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Ver. 24.11
November 5th, 2024

046.0000 Family Athyriaceae Alston; Taxon 5: 25 (1956)
6-7 genera, ca. 760 spp. Circumscription following Rothfels et al. (2012). According to Fraser-Jenkins et al. (2018) it should be included in Woodsiaceae s. lat.
Species listed in database: 758; hybrids: 102
= Athyriinae Milde; Bot. Zeitung (Berlin) 28: 332 (1870)
= Athyrioideae B.K.Nayar; Taxon 19: 235 (1970)
= Physematieae J.Sm.; (1875)
= Deparioideae M.L.Wang & Y.T.Hsieh; Acta Phytotax. Sin. 42: 526 (2004)
= Diplazioideae Shmakov; Turczaninowia 4(1-2): 59 (2001), nom. superfl.
= Diplazioideae Ching & Z.R.Wang; Acta Phytotax. Sin. 35: 35 (1997)
= Diplazieae Pfeiff.; Nomencl. Bot. [Pfeiffer] 1(1): 628 (1873)
= Diplaziinae Payer; Bot. Crypt.: 198 (1850)
046.0004 Genus Cornopteris Nakai; Bot. Mag. (Tokyo) 44: 7 (1930)
Distribution: China, E-Asia, N-trop. Asia. Fraser-Jenkins et al. (2018) make a strong argument to keep the genus separate from Athyrium s. lat., supported by molecular results (Wei et al. 2017)
Species listed in database: 13; hybrids: 2
= Athyrium subgen.Cornopteris (Nakai) Seriz.; J. Jap. Bot. 56(7): 194 (1981)
species :
Cornopteris approximata W.M.Chu; Acta Bot. Yunnan. 3(3): 342 (1981)
Distribution: China (NE-Yunnan)
36 CHI
Synonyms: hidden
= Athyrium approximatum (W.M.Chu) Christenh.; Global Fl. 1(4): 23 (2018)
species :
Cornopteris atroviridis (Alderw.) M.Kato; Acta Phytotax. Geobot. 30(4-6): 113 (1979)
Distribution: peninsular Malaysia (Cameron Highlands, Pahang); Sumatra; Borneo (Mt. Kinabalu, Sabah); Java; Moluccas (Ambon)
42 BOR JAW MLY MOL SUM
Remarks: not in India
Synonyms: hidden
= Asplenium amplissimum Baker; Ann. Bot. (Oxford) 5: 310 (1891)
= Athyrium amplissimum (Baker) Holttum; Rev. Fl. Malaya 2: 569, f. 338 (1955)
= Athyrium atroviride (Alderw.) Seriz.; J. Jap. Bot. 56(7): 195 (1981)
= Diplazium amplissimum (Baker) Diels; Nat. Pflanzenfam. [Engler & Prantl] I, 4: 227 (1899)
= Diplazium atroviride (Alderw.) C.V.Morton; Contr. U. S. Natl. Herb. 38: 254 (1973)
= Dryopteris atroviridis Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 16: 26 (1914), pro syn.
= Phegopteris atroviridis Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 16: 26 (1914)
species :
Cornopteris badia Ching; Bull. Fan Mem. Inst. Biol. Bot. 11: 58 (1941)
Distribution: China (Yunnan); Nepal; Bhutan; N-India (Arunachal Pradesh, Sikkim, Uttarakhand, West Bengal); Myanmar (Kachin, etc.)
36 CHI 40 BHU IND NEP 41 MYA
Remarks: inclusion of C. quadripinnatifida fide Fraser-Jenkins et al. (2015, 2018)
Synonyms: hidden
= Athyrium badium (Ching) Christenh.; Global Fl. 1(4): 23 (2018)
= Athyrium quadripinnatifidum (M.Kato) Seriz.; J. Jap. Bot. 56(7): 196 (1981)
= Cornopteris badia f. quadripinnatifida (M.Kato) W.M.Chu; Fl. Reipubl. Popularis Sin. [Fl. China Ed. Committee] 3(2): 365 (1999)
= Cornopteris badia var. pubescens Z.R.Wang; Vasc. Pl. Hengduan Mount. 1: 88 (1993) (without latin descr. or type), et Acta Phytotax. Sin. 32(1): 84 (1994)
= Cornopteris quadripinnatifida M.Kato; Acta Phytotax. Geobot. 30(4-6): 114 (1979)
species :
Cornopteris banajaoensis (C.Chr.) K.Iwats. & M.G.Price; S. E. Asian Stud. 14(4): 564 (1977) (banahaoensis auct.)
Distribution: S-Japan (Yakushima Isl.); Volcano Isl. (Minami-Iwojima); Taiwan; Nepal; Bhutan; India (Sikkim, West Bengal, Darjeeling); Sulawesi; Philippines (Luzon)
38 JAP KZN NNS TAI 40 BHU IND NEP 42 PHI SUL
Synonyms: hidden
= Athyrium banajaoense (C.Chr.) Christenh.; Global Fl. 1(4): 23 (2018)
= Athyrium fluviale (Hayata) C.Chr.; Index Filic., Suppl. Tert.: 41 (1934)
= Athyrium nudum Copel.; Fern Fl. Philipp. 3: 391 (1958), nom. nov. for Dryopteris banajaoensis C. Chr.
= Athyrium tagawae C.Chr.; Index Filic., Suppl. Tert.: 44 (1934), nom. nov. for Cornopteris tashiroi Tagawa
= Cornopteris fluvialis (Hayata) Tagawa; Acta Phytotax. Geobot. 1: 158 (1932)
= Cornopteris tashiroi Tagawa; Acta Phytotax. Geobot. 1: 159 (1932) (non Athyrium tashiroi Tagawa)
= Dryopteris athyriiformis Rosenst.; Hedwigia 66: 344 (1915)
= Dryopteris banajaoensis C.Chr.; Index Filic., Suppl. 2 (1906-1912): 30 (1913) (non Athyrium banahaoense Copel.)
= Dryopteris fluvialis Hayata; Icon. Pl. Formosan. 4: 152-154, f. 94 (1914)
= Dryopteris tenerrima Copel.; Philipp. J. Sci. 4: 111 (1909) (non Rosenst. (1906))
= Phegopteris banajaoensis (C.Chr.) Alderw.; Malayan Ferns Fern Allies, Suppl. 317 (1917)
species :
Cornopteris christenseniana (Koidz.) Tagawa; Acta Phytotax. Geobot. 2(3): 195 (1933)
Distribution: China (Zhejiang); South Korea (incl. Cheju Isl.); Japan; ?Taiwan [probable]
36 CHI 38 JAP KOR
Synonyms: hidden
= Athyrium christensenianum (Koidz.) Seriz.; J. Jap. Bot. 56(7): 195 (1981)
= Athyrium crenulatoserrulatum f. hakonense Makino; Bot. Mag. (Tokyo) 13: 27 (1899)
= Athyrium hakonense (Makino) Ohwi; Bull. Natl. Sci. Mus., Tokyo, 3: 100 (1956), nom. illeg.
= Athyrium hakonense (Nakai) C.Chr.; Index Filic., Suppl. Tert.: 42 (1934) (non (Makino) Ohwi)
= Cornopteris hakonensis Nakai; Bot. Mag. (Tokyo) 45: 94 (1931)
= Diplazium christensenianum Koidz.; Bot. Mag. (Tokyo) 88: 105 (1924)
species :
Cornopteris crenulatoserrulata (Makino) Nakai; Bot. Mag. (Tokyo) 46: 95 (1931)
Distribution: Russian Far East (Khabarovsk, Primorye); China (Heilongjiang, Henan, Jilin, Liaoning, Shaanxi); North Korea; South Korea (incl. Cheju Isl.); Japan
31 RFE 36 CHI 38 JAP KOR NKO
Remarks: not in India
Synonyms: hidden
= Athyrium austroussuriense (Kom.) Fomin; Fl. Sibir. Orient. Extremi 5: 122, fig. (1930)
= Athyrium crenulatoserrulatum Makino; Bot. Mag. (Tokyo) 18: 26 (1899)
= Athyrium koryoense C.Chr.; Index Filic., Suppl. Tert.: 42 (1934), nom. nov. for Cornopteris coreana Nakai
= Cornopteris coreana Nakai; Bot. Mag. (Tokyo) 45: 96 (1931)
= Cornopteris crenulatoserrulata f. coreana (Nakai) H.Itô; Nov. Fl. Jap. [Nakai & Honda] 4: 99 (1939)
= Cornopteris koryoensis Nakai; Bot. Mag. (Tokyo) 46: 96 (1931) (non Athyrium koryoense Christ (1902))
= Dryopteris austroussuriensis Kom.; Izv. Imp. Bot. Sada Petra Velikago 16: 147 (1916)
= Dryopteris crenulatoserrulata (Makino) C.Chr.; Index Filic. 259 (1905)
= Neoathyrium crenulatoserrulatum (Makino) Ching & Z.R.Wang; Acta Phytotax. Sin. 20(1): 77 (1982)
= Neoathyrium crenulatoserrulatum var. coreanum (Nakai) S.X.Li & J.Z.Wang; Clav. Pl. Chin. Bor.-Or., ed. 2 (ed. P. Y. Fu): 39 (1995)
= Phegopteris austroussuriensis (Kom.) Kom. & Aliss.; Malyi Opred. Rast. Dal´ne-Vost. Kraja: 61 (1925)
= Phegopteris crenulatoserrulata (Makino) Makino; Bot. Mag. (Tokyo) 17: 78 (1903)
= Pseudathyrium crenulatoserrulatum (Makino) Nakai; Rep. Veg. Daisetsusan Mountains 44 (1934)
species :
Cornopteris decurrentialata (Hook.) Nakai; Bot. Mag. (Tokyo) 44: 8 (1930)
Distribution: China (Guizhou, Hunan, Jiangxi, Sichuan, Yunnan, Zhejiang); Tibet; South Korea (Cheju Isl.); Japan; Taiwan; Nepal; Bhutan; India (Arunachal Pradesh, Meghalaya, Mizoram, Nagaland, Sikkim, West Bengal); Myanmar (Kachin, etc.); Vietnam; Philippines
36 CHI TIB 38 JAP KOR TAI 40 BHU IND NEP 41 MYA VIE 42 PHI
Synonyms: hidden
= Athyrium birii (Ching ex Bir) Seriz.; J. Jap. Bot. 56(7): 195 (1981)
= Athyrium decurrentialatum f. platyphyllum (H.Itô) Seriz.; J. Jap. Bot. 56: 195 (1981)
= Athyrium decurrentialatum (Hook.) Copel.; Philipp. J. Sci. 8: 279 (1909)
= Athyrium filix-incantatrix Christenh.; Global Fl. 1(4): 23 (2018), nom. nov. for Cornopteris latibasis W. M. Chu
= Athyrium yunnanense Christenh.; Global Fl. 1(4): 25 (2018), nom. nov. for C. latiloba Ching
= Cornopteris birii Ching ex Bir; Nova Hedwigia 7: 502, t. 34, 35 (1964) (fide Fraser-Jenkins 2008)
= Cornopteris decurrentialata f. angustifrons H.Itô; Nov. Fl. Jap. [Nakai & Honda] 4: 97 (1939)
= Cornopteris decurrentialata f. latifrons H.Itô; Nov. Fl. Jap. [Nakai & Honda] 4: 97 (1939)
= Cornopteris decurrentialata f. musashiensis (Nakai) Nakaike; J. Nippon Fernist Club 3(Suppl. 2): 46 (2004)
= Cornopteris decurrentialata f. platyphylla H.Itô; Nov. Fl. Jap. [Nakai & Honda] 4: 98 (1939)
= Cornopteris decurrentialata f. stenophylla H.Itô; Nov. Fl. Jap. [Nakai & Honda] 4: 98 (1939)
= Cornopteris decurrentialata var. glabra H.Itô; Nov. Fl. Jap. [Nakai & Honda] 4: 97 (1939)
= Cornopteris latibasis W.M.Chu; Acta Bot. Yunnan. 3(3): 344 (1981)
= Cornopteris latiloba Ching; Fl. Xizangica 1: 144 (1983)
= Cornopteris tsangii Ching; Lingnan Sci. J. 21: 32 (1945)
= Diplazium decurrentialatum (Hook.) C.Chr.; Bull. Acad. Int. Géogr. Bot. 70 (1911)
= Diplazium hookerianum Koidz.; Bot. Mag. (Tokyo) 38: 105 (1924), nom. superfl.
= Dryopteris decurrentialata (Hook.) C.Chr.; Index Filic. 261 (1905)
= Gymnogramma decurrentialata Hook.; Sp. Fil. [W. J. Hooker] Fil. [W. J. Hooker] 5: 142, t. 294 (1864)
= Leptogramma decurrentialata (Hook.) J.Sm.; Hist. Fil. 232 (1875)
= Nephrodium decurrentialatum (Hook.) Diels; Nat. Pflanzenfam. [Engler & Prantl] I, 4: 171 (1899)
= Phegopteris decurrentialata (Hook.) Christ; Farnkr. Erde 274 (1897)
= Thelypteris decurrentialata (Hook.) C.F.Reed; Phytologia 17: 271 (1968)
form :
f. pilosella (H.Itô) W.M.Chu; Fl. Reipubl. Popularis Sin. [Fl. China Ed. Committee] 3(2): 358 (1999)
Distribution: China (Guizhou, Hunan, Jiangxi, Sichuan, Yunnan, Zhejiang); ?Japan; Taiwan
36 CHI 38 TAI
Synonyms: hidden
= Athyrium decurrentialatum var. pilosellum (H.Itô) Ohwi; Fl. Jap. [Ohwi] 110 (1957)
= Athyrium musashiense (Nakai) C.Chr.; Index Filic., Suppl. Tert.: 43 (1934)
= Athyrium unifurcatum var. harryanum C.Chr.; (1924)
= Cornopteris decurrentialata var. pilosella H.Itô; Bot. Mag. (Tokyo) 52: 588 (1938)
= Cornopteris glandulosopilosa S.F.Wu; Keys Vasc. Pl. Wuling Mts. 565 (1995)
= Cornopteris musashiensis Nakai; Bot. Mag. (Tokyo) 44: 8 (1930)
species :
Cornopteris major W.M.Chu; Acta Bot. Yunnan. 3(3): 348 (1981)
Distribution: China (Yunnan)
36 CHI
Synonyms: hidden
= Athyrium majus (W.M.Chu) Christenh.; Global Fl. 1(4): 23 (2018)
species :
Cornopteris omeiensis Ching; Bull. Fan Mem. Inst. Biol. Bot., ser. 2, 1: 287 (1949) (omeigensis)
Distribution: China (Guizhou, Sichuan)
36 CHI
Synonyms: hidden
= Diplazium japonicum var. nudisorum C.Chr.; (1924)
= Athyrium narthex Christenh.; Global Fl. 1(4): 23 (2018), nom. nov.
species :
Cornopteris opaca (D.Don) Tagawa; Acta Phytotax. Geobot. 8: 92 (1939)
Distribution: China (Fujian, Guangxi, Jiangxi, Yunnan); Japan; Taiwan; Nepal; Bhutan; N-India (Arunachal Pradesh, Meghalaya, Sikkim, West Bengal, Darjeeling); Myanmar (Kachin, etc.); N-Thailand; SW-Thailand; Laos; Vietnam; peninsular Malaysia (Pahang); Borneo (Sabah, S-Kalimantan); Sulawesi; Philippines; Java; Lesser Sunda Isl. (Bali); Moluccas (Seram)
36 CHI 38 JAP TAI 40 BHU IND NEP 41 LAO MYA THA VIE 42 BOR JAW LSI MLY MOL PHI SUL
Synonyms: hidden
= Athyrium gymnocarpum Copel.; Philipp. J. Sci. 40: 301, t. 4 (1929)
= Athyrium opacum (D.Don) Copel.; Philipp. J. Sci. 3: 279 (1908)
= Cornopteris likiangensis Ching; Lingnan Sci. J. 21: 32 (1945)
= Diplazium opacum (D.Don) Christ; Bull. Acad. Int. Géogr., sér. 3, 16: 242 (1906)
= Dryopteris bankinsinensis Hayata; Icon. Pl. Formosan. 8: 146, f. 73-74 (1919)
= Dryopteris opaca (D.Don) C.Chr.; Index Filic. 280 (1905)
= Dryopteris succulentipes Hayata; Icon. Pl. Formosan. 8: 149, f. 77-78 (1919)
= Grammitis obtusata (Blume) C.Presl; Tent. Pterid. 209 (1836)
= Gymnogramma obtusata Blume; Enum. Pl. Javae 2: 113 (1828)
= Gymnogramma opaca (D.Don) Spreng.; Syst. Veg., ed. 16 [Sprengel] 4(1): 39 (1827)
= Hemionitis opaca D.Don; Prodr. Fl. Nepal. 13 (1825)
= Leptogramma obtusata (Blume) J.Sm.; Hist. Fil. 232 (1875)
= Leptogramma opaca (D.Don) Bedd.; Ferns Brit. India 379 (1883)
= Nephrodium obtusatum (Blume) Diels; Nat. Pflanzenfam. [Engler & Prantl] I, 4: 171 (1899), nom. illeg.
= Phegopteris obtusata (Blume) Christ; Farnkr. Erde 274 (1897)
= Phegopteris opaca (D.Don) Mett.; Abh. Senckenberg. Naturf. Ges. 2(2) [Pheg. u. Asp.]: 299 (1858)
= Thelypteris opaca (D.Don) C.F.Reed; Phytologia 17: 299 (1968)
form :
f. glabrescens Sa.Kurata; J. Geobot. 12: 41 (1963)
Distribution: China (Fujian, Guangxi, Yunnan); Japan; Taiwan; Vietnam
36 CHI 38 JAP TAI 41 VIE
Synonyms: hidden
= Athyrium opacum f. glabrescens (Sa.Kurata) Seriz.; J. Jap. Bot. 56(7): 196 (1981)
= Athyrium opacum var. glabrescens (Sa.Kurata) Seriz.; Shidekobushi 3(1): 53 (2015)
species :
Cornopteris philippinensis M.Kato; Acta Phytotax. Geobot. 30(4-6): 110 (1979)
Distribution: Philippines; Moluccas (Seram)
42 MOL PHI
Remarks: Taiwanese records refer probably to an extreme form of C. opaca fide Knapp (Ferns of Taiwan, 2011 ff.)
Synonyms: hidden
= Athyrium philippinense (M.Kato) Christenh.; Global Fl. 1(4): 25 (2018), nom. illeg.
species :
Cornopteris pseudofluvialis Ching & W.M.Chu; Acta Bot. Yunnan. 3(3): 345 (1981)
Distribution: China (SE-Yunnan)
36 CHI
Synonyms: hidden
= Athyrium pseudofluviale (Ching & W.M.Chu) Christenh.; Global Fl. 1(4): 25 (2018)
species :
Cornopteris seramensis M.Kato; J. Jap. Bot. 61(8): 233 (1986)
Distribution: Moluccas (Seram)
42 MOL
Synonyms: hidden
= Athyrium seramense (M.Kato) Christenh.; Global Fl. 1(4): 25 (2018)
species :
Cornopteris ×masachikana Sa.Kurata; J. Geobot. 12: 42 (1963)
Distribution: Japan
38 JAP
Remarks: Cornopteris banajaoensis x opacum
Synonyms: hidden
= Athyrium ×masachikanum (Sa.Kurata) Seriz.; J. Jap. Bot. 56: 196 (1981)
species :
Cornopteris ×undulatipinnula (Seriz.) Nakaike; New Fl. Japan, Pterid. 840 (1992)
Distribution: Japan
38 JAP
Remarks: Cornopteris decurrentialata x banajaoensis (fluviale)
Synonyms: hidden
= Athyrium ×undulatipinnulum Seriz.; J. Jap. Bot. 56(7): 196 (1981)
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A complete, synonymic checklist of the Higher Plants of the World

Version 24.11 (November 5th, 2024)

Covering 355,576 accepted species,
8,715 hybrids, 47,863 accepted infraspecific taxa and 1,000,581 synonyms,
in total 1,412,785* names

[*about 25,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus" are deliberately omitted]

With complete, detailed distribution data (countries and regions) and literature references

Arranged in linear phylogenetic sequence reflecting true relationships

Fully searchable and continously updated

With 38,209 pictures of 15,787 species

Assembled through 40 years independently from other databases

In cooperation with Catalogue of Life