030.0000
Family
Pteridaceae E.D.M.Kirchn.; Schul-Bot. 109 (1831)
48 genera, ca. 1350 spp. The family has been split extensively in smaller subgroups, but Smith et al. (2006) and PPG-I argue for keeping them together.
Species listed in database: 1406; hybrids: 68
= Acrostichaceae Mett. ex A.B.Frank; Syn. Pflanzenk., ed. 2, 3: 1458 (1877), nom. rej.
= Actiniopteridaceae Pic.Serm.; Webbia 17: 5 (1962)
= Adiantaceae Newman; Hist. Brit. Ferns 5 (1840) nom. cons. over Parkeriaceae (1825)
= Anopteraceae Doweld; Tent. Syst. Pl. Vasc. (Tracheophyta) 51 (2001)
= Antrophyaceae (Link) Ching; Acta Phytotax. Sin. 16: 11 (1978)
= Ceratopteridaceae Underw.; Native Ferns, ed. 6: 65, 78 (1900)
= Cheilanthaceae B.K.Nayar; Taxon 19: 233 (1970)
= Cryptogrammaceae Pic.Serm.; Webbia 17: 299 (1963)
= Hemionitidaceae Pic.Serm.; Webbia 21: 487 (1966)
= Negripteridaceae Pic.Serm.; Nuovo Giorn. Bot. Ital., n. s. 53: 160 (1946)
= Parkeriaceae Hook.; Exot. Fl. 2: t. 147 (1825), nom. rej. in favor of Adiantaceae
= Platyzomataceae Nakai; Bull. Natl. Sci. Mus., Tokyo, 29: 4 (1950)
= Pteridaceae Reichenb.; Handb. Nat. Pfl.-Syst. 138 (1837)
= Sinopteridaceae Koidz.; Acta Phytotax. Geobot. 3: 50 (1934)
= Taenitidaceae (C.Presl) Pic.Serm.; Webbia 29: 1 (1974)
= Vittariaceae Ching; Sunyatsenia 5: 232 (1940)
030.6017
Genus
Aleuritopteris Fée; Mém. Foug. 5 (Gen. Filic.) 148 (1852)
Distribution: Mostly China, India, Indochina, 3 Africa, few neotropics. Allosorus was wrongly lectotypified and was rejected as name. Inclusion of Oeosporangium and Mildella fide molecular results of Schuettpelz et al. 2025.
Species listed in database: 78; hybrids: 21
= Allosorus Bernh.; Neues J. Bot. 1(2): 36 (1805), nom. rej.
= Gymnia Hamilton ex D.Don; Prodr. Fl. Nepal. 16 (1825)
= Leptolepidium K.H.Shing & S.K.Wu; Acta Bot. Yunnan. 1(1): 115 (1979)
= Mildella Trevis.; Rendiconti Reale Ist. Lombardo Sci., Ser. 2, 9(20): 810 (1876)
= Negripteris Pic.Serm.; Nuovo Giorn. Bot. Ital., n. s. 53: 130, t. 14-16 (1946) (fide Fraser-Jenkins 2008)
= Oeosporangium Vis.; Atti Reale Ist. Veneto Sci. Lett. Arti, Ser. 3, 12: 656 (1867)
= Parallosorus Keyserl.; Polyp. Herb. Bunge 5 (1873)
= Sinopteris C.Chr. & Ching; Bull. Fan Mem. Inst. Biol., Bot. 4: 359 (1933)
Search for: Aleuritopteris argentea-var.-obscura
Nothing found!
species :
Aleuritopteris acrostica
(Balb.) Windham & Schuettp.; Taxon 74(2), p. 11 of 32 (2025)
Distribution: Portugal; Spain; Andorra; Gibraltar; Baleares; Croatia; Greece (widespread, but not North, incl. Sporades, Kiklades); Crete (widespread); Morocco; Algeria; Tunisia; Libya; Cape Verde Isl. (Fogo Isl.); ?Ethiopia; Turkmenistan; Tajikistan; East Aegaean Isl. (widespread, incl. Rhodos); Afghanistan; Pakistan (Baluchistan, NW-Pakistan, Chitral, Murree, Hazara); India (Himachal Pradesh, Jammu & Kashmir)
12 ADO BAL GIB POR SPA 13 CRO GRC KRI 20 ALG LBY MOR TUN 21 CVI 32 TKM TZK 34 AFG EAI 40 IND PAK
Remarks: allotetraploid between A. pteridioides and A. persicum, therefore best to be recognized at species level. For a long time known as Cheilanthes pteridioides, while Cheilanthes maderensis in the previous sense is todays A. pteridioides
Synonyms: hidden
= Adiantum odorum (Sw.) DC.; Fl. Franç. [de Candolle & Lamarck], éd. 3, 6: 237 (1815)
= Allosorus acrosticus (Balb.) Christenh.; Willdenowia 42(2): 284 (2012)
= Allosorus pusillus Bernh.; Neues J. Bot. 12: 36 (1806), nom. superfl.
= Cheilanthes acrostica (Balb.) Tod.; Giorn. Sci. Nat. Econ. Palermo 1: 215 (seors 10) (1866)
= Cheilanthes fragrans var. acrostica (Balb.) Fiori; Fl. Ital. Crypt. 5: 267 (1943)
= Cheilanthes fragrans var. paleacea Pamp.; Pl. Tripolitanae 11 (1914)
= Cheilanthes odora Sw.; Syn. Fil. (Kiliae) [O.Swartz] 127, 327 (1806), nom. superfl.
