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Ver. 25.01
Jan. 21th, 2024

404.0000 Family Cactaceae Juss.; Gen. Pl. [Jussieu] 310 (1789), nom. cons.
133 genera, ca. 1700-1800 spp. There has been excessive splitting, leading to over 3000 species in some collectors view. The phylogeny is still somewhat in flux, but is getting more stabilized now. It should be good practice in the future that each ´new species´ description is confirmed by molecular genetics, not only by morphology.
Species listed in database: 1796; hybrids: 77
= Cereaceae DC. & Spreng.(transl.Jameson); Elem. Philos. Pl. 142 (1821)
= Nopaleaceae Schmid & Curtman; Pflanzenr. 2: 135 (1856)
= Opuntiaceae Desv.; S. Gérardin de Mirecourt & N. A. Desvaux, Dict. Rais. Bot. 52, 385 (1817)
404.5869 Genus Borzicactus Riccob.; Boll. Reale Orto Bot. Palermo 8: 261 (1909) (4)
Distribution: Andes. Valid genus according to molecular results. Loxanthocereus is frequently kept separate.
Species listed in database: 23
= Bolivicereus Cárdenas; Cact. Succ. J. (Los Angeles) 23: 91 (1951)
= Borzicactella Ritter; Kakteen
= Binghamia Britton & Rose; Cactaceae [Britton & Rose] 2: 16 (1920)
= Clistanthocereus Backeb.; Jahrb. Deutsch. Kakteen-Ges. Teil 1 (Mai) Blatt 24, in adnot. (1937)
= Demnosa Fric; Möller´s Deutsche Gärtn.-Zeitung 44: 170 (1929)
= Hildewintera Ritter; Kakteen And. Sukk. 17: 11 (1966)
= Loxanthocereus Backeb.; Cactaceae (Berlin) 1937(1): 24 (1937)
= Maritimocereus Akers & Buining; Succulenta (Netherlands) 49 (1950)
= Winteria Ritter; Kakteen And. Sukk. 13: 4 (1962) (non Murray)
= Winterocereus Backeb.; 455 (1966)
species :
Borzicactus acanthurus (Vaupel) Britton & Rose; Cactaceae [Britton & Rose] 2: 161 (1920)
subspecies :
subsp. acanthurus
Distribution: Peru (near Lima)
83 PER
Synonyms: hidden
= Binghamia acanthura (Vaupel) Borg; Cacti, ed. 2 [Borg] 185 (1951)
= Binghamia eriotricha (Werderm. & Backeb.) Borg; Cacti, ed. 2 [Borg] 185 (1951)
= Borzicactus eriotrichus (Werderm. & Backeb.) Backeb.; Kakteen-Freund 3: 121 (1934)
= Borzicactus keller-badensis (Backeb. & Krainz) Krainz; Städtische Sukkulentensammlung Zürich, Kat. Kult. Stehenden Art. 39 (1967)
= Cereus acanthurus Vaupel; Bot. Jahrb. Syst. 50, Beibl. 111: 13 (1913)
= Cereus eriotrichus Werderm. & Backeb.; Neue Kakteen 80 (1931), et Repert. Spec. Nov. Regni Veg. 30: 61 (1932)
= Cleistocactus acanthurus (Vaupel) D.R.Hunt; Bradleya 5: 92 (1987)
= Cleistocactus acanthurus subsp. canetensis (Rauh & Backeb.) Ostolaza; CactusWorld 25: 218 (2007)
= Cleistocactus erectispinus (Rauh & Backeb.) Ostolaza; Brit. Cact. Succ. J. 21: 92 (2003)
= Cleistocactus erigens (Rauh & Backeb.) Ostolaza; CactusWorld 25(4): 219 (2007)
= Echinopsis acanthura (Vaupel) Molinari; Succulentopia 13: 18 (2015)
= Echinopsis acanthura subsp. canetensis (Rauh & Backeb.) Molinari; Succulentopia 13: 18 (2015)
= Echinopsis erectispina (Rauh & Backeb.) Molinari; Succulentopia 13: 19 (2015)
= Echinopsis erigens (Rauh & Backeb.) Molinari; Succulentopia 13: 19 (2015)
= Haageocereus convergens F.Ritter; (nomen)
= Haageocereus eremiticus F.Ritter; (nomen)
= Haageocereus imperialensis F.Ritter; (nomen)
= Haageocereus neglectus F.Ritter; (nomen)
= Haageocereus pacaranensis F.Ritter; (nomen)
= Haageocereus pacaranensis var. tenuispinus F.Ritter; (nomen)
= Haageocereus paradoxus Rauh & Backeb.; Descr. Cact. Nov. [Backeberg] 1: 21 (1956 publ. 1957)
= Loxanthocereus acanthurus (Vaupel) Backeb.; Cactaceae (Berlin) 1937(1): 24 (1937)
= Loxanthocereus acanthurus var. ferox Backeb.; Cact. Succ. J. (Los Angeles) 23: 19 (1951)
= Loxanthocereus bicolor F.Ritter; Kakteen Südamerika 4: 1455 (1981)
= Loxanthocereus canetensis Rauh & Backeb.; Descr. Cact. Nov. [Backeberg] 1: 16 (1956 publ. 1957)
= Loxanthocereus cantaensis Rauh & Backeb.; Sitzungsber. Heidelberger Akad. Wiss., Math.-Naturwiss. Kl. 1958: 314 (1958)
= Loxanthocereus convergens F.Ritter; Kakteen Südamerika 4: 1456 (1981)
= Loxanthocereus cullmannianus Backeb.; Cactaceae [Backeberg] 6: 3674 (1962)
= Loxanthocereus erectispinus Rauh & Backeb.; Descr. Cact. Nov. [Backeberg] 1: 15 (1956 publ. 1957)
= Loxanthocereus eremiticus F.Ritter; Kakteen Südamerika 4: 1459 (1981)
= Loxanthocereus erigens Rauh & Backeb.; Descr. Cact. Nov. [Backeberg] 1: 16 (1956 publ. 1957)
= Loxanthocereus eriotrichus (Werderm. & Backeb.) Backeb.; Blätt. Kakteenf. 4(11): 4 (1937)
= Loxanthocereus eulalianus Rauh & Backeb.; Descr. Cact. Nov. [Backeberg] 1: 14 (?1956 publ. 1957), sphalm.
= Loxanthocereus eulaliensis Rauh & Backeb.; Descr. Cact. Nov. [Backeberg] 14 (1957) (eulalianus), et Rauh, Beitr. Kenntn. Peruan. Kakteenveg. (Sitz. Heidelb. Akad. Wiss., Math.-Nat., Abh. 1) 317 (1958)
= Loxanthocereus gracilispinus Rauh & Backeb.; Descr. Cact. Nov. [Backeberg] 1: 16 (1956 publ. 1957)
= Loxanthocereus keller-badensis Backeb. & Krainz; Sukkulentenkunde 2: 22 (1948)
= Loxanthocereus multifloccosus Rauh & Backeb.; Descr. Cact. Nov. [Backeberg] 16 (1957)
= Loxanthocereus neglectus F.Ritter; Taxon 13: 116 (1964)
= Loxanthocereus neglectus var. chimbotensis F.Ritter; Taxon 13: 116 (1964)
= Loxanthocereus pacaranensis F.Ritter; Kakteen Südamerika 4: 1468 (1981)
subspecies :
subsp. faustianus (Backeb.) G.J.Charles; Cactaceae Syst. Init. 2: 14 (2012)
Distribution: Peru (Rimac Valley, Santa Eulalia Valley)
83 PER
Synonyms: hidden
= Borzicactus faustianus Backeb.; Desert 5: 4 (1933), Issue 140, Backeb. in Backeb. & F. M. Knuth, Kaktus-ABC [Backeberg & Knuth] 193, 411 (1935)
= Cleistocactus acanthurus subsp. faustianus (Backeb.) Ostolaza; Cactaceae Consensus Init. 6: 8 (1998)
= Cleistocactus acanthurus var. faustianus (Backeb.) Ostolaza; Brit. Cact. Succ. J. 14(4): 171 (1996)
= Echinopsis acanthura subsp. faustiana (Backeb.) Molinari & Mayta; Succulentopia 14: 19 (2015)
= Haageocereus faustianus (Backeb.) F.Ritter
= Loxanthocereus faustianus (Backeb.) Backeb.; Stachlige Wildnis 187 (1942)
subspecies :
subsp. pullatus (Rauh & Backeb.) G.J.Charles; Cactaceae Syst. Init. 26: 14 (2012)
Distribution: Peru (Jicamarca, Santa Clara, Manchay, Picapidera, Caracoles)
83 PER
Synonyms: hidden
= Cleistocactus acanthurus subsp. pullatus (Rauh & Backeb.) Ostolaza; Cactaceae Consensus Init. 6: 8 (1998)
= Cleistocactus acanthurus var. pullatus (Rauh & Backeb.) Ostolaza; Brit. Cact. Succ. J. 14(4): 171 (1996)
= Echinopsis acanthura subsp. pullata (Rauh & Backeb.) Molinari; Succulentopia 13: 18 (2015)
= Loxanthocereus acanthurus subsp. pullatus (Rauh & Backeb.) Ostolaza; CactusWorld 27(1): 46 (2009)
= Loxanthocereus pullatus Rauh & Backeb.; Descr. Cact. Nov. [Backeberg] 1: 14 (1956 publ. 1957)
= Loxanthocereus pullatus var. brevispinus Rauh & Backeb.; Descr. Cact. Nov. [Backeberg] 1: 15 (1956 publ. 1957)
= Loxanthocereus pullatus var. fulviceps Rauh & Backeb.; Descr. Cact. Nov. [Backeberg] 1: 15 (1956 publ. 1957)
species :
Borzicactus aureispinus (F.Ritter) G.D.Rowley; Repert. Pl. Succ. (I. O. S.) 24: 6 (1973 publ. 1975)
Distribution: Bolivia (Santa Cruz, Florida Prov.)
83 BOL
Synonyms: hidden
= Bolivicactus aureispinus (F.Ritter) Lodé; Cact.-Advent. Int. (Almeria) 36: 131 (2024)
= Bolivicereus colademononis (Diers & Krahn) Lodé; Cact.-Advent. Int. (Almeria) 35(122): 173 (2023)
= Bolivicereus simius-cauda Lodé; Cact.-Advent. Int. (Almeria) 34(121): 91 (2022), nom. nov. illeg. for Hildewintera colademononis
= Borzicactus colademononis (Diers & Krahn) Lodé; Cact.-Advent. Int. (Almeria) 97: 2 (2013 publ. 2012)
= Cleistocactus aureispinus (F.Ritter) D.R.Hunt; Bradleya 5: 92 (1987), non Fric (1928)
= Cleistocactus colademononis (Diers & Krahn) Mottram; CactusWorld 24(1): 21 (2006)
= Cleistocactus winteri D.R.Hunt; Bradleya 6: 100 (1988), nom. nov.
= Cleistocactus winteri subsp. colademono D.R.Hunt; Cactaceae Syst. Init. 20: 14 (2005), nom. nov.
= Hildewintera aureispina (F.Ritter) F.Ritter ex G.D.Rowley; Regnum Veg. 54: 15 (1968)
= Hildewintera colademononis Diers & Krahn; Kakteen And. Sukk. 54(8): 221 (2003)
= Loxanthocereus aureispinus (F.Ritter) Buxb.; Kakteen [H.Krainz] 57: CV b (1974), no basionym page.
= Winteria aureispina F.Ritter; Kakteen And. Sukk. 13: 4 (1962)
= Winterocereus aureispinus (F.Ritter) Backeb.; Kakteenlexikon 455 (1966)
= Winterocereus colademononis (Diers & Krahn) Metzing & R.Kiesling; Taxon 56(1): 227 (2007)
species :
Borzicactus fieldianus Britton & Rose; Cactaceae [Britton & Rose] 4: 278 (1923)
subspecies :
subsp. fieldianus
Distribution: Peru (Ancash)
83 PER
Synonyms: hidden
= Borzicactus cajamarcensis F.Ritter; Kakteen Südamerika 4: 1375 (1981)
= Borzicactus calviflorus F.Ritter; Taxon 13: 118 (1964)
= Borzicactus fieldianus subsp. cajamarcensis (F.Ritter) Lodé; Int. Cact. Advent. 31(1): 20 (2019)
= Cleistocactus fieldianus (Britton & Rose) D.R.Hunt; Bradleya 5: 92 (1987)
= Clistanthocereus calviflorus (F.Ritter) Backeb.; Kakteenlexikon 91 (1966)
= Clistanthocereus fieldianus (Britton & Rose) Backeb.; Jahrb. Deutsch. Kakteen-Ges. Teil 1 (Mai) Blatt 24, in adnot. (1937)
= Echinopsis fieldiana (Britton & Rose) Molinari; Succulentopia 13: 18 (2015)
subspecies :
subsp. tessellatus (Akers & Buining) G.J.Charles; Cactaceae Syst. Init. 2: 14 (2012)
Distribution: NC-Peru
83 PER
Synonyms: hidden
= Borzicactus fieldianus var. tessellatus (Akers & Buining) Krainz ex F.Ritter; Kakteen Südamerika 4: 1377 (1981)
= Borzicactus tessellatus Akers & Buining; Succulenta (Netherlands) 81 (1954)
= Cleistocactus fieldianus subsp. tessellatus (Akers & Buining) Ostolaza; Cactaceae Consensus Init. 6: 8 (1998)
= Clistanthocereus tessellatus (Akers & Buining) Backeb.; Cactaceae [Backeberg] 2: 939 (1959)
= Echinopsis fieldiana subsp. tessellata (Akers & Buining) Molinari & Mayta; Succulentopia 14: 19 (2015)
species :
Borzicactus hoffmannii (G.J.Charles) G.J.Charles; Cactaceae Syst. Init. 2: 14 (2012)
Distribution: Peru
83 PER
Synonyms: hidden
= Cleistocactus hoffmannii G.J.Charles; Cactaceae Syst. Init. 20: 12 (2005)
= Echinopsis hoffmannii (G.J.Charles) Molinari; Succulentopia 13: 19 (2015)
= Loxanthocereus hoffmannii F.Ritter; Kakteen Südamerika 4: 1461 (1981)
species :
Borzicactus hoxeyi G.J.Charles; Bradleya 31: 27 (2013)
Distribution: Peru
83 PER
Synonyms: hidden
= Loxanthocereus hoxeyi (G.J.Charles) Lodé; Cact.-Advent. Int. (Almeria) 101: 14 (2014)
= Loxanthocereus hoxeyi subsp. grandis Hoxey; Bradleya 38: 18 (2020)
species :
Borzicactus hutchisonii G.J.Charles; Bradleya 28: 3 (2010)
Distribution: Peru
83 PER
species :
Borzicactus hystrix (Rauh & Backeb.) G.J.Charles; Cactaceae Syst. Init. 2: 14 (2012)
Distribution: Peru
83 PER
Synonyms: hidden
= Cleistocactus clavispinus (Rauh & Backeb.) Ostolaza; Brit. Cact. Succ. J. 16(3): 133 (1998)
= Cleistocactus hystrix (Rauh & Backeb.) Ostolaza; Brit. Cact. Succ. J. 16(3): 133 (1998)
= Cleistocactus peculiaris (Rauh & Backeb.) Ostolaza; Brit. Cact. Succ. J. 16(3): 129 (1998)
= Echinopsis hystrix (Rauh & Backeb.) Anceschi & Magli; Cactus in Habitat S. Amer. 2013/2021: 38 (2021)
= Echinopsis peculiaris (Rauh & Backeb.) Molinari & Mayta; Succulentopia 14: 20 (2015)
= Haageocereus andinus F.Ritter; (nomen)
= Haageocereus deserticola F.Ritter; (nomen)
= Haageocereus hystrix F.Ritter; (nomen)
= Haageocereus montanus F.Ritter; (nomen)
= Haageocereus peculiaris (Rauh & Backeb.) F.Ritter; Backeberg, Descr. Erört. Taxa Fragen Sp. 14 (1958)
= Loxanthocereus clavispinus Rauh & Backeb.; Descr. Cact. Nov. [Backeberg] 15 (1957)
= Loxanthocereus hystrix Rauh & Backeb.; Descr. Cact. Nov. [Backeberg] 15 (1957)
= Loxanthocereus hystrix var. brunnescens Rauh; Sitzungsber. Heidelberger Akad. Wiss., Math.-Naturwiss. Kl. 1958(1): 312 (1958)
= Loxanthocereus peculiaris Rauh & Backeb.; Descr. Cact. Nov. [Backeberg] 1: 16 (1956 publ. 1957)
= Loxanthocereus ×deserticola F.Ritter; Kakteen Südamerika 4: 1458 (1981)
= Loxanthocereus ×ferrugineus Rauh & Backeb.; Descr. Cact. Nov. [Backeberg] 15 (1957)
= Loxanthocereus ×montanus F.Ritter; Kakteen Südamerika 4: 1463 (1981)
species :
Borzicactus icosagonus (Kunth) Britton & Rose; Cactaceae [Britton & Rose] 2: 160 (1920)
subspecies :
subsp. icosagonus
Distribution: S-Ecuador; N-Peru
83 ECU PER
Remarks: often included in Matucana
Synonyms: hidden
= Binghamia aurivilla (Britton & Rose) Backeb.; Blätt. Kakteenf. 1: 7 (1935)
= Binghamia icosagona (Kunth) Backeb.; Kaktus-ABC [Backeberg & Knuth] 197 (1935)
= Borzicactus aurivillus (K.Schum.) Britton & Rose; Cactaceae [Britton & Rose] 2: 163, 226 (1920)
= Borzicactus crassiserpens (Rauh & Backeb.) Lodé; Cact.-Advent. Int. (Almeria) 98(Suppl.): 3 (2013)
= Borzicactus ×crassiserpens (Rauh & Backeb.) Ostolaza; Cactus Perú 266 (2014)
= Borzicactus icosagonus f. aurantiaciflorus (Backeb.) Krainz; Kakteen [H.Krainz] 15: C Vb (1960)
= Cactus icosagonus Kunth; Nov. Gen. Sp. [H.B.K.] 6: 67 (1823)
= Cereus aurivillus K.Schum.; Monatsschr. Kakteenk. 13: 67 (1903)
= Cereus icosagonus (Kunth) DC.; Prodr. [A.P. de Candolle] 3: 467 (1828)
= Cereus isogonus K.Schum.; Gesamtbeschr. Kakt. 102 (1897)
= Cleistocactus ×crassiserpens Rauh & Backeb.; Descr. Cact. Nov. [Backeberg] 1: 17 (1956 publ. 1957)
= Cleistocactus icosagonus (Kunth) F.A.C.Weber; Cleistocactus 6 (1904)
= Loxanthocereus crassiserpens (Rauh & Backeb.) Backeb.; Cactaceae [Backeberg] 6: 3674 (1962)
= Loxanthocereus trujilloensis F.Ritter; Kakteen Südamerika 4: 1472 (1981)
= Matucana icosagona (Kunth) Buxb.; Kakteen [H.Krainz] 54: CVb (1973)
= Seticereus aurivillus Backeb.; Cactaceae (Berlin) 1937(2): 14 (1937)
= Seticereus ferrugineus Backeb.; Jahrb. Deutsch. Kakteen-Ges. Teil 2 (Okt.), Blatt 14, Kakt. u. Sukkulent. 1937, 185 (1937)
= Seticereus icosagonus (K.Schum.) Backeb.; Cactaceae (Berlin) 1937(2): 14 (1937)
= Seticereus icosagonus var. aurantiaciflorus Backeb.; Kakteenkunde 1943: 31 (1943)
= Seticereus icosagonus var. ferruginea (Backeb.) Backeb.; 406 (1966)
= Seticereus icosagonus var. oehmeanus (Backeb.) Backeb.; Cactaceae [Backeberg] 2: 981 (1959)
= Seticereus oehmeanus Backeb.; Jahrb. Deutsch. Kakteen-Ges. Teil 2 (Okt.), Blatt 14, Kakt. u. Sukkulent. 1937, 186 (1937)
= Seticereus oehmeanus var. ferrugineus (Backeb.) Backeb.; Kakteenkunde 1943: 31 (1943)
subspecies :
subsp. humboldtii (Kunth) G.J.Charles; Bradleya 30: 101 (2012)
Distribution: Peru
83 PER
Synonyms: hidden
= Binghamia humboldtii (Kunth) Backeb.; Blätt. Kakteenf. 10: 8 (1934)
= Borzicactus humboldtii (Kunth) Britton & Rose; Cactaceae [Britton & Rose] 2: 163 (1920)
= Cactus humboldtii Kunth; Nov. Gen. Sp. [H.B.K.] 6: 66 (1823)
= Cereus humboldtii (Kunth) DC.; Prodr. [A.P. de Candolle] 3: 467 (1828)
= Cleistocactus humboldtii (Kunth) F.A.C.Weber; Roland Goss. Cleistocactus 5 (1904)
= Matucana humboldtii (Kunth) Buxb.; Kakteen [H.Krainz] 54: CVb (1973)
= Seticereus humboldtii (Kunth) Backeb.; Blätt. Kakteenf. No. 11, p. 6 (1937)
subspecies :
subsp. roseiflorus (Buining) G.J.Charles; Bradleya 30: 101 (2012)
Distribution: Ecuador
83 ECU
Synonyms: hidden
= Akersia roseiflora Buining; Succulenta (Netherlands) 1961: 25 (1961)
= Borzicactus roseiflorus (Buining) Ostolaza; Cactus Perú 264 (2014)
= Cleistocactus roseiflorus (Buining) G.D.Rowley; Brit. Cact. Succ. J. 18: 208 (2000)
= Matucana roseiflora (G.J.Charles) G.J.Charles & Hoxey; CactusWorld 40(2): 82 (2022)
species :
Borzicactus jajoianus (Backeb.) Backeb.; Kaktus-ABC [Backeberg & Knuth] 194, 411 (1935)
Distribution: Ecuador
83 ECU
Synonyms: hidden
= Cereus jajoianus Backeb.; Cact. J. (Croydon) 2: 4 (1933)
= Cleistocactus jajoianus (Backeb.) G.J.Charles; Cactaceae Syst. Init. 20: 13 (2005)
= Echinopsis kellyana Mayta; Succulentopia 14: 14 (2015), nom. nov. for Cereus jajoanus Backeb.
= Loxanthocereus jajoianus (Backeb.) Backeb.; Stachlige Wildnis 187 (1942)
= Loxanthocereus riomajensis Rauh & Backeb.; Descr. Cact. Nov. [Backeberg] 1: 15 (1956 publ. 1957)
species :
Borzicactus leonensis (Madsen) G.J.Charles; Cactaceae Syst. Init. 2: 14 (2012)
Distribution: Ecuador
83 ECU
Synonyms: hidden
= Borzicactus sepium subsp. leonensis (Madsen) Guiggi; Cactology 5(Suppl. II): 5 (2016)
= Cleistocactus leonensis Madsen; Fl. Ecuador 35(45): 18 (1989)
species :
Borzicactus neoroezlii F.Ritter; Kakteen And. Sukk. 12: 55 (1961)
Distribution: ?E-Ecuador; Peru (Olmos, Lambayeque)
83 PER
Synonyms: hidden
= Cleistocactus neoroezlii (F.Ritter) Buxb.; Kakteen [H.Krainz] 57: CVb (1974), without basionym vol. or page.
species :
Borzicactus pachycladus (Rauh & Backeb.) G.J.Charles; Cactaceae Syst. Init. 2: 14 (2012)
Distribution: C-Peru
83 PER
Synonyms: hidden
= Borzicactus piscoensis (Rauh & Backeb.) Rauh & Backeb.; Sitzungsber. Heidelberger Akad. Wiss., Math.-Naturwiss. Kl., Abh. 1 (Beitr. Peruan. Kakt.): 291 (1958)
= Cleistocactus pachycladus (Rauh & Backeb.) Ostolaza; Cactaceae Consensus Init. 6: 8 (1998)
= Echinopsis pachyclada (Rauh & Backeb.) Molinari; Succulentopia 13: 19 (2015)
= Loxanthocereus pachycladus Rauh & Backeb.; Descr. Cact. Nov. [Backeberg] 16 (1957)
= Loxanthocereus piscoensis Rauh & Backeb.; Descr. Cact. Nov. [Backeberg] 1: 16 (1956 publ. 1957)
= Loxanthocereus yauyosensis F.Ritter; Kakteen Südamerika 4: 1476 (1981)
= Oreocereus piscoensis (Rauh & Backeb.) F.Ritter; Kakteen Südamerika 4: 1365 (1981)
species :
Borzicactus plagiostoma (Vaupel) Britton & Rose; Cactaceae [Britton & Rose] 2: 163 (1920)
Distribution: Peru (Ancash to Cajamarca)
83 PER
Synonyms: hidden
= Binghamia plagiostoma (Vaupel) Backeb.
= Borzicactus fieldianus subsp. samnensis (F.Ritter) Ostolaza; Cactus Perú 260 (2014)
= Borzicactus purpureus F.Ritter; Kakteen Südamerika 4: 1383 (1981)
= Borzicactus samnensis F.Ritter; Taxon 13: 118 (1964)
= Cereus plagiostoma Vaupel; Bot. Jahrb. Syst. 50, Beibl. 111: 20 (1913)
= Cleistocactus fieldianus subsp. samnensis (F.Ritter) Ostolaza; Cactaceae Consensus Init. 6: 8 (1998)
= Cleistocactus plagiostoma (Vaupel) D.R.Hunt; Cactaceae Consensus Init. 2: 3 (1997)
= Clistanthocereus samnensis (F.Ritter) Backeb.; Kakteenlexikon 91 (1966)
= Echinopsis plagiostoma (Vaupel) Molinari & Mayta; Succulentopia 14: 20 (2015)
species :
Borzicactus samaipatanus (Cárdenas) Kimnach; Cact. Succ. J. (Los Angeles) 32: 93 (1960)
Distribution: Bolivia (Santa Cruz)
83 BOL
Synonyms: hidden
= Bolivicereus brevicaulis F.Ritter; Kakteen Südamerika 2: 703 (1980)
= Bolivicereus croceus F.Ritter; Kakteen Südamerika 2: 704 (1980)
= Bolivicereus rufus F.Ritter; Kakteen Südamerika 2: 703 (1980)
= Bolivicereus samaipatanus Cárdenas; Cact. Succ. J. (Los Angeles) 23: 91 (1951)
= Bolivicereus samaipatanus subsp. divi-miseratus (Cárdenas) Guiggi; Cactology 3: 6 (2012)
= Bolivicereus samaipatanus var. divi-miseratus Cárdenas ex Backeb.; Cactaceae [Backeberg] 2: 973 (1959)
= Bolivicereus samaipatanus var. multiflorus Cárdenas; Cact. Succ. J. (Los Angeles) 23(3): 93 (1951)
= Borzicactus samaipatanus var. divi-miseratus (Cárdenas ex Backeb.) Krainz; Kakteen [H.Krainz] 21: C Vb (1962)
= Borzicactus samaipatanus var. multiflorus (Cárdenas) Krainz; Kakteen [H.Krainz] 21: C Vb (1962)
= Cleistocactus samaipatanus (Cárdenas) D.R.Hunt; Bradleya 5: 92 (1987)
= Cleistocactus samaipatanus subsp. divi-miseratus (Cárdenas ex Backeb.) M.Lowry; Bradleya 34: 171 (2016)
= Echinopsis samaipatana (Cárdenas) Anceschi & Magli; Cactus in Habitat S. Amer. 2011/2013: 39 (2013)
species :
Borzicactus sepium (Kunth) Britton & Rose; Cactaceae [Britton & Rose] 2: 160 (1920)
Distribution: Colombia (Nariño); C-Ecuador; N-Ecuador
82 CLM 83 ECU
Synonyms: hidden
= Borzicactus aequatorialis Backeb.; Descr. Cact. Nov. [Backeberg] 1: 17 (1956 publ. 1957)
= Borzicactus roezlii (F.Haage ex Schum.) Backeb.; Blätt. Kakteenf. No. 12, p. 6 (1937)
= Borzicactus sepium subsp. ventimigliae (Riccob.) Lodé; Int. Cact. Advent. 31(1): 23 (2019)
= Borzicactus sepium subsp. websterianus (Backeb.) Lodé; Int. Cact. Advent. 31(1): 25 (2019)
= Borzicactus sepium var. roezlii (F.Haage ex K.Schum.) Borg; Cacti [Borg] 147 (1937)
= Borzicactus ventimigliae Riccob.; Boll. Reale Orto Bot. Palermo 8: 262 (1909)
= Borzicactus websterianus Backeb.; Blätt. Kakteenf. No. 11, P. 2 (1937)
= Borzicactus websterianus var. rufispinus Backeb.; Blätt. Kakteenf. 11: 2 (1937)
= Cactus sepium Kunth; Nov. Gen. Sp. [H.B.K.] 6: 66 (1823)
= Cereus chotaensis (F.A.C.Weber) Vaupel; Monatsschr. Kakteenk. 23: 25, 38 (1913)
= Cereus roezlii Backeb.; Cact. J. (Croydon) 2: 33 (1933), nom. illeg.
= Cereus roezlii F.Haage ex K.Schum.; Gesamtbeschr. Kakt. 64 (1897)
= Cereus sepium (Kunth) DC.; Prodr. [A.P. de Candolle] 3: 467 (1828)
= Cereus ventimigliae (Riccob.) Vaupel; Monatsschr. Kakteenk. 23: 13, 37 (1913)
= Cleistocactus chotaensis F.A.C.Weber; Roland Goss. Cleistoc. 19 (1904)
= Cleistocactus roezlii (F.Haage ex K.Schum.) Backeb.; Kaktus-ABC [Backeberg & Knuth] 190 (1936)
= Cleistocactus sepium (Kunth) F.A.C.Weber; Rol.-Goss. Cleistoc. 8 (1904)
= Cleistocactus sepium var. ventimigliae (Riccob.) Madsen; Fl. Ecuador 35(45): 22 (1989)
= Seticereus roezlii (F.Haage ex K.Schum.) Backeb.; Sukkulentenkunde 3: 5 (1949)
variant :
var. morleyanus (Britton & Rose) Krainz; Kakteen [H.Krainz] 46-47: CVb. (1971)
Distribution: Ecuador
83 ECU
Synonyms: hidden
= Borzicactus morleyanus Britton & Rose; Cactaceae [Britton & Rose] 2: 160 (1920)
= Borzicactus sepium subsp. morleyanus (Britton & Rose) Lodé; Int. Cact. Advent. 31(1): 22 (2019)
= Cleistocactus sepium var. morleyanus (Britton & Rose) Madsen; Fl. Ecuador 35(45): 21 (1989)
species :
Borzicactus serpens (Kunth) Kimnach; Cact. Succ. J. (Los Angeles) 32: 95 (1960)
Distribution: Peru
83 PER
Synonyms: hidden
= Bolivicereus serpens (Kunth) Backeb.; Cactaceae [Backeberg] 6: 3476 (1962)
= Borzicactella serpens (Kunth) F.Ritter; Kakteen Südamerika 4: 1387 (1981)
= Borzicactella serpens subsp. erecta (G.J.Charles) Guiggi; Cactology 5(Suppl. II): 1 (2016)
= Borzicactus longiserpens (Leuenb.) G.J.Charles; Bradleya 28: 7 (2010)
= Borzicactus longiserpens subsp. erectus G.J.Charles; Bradleya 28: 8 (2010)
= Borzicactus serpens subsp. erectus (G.J.Charles) Ostolaza; Cactus Perú 266 (2014)
= Cactus serpens Kunth; Nov. Gen. Sp. [H.B.K.] 6: 68 (1823)
= Cereaster sepens (Kunth) F.Berge; Buch Welt, ed. 2: 344 (1843)
= Cereus serpens (Kunth) DC.; Prodr. [A.P. de Candolle] 3: 470 (1828)
= Cleistocactus longiserpens Leuenb.; Bot. Jahrb. Syst. 124(1): 23 (2002), nom. nov.
= Cleistocactus serpens (Kunth) F.A.C.Weber; Cleistocactus 11 (1904)
= Echinopsis serpens (Kunth) Molinari; Succulentopia 13: 19 (2015)
species :
Borzicactus sextonianus (Backeb.) Kimnach; Cact. Succ. J. (Los Angeles) 32: 95 (1960)
Distribution: W-Peru
83 PER
Synonyms: hidden
= Borzicactus gracilis (Akers & Buining) Buxb. & Krainz; Kakteen [H.Krainz] 21: C vb. (1962)
= Borzicactus gracilis var. aticensis (Rauh & Backeb.) Krainz; Kakteen [H.Krainz] 21: C Vb (1962)
= Borzicactus gracilis var. camanaensis (Rauh & Backeb.) Krainz; Kakteen [H.Krainz] 21: C Vb (1962)
= Cereus sextonianus Backeb.; Cact. J. (Croydon) 2: 4 (1933), nomen
= Cleistocactus sextonianus (Backeb.) D.R.Hunt; Bradleya 9: 86 (1991), et Cactaceae Syst. Init. 15: 8 (2003)
= Echinopsis sextoniana (Backeb.) Mayta; Succulentopia 14: 15 (2015)
= Erdisia sextoniana (Backeb.) Backeb.; Cact. J. (Croydon) 2: 4 (1933), et Backeb. & Knuth, Kaktus-ABC,174, 411 (1935), cum descr.
= Loxanthocereus aticensis Rauh & Backeb.; Descr. Cact. Nov. [Backeberg] 1: 15 (1956 publ. 1957)
= Loxanthocereus camanaensis Rauh & Backeb.; Descr. Cact. Nov. [Backeberg] 15 (1957)
= Loxanthocereus gracilis (Akers & Buining) Backeb.; Sitzungsber. Heidelberger Akad. Wiss., Math.-Naturwiss. Kl., Abh. 1 (Beitr. Peruan. Kakt.): 300 (1958)
= Loxanthocereus nanus Backeb.; Descr. Cact. Nov. [Backeberg] 17 (1957)
= Loxanthocereus puquiensis F.Ritter; Kakteen Südamerika 4: 1470 (1981)
= Loxanthocereus sextonianus (Backeb.) Backeb.; Stachlige Wildnis 187 (1942)
= Loxanthocereus splendens Backeb.; Descr. Cact. Nov. [Backeberg] 14 (1957)
= Loxanthocereus variabilis F.Ritter; Kakteen Südamerika 4: 1473 (1981)
= Maritimocereus gracilis Akers & Buining; Succulenta (Netherlands) 50 (1950)
species :
Borzicactus silvaticus Janke; Kakteen And. Sukk. 70(2): 50 (2019)
Distribution: Peru
83 PER
Remarks: provisionally accepted name, status not confirmed by molecular data
species :
Borzicactus strausii (Heese) A.Berger; Entwicklungslin. Kakt. 53 (1926)
Distribution: Bolivia (Tarija)
83 BOL
Synonyms: hidden
= Cephalocereus strausii (Heese) Houghton; J. Cact. Succ. Soc. Amer. 1: 170 (1930)
= Cereus strausii (Heese) Vaupel; Monatsschr. Kakteenk. 23: 37 (1913)
= Cleistocactus strausii (Heese) Backeb.; Kakteen-Freund 3: 121 (Nov. 1934), Issue 146, Backeb. in Cact. J. 3: 25 (Dec. 1934)
= Denmoza strausii (Heese) Fric; Möller´s Deutsche Gärtn.-Zeitung 44: 170 (1929)
= Denmoza strausii var. luteispina Fric; Möller´s Deutsche Gärtn.-Zeitung 44: 170 (1929)
= Echinopsis nothostrausii Anceschi & Magli; Cactus in Habitat S. Amer. 2011/2013: 39 (2013)
= Echinopsis strausii (Heese) Anceschi & Magli; Cactus in Habitat S. Amer. 2013/2021: 39 (2021)
= Pilocereus strausii f. cristatus Dörfl.; Monatsschr. Deutsch. Kakteen-Ges. 2: 54 (1930)
= Pilocereus strausii Heese; Gartenflora 56: 410 (1907)
species :
Borzicactus sulcifer (Rauh & Backeb.) Kimnach; Cact. Succ. J. (Los Angeles) 32: 95 (1960)
Distribution: S-Ecuador; N-Peru
83 ECU PER
Synonyms: hidden
= Borzicactus pseudothelegonus (Rauh & Backeb.) Rauh & Backeb.; Descr. Cact. Nov. [Backeberg] 3: 5 (1963)
= Cereus pseudothelegonus Rauh & Backeb.; Cactaceae [Backeberg] 2: 1136 (1959)
= Cleistocactus granditessellatus (Rauh & Backeb.) Leuenb.; Bot. Jahrb. Syst. 124(1): 20 (2002)
= Cleistocactus sulcifer (Rauh & Backeb.) Leuenb.; Bot. Jahrb. Syst. 124(1): 22 (2002)
= Loxanthocereus granditessellatus Rauh & Backeb.; Descr. Cact. Nov. [Backeberg] 1: 15 (1956 publ. 1957)
= Loxanthocereus otuscensis F.Ritter; Kakteen Südamerika 4: 1467 (1981)
= Loxanthocereus parvitesselatus F.Ritter; Kakteen Südamerika 4: 1469 (1981)
= Loxanthocereus sulcifer Rauh & Backeb.; Descr. Cact. Nov. [Backeberg] 1: 14 (1956 publ. 1957)
= Loxanthocereus sulcifer var. longispinus Rauh & Backeb.; Descr. Cact. Nov. [Backeberg] 1: 14 (1956 publ. 1957)
species :
Borzicactus tenuiserpens (Rauh & Backeb.) Kimnach; Cact. Succ. J. (Los Angeles) 32: 95 (1960)
Distribution: N-Peru
83 PER
Synonyms: hidden
= Bolivicereus tenuiserpens (Rauh & Backeb.) Backeb.; Cactaceae [Backeberg] 6: 3677 (1962)
= Borzicactella tenuiserpens (Rauh & Backeb.) F.Ritter; Kakteen Südamerika 4: 1387 (1981)
= Cleistocactus tenuiserpens Rauh & Backeb.; Descr. Cact. Nov. [Backeberg] 1: 17 (1956 publ. 1957)
species :
Borzicactus tominensis (Weing.) Borg; Cacti [Borg] 147 (1937)
Distribution: Bolivia (Chuquisaca, Azurduy Prov.)
83 BOL
Synonyms: hidden
= Cereus tominensis Weing.; Monatsschr. Deutsch. Kakteen-Ges. 3: 117 (1931)
= Cleistocactus capadalensis F.Ritter; Kakteen Südamerika 2: 677 (1980)
= Cleistocactus clavicaulis Cárdenas; Cactus (Paris) No. 80-81, 20 (1964)
= Cleistocactus crassicaulis Cárdenas; Cact. Succ. J. (Los Angeles) 33: 77 (1961)
= Cleistocactus crassicaulis var. paucispinus F.Ritter; Kakteen Südamerika 2: 674 (1980)
= Cleistocactus mendozae Cárdenas; Cact. Succ. J. (Los Angeles) 35: 202 (1963)
= Cleistocactus micropetalus F.Ritter; Kakteen Südamerika 2: 675 (1980)
= Cleistocactus tominensis (Weing.) Backeb.; Kaktus-ABC [Backeberg & Knuth] 190 (1935)
= Cleistocactus tominensis subsp. micropetalus (F.Ritter) Mottram; Cactaceae Syst. Init. 13: 12 (2002)
= Cleistocactus viridialabastri Cárdenas; Cact. Succ. J. (Los Angeles) 35: 201 (1963)
= Echinopsis micropetala (F.Ritter) Anceschi & Magli; Cactus in Habitat S. Amer. 2013/2021: 39 (2021)
= Echinopsis tominensis (Weing.) Anceschi & Magli; Cactus in Habitat S. Amer. 2011/2013: 40 (2013)
species :
Borzicactus xylorhizus (F.Ritter) G.J.Charles; Cactaceae Syst. Init. 2: 14 (2012)
Distribution: Peru (Lima)
83 PER
Synonyms: hidden
= Cleistocactus xylorhizus (F.Ritter) Ostolaza; Brit. Cact. Succ. J. 14(4): 170 (1996)
= Echinopsis xylorhiza (F.Ritter) Molinari; Succulentopia 13: 19 (2015)
= Loxanthocereus xylorhizus F.Ritter; Kakteen Südamerika 4: 1475 (1981)
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A complete, synonymic checklist of the Higher Plants of the World


Version 25.01 (January 21th, 2025)

Covering 355,968 accepted species,
8,748 hybrids, 47,863 accepted infraspecific taxa and 1,002,228 synonyms,
in total 1,418,722* names

[*about 25,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus" are deliberately omitted]


With complete, detailed distribution data (countries and regions) and literature references

Arranged in linear phylogenetic sequence reflecting true relationships

Fully searchable and continously updated

With 38,209 pictures of 15,787 species

Assembled through 40 years independently from other databases

In cooperation with Catalogue of Life