN
Synonyms: hidden
= Cousinia ecbatanensis f. inermis Bornm.
species :
Cousinia pinarocephala
Boiss.; Diagn. Pl. Orient., ser. 1, 6: 121 (1846)
Distribution: Iran (EC-Iran, N-Iran, Iranian Aserbaijan)
34 IRN
Synonyms: hidden
= Arctium pinarocephalum (Boiss.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Cousinia declinata Parsa; Kew Bull. 3(2): 204 (1948)
= Cousinia pinarocephala var. totschalensis Bornm.; (1907)
species :
Cousinia pineticola
Rech.f. & Gilli; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 138 (1955)
Distribution: Afghanistan (Baghlan, Ghazni, Laghman, Orozgan / Daykundi, Paktia / Khost, Parwan); Pakistan (Chitral)
34 AFG 40 PAK
species :
Cousinia piptocephala
Bunge; Mém. Acad. Sci. Petersb. 7 sér., 9 2: 26 (1865)
Distribution: Iran (EC-Iran, NE-Iran: Mts., S-Iran, SW-Iran)
34 IRN
Synonyms: hidden
= Arctium piptocephalum (Bunge) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia platyacantha
Bunge; Mém. Acad. Sci. Petersb. 7 sér., 9 2: 37 (1865)
Distribution: Iran (NE-Iran: Mts.)
34 IRN
Synonyms: hidden
= Cousinia centauroides Regel; Bull. Soc. Imp. Naturalistes Moscou 40: (1867) II. 152
species :
Cousinia platylepis
Schrenk ex Fisch. & C.A.Mey.; Index Sem. (St. Petersburg) 9: 10 (1843)
Distribution: Kazakhstan; Uzbekistan; Kyrgyzstan; Tajikistan; China (NW-Xinjiang)
32 KAZ KGZ TZK UZB 36 CHI
Synonyms: hidden
= Arctium platylepis (Schrenk ex Fisch. & C.A.Mey.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia platyptera
Bornm. & C.Winkl.; Oesterr. Bot. Z. 62: 181, t. 3, f. 1 (1912)
Distribution: Iran (S-Iran)
34 IRN
species :
Cousinia platyraphis
Kult.; Acta Univ. As. Med., ser. 8b 7: 1 (1929)
Distribution: Iran (E-Iran)
34 IRN
species :
Cousinia platystegia
Tscherneva; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Uzb. SSR 17: 103 (1962)
Distribution: Uzbekistan
32 UZB
species :
Cousinia podlechii
Rech.f.; Anz. Österr. Akad. Wiss., Math.-Nat., ciii. 286 (1966)
Distribution: Afghanistan (Baghlan, Parwan)
34 AFG
species :
Cousinia podophylla
Tscherneva; Not. Syst. Herb. Inst. Bot. Acad. Sci. . 15: 43 (1959)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia polycephala
Rupr.; Mem. Acad. Imp. Sci. Saint Petersbourg (Ser. 7) 14(4): 54 (1869)
Distribution: Kazakhstan; Turkmenistan; Uzbekistan; Kyrgyzstan; Tajikistan; China (Xinjiang)
32 KAZ KGZ TKM TZK UZB 36 CHI
Synonyms: hidden
= Arctium polycephalum (Rupr.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia polyneura
Rech.f.; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 140 (1955)
Distribution: Afghanistan (Kabul)
34 AFG
species :
Cousinia polytimetica
Tscherneva; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Uzb. SSR 17: 103 (1962)
Distribution: Uzbekistan
32 UZB
species :
Cousinia porphyrochrysea
Rech.f.; Fl. Iranica 90: 278 (1972)
Distribution: Afghanistan (Badakshan, Takhar)
34 AFG
species :
Cousinia porphyrostephana
Rech.f.; Fl. Iranica 90: 266 (1972)
Distribution: Afghanistan (Baghlan, Parwan)
34 AFG
species :
Cousinia postiana
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 12: 223 (1892)
Distribution: Lebanon (Antilebanon); Syria (C-Syrian Desert); Jordania (S-Jordania)
34 JOR LEB SYR
species :
Cousinia praestans
Tscherneva & Vved.; Not. Syst. Herb. Inst. Bot. Acad. Sci. . 16: 57 (1961)
Distribution: Uzbekistan
32 UZB
species :
Cousinia prasina
Jaub. & Spach; Ill. Pl. Or. 2: 86, t. 168 (1846)
Distribution: Iran (EC-Iran, W-Iran)
34 IRN
Synonyms: hidden
= Arctium prasinum (Jaub. & Spach) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia princeps
Franch.; Ann. Sci. Nat. Ser. VI. 16: 324 (1888)
Distribution: Tajikistan
32 TZK
Synonyms: hidden
= Arctium princeps (Franch.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia prolifera
Jaub. & Spach; Ill. Pl. Or. 2: 75, t. 157 (1845)
Distribution: Kazakhstan; Turkmenistan; Uzbekistan; Tajikistan; Iraq (NW-Iraq); Iran (EC-Iran, E-Iran, NE-Iran: Mts., Iranian Aserbaijan, S-Iran); Afghanistan (Balkh, Farah, Helmand, Herat, Kabul, Kandahar, Kunar / Nuristan, Nangarhar, Paktia / Khost, Parwan); Pakistan (Baluchistan, Waziristan, Kurram, Pakistani Punjab, Peshawar, Jhelum, Rawalpindi, Swat, Hazara); NW-India (Jammu & Kashmir, Rajasthan); Myanmar
32 KAZ TKM TZK UZB 34 AFG IRN IRQ 40 IND PAK 41 MYA
Synonyms: hidden
= Arctium minutum (Boiss.) Kuntze; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 197 (1887)
= Arctium proliferum (Jaub. & Spach) Kuntze; Revis. Gen. Pl. 1: 367 (1891)
= Cousinia aralensis Bunge; Mem. Sav. Etr. Petersb. 7: 354 (1851), et Beitr. Fl. Russl.: 178 (1852)
= Cousinia calcitrapiformis Jaub. & Spach; Illustr. 2: 76, t. 158 (1845)
= Cousinia minuta Boiss.; Diagn. Pl. Orient., ser. 1, 6: 117 (1846)
species :
Cousinia proxima
Juz.; Acta Inst. Bot. Acad. Sci. U.R.S.S., ser. I. Fasc. 3, 301 (1937)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia psammophila
Kult.; Acta Univ. As. Med. Ser. VIII. Bot. Fasc. VI. 15 (1929)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia pseudactinia
Rech.f.; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 142 (1955)
Distribution: Afghanistan (Samangan)
34 AFG
species :
Cousinia pseudaffinis
Kult.; Trans. Sci. Soc. Turk. 1: 27 (1923), et in Acta Univ. As. Med. Ser. VIII b, Bot. Fasc. 3, 105 (1928)
Distribution: Kazakhstan; Uzbekistan; Kyrgyzstan
32 KAZ KGZ UZB
species :
Cousinia pseudobonvalotii
Juz.; Trudy sk. Bazy 8: 525 (1940)
Distribution: ?Turkmenistan; Uzbekistan; Kyrgyzstan; Tajikistan
32 KGZ TZK UZB
species :
Cousinia pseudocandolleana
Assadi; Iranian J. Bot. 15(1): 42 (2009)
Distribution: Iran
34 IRN
species :
Cousinia pseudocirsium
Rech.f.; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 144 (1955)
Distribution: Afghanistan (Badakshan)
34 AFG
species :
Cousinia pseudodshizakensis
Tschern. & Vved.; Not. Syst. Herb. Inst. Bot. Acad. Sci. . 16: 59 (1961)
Distribution: Uzbekistan; Kyrgyzstan; Tajikistan
32 KGZ TZK UZB
species :
Cousinia pseudolanata
Popov ex Tscherneva; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Uzb. SSR 17: 104 (1962)
Distribution: Uzbekistan
32 UZB
species :
Cousinia pseudomollis
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 470 (1887)
Distribution: Kazakhstan; Uzbekistan; Kyrgyzstan (Pamir-Alai, Tianshan); Tajikistan; Afghanistan (Badakshan, Baghlan, Balkh, Samangan, Takhar)
32 KAZ KGZ TZK UZB 34 AFG
Synonyms: hidden
= Arctium pseudomolle (C.Winkl.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Cousinia pseudomollis var. platyacantha Bornm.; (1916)
species :
Cousinia pseudostenolepis
Rech.f. & Edelb.; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 147 (1955)
Distribution: Afghanistan (Badakshan, Baghlan, Bamyan, Orozgan / Daykundi, Samangan, Takhar)
34 AFG
species :
Cousinia pterocarpa
Boiss.; Fl. Orient. 3: 501 (1875)
Distribution: Afghanistan (Ghazni, Wardak)
34 AFG
Synonyms: hidden
= Arctium pterocarpum (Boiss.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia pterocaulos
(C.A.Mey.) Rech.f.; Fl. Iranica 90: 95 (1972)
Distribution: Azerbaijan (Talysh); Iran (N-Iran, Iranian Aserbaijan, W-Iran)
33 AZE 34 IRN
Synonyms: hidden
= Arctium hystrix (C.A.Mey.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Carthamus pterocaulos C.A.Mey.; Verz. Pfl. Caucasus 67 (1831)
= Cousinia hystrix C.A.Mey.; Ind. Sem. Horti Petrop. 2: 34 (1835)
species :
Cousinia pugionifera
Jaub. & Spach; Ill. Pl. Or. 2: 92, t. 174 (1846)
Distribution: Iran (EC-Iran, SW-Iran)
34 IRN
Synonyms: hidden
= Arctium pugioniferum (Jaub. & Spach) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia pulchella
Bunge; Del. Sem. Hort. Dorpat. (1845) 3
Distribution: Uzbekistan; Kyrgyzstan; Tajikistan (Pamir-Alai); Afghanistan (Badakshan, Baghlan, Orozgan / Daykundi)
32 KGZ TZK UZB 34 AFG
Synonyms: hidden
= Arctium pulchellum (Bunge) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia pulcherantha
Attar & Mirtadz.; Nordic J. Bot. 32(3): 281 (2014)
Distribution: Iran
34 IRN
species :
Cousinia pulchra
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 478 (1887)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
Synonyms: hidden
= Arctium pulchrum (C.Winkl.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia pulvinaris
Rech.f. & Köie; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 147 (1955)
Distribution: Afghanistan (Farah)
34 AFG
species :
Cousinia pungens
Juz.; Trans. Rubb. & Guttap. Inst., Fasc. 4, 19 (1932), et Acta Inst. Bot. Acad. Sci. U.R.S.S., ser. I. Fasc. 3, 306, 308 (1937)
Distribution: Uzbekistan; Kyrgyzstan
32 KGZ UZB
species :
Cousinia purpurea
C.A.Mey.; Prodr. [A. P. de Candolle] 6: 555 (1838)
Distribution: Northern Caucasus; Armenia; Azerbaijan (incl. Nachichevan); Iran (Iranian Aserbaijan)
33 ARM AZE NCS 34 IRN
Synonyms: hidden
= Arctium purpureum (C.A.Mey.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia pusilla
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 468 (1887)
Distribution: Turkmenistan; Uzbekistan; Tajikistan
32 TKM TZK UZB
Synonyms: hidden
= Arctium pusillum (C.Winkl.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia pycnocephala
Rech.f.; Fl. Iranica 90: 195 (1972)
Distribution: Tajikistan; Afghanistan (Faryab)
32 TZK 34 AFG
species :
Cousinia pycnoloba
Boiss.; Diagn. Ser. II. 3: 57 (1856)
Distribution: Afghanistan (Ghazni, Wardak, Orozgan / Daykundi, Paktia / Khost); Pakistan (Baluchistan, Kurram)
34 AFG 40 PAK
Synonyms: hidden
= Cousinia scala Aitch. & Hemsl.; J. Linn. Soc., Bot. 19: 172 (1882)
species :
Cousinia pygmaea
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 467 (1887)
Distribution: Turkmenistan; Uzbekistan; Tajikistan; Afghanistan (Baghlan, Ghazni)
32 TKM TZK UZB 34 AFG
Synonyms: hidden
= Arctium pygmaeum (C.Winkl.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia qaisarensis
Rech.f.; Fl. Iranica 90: 261 (1972)
Distribution: Afghanistan (Faryab)
34 AFG
species :
Cousinia qaradaghensis
Rech.f.; Fl. Iranica 90: 258 (1972)
Distribution: Armenia; Iran (Iranian Aserbaijan)
33 ARM 34 IRN
species :
Cousinia qarehbilensis
Rech.f.; Fl. Iranica 139a: 131, t. 123 (1979)
Distribution: Iran (N-Iran)
34 IRN
species :
Cousinia quettensis
Rech.f.; Anz. Österr. Akad. Wiss., Math.-Nat., ciii. 287 (1966)
Distribution: S-Afghanistan; Pakistan
34 AFG 40 PAK
species :
Cousinia racemosa
Boiss.; Diagn. Ser. II. 3: 56 (1856)
Distribution: Afghanistan (Bamyan, Ghorat, Kabul, Wardak, Parwan); Pakistan (Chitral, Drosh)
34 AFG 40 PAK
Synonyms: hidden
= Arctium racemosum (Boiss.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia raddeana
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: (1887) 473 in obs., et 11: (1889) 132 t. 1
Distribution: Turkmenistan; Tajikistan (S-Pamir-Alai); Afghanistan (Badghys, Balkh, Faryab, Samangan)
32 TKM TZK 34 AFG
Synonyms: hidden
= Arctium raddeanum (C.Winkl.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Cousinia nobilis Rech.f.; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 132 (1955)
species :
Cousinia radians
Bunge; Del. Sem. Hort. Dorpat. (1845) 3
Distribution: Turkmenistan; Uzbekistan; Kyrgyzstan; Tajikistan
32 KGZ TKM TZK UZB
Synonyms: hidden
= Arctium radians (Bunge) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Arctium stenolepis (Regel & Schmalh.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Cousinia interrupta C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 12: 229 (1892)
= Cousinia stenolepis Regel & Schmalh.; Izv. Imp. Obshch. Lyubit. Estestv. Moskovsk. Univ. 34(2): 49 (1882)
species :
Cousinia ramosissima
DC.; Predr. 6: 552 (1838)
Distribution: Turkey (Inner Anatolia, S-Anatolia); Lebanon (Antilebanon, C-Lebanon); Syria (C-Syrian Desert, NW-Syria, Jbel Druze)
34 LEB SYR TUR
Synonyms: hidden
= Arctium ramosissimum (DC.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Cousinia drusorum Bornm. & Sam.; Repert. Spec. Nov. Regni Veg. 40: 342 (1936)
species :
Cousinia ramulosa
Rech.f.; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 149 (1955)
Distribution: Tajikistan (Pamir); Afghanistan (Badakshan, Kunar / Nuristan, Parwan)
32 TZK 34 AFG
species :
Cousinia raphiostegia
Rech.f.; Fl. Iranica 139a: 128, t. 120 (1979)
Distribution: Iran (EC-Iran)
34 IRN
species :
Cousinia rava
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 14: 227 (1895)
Distribution: Tajikistan
32 TZK
species :
Cousinia rawanduzensis
Mehregan; Iranian J. Bot. 17(2): 144 (2011)
Distribution: Iraq
34 IRQ
species :
Cousinia rechingerae
Bornm.; Repert. Spec. Nov. Regni Veg. 48: 133, t. 309, f. 1 (1940)
Distribution: Iran (NE-Iran: Mts., N-Iran)
34 IRN
Synonyms: hidden
= Cousinia ahmed-adlii Bornm. & Gauba; Repert. Spec. Nov. Regni Veg. 49: 263 (1940)
= Cousinia kandewanensis var. golydaghi Parsa; (1948)
species :
Cousinia rechingerorum
Bornm.; Repert. Spec. Nov. Regni Veg. 48: 145, t. 313, f. 1 (1940)
Distribution: Iran (E-Iran)
34 IRN
Synonyms: hidden
= Cousinia malekiana Parsa; Fl. Iran 3: 539 (1949)
species :
Cousinia recurvata
DC.; Prodr. [A. P. de Candolle] 6: 555 (1838)
Distribution: Iran (NE-Iran: Mts., SW-Iran)
34 IRN
Synonyms: hidden
= Arctium recurvatum (DC.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia regelii
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 14: 229 (1895)
Distribution: Kazakhstan; Uzbekistan; Kyrgyzstan; Tajikistan
32 KAZ KGZ TZK UZB
species :
Cousinia renominata
Rech.f.; Fl. Iranica 90: 163 (1972)
Distribution: Iran (NE-Iran: Mts.)
34 IRN
Synonyms: hidden
= Cousinia turkmenorum Bornm. & Gauba; (1939) (nec Bornm. 1916
species :
Cousinia resinosa
Juz.; Trans. Rubb. & Guttap. Inst., Fasc. 4, 24 (1932), et Acta Inst. Bot. Acad. Sci. U.R.S.S., ser. I. Fasc. 3, 306 (1937)
Distribution: Kazakhstan; Turkmenistan; Uzbekistan; Kyrgyzstan; Tajikistan
32 KAZ KGZ TKM TZK UZB
species :
Cousinia rhabdodes
Bornm. & Rech.f.; Repert. Spec. Nov. Regni Veg. 48: 137, t. 310, f. 1 (1940)
Distribution: Iran (EC-Iran)
34 IRN
species :
Cousinia rhaphiocephala
Rech.f.; Fl. Iranica 90: 319 (1972)
Distribution: Iran (SW-Iran)
34 IRN
species :
Cousinia rhodantha
Kult.; Acta Univ. As. Med. Ser. VIII. Bot. Fasc. VI. 22 (1929)
Distribution: Uzbekistan
32 UZB
species :
Cousinia rhombiformis
C.Winkl. & Strauss; Trudy Imp. S.-Peterburgsk. Bot. Sada 14: 232 (1897)
Distribution: Iran (SW-Iran)
34 IRN
species :
Cousinia rigida
Kult.; Acta Univ. As. Med. Ser. VIII. Bot. Fasc. VI. 26 (1929)
Distribution: Kazakhstan; Kyrgyzstan
32 KAZ KGZ
species :
Cousinia rigidissima
Rech.f.; Fl. Iranica 90: 200 (1972)
Distribution: Afghanistan (Ghorat)
34 AFG
species :
Cousinia robusta
Rech.f.; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 151 (1955)
Distribution: Afghanistan (Bamyan)
34 AFG
species :
Cousinia rosea
Kult.; Acta Univ. As. Med. Ser. VIII b, Bot. Fasc. 3, 105 (1928), et Acta Univ. As. Med. Ser. VIII. Bot. Fasc. VI. 17 (1929)
Distribution: Turkmenistan; Uzbekistan
32 TKM UZB
species :
Cousinia rotundifolia
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 579 (1887)
Distribution: Turkmenistan; Uzbekistan; Tajikistan
32 TKM TZK UZB
Synonyms: hidden
= Cousinia kultiasowii Juz.; Trudy Bot. Inst. Akad. Nauk S.S.S.R., ser. 1, Fl. Sist. Vyssh. Rast. 3: 317 (1937)
= Cousinia rubrarenosa Kult.; Trudy Sredne-Aziatsk. Gosud. Univ., ser. 8b, Bot. 6: 18 (1929)
species :
Cousinia rubiginosa
Kult.; Acta Univ. As. Med. Ser. VIII. Bot. Fasc. VI. 23 (1929)
Distribution: Tajikistan
32 TZK
species :
Cousinia rudis
Rech.f.; Fl. Iranica 90: 42 (1972)
Distribution: Afghanistan (Balkh)
34 AFG
species :
Cousinia rufidula
Bornm.; Beih. Bot. Centralbl. 34(2): 270 (1916)
Distribution: Afghanistan (Ghorat, Herat)
34 AFG
species :
Cousinia sabalanica
Attar, Ghahr. & Assadi; Nordic J. Bot. 20(6): 699 (2000 publ. 2001)
Distribution: Iran
34 IRN
species :
Cousinia sabzevarensis
Rech.f.; Fl. Iranica 139a: 134, t. 126 (1979)
Distribution: Iran (NE-Iran: Mts., N-Iran)
34 IRN
species :
Cousinia sagittata
C.Winkl. & Strauss; Trudy Imp. S.-Peterburgsk. Bot. Sada 14: 233 (1897)
subspecies :
subsp. sagittata
Distribution: Iran (SW-Iran)
34 IRN
subspecies :
subsp. iranica
(C.Winkl. & Strauss) Mehregan; Willdenowia 38(2): 328 (2008)
Distribution: Iran (SW-Iran)
34 IRN
Synonyms: hidden
= Cousinia iranica C.Winkl. & Strauss; Trudy Imp. S.-Peterburgsk. Bot. Sada 14: 233 (1897)
species :
Cousinia sahandica
Attar & Djavadi; Iranian J. Bot. 16(1): 161 (2010)
Distribution: Iran
34 IRN
species :
Cousinia sakawensis
Boiss. & Hausskn.; Boiss., Fl. Orient. 3: 489 (1875)
Distribution: Iran (SW-Iran)
34 IRN
Synonyms: hidden
= Arctium sakawense (Boiss. & Hausskn.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia salangensis
Rech.f.; Anz. Österr. Akad. Wiss., Math.-Nat., ciii. 288 (1966)
Distribution: Afghanistan (Baghlan, Parwan)
34 AFG
species :
Cousinia saloukensis
Mehregan; Willdenowia 41(2): 263 (2011)
Distribution: Iran
34 IRN
species :
Cousinia sanandajensis
Rech.f.; Fl. Iranica 90.235, t. 118 (1972)
Distribution: Iran (SW-Iran)
34 IRN
species :
Cousinia sarawschanica
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 9: 424 (1886)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia sarzehensis
Attar, Ghahr. & Assadi; Nordic J. Bot. 20(6): 697 (2000 publ. 2001)
Distribution: Iran
34 IRN
species :
Cousinia scabrida
Juz.; Acta Inst. Bot. Acad. Sci. U.R.S.S., ser. I. Fasc. 3, 306 (1937)
Distribution: Kazakhstan; Kyrgyzstan
32 KAZ KGZ
species :
Cousinia scariosa
Regel; Bull. Soc. Imp. Naturalistes Moscou 40: (1867) II. 154
Distribution: Afghanistan (Ghorat, Jawzjan / Sar-e-Pol, Wardak); Pakistan (Chitral)
34 AFG 40 PAK
Synonyms: hidden
= Arctium scariosum (Regel) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Cousinia lamprophyllaria Rech.f.; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 114 (1955)
species :
Cousinia scheibeana
Bornm.; Repert. Spec. Nov. Regni Veg., Beih. 108: 43 (1938)
Distribution: Afghanistan (Badakshan, Bamyan, Kunar / Nuristan, Parwan); Pakistan (Chitral); NW-India (Jammu & Kashmir)
34 AFG 40 IND PAK
species :
Cousinia schepsaica
Karmysch.; Fl. Kazakhst. 9: 244, 569 (1966)
Distribution: Kazakhstan
32 KAZ
species :
Cousinia schindleriana
Bornm. & Gauba; Mitt. Thür. Bot. Ver. 47: 124, t. 11, f. 2 (1941)
Distribution: Iran (NE-Iran: Mts.)
34 IRN
species :
Cousinia schiraziana
Attar; Iranian J. Bot. 16(1): 139 (2010)
Distribution: Iran
34 IRN
species :
Cousinia schischkinii
Juz.; Acta Inst. Bot. Acad. Sci. U.R.S.S., ser. I. Fasc. 3, 316 (1937)
Distribution: Kyrgyzstan (Tianshan)
32 KGZ
species :
Cousinia schistoptera
Juz.; Acta Inst. Bot. Acad. Sci. U.R.S.S., ser. I. Fasc. 3, 301 (1937)
Distribution: Kazakhstan; Turkmenistan; Uzbekistan
32 KAZ TKM UZB
species :
Cousinia schistosa
Rech.f. & Edelb.; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 154 (1955)
Distribution: Afghanistan (Kunar / Nuristan)
34 AFG
species :
Cousinia schtschurowskiana
Regel & Schmalh. ex Regel; Pl. Nov. Fedsch. 50 (1882)
Distribution: Uzbekistan
32 UZB
Synonyms: hidden
= Arctium schtschurowskianum (Regel & Schmalh.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia scleracantha
Kult. ex Tscherneva; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Uzb. SSR 17: 104 (1962)
Distribution: Uzbekistan; Kyrgyzstan
32 KGZ UZB
species :
Cousinia sclerolepis
C.Shih; Bull. Bot. Res., Harbin 4(2): 60-61, photo. 1 (1984)
Distribution: China (W-Xinjiang)
36 CHI
species :
Cousinia sclerophylla
Juz.; Acta Inst. Bot. Acad. Sci. U.R.S.S., ser. I. Fasc. 3, 308 (1937)
Distribution: Tajikistan
32 TZK
species :
Cousinia sefidiana
(Pau) Rech.f.; An. Jard. Bot. Madrid, 47(2): 365 (1989 publ. (1990)
Distribution: Iran
34 IRN
Synonyms: hidden
= Cousinia cymbolepis var. sefidiana Pau; Trab. Mus. Nac. Ci. Nat., ser. Bot. 14: 40 (1918)
species :
Cousinia seidlitzii
Bunge; Mém. Acad. Sci. Pétersb. 7 sér., 9 2: 20 (1865)
Distribution: Iran (EC-Iran, Iranian Aserbaijan, W-Iran)
34 IRN
Synonyms: hidden
= Arctium seidlitzii (Bunge) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia semilacera
Juz.; Acta Inst. Bot. Acad. Sci. U.R.S.S., ser. I. Fasc. 3, 322 (1937)
Distribution: Tajikistan (Pamir-Alai); Afghanistan (Badakshan)
32 TZK 34 AFG
Synonyms: hidden
= Cousinia jacea Rech.f. & Edelb.; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 104 (1955)
species :
Cousinia semnanensis
Attar, Mirtadz. & Sotoodeh; Nova Biol. Reperta 9(20: 147 (2022)
Distribution: Iran
34 IRN
species :
Cousinia serratuloides
Boiss.; Diagn. Pl. Orient., ser. 1, 6: 118 (1846)
Distribution: Iran
34 IRN
Synonyms: hidden
= Arctium serratulodes (Boiss.) Kuntze; Revis. Gen. Pl. 1: 367 (1891)
= Cousinia olivieri Jaub. & Spach; Illustr. t. 172 (1846) (non DC. 1838)
species :
Cousinia sewerzovii
Regel; Bull. Soc. Imp. Naturalistes Moscou 40: (1867) II. 151 (sewerzowii)
Distribution: Kazakhstan; Uzbekistan; Tajikistan; Afghanistan (Parwan)
32 KAZ TZK UZB 34 AFG
Synonyms: hidden
= Arctium sewerzovii (Regel) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia shahrestanica
Rech.f.; Anz. Österr. Akad. Wiss., Math.-Nat., 105: 7 (1969)
Distribution: Afghanistan (Orozgan / Daykundi)
34 AFG
species :
Cousinia shahuensis
Attar; Taxon. Biosyst. 3(6): 61 (2011)
Distribution: Iran
34 IRN
species :
Cousinia shahvarica
Rech.f.; Ann. Naturhist. Mus. Wien 57: 71 (1950)
Distribution: Iran (EC-Iran, N-Iran)
34 IRN
species :
Cousinia sharifii
Attar & Amini Rad; Phytotaxa 395(4): 287 (2019)
Distribution: Iran
34 IRN
species :
Cousinia shebliensis
Ghahr., Iranshahr & Attar; Iranian J. Bot. 8(1): 19 (1999)
Distribution: Iran
34 IRN
species :
Cousinia sheidaii
Attar, Ghahr. & Mahdigholi; Novon 17(1): 5 (2007)
Distribution: Iran
34 IRN
species :
Cousinia shibarensis
Rech.f.; Fl. Iranica 90: 271 (1972)
Distribution: Afghanistan (Parwan)
34 AFG
species :
Cousinia shorlughensis
Rech.f.; Fl. Iranica 139a: 116 (1979)
Distribution: E-Afghanistan
34 AFG
species :
Cousinia shugnanica
Juz.; Trudy Tadz. Bazy Akad. Nauk 8: 525, 564 (1940)
Distribution: Tajikistan; Afghanistan (Badakshan, Wakhan, Kunar / Nuristan, Takhar)
32 TZK 34 AFG
Synonyms: hidden
= Cousinia lindbergii Rech.f.; Ann. Naturhist. Mus. Wien 65: 36 (1962)
species :
Cousinia shulabadensis
Attar & Ghahr.; Iranian J. Bot. 9(2): 162 (2002)
Distribution: Iran
34 IRN
species :
Cousinia sicigera
C.Winkl. & Bornm.; Bull. Herb. Boiss. 5: 166, t. 7 (1897)
Distribution: Iran (EC-Iran, SW-Iran)
34 IRN
species :
Cousinia silvanica
Attar; Iranian J. Bot. 16(1): 153 (2010)
Distribution: Iran
34 IRN
species :
Cousinia silyboides
Jaub. & Spach; Illustr. 2: 94, t. 176 (1846)
subspecies :
subsp. silyboides
Distribution: Iran
34 IRN
Synonyms: hidden
= Arctium silybodes (Jaub. & Spach) Kuntze; Revis. Gen. Pl. 1: 307 et 308 (1891)
subspecies :
subsp. disfulensis
(Bornm.) Mehregan; Willdenowia 38(2): 311 (2008)
Distribution: Iran (SW-Iran)
34 IRN
Synonyms: hidden
= Cousinia disfulensis Bornm.; Köie, Danish Scient. Invest. Iran 4, Beitr. Sw. Iran 1: 23, f. 3 (1945)
subspecies :
subsp. zardkuhensis
(Attar & Ghahr.) Mehregan; Willdenowia 38(2): 310 (2008)
Distribution: Iran
34 IRN
Synonyms: hidden
= Cousinia zardkuhensis Attar & Ghahr.; Iranian J. Bot. 9(2): 164 (2002)
species :
Cousinia simulatrix
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 573 (1887)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia singularis
Rech.f.; Anz. Österr. Akad. Wiss., Math.-Nat., 101: 350 (1964)
Distribution: Afghanistan (Ghazni)
34 AFG
species :
Cousinia sintenisii
Freyn; Oesterr. Bot. Z. 42 6: 205 (1892)
Distribution: Turkey (E-Anatolia, SE-Anatolia)
34 TUR
species :
Cousinia sivasica
Hub.-Mor.; Notes Roy. Bot. Gard. Edinburgh 32: 53 (1972)
Distribution: Turkey (Inner Anatolia)
34 TUR
species :
Cousinia smirnowii
Trautv.; Trudy Imp. S.-Peterburgsk. Bot. Sada 8: 488 (1883)
Distribution: Turkmenistan; Iran (EC-Iran, NE-Iran: Mts., N-Iran)
32 TKM 34 IRN
Synonyms: hidden
= Arctium smirnowii (Trautv.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Cousinia fulgens Bornm. & Rchb.f.; Repert. Spec. Nov. Regni Veg. 48: 139 (1940)
= Cousinia globosa (Kult.) Bornm.; (1940)
= Cousinia russeola Juz.; Trudy Bot. Inst. Akad. Nauk S. S. S. R., ser. 1, Fl. Sist. Vyssh. Rast. 3: 319 (1937)
= Cousinia smirnowii subsp. sintenisiana Bornm.; (1940)
= Cousinia smirnowii subsp. trautvetteriana Bornm.; (1940)
= Cousinia smirnowii var. globosa Kult.; (1929)
species :
Cousinia sogdiana
Bornm.; Beih. Bot. Centralbl. 34(2): 154 (1916)
Distribution: Turkmenistan; Uzbekistan
32 TKM UZB
species :
Cousinia sororia
Juz.; Trudy sk. Bazy 8: 525 (1940)
Distribution: Tajikistan
32 TZK
species :
Cousinia spathulata
Kult.; Acta Univ. As. Med., ser. 8b 7: 3, f. 2 (1929)
Distribution: Iran (E-Iran)
34 IRN
species :
Cousinia speciosa
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 579 (1887)
Distribution: Kyrgyzstan; Tajikistan
32 KGZ TZK
species :
Cousinia sphaerocephala
Jaub. & Spach; Ill. Pl. Or. 2: 79 (1845)
Distribution: Iran (EC-Iran, N-Iran)
34 IRN
Synonyms: hidden
= Arctium sphaerocephalum (Jaub. & Spach) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia spiridonovii
Juz.; Trans. Rubb. & Guttap. Inst., Fasc. 4, 12 (1932)
Distribution: Kazakhstan; Uzbekistan; Tajikistan
32 KAZ TZK UZB
species :
Cousinia splendida
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 13: 240 (1894)
Distribution: Tajikistan
32 TZK
Synonyms: hidden
= Cousinia lipskyi C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 14: 237 (1895)
species :
Cousinia sporadocephala
Juz.; Acta Inst. Bot. Acad. Sci. U.R.S.S., ser. I. Fasc. 3, 307 (1937)
Distribution: Uzbekistan; Kyrgyzstan
32 KGZ UZB
species :
Cousinia spryginii
Kult.; Korovin, Kult. & Popov, Descr. Pl. Nov. Turkmenistan. (Bodenk. u. Pflanzengeogr. Beob. u. Unters. 1: Flussgeb. d. Amu Darya u. Syr Darya, II.), 78 (1916)
Distribution: Turkmenistan; Uzbekistan
32 TKM UZB
species :
Cousinia stahliana
Bornm. & Gauba; Mitt. Thür. Bot. Ver. 47: 117, t. 13 (1941)
Distribution: Turkmenistan; Iran (NE-Iran: Mts.)
32 TKM 34 IRN
species :
Cousinia stapfiana
Freyn & Sint.; Oesterr. Bot. Z. 44: 218 (1894)
Distribution: Turkey (Inner Anatolia, N-Anatolia)
34 TUR
Synonyms: hidden
= Cousinia stapfiana var. ankarensis Wagenitz; Willdenowia 3 2: 276 (1962)
species :
Cousinia stechmanniopsis
Rech.f.; Fl. Iranica 90: 52 (1972)
Distribution: Afghanistan (Herat)
34 AFG
species :
Cousinia stellaris
Bornm.; Beih. Bot. Centralbl. 34(2): 161 (1916)
Distribution: Uzbekistan; Kyrgyzstan; Tajikistan
32 KGZ TZK UZB
species :
Cousinia stenocalathia
Rech.f.; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 156 (1955)
Distribution: Afghanistan (Badakshan, Baghlan, Takhar)
34 AFG
species :
Cousinia stenocephala
Boiss.; Diagn. Pl. Orient., ser. 1, 6: 115 (1846)
Distribution: Turkey (SE-Anatolia, SE-Anatolia: Mesopotamian Anatolia); Iraq (NE-Iraq, NW-Iraq); Iran (W-Iran); Syria (Jazira)
34 IRN IRQ SYR TUR
Synonyms: hidden
= Arctium involucratum (Boiss.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Arctium stenocephalum (Boiss.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Cousinia involucrata Boiss.; Diagn. 1 6: 114 (1846)
species :
Cousinia stenophylla
Kult.; Acta Univ. As. Med. Ser. VIII. Bot. Fasc. VI. 7 (1929)
Distribution: Turkmenistan
32 TKM
species :
Cousinia stephanophora
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 575 (1887)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia stereolepis
Rech.f.; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 158 (1955)
Distribution: Afghanistan (Ghorat, Jawzjan / Sar-e-Pol)
34 AFG
species :
Cousinia stereoneura
Rech.f.; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 159 (1955)
Distribution: Afghanistan (Bamyan)
34 AFG
species :
Cousinia stocksii
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 12: 216 (1893)
Distribution: Iran (S-Iran, SW-Iran); Afghanistan (Farah, Helmand, Kandahar, Orozgan / Daykundi); Pakistan (Baluchistan)
34 AFG IRN 40 PAK
species :
Cousinia stricta
Tscherneva; Not. Syst. Herb. Inst. Bot. Acad. Sci. . 16: 60 (1961)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia strobilocephala
Tscherneva & Vved.; Not. Syst. Herb. Inst. Bot. Acad. Sci. . 16: 62 (1961)
Distribution: Uzbekistan
32 UZB
species :
Cousinia stroterolepis
Rech.f.; Fl. Iranica 90: 250 (1972)
Distribution: Iran (SW-Iran)
34 IRN
species :
Cousinia subappendiculata
Kult.; Not. Syst. Herb. Hort. Petrop. 5: 140 (1924), et Not. Syst. Herb. Hort. Bot. Russia 6: 27 (1926)
Distribution: Kyrgyzstan
32 KGZ
species :
Cousinia subcandicans
Tscherneva; Not. Syst. Herb. Inst. Bot. Acad. Sci. . 15: 41 (1959)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia subinflata
Bornm.; Oesterr. Bot. Z. 63: 291 (1913)
Distribution: Iran (SW-Iran)
34 IRN
species :
Cousinia submutica
Franch.; Ann. Sci. Nat. Ser. VI. 16: 314 (1883)
Distribution: Uzbekistan; Kyrgyzstan; Tajikistan
32 KGZ TZK UZB
Synonyms: hidden
= Arctium submuticum (Franch.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia subpectinata
Mirtadz., Attar & Assadi; Iranian J. Bot. 10(2): 144 (2004)
Distribution: Iran
34 IRN
species :
Cousinia subscaposa
Rech.f.; Fl. Iranica 90: 270 (1972)
Distribution: Afghanistan (Badakshan, Baghlan, Takhar); Pakistan (Chitral)
34 AFG 40 PAK
species :
Cousinia sylvicola
Bunge; Del. Sem. Hort. Dorp. (1845) 3
Distribution: Uzbekistan
32 UZB
species :
Cousinia tabrisiana
Bunge; Mém. Acad. Sci. Pétersb. 7 sér., 9 2: 16 (1865)
Distribution: Iran (Iranian Aserbaijan)
34 IRN
Synonyms: hidden
= Arctium tabrisianum (Bunge) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia takharensis
Rech.f.; Fl. Iranica 90: 116 (1972)
Distribution: Afghanistan (Wakhan Kunar / Nuristan, Takhar)
34 AFG
species :
Cousinia talassica
(Kult.) Juz. ex Tscherneva; Komolrov, Fl. U.R.S.S. 27: 154 (1962)
Distribution: Kazakhstan; Kyrgyzstan
32 KAZ KGZ
Synonyms: hidden
= Cousinia affinis var. talassica Kult.
species :
Cousinia tamarae
Juz.; Acta Inst. Bot. Acad. Sci. U.R.S.S., ser. I. Fasc. 3, 297 (1937)
Distribution: Kyrgyzstan
32 KGZ
Synonyms: hidden
= Cousinia krauseana var. macrocephala C.Winkl.
species :
Cousinia tashkurghanica
Rech.f.; Fl. Iranica 90: 42 (1972)
Distribution: Afghanistan (Samangan)
34 AFG
species :
Cousinia taybadensis
Djavadi & Attar; Rostaniha 7(Suppl. 2): 168 (2007)
Distribution: Iran
34 IRN
species :
Cousinia tedshenica
Tscherneva; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Uzb. SSR 17: 105 (1962)
Distribution: Turkmenistan; Uzbekistan
32 TKM UZB
species :
Cousinia tenella
Fisch. & C.A.Mey.; Ind. Sem. Horti Petrop. 1: 25 (1834), et Linnaea 10: Litt.-Ber. 86 (1836)
Distribution: Germany [I]; Latvia [I]; Kazakhstan; Turkmenistan; Uzbekistan; Kyrgyzstan; Tajikistan; Armenia; Azerbaijan (incl. Nachichevan); Iran (EC-Iran, NE-Iran: Mts., N-Iran, Iranian Aserbaijan); Afghanistan (Badghys, Baghlan, Faryab, Qunduz, Samangan); Pakistan (Baluchistan)
11 ger 14 lat 32 KAZ KGZ TKM TZK UZB 33 ARM AZE 34 AFG IRN 40 PAK
Synonyms: hidden
= Arctium tenellum (Fisch. & C.A.Mey.) Kuntze; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 197 (1887)
= Lappa tenella (Fisch. & C.A.Mey.) DC.; Prodr. [A. P. de Candolle] 6: 662 (1838)
species :
Cousinia tenuifolia
C.A.Mey.; Prodr. [A. P. de Candolle] 6: 554 (1838)
Distribution: Armenia; Iran (Iranian Aserbaijan)
33 ARM 34 IRN
Synonyms: hidden
= Arctium tenuifolium (C.A.Mey.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia tenuiramula
Rech.f.; Fl. Iranica 139a: 124, t. 113 (1979)
Distribution: Iran (SW-Iran)
34 IRN
species :
Cousinia tenuisecta
Juz.; Acta Inst. Bot. Acad. Sci. U.R.S.S., ser. I. Fasc. 3, 306 (1937)
Distribution: Kazakhstan; Kyrgyzstan
32 KAZ KGZ
species :
Cousinia tenuispina
Rech.f.; Fl. Iranica 90: 83 (1972), nom. nov.
Distribution: Afghanistan (Bamyan, Ghazni, Ghorat, Herat, Wardak)
34 AFG
Synonyms: hidden
= Cousinia leptacantha Rech.f. & Köie; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 116 (1955)
species :
Cousinia termei
Rech.f.; Fl. Iranica 139a: 150, t. 146 (1979)
Distribution: Iran (NE-Iran: Mts.)
34 IRN
species :
Cousinia tetanocephala
Bornm. & Gauba; Mitt. Thür. Bot. Ver. 47: 119, t. 14 (1941)
Distribution: Iran (E-Iran)
34 IRN
species :
Cousinia thamnodes
Boiss. & Hausskn.; Boiss., Fl. Orient. 3: 471 (1875)
Distribution: Iran (S-Iran, SW-Iran)
34 IRN
Synonyms: hidden
= Arctium thamnodes (Boiss. & Hausskn.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia thomsonii
C.B.Clarke; Compos. Ind. 213 (1876)
Distribution: Afghanistan (Kunar / Nuristan, Laghman); S-Tibet; Pakistan (Swat, Dir, Chitral, Gilgit, Astor); Nepal; N-India (Dras, Ladakh, Jammu & Kashmir, Zanskar, Himachal Pradesh, Uttarakhand)
34 AFG 36 TIB 40 IND NEP PAK
Synonyms: hidden
= Arctium thomsonii (C.B.Clarke) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Cousinia brandisii C.B.Clarke; Compos. Ind. 214 (1876)
= Cousinia elasoonensis Rech.f. & Edelb.; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 91 (1955)
= Cousinia xanthophoenicea Bornm.; Repert. Spec. Nov. Regni Veg., Beih. 108: 46 (1938)
species :
Cousinia tianschanica
Kult.; Bull. Inst. Pedol. & Geobot. Univ. As. Centr. 1: 113 (1925)
Distribution: Kazakhstan; Uzbekistan; Kyrgyzstan; Tajikistan
32 KAZ KGZ TZK UZB
species :
Cousinia tirinensis
Rech.f.; Anz. Österr. Akad. Wiss., Math.-Nat., 105: 8 (1969)
Distribution: Afghanistan (Kandahar, Orozgan / Daykundi)
34 AFG
species :
Cousinia touchalensis
Attar; Taxon. Biosyst. 3(6): 65 (2011)
Distribution: Iran
34 IRN
species :
Cousinia trachylepis
Bunge; Mém. Acad. Sci. Pétersb. 7 sér., 9 2: 22 (1865)
Distribution: Iran (EC-Iran)
34 IRN
Synonyms: hidden
= Arctium trachylepis (Bunge) Kuntze; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 197 (1887)
= Cousinia masandarana Bornm.; Oesterr. Bot. Z. 75: 232 (1926)
= Cousinia meyameyensis Parsa; Kew Bull. 3(2): 206 (1948)
= Cousinia masanderana var. mallophora Bornm. & Gauba; (1944)
species :
Cousinia trachyphylla
Juz.; Trudy sk. Bazy 8: 524 (1940)
Distribution: Tajikistan
32 TZK
species :
Cousinia trachyphyllaria
Bornm. & Rech.f.; Repert. Spec. Nov. Regni Veg. 48: 143, t. 312, f. 2 (1940)
Distribution: Iran (NE-Iran: Mts.)
34 IRN
species :
Cousinia tragacanthoides
Rech.f. & Gilli; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 161 (1955)
Distribution: Afghanistan (Bamyan)
34 AFG
Synonyms: hidden
= Tiarocarpus tragacanthoides (Rech.f. & Gilli) Rech.f.; Anz. Österr. Akad. Wiss., Mat. Naturwiss., 108: 6 (1971 publ. 1972)
species :
Cousinia transiliensis
Juz.; Acta Inst. Bot. Acad. Sci. U.R.S.S., ser. I. Fasc. 3, 298 (1937)
Distribution: Kazakhstan; Kyrgyzstan
32 KAZ KGZ
species :
Cousinia transoxana
Tscherneva; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Uzb. SSR 17: 105 (1962)
Distribution: Turkmenistan; Uzbekistan
32 TKM UZB
species :
Cousinia triceps
Kult.; Acta Univ. As. Med. Ser. VIII. Bot. Fasc. VI. 8 (1929)
Distribution: Turkmenistan; Uzbekistan
32 TKM UZB
species :
Cousinia trichophora
Kult.; Acta Univ. As. Med. Ser. VIII. Bot. Fasc. VI. 26 (1929)
Distribution: Uzbekistan
32 UZB
species :
Cousinia tricolor
Rech.f.; Anz. Österr. Akad. Wiss., Math.-Nat., 101: 351 (1964)
Distribution: Afghanistan (Ghorat)
34 AFG
species :
Cousinia trollii
Bornm.; Repert. Spec. Nov. Regni Veg., Beih. 108: 45 (1938)
Distribution: Afghanistan (Kunar / Nuristan); Pakistan (Chitral)
34 AFG 40 PAK
species :
Cousinia tscherneviae
Berdyev; Novit. Syst. Pl. Vasc., Akad. Sci. U.R.S.S. 6: 200 (1970)
Distribution: Turkmenistan
32 TKM
species :
Cousinia turcomanica
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 11: 133 (1889)
Distribution: Turkmenistan; Iran (EC-Iran, E-Iran, NE-Iran: Mts., N-Iran)
32 TKM 34 IRN
Synonyms: hidden
= Arctium turcomanicum (C.Winkl.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Cousinia turcomanica var. leiophylla Bornm. & Sint.; (1912)
= Serratula microcephala Trautv.; Trudy Imp. S.-Peterburgsk. Bot. Sada 9: 458 (1886)
species :
Cousinia turkestanica
(Regel) Juz.
Distribution: Kazakhstan; Turkmenistan; Uzbekistan; Kyrgyzstan
32 KAZ KGZ TKM UZB
Synonyms: hidden
= Cousinia decurrens var. turkestanica Regel
= Cousinia syrdarjensis Kult.; Acta Univ. As. Med. Ser. VIII b, Bot. Fasc. 3, 106 (1928)
species :
Cousinia turkmenorum
Bornm.; Beih. Bot. Centralbl. 34 2: 153 (1917)
Distribution: Turkmenistan; Iran (EC-Iran, E-Iran, NE-Iran: Mts., N-Iran); Afghanistan (Farah, Herat)
32 TKM 34 AFG IRN
Synonyms: hidden
= Cousinia dissecta var. microcephala Trautv.; Trudy Imp. S.-Peterburgsk. Bot. Sada 9: 456 (1886)
= Cousinia dissecta var. simplicior Trautv.; Trudy Imp. S.-Peterburgsk. Bot. Sada 9: 456 (1886)
= Cousinia simplicior Juz.; Not. Syst. Herb. Hort. Petrop. 4: 77 (1923), in adnot.
= Cousinia stenoptera Juz.; Not. Syst. Herb. Hort. Petrop. 2: 59 (1921)
species :
Cousinia ulotoma
Bornm.; Beih. Bot. Centralbl. 34(2): 148 (1916)
Distribution: Tajikistan
32 TZK
species :
Cousinia umbilicata
Juz.; Acta Inst. Bot. Acad. Sci. U.R.S.S., ser. I. Fasc. 3, 306 (1937)
Distribution: Kazakhstan; Uzbekistan
32 KAZ UZB
species :
Cousinia unaiensis
Rech.f.; Fl. Iranica 90: 136 (1972)
Distribution: Afghanistan (Ghazni, Wardak)
34 AFG
species :
Cousinia urumiensis
Bornm.; Verh. K. K. Zool.-Bot. Ges. Wien 60: 135 (1910)
Distribution: Turkey (E-Anatolia); Iran (Iranian Aserbaijan)
34 IRN TUR
Synonyms: hidden
= Cousinia azerbaijanensis Parsa; Kew Bull. 3(2): 202 (1948)
= Cousinia azerbaijanensis var. elongata Parsa; (1948)
species :
Cousinia verbascifolia
Bunge; Mém. Acad. Sci. Pétersb. 7 s&., 9 2: 52 (1865)
Distribution: Iran (EC-Iran, NE-Iran: Mts.)
34 IRN
Synonyms: hidden
= Arctium verbascifolium (Bunge) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia verticillaris
Bunge; Mem. Sav. Etr. Petersb. 6: 357 (1851), et Beitr. Fl. Russl.: 181 (1852)
Distribution: Uzbekistan; Kyrgyzstan; Tajikistan
32 KGZ TZK UZB
Synonyms: hidden
= Arctium verticillare (Bunge) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Cousinia hissarica C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 93 (1887)
= Cousinia poliothrix C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 577 (1887)
species :
Cousinia vicaria
Kult.; Trans. Sci. Soc. Turkmenistan. 1: 113 (1923), et in Bull. Univ. As. Centr. 7: Suppl. 32 (1924)
Distribution: Kazakhstan; Uzbekistan
32 KAZ UZB
species :
Cousinia volkii
Rech.f.; Anz. Österr. Akad. Wiss., Math.-Nat., 101: 352 (1964)
Distribution: Afghanistan (Bamyan)
34 AFG
species :
Cousinia vvedenskyi
Tscherneva; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Uzb. SSR 17: 106 (1962)
Distribution: Uzbekistan
32 UZB
species :
Cousinia waldheimiana
Bornm.; Beih. Bot. Centralbl. 34(2): 140 (1916)
Distribution: Kyrgyzstan
32 KGZ
species :
Cousinia wendelboi
Rech.f.; Fl. Iranica 90: 167, t. 75 (1972)
Distribution: Iran (N-Iran)
34 IRN
species :
Cousinia wesheni
Post; Pl. Postianae 2: 13 (1891)
Distribution: Turkey (SE-Anatolia: Mesopotamian Anatolia); Iraq (NE-Iraq, NW-Iraq); Syria (C-Syrian Desert, Jazira)
34 IRQ SYR TUR
Synonyms: hidden
= Cousinia chaborasica Bornm. & Hand.-Mazz.; Oesterr. Bot. Z. 62: 183 (1912)
species :
Cousinia wilhelminae
Rech.f.; Fl. Iranica 90: 142 (1972)
Distribution: Iran (Iranian Aserbaijan)
34 IRN
species :
Cousinia winkleriana
Aitch. & Hemsl.; Trans. Linn. Soc. Ser. II. 3: 79 (1886)
Distribution: Afghanistan (Herat)
34 AFG
species :
Cousinia woronowii
Bornm.; Oesterr. Bot. Z. 63: 290 (1913)
Distribution: Turkey (E-Anatolia, NE-Anatolia)
34 TUR
species :
Cousinia xanthacantha
Regel; Bull. Soc. Imp. Naturalistes Moscou 11 2: 156 (1867)
Distribution: Afghanistan (Baghlan, Bamyan, Kabul, Wardak, Orozgan / Daykundi, Paktia / Khost, Parwan)
34 AFG
Synonyms: hidden
= Arctium xanthacanthum (Regel) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Cousinia neurophylla Rech.f. & Köie; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 130 (1955)
species :
Cousinia xanthina
Bornm.; Beih. Bot. Centralbl. 34(2): 190 (1916)
Distribution: Uzbekistan; Kyrgyzstan; Tajikistan
32 KGZ TZK UZB
Synonyms: hidden
= Cousinia curviloba Juz.; Trudy Bot. Inst. Akad. Nauk S.S.S.R., ser. 1, Fl. Sist. Vyssh. Rast. 3: 321 (1937)
species :
Cousinia xanthiocephala
Tscherneva; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Uzb. SSR 17: 107 (1962)
Distribution: Kazakhstan
32 KAZ
species :
Cousinia xanthothyrsa
Rech.f. & Gilli; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 165 (1955)
Distribution: Afghanistan (Ghazni, Kabul, Logar, Wardak)
34 AFG
species :
Cousinia xanthula
Rech.f.; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 167 (1955)
Distribution: Afghanistan (Bamyan)
34 AFG
species :
Cousinia xiphiolepis
Boiss.; Diagn. Pl. Orient., ser. 1, 6: 121 (1846)
Distribution: Iran (EC-Iran)
34 IRN
Synonyms: hidden
= Arctium actinocephalum (Jaub. & Spach) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Arctium xiphiolepis (Boiss.) Kuntze; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 197 (1887)
= Cousinia actinocephala Jaub. & Spach; Illustr. 2: 93, t. 175 (1846)
species :
Cousinia yasujensis
Attar; Iranian J. Bot. 17(2): 153 (2011)
Distribution: Iran
34 IRN
species :
Cousinia zagrica
Attar, Ghahr. & Assadi; Sendtnera 8: 5 (2002)
Distribution: Iran
34 IRN
species :
Cousinia ×androssovii
Juz.; Acta Inst. Bot. Acad. Sci. U.R.S.S., ser. I. Fasc. 3, 305 (1937)
Distribution: Turkmenistan; Tajikistan; Afghanistan (Badakshan, Baghlan, Takhar)
32 TKM TZK 34 AFG
species :
Cousinia ×atripurpurea
Juz.; Acta Inst. Bot. Acad. Sci. U.R.S.S., ser. I. Fasc. 3, 313 (1937)
Distribution: Tajikistan
32 TZK
Remarks: Cousinia coronata Franch. x Cousinia microcarpa Boiss.
species :
Cousinia ×cana
Juz.; Acta Inst. Bot. Acad. Sci. U.R.S.S., ser. I. Fasc. 3, 316 (1937)
Distribution: Kazakhstan
32 KAZ
Remarks: Cousinia ninae Juz. x Cousinia sewerzowii Regel
species :
Cousinia ×heterogenetos
Bornm.; Beih. Bot. Centralbl. 34: 11. 163 (1916)
Distribution: Tajikistan
32 TZK
Remarks: Cousinia outichaschensis Franch. x Cousinia pulchella Bunge
species :
Cousinia ×heteromorpha
Bornm.
Distribution: C-Asia
Remarks: Cousinia alpestris (Bornm.) Juz. x Cousinia submutica Franch.
species :
Cousinia ×hybrida
Rech.f.; Ann. Naturhist. Mus. Wien 57: 83 (1950)
Distribution: Iran (N-Iran)
34 IRN
Remarks: Cousinia crispa x sphaerocephala
species :
Cousinia ×iskanderi
Bornm.; Beih. Bot. Centralbl. 34: 11. 168 (1916)
Distribution: Tajikistan
32 TZK
Remarks: Cousinia outichaschensis Franch. x Cousinia submutica Franch.
species :
Cousinia ×kamarbandensis
Rech.f.; Ann. Naturhist. Mus. Wien 57: 82 (1950)
Distribution: Iran (N-Iran)
34 IRN
Remarks: Cousinia crispa x gmelini
species :
Cousinia ×kurubasgecidiensis
Ilçim & Özçelik; Biol. Diversity Conservation 6(1): 71 (2013)
Distribution: Turkey
34 TUR
species :
Cousinia ×mesomorpha
Rech.f.; Candollea 31: 87, f. 1 (1976)
Distribution: Iran (EC-Iran)
34 IRN
Remarks: Cousinia behboudiana x belangeri
species :
Cousinia ×paraatripurpurea
Kamelin; Bot. Zhurn. (Moscow & Leningrad) 58(5): 632 (1973), without latin descr.
Distribution: C-Asia
Remarks: Cousinia microcarpa Boiss. x Cousinia radians Bunge
species :
Cousinia ×triacantha
Kult.; Acta Univ. As. Med. Ser. VIII. Bot. Fasc. VI. 2 (1929)
Distribution: Turkmenistan
32 TKM
Remarks: Cousinia alberti x umbrosa