= Cheilanthes pteridioides f. paleacea (Pamp.) Maire & Weiller; Encycl. Biol. 33: 71 (1952)
= Cheilanthes pteridioides subsp. acrostica (Balb.) O.Bolòs, Vigo, Masalles & Ninot; Fl. Man. Paisos Catalans 1213 (1990)
= Hemionitis acrostica (Balb.) Mosyakin; Phytotaxa 373(2): 164-168 (2018)
= Hemionitis atreyu Christenh.; Global Fl. 4: 10 (2018), nom. nov. superfl. for Cheilanthes acrostica Tod.
= Oeosporangium acrosticum (Balb.) L.Sáez & Aymerich; Orsis 31: 33 (2017)
= Oeosporangium pteridioides subsp. acrosticum (Balb.) Fraser-Jenk. & Pariyar; Annot. Checkl. Ind. Pterid. 1: 259 (2016)
= Pteris acrostica Balb.; Elenco 98 (1801)
species :
Aleuritopteris agetae
Saiki; J. Phytogeogr. Taxon. 32(1): 1 (1984)
Distribution: Taiwan
38 TAI
Remarks: included in A. krameri by Flora of China, but kept separate by Knapp (Ferns of Taiwan)
Synonyms: hidden
= Cheilanthes agetae (Saiki) C.M.Kuo; Taiwania 30: 57 (1985)
= Hemionitis agetae (Saiki) Christenh.; Global Fl. 4: 9 (2018)
species :
Aleuritopteris albofusca
(Baker) Pic.Serm.; Nuovo Giorn. Bot. Ital., n. s. 53: 154 (1946)
Distribution: China (Gansu, Guizhou, Hebei, S-Hunan, Sichuan, Yunnan); Tibet
36 CHI TIB
Synonyms: hidden
= Aleuritopteris mairei (Brause) Ching; Bull. Fan Mem. Inst. Biol., Bot., Ser. 2, 1: 272 (1949)
= Cheilanthes albofusca Baker; Bull. Misc. Inform. Kew 1896: 54 (1896)
= Cheilanthes mairei Brause; Hedwigia 64: 202, t. 4 D (1914)
= Hemionitis albofusca (Baker) Christenh.; Global Fl. 4: 9 (2018)
= Sinopteris albofusca (Baker) Ching; Sunyatsenia 6: 11 (1941)
= Sinopteris hopeiensis C.Chr. & Ching; Bull. Fan Mem. Inst. Biol., Bot. 4: 361 (1933)
species :
Aleuritopteris albomarginata
(C.B.Clarke) Ching; Hong Kong Naturalist 10: 199 (1941)
Distribution: China (Guangxi, Guizhou, Sichuan, Yunnan); Tibet; Pakistan; Nepal; Bhutan; India (Jammu & Kashmir, Arunachal Pradesh, Himachal Pradesh, Maharashtra, Manipur, Meghalaya, Nagaland, Odisha, Sikkim, Uttar Pradesh, Uttarakhand, West Bengal, Darjeeling); Myanmar; Thailand; Vietnam; Philippines
36 CHI TIB 40 BHU IND NEP PAK 41 MYA THA VIE 42 PHI
Synonyms: hidden
= Aleuritopteris dalhousieae (Hook.) Ching; Hong Kong Naturalist 10: 203 (1941), nom. rej.
= Allosorus albomarginatus (C.B.Clarke) Farw.; Amer. Midl. Naturalist 12: 284 (1931)
= Cheilanthes albomarginata C.B.Clarke; Trans. Linn. Soc. London, Bot., Ser. 2, 1: 456, t. 52 (1880)
= Cheilanthes dalhousiae Hook.; Sp. Fil. [W.J.Hooker] 2: 80 (1852), excl. fig. 78B, nom. rej.
= Cheilanthes farinosa var. albomarginata (C.B.Clarke) Bedd.; Ferns Brit. India Suppl. 22 (1892)
= Cheilanthes farinosa var. dalhousiae (Hook.) C.B.Clarke; Trans. Linn. Soc. London, Bot., Ser. 2, 1: 457, t. 51 (1880)
= Hemionitis albomarginata (C.B.Clarke) Christenh.; Global Fl. 4: 9 (2018)
= Hemionitis dalhousiae (Hook.) Christenh.; Global Fl. 4: 12 (2018)
= Leptolepidium dalhousiae (Hook.) K.H.Shing & S.K.Wu; Acta Bot. Yunnan. 1(1): 116 (1979)
species :
Aleuritopteris anceps
(Blanf.) Panigrahi; Bull. Bot. Surv. India 2: 321 (1961)
Distribution: China (Fujian, Guangdong, Guangxi, Guizhou, Hunan, Jiangxi, Sichuan, Yunnan, Zhejiang); Tibet; Pakistan; Nepal; India (Jammu & Kashmir, Assam State, Chandigarh, ?Chhattisgarh, Himachal Pradesh, Karnataka, Kerala, Madhya Pradesh, Maharashtra, Meghalaya, Nagaland, Odisha, Rajasthan, Sikkim, Tamil Nadu, Uttarakhand, West Bengal); Sri Lanka; Bangladesh; Myanmar; Thailand; Java; Lesser Sunda Isl. (Bali, Lombok, Timor)
36 CHI TIB 40 BAN IND NEP PAK SRL 41 MYA THA 42 JAW LSI
Synonyms: hidden
= Aleuritopteris calicicola Ching; Sporae Pterid. Sin. 157, t. 33, f. 20-21 (1976)
= Aleuritopteris farinosa var. anceps (Blanf.) Ching; Hong Kong Naturalist 10: 194 (1941)
= Aleuritopteris javanensis Saiki; J. Phytogeogr. Taxon. 32(1): 7 (1984), non Cheilanthes javanensis (Willd.) T. Moore (fide Fraser-Jenkins 1997)
= Aleuritopteris pseudofarinosa Ching & S.K.Wu; Acta Phytotax. Sin. 19(1): 72 (1981) (fide Fraser-Jenkins 1997)
= Aleuritopteris pseudofarinosa var. glandulosa H.G.Zhou; Guihaia 12(4): 323 (1992) (fide Fraser-Jenkins 1997)
= Aleuritopteris rigidula Saiki; J. Phytogeogr. Taxon. 32(2): 97 (1984) (as "(Wall.) Saiki"), nom. nud. inval.
= Aleuritopteris wuyishanensis Ching; Wuyi Sci. J. 1: 2 (1981) (fide Fl. China)
= Cheilanthes anceps Blanf.; J. Simla Naturalists´ Soc. 1: 21 (1886)
= Cheilanthes candida Zoll.; Natuur- Geneesk. Arch. Ned.-Indië 2: 203 (1845), non M. Martens & Galeotti 1842
= Cheilanthes farinosa var. anceps (Blanf.) Blanf.; J. Asiat. Soc. Bengal, Pt. 2, Nat. Hist. 57, pt. 2(4): 249 (1888)
= Cheilanthes pseudofarinosa (Ching & S.K.Wu) K.Iwats.; Fl. Thailand 3(4): 618 (1989) (fide Fraser-Jenkins 1997)
= Hemionitis anceps (Blanf.) Christenh.; Global Fl. 4: 9 (2018)
species :
Aleuritopteris argentea
(S.G.Gmel.) Fée; Mém. Foug. 5 (Gen. Filic.) 154 (1852)
Name: Silver Cloak fern
Distribution: Siberia (Altai, Buryatia, Chita, Irkutsk, Krasnoyarsk, Tuva, Yakutia); Russian Far East (Amur, Khabarovsk, Primorye); China (Anhui, Beijing, Fujian, Gansu, Guangdong, Guangxi, Guizhou, Hainan, Henan, Heilongjiang, Hebei, Hongkong, Hubei, Hunan, Jiangsu, Jiangxi, Jilin, Liaoning, Macao, Nei Mongol, Ningxia, Qinghai, Shaanxi, Shandong, Shanghai, Shanxi, Sichuan, Xinjiang, Yunnan, Zhejiang); Tibet; Mongolia; North Korea; South Korea (incl. Cheju Isl.); Japan; Ryukyu Isl.; Taiwan; Nepal; Bhutan; India (Sikkim); Myanmar; N-Thailand; South China Sea Isl.
30 SIB 31 RFE 36 CHI TIB 37 MON 38 JAP KOR NKO NNS TAI 40 BHU IND NEP 41 MYA THA 42 SCS
Conservation Status: IUCN CR for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Aleuritopteris argentea var. flava Ching & S.K.Wu; Acta Phytotax. Sin. 19(1): 67 (1981)
= Aleuritopteris argentea var. geraniifolia Ching & S.K.Wu; Acta Phytotax. Sin. 19(1): 67 (1981)
= Aleuritopteris argentea var. major Ching; Hong Kong Naturalist 10(3-4): 197 (1941)
= Aleuritopteris flava (Ching & S.K.Wu) S.R.Ghosh; Pterid. Fl. E. India 388 (2004), nom. illeg.
= Aleuritopteris geraniifolia Ching; Sporae Pterid. Sin. (1976)
= Aleuritopteris michelii (Christ) Ching; Hong Kong Naturalist 10: 198 (1941)
= Aleuritopteris qianguiensis W.M.Chu & H.G.Zhou; Guihaia 12(4): 321 (1992)
= Allosorus argenteus (S.G.Gmel.) C.Presl; Tent. Pterid. 153 (1836)
= Cassebeera argentea (S.G.Gmel.) J.Sm.; J. Bot. [Hooker] 4: 159 (1841)
= Cheilanthes argentea (S.G.Gmel.) Kunze; Linnaea 23: 242 (1852)
= Cheilanthes argentea var. sulphurea Hook.; Fil. Exot., t. 95, f. 3 (1859)
= Doryopteris argentea (S.G.Gmel.) Christ; Bull. Herb. Boissier, Sér. 2, 2: 831 (1902)
= Doryopteris michelii Christ; Bull. Acad. Int. Géogr. Bot., Sér. 3, 20: 14 (1910)
= Dryopteris argentea (S.G.Gmel.) Christ; Bull. Acad. Int. Géogr. Bot., Sér. 3, 11: 231 (1902)
= Hemionitis amarganthica Christenh.; Global Fl. 4: 9 (2018), nom. nov. for P. argentea S. G. Gmel.
= Hemionitis michelii (Christ) Christenh.; Global Fl. 4: 17 (2018)
= Hemionitis qianguiensis (W.M.Chu & H.G.Zhou) Christenh.; Global Fl. 4: 20 (2018)
= Pteris argentea S.G.Gmel.; Novi Comment. Acad. Sci. Imp. Petrop. 12: 519 (1768)
variant :
var. obscura
(Christ) Ching; Hong Kong Naturalist 10: 198 (1941)
Distribution: China (Gansu, Guizhou, Hebei, Henan, Liaoning, Qinghai, Shandong, Shaanxi, Shanxi, Sichuan, Yunnan)
36 CHI
Remarks: A. shensiensis might be a bona sp.
Synonyms: hidden
= Aleuritopteris nuda Ching; Fl. Tsinling. 2: 207 (1974)
= Aleuritopteris shensiensis Ching; Fl. Tsinling. 2: 207 (1974)
= Cheilanthes argentea f. obscura (Christ) Kitag.; J. Jap. Bot. 41: 368 (1966)
= Cheilanthes argentea var. obscura Christ; Nuovo Giorn. Bot. Ital., n. s. 4(1) (1897)
= Hemionitis nuda (Ching) Christenh.; Global Fl. 4: 18 (2018)
= Hemionitis shensiensis (Ching) Christenh.; Global Fl. 4: 21 (2018)
species :
Aleuritopteris belangeri
(Bory) Windham & Schuettp.; Taxon 74(2), p. 12 of 32 (2025)
Name: Belangers spleenwort
Distribution: China (Hainan); Nepal; India (Assam State, Meghalaya, Mizoram, Tripura, West Bengal); Bangladesh; Myanmar; Thailand; Laos; Cambodia; Vietnam; peninsular Malaysia; Borneo (Mt. Kinabalu, Sabah); Philippines (Luzon)
36 CHI 40 BAN IND NEP 41 CBD LAO MYA THA VIE 42 BOR MLY PHI
Synonyms: hidden
= Allosorus caespitosus C.Presl; Tent. Pterid. 152 (1836)
= Allosorus cambodiensis (Baker) Kuntze; Revis. Gen. Pl. 2: 806 (1891)
= Cheilanthes belangeri (Bory) C.Chr.; Index Filic. 172 (1905)
= Cheilanthes griffithiana Fée; Mém. Foug. 5 (Gen. Filic.) 157 (1852)
= Cheilanthes malaccensis Fée; Mém. Foug. 5 (Gen. Filic.) 167 (1852)
= Cheilanthes varians Wall. ex Hook.; Sp. Fil. [W.J.Hooker] 2: 89, t. 103 A (1852)
= Cheilosoria belangeri (Bory) Ching & K.H.Shing; Glossary Lycopods Ferns 39 (1982)
= Hemionitis belangeri (Bory) Christenh.; Global Fl. 4: 10 (2018)
= Oeosporangium belangeri (Bory) Fraser-Jenk.; Annot. Checkl. Ind. Pterid. 1: 255 (2016)
= Pellaea cambodiensis Baker; Ann. Bot. (Oxford) 5: 213 (1891)
= Pteridella belangeri (Bory) Mett. ex Kuhn; Reisen Ost-Afr. [Decken] 3(3): 16 (1879)
= Pteris belangeri Bory; Voy. Indes Or. [Bélanger] 2: 44 (1833)
= Pteris caespitosa Wall.; Numer. List [Wallich] no. 90 (1829), nomen
= Pteris varians Wall.; Numer. List [Wallich] no. 86 (1829), nomen
species :
Aleuritopteris belensis
(Weath. ex Copel.) H.Schneid.; Guide Alpine Subalpine Fl. Mount Jaya [R.J.Johns et al.] 43 (2006)
Distribution: New Guinea (Papua New Guinea, Irian Jaya)
42 NWG
Synonyms: hidden
= Cheilanthes belensis Weath. ex Copel.; Univ. Calif. Publ. Bot. 18: 218 (1942)
= Hemionitis belensis (Weath. ex Copel.) Christenh.; Global Fl. 4: 10 (2018)
species :
Aleuritopteris bicolor
(Roxb.) Fraser-Jenk.; Fern Gaz. 18(5): 223 (2009)
Distribution: Pakistan; Nepal; Bhutan; India (Jammu & Kashmir, Andhra Pradesh, Arunachal Pradesh, Assam State, Bihar, Chandigarh, Chhattisgarh, Goa, Gujarat, Haryana, Himachal Pradesh, Jharkhand, Karnataka, Kerala, Madhya Pradesh, Maharashtra, Manipur, Meghalaya, Mizoram, Odisha, ?Punjab, Rajasthan, Sikkim, Tamil Nadu, Tripura, Uttarakhand, Uttar Pradesh, West Bengal); Sri Lanka; Bangladesh; Myanmar; ?Thailand; Laos; peninsular Malaysia (Bt Baling, Kedah, Bt Chintamani, Pahang); Java; Costa Rica [I]
40 BAN BHU IND NEP PAK SRL 41 LAO MYA 42 JAW MLY 80 cos
Synonyms: hidden
= Aleuritopteris bicolor (Roxb.) Kholia & Punetha; (1995) (comb. inval., sine basionym, fide Fraser-Jenkins 1997)
= Aleuritopteris kathmanduensis Ching & S.K.Wu; Acta Bot. Yunnan. 5(2): 167 (1983) (fide Fraser-Jenkins 1997)
= Ceropteris monosticha Fée; Mém. Foug. 7 (Icon. Esp. Nouv.): 44, t. 22, f. 2 (1857)
= Cheilanthes bicolor (Roxb.) Griff. ex Fraser-Jenk.; Pakistan Syst. 5(1-2): 94 (1992)
= Cheilanthes bullosa Bedd.; Ferns S. India, t. 192 (1863)
= Cheilanthes discolor Griff.; Calcutta J. Nat. Hist. 4: 466 (1844)
= Cheilanthes farinosa var. tenera C.B.Clarke & Baker; J. Linn. Soc., Bot. 25: 441 (1889) (fide Fraser-Jenkins et al. 2015)
= Cheilanthes pulveracea Spreng.; Syst. Veg., ed. 16 [Sprengel] 4(1): 115 (1827)
= Hemionitis bicolor (Roxb.) Christenh.; Global Fl. 4: 10 (2018)
= Pteris argentea Bory; Voy. Îles Afrique 1: 327 (1804), nom. illeg.
= Pteris bicolor Roxb.; Calcutta J. Nat. Hist. 4: 507 (1844)
species :
Aleuritopteris bullosa
(Kunze) Ching; Hong Kong Naturalist 10: 202 (1941)
Distribution: India (Karnataka, Kerala, Tamil Nadu); Sri Lanka
40 IND SRL
Synonyms: hidden
= Aleuritopteris flaccida (Bedd.) B.K.Nayar & S.Kaur; Companion Handb. Ferns Brit. India [Nayar & Kaur] 25 (1974) (comb. inval. fide Fraser-Jenkins (1997))
= Aleuritopteris indica Fée; Mém. Foug. 5 (Gen. Filic.) 154 (1852)
= Cheilanthes bullata Baker; Syn. Fil. [W.J.Hooker & Baker], ed. 1: 137 (1867)
= Cheilanthes bullosa Kunze; Linnaea 24: 274 (1851)
= Cheilanthes farinosa var. bullosa Christ; (1898)
= Cheilanthes farinosa var. flaccida Bedd.; Ferns Brit. India 93 (1883)
= Cheilanthes flaccida (Bedd.) Mehra & Bir; Res. Bull. Punjab Univ., New Ser., Sci. 15: 110 (1964)
= Hemionitis bullosa (Kunze) Christenh.; Global Fl. 4: 11 (2018)
species :
Aleuritopteris chinensis
(Baker) Windham & Schuettp.; Taxon 74(2), p. 12 of 32 (2025)
Distribution: China (Chongqing, N-Guangxi, NE-Guizhou, W-Hubei, Sichuan)
36 CHI
Synonyms: hidden
= Oeosporangium chinense (Baker) Fraser-Jenk.; Annot. Checkl. Ind. Pterid. 2: 451 (2018)
= Cheilanthes chinensis (Baker) Domin; Biblioth. Bot. 86: 133 nota (1913)
= Hemionitis chinensis (Baker) Christenh.; Global Fl. 4: 11 (2018)
= Notholaena chinensis Baker; Gard. Chron., Ser. 2, 14: 494 (1880)
species :
Aleuritopteris chrysophylla
(Hook.) Ching; Hong Kong Naturalist 10: 201 (1941)
Distribution: China (Guangxi, Yunnan, Hainan); Tibet; Nepal; Bhutan; India (Arunachal Pradesh, ?Himachal Pradesh, Meghalaya, Sikkim, Uttarakhand, West Bengal); Myanmar; Thailand; Vietnam
36 CHI TIB 40 BHU IND NEP 41 MYA THA VIE
Synonyms: hidden
= Aleuritopteris flavopygmaea S.R.Ghosh; Pterid. Fl. E. India 388 (2004) (fide Fraser-Jenkins 2008)
= Aleuritopteris humatifolia X.C.Zhang & L.Shi; Acta Phytotax. Sin. 32(1): 93 (1994)
= Cheilanthes argentea var. chrysophylla (Hook.) Hook.; Fil. Exot., t. 95, f. 1, 6-8 (1859), et Syn. Fil. [W.J.Hooker & Baker] 142 (1865)
= Cheilanthes argentea var. sulphurea C.B.Clarke; Trans. Linn. Soc. London, Bot., Ser. 2, 1: ? (1880), non Hook.
= Cheilanthes chrysophylla Hook.; Sp. Fil. [W.J.Hooker] 2: 113 (1852)
= Cheilanthes farinosa var. chrysophylla (Hook.) Bedd.; Ferns Brit. India 93 (1883), isonym
= Cheilanthes farinosa var. chrysophylla (Hook.) C.B.Clarke; Trans. Linn. Soc. London, Bot., Ser. 2, 1: 458 (1880)
= Hemionitis chrysophylla (Hook.) Christenh.; Global Fl. 4: 11 (2018)
species :
Aleuritopteris chusana
(Hook.) Windham & Schuettp.; Taxon 74(2), p. 12 of 32 (2025)
Distribution: China (Anhui, Chongqing, Fujian, Gansu, Guangdong, Guangxi, Guizhou, Henan, Hongkong, Hubei, Hunan, Jiangsu, Jiangxi, Shaanxi, Sichuan, Zhejiang); South Korea (Cheju Isl.); Japan; Taiwan; Laos; Vietnam; Sulawesi; Philippines (Luzon)
36 CHI 38 JAP KOR TAI 41 LAO VIE 42 PHI SUL
Synonyms: hidden
= Adiantopsis fordii (Baker) C.Chr.; Index Filic. 22 (1905)
= Cheilanthes bockii Diels; Bot. Jahrb. Syst. 29: 199 (1900)
= Cheilanthes boltonii Copel.; Fragm. Fl. Philipp. [Perkins] 192 (1905)
= Cheilanthes chusana Hook.; Sp. Fil. [W.J.Hooker] 2: 95, t. 106 B (1852)
= Cheilanthes fordii Baker; J. Bot. 17: 304 (1879)
= Cheilanthes mysurensis var. chusana (Hook.) C.Chr.; Index Filic. 293 (1901)
= Cheilanthes mysurensis var. giraldii Christ; Notul. Syst. (Paris) 1(2): 51 (1909)
= Cheilanthes tenuifolia Hook.; Five Months Yang-Tsze [Blakiston] 362 (1862) (non Sw.)
= Cheilosoria chusana (Hook.) Ching; J. Arnold Arbor. 64(1): 19 (1983)
= Hemionitis chusana (Hook.) Christenh.; Global Fl. 4: 11 (2018)
= Oeosporangium chusanum (Hook.) Fraser-Jenk.; Annot. Checkl. Ind. Pterid. 1: 256 (2016)
species :
Aleuritopteris coriacea
(Decne.) Windham & Schuettp.; Taxon 74(2), p. 12 of 32 (2025)
Distribution: SE-Egypt; Niger; Chad; Sudan; Ethiopia; Eritrea; Djibouti; Somalia; Tanzania; SW-Iran; Saudi Arabia; Oman; Yemen (N-Yemen, S-Yemen)
20 EGY 22 NGR 24 CHA DJI ERI ETH SOM SUD 25 TAN 34 IRN 35 OMA SAU YEM
Synonyms: hidden
= Allosorus coriaceus (Decne.) Christenh.; Willdenowia 42(2): 284 (2012)
= Cheilanthes arabica (Decne.) Buchinger; Flora 303 (1846)
= Cheilanthes coriacea Decne.; Arch. Mus. Hist. Nat. 2: 190 (1841)
= Cheilanthes decaisnei Kunze; Index Sem. (Leipzig) 1850: 23, 307 (1850)
= Hemionitis morla Christenh.; Global Fl. 4: 18 (2018), nom. nov. for Cheilanthes coriacea Decne.
= Oeosporangium coriaceum (Decne.) Fraser-Jenk. & Pariyar; Annot. Checkl. Ind. Pterid. 1: 264 (2016)
= Pellaea abyssinica Fée; Mém. Foug. 6 (Icon. Esp. Nouv.): sub t. 3 (1853)
= Pellaea arabica Fée; Mém. Foug. 5 (Gen. Filic.) 130 (1852)
= Pellaea lomarioides Baker; Bull. Misc. Inform. Kew 1895: 229 (1895)
species :
Aleuritopteris dealbata
(C.Presl) Fée; Mém. Foug. 5 (Gen. Filic.) 154, t. 12 B, f. 2 (1852)
Distribution: China (Guizhou, Yunnan); Nepal; Bhutan; India (Assam State, Meghalaya, Sikkim, Uttarakhand, West Bengal); Myanmar; Thailand
36 CHI 40 BHU IND NEP 41 MYA THA
Synonyms: hidden
= Aleuritopteris doniana S.K.Wu; Acta Bot. Yunnan. 5(2): 169 (1983) (nom. nov. superfl.) ex Ching, Indian Fern J. 1(1-2): 49 (1984 publ. 1985)
= Aleuritopteris gymnocarpa Ching; Sporae Pterid. Sin. 158 (1976), nom. nud.
= Aleuritopteris interrupta Saiki; J. Phytogeogr. Taxon. 32(1): 5 (1984) (fide Fraser-Jenkins et al. 2015)
= Aleuritopteris sikkimensis S.R.Ghosh; Pterid. Fl. E. India 395 (2004) (fide Fraser-Jenkins 2008)
= Allosorus dealbatus C.Presl; Tent. Pterid. 153 (1836)
= Cheilanthes dealbata D.Don; Prodr. Fl. Nepal. 16 (1825) (non Pursh (1814))
= Cheilanthes doniana (S.K.Wu) Fraser-Jenk. & Khullar; Ill. Fern Fl. W. Himalaya 1: 200 (1994), nom. nov.
= Cheilanthes farinosa var. dealbata (C.Presl) C.Chr.; Index Filic. 37 (1905)
= Hemionitis gymnocarpa (Ching) Christenh.; Global Fl. 4: 15 (2018), nom. nud.
species :
Aleuritopteris delicatula
(Tagawa & K.Iwats.) Windham & Schuettp.; Taxon 74(2), p. 12 of 32 (2025)
Distribution: N-Thailand (Chiang Mai)
41 THA
Conservation Status: DD in Thailand
Synonyms: hidden
= Cheilanthes delicatula Tagawa & K.Iwats.; Acta Phytotax. Geobot. 25: 19 (1971)
= Hemionitis delicatessa Christenh.; Global Fl. 4: 12 (2018), nom. nov.
= Oeosporangium delicatulum (Tagawa & K.Iwats.) Fraser-Jenk. & Pariyar; Annot. Checkl. Ind. Pterid. 1: 265 (2016)
species :
Aleuritopteris dubia
(C.Hope) Ching; Hong Kong Naturalist 10: 200 (1941)
Distribution: China (Guizhou, Sichuan, Yunnan); Tibet; Nepal; Bhutan; India (Jammu & Kashmir, Arunachal Pradesh, Himachal Pradesh, Maharashtra, Sikkim, Uttarakhand, West Bengal); Myanmar; Thailand; Vietnam; Philippines
36 CHI TIB 40 BHU IND NEP 41 MYA THA VIE 42 PHI
Synonyms: hidden
= Aleuritopteris humatoides Saiki; J. Phytogeogr. Taxon. 32(1): 5 (1984) (fide Fraser-Jenkins et al. 2015)
= Aleuritopteris subrufa (Baker) Ching; Hong Kong Naturalist 10: 201 (1941) (fide Fraser-Jenkins 1997)
= Cheilanthes dubia C.Hope; J. Bombay Nat. Hist. Soc. 12: 528 t 2 (1899)
= Cheilanthes leveillei Christ; Bull. Acad. Int. Géogr. Bot. 149 (1907)
= Cheilanthes subrufa Baker; Bull. Misc. Inform. Kew 1906: 8 (1906) (fide Fraser-Jenkins 1997)
= Cheilanthes wusukungii Miyam. & H.Ohba; J. Jap. Bot. 72: 168 (1997) (fide Fraser-Jenkins 1997)
= Hemionitis dubia (C.Hope) Christenh.; Global Fl. 4: 13 (2018)
= Hemionitis humatoides (Saiki) Christenh.; Global Fl. 4: 15 (2018)
species :
Aleuritopteris duclouxii
(Christ) Ching; Hong Kong Naturalist 10: 199 (1941)
Distribution: China (Guangxi, Guizhou, Hunan, Sichuan, Yunnan); NE-India (Arunachal Pradesh); Myanmar
36 CHI 40 IND 41 MYA
Conservation Status: IUCN CR for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Aleuritopteris duclouxii var. sulphurea (Ching) Ching; Hong Kong Naturalist 10(3-4): 199-200 (1941)
= Cheilanthes duclouxii (Christ) Ching; Index Filic., Suppl. Tert. 54 (1934)
= Cheilanthes duclouxii var. sulphurea Ching; Icon. Filic. Sin. 3: 133 (1935)
= Doryopteris duclouxii Christ; Bull. Acad. Int. Géogr. Bot., Sér. 3, 11: 231 (1902)
= Doryopteris mairei Brause; Hedwigia 64: 206, t. 4 (1914)
= Doryopteris muralis Christ; Bull. Acad. Int. Géogr. Bot., Sér. 3, 13: 111 (1904)
= Hemionitis duclouxii (Christ) Christenh.; Global Fl. 4: 13 (2018)
species :
Aleuritopteris duthiei
(Baker) Ching; Bull. Fan Mem. Inst. Biol., Bot., Ser. 2, 1: 271 (1949)
Distribution: China (SW-Sichuan); Tibet; Nepal; Bhutan; India (Uttarakhand, Sikkim)
36 CHI TIB 40 BHU IND NEP
Conservation Status: IUCN CR for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Cheilanthes duthiei Baker; Ann. Bot. (Oxford) 5: 210 (1891)
= Hemionitis duthiei (Baker) Christenh.; Global Fl. 4: 13 (2018)
= Leptolepidium duthiei (Baker) X.C.Zhang & G.M.Zhang; Acta Phytotax. Sin. 40: 286 (2002)
= Oeosporangium duthiei (Baker) Fraser-Jenk. & Ph.Wangdi; Pterid. Bhutan 132 (2023)
species :
Aleuritopteris ebenipes
X.C.Zhang; Acta Phytotax. Sin. 32(1): 94 (1994)
Distribution: China (Guizhou)
36 CHI
Synonyms: hidden
= Hemionitis ebenipes (X.C.Zhang) Christenh.; Global Fl. 4: 13 (2018)
species :
Aleuritopteris elegans
(Poir.) Windham & Schuettp.; Taxon 74(2), p. 12 of 32 (2025)
Distribution: China (Fujian, Guangdong, Hainan, Zhejiang); India (Andhra Pradesh, Karnataka, Kerala, Odisha, Tamil Nadu); Sri Lanka; Myanmar; Thailand; Laos; Vietnam
36 CHI 40 IND SRL 41 LAO MYA THA VIE
Remarks: not in Taiwan fide Knapp (Ferns of Taiwan, 2011 ff.). E-Asian records for this species frequently refer to the closely related C. chusana
Synonyms: hidden
= Adiantum scandicinum Willd.; Syn. Fil. (Kiliae) [O.Swartz] 326 (1806) (in obs.)
= Cheilanthes cadieri Christ; J. Bot. [Morot] 19: 70 (1905)
= Cheilanthes melanocoma Bory; Voy. Indes Or. [Bélanger] 2(1): 71 (1833)
= Cheilanthes mysurensis Wall. ex Hook.; Sp. Fil. [W.J.Hooker] 2: 94, 100A (1852) (non Asplenium mysurense B. Heyne ex Roth (1821))
= Cheilanthes opposita Kaulf.; Enum. Filic. [Kaulfuss] 211 (1824)
= Cheilanthes swartzii Webb & Berthel.; Hist. Nat. Iles Canaries (Phytogr.) 3: 453 (1849)
= Cheilosoria mysuriensis (Wall. ex Hook.) Ching & K.H.Shing; Fl. Fujianica 1: 84 (1982)
= Hemionitis opposita (Kaulf.) Christenh.; Global Fl. 4: 18 (2018)
= Oeosporangium elegans (Poir.) Fraser-Jenk. & Pariyar; Annot. Checkl. Ind. Pterid. 1: 265 (2016)
= Pteris elegans Poir.; Encycl. [J.Lamarck et al.] 5: 718 (1804), non Sw. 1817, nec al., nec Cheilanthes elegans Desv. 1811.
species :
Aleuritopteris erythraea
(Pic.Serm.) Windham & Schuettp.; Taxon 74(2), p. 12 of 32 (2025)
Distribution: Ethiopia; Eritrea
24 ERI ETH
Synonyms: hidden
= Cheilanthes erythraea Pic.Serm.; Webbia 27(2): 420, 422, fig. 10, t. A-E (1972 publ. 1973)
= Hemionitis erythraea (Pic.Serm.) Christenh.; Global Fl. 4: 13 (2018)
species :
Aleuritopteris fallax
(M.Martens & Galeotti) Windham & Schuettp.; Taxon 74(2), p. 12 of 32 (2025)
Distribution: Mexico (Aguascalientes, Chiapas, Chihuahua, Coahuila, Colima, Ciudad de Mexico, Guerrero, Hidalgo, Jalisco, Mexico State, Michoacan, Morelos, Nuevo Leon, Oaxaca, Puebla, Queretaro, San Luis Potosi, Sinaloa, Sonora, Tamaulipas, Tlaxcala, Veracruz, Zacatecas); Guatemala
79 MEX 80 GUA
Synonyms: hidden
= Mildella fallax (M.Martens & Galeotti) G.L.Nesom; Phytologia 75(5): 384 (1993)
= Mildella intramarginalis var. serratifolia (Hook. & Baker) C.C.Hall & Lellinger; Amer. Fern J. 57: 124 (1967)
= Pellaea intramarginalis var. serratifolia Hook. & Baker; Syn. Fil. [W.J.Hooker & Baker] 149 (1867)
= Pteris fallax M.Martens & Galeotti; Nouv. Mém. Acad. Roy. Sci. Bruxelles 15 (Mém. Lycopod.): 53, t. 14, f. 2 (1842)
species :
Aleuritopteris farinosa
(Forssk.) Fée; Mém. Foug. 5 (Gen. Filic.) 154, t. 12 B, f. 1 (1852)
Name: Floury Cloak fern
Distribution: Senegal; Guinea; Sierra Leone; Nigeria; Cameroon; Bioko Isl. [Fernando Poo]; D.R.Congo [Zaire]; Rwanda; Burundi; Sudan; Ethiopia; Eritrea; Djibouti; Somalia; Soqotra; Uganda; Kenya; Tanzania; Angola; Zambia; Malawi; Zimbabwe; Botswana; Madagascar; Comores (Grande Comore, Anjouan, Mayotte); Mauritius; La Réunion; Saudi Arabia; Oman; Yemen (N-Yemen)
22 GUI NGA SEN SIE 23 BIO BUR CMN RWA ZAI 24 DJI ERI ETH SOC SOM SUD 25 KEN TAN UGA 26 ANG MLW ZAM ZIM 27 BOT 29 COM MAU MDG REU 35 OMA SAU YEM
Remarks: A. farinosa is a bona sp. fide Fraser-Jenkins (1997). The neotropical A. mexicana Fée is sometimes included here.
Synonyms: hidden
= Aleuritopteris aethiopica Saiki; J. Phytogeogr. Taxon. 32(2): 81 (1984)
= Aleuritopteris afra Pic.Serm.; Webbia 27: 415 (1972 publ. 1973)
= Aleuritopteris argyrophylla (Sw.) Fée; Mém. Foug. 5 (Gen. Filic.) 154 (1852)
= Aleuritopteris centrafricana Saiki; J. Phytogeogr. Taxon. 32: 82 (1984)
= Aleuritopteris decursiva (Forssk.) Saiki; J. Phytogeogr. Taxon. 32(2): 85 (1984)
= Aleuritopteris flava Saiki; J. Phytogeogr. Taxon. 32(2): 86 (1984), non A. argentea var. flava Ching 1981
= Aleuritopteris gigantea Saiki; J. Phytogeogr. Taxon. 32(2): 86 (1984)
= Aleuritopteris leptophylla Saiki; J. Phytogeogr. Taxon. 32(2): 88 (1984)
= Aleuritopteris longifrons Saiki; J. Phytogeogr. Taxon. 32(2): 88 (1984)
= Aleuritopteris vandervekenii Pic.Serm.; Bull. Jard. Bot. Natl. Belg. 53(1-2): 210 (1983)
= Allosorus argyrophyllus (Sw.) C.Presl; Tent. Pterid. 153 (1836)
= Allosorus farinosus (Forssk.) C.Presl; Tent. Pterid. 153 (1836)
= Cassebeera farinosa (Forssk.) J.Sm.; J. Bot. [Hooker] 4: 159 (1841)
= Cheilanthes afra (Pic.Serm.) Kornas; Distr. Ecol. Pterid. Zambia 48 (1979)
= Cheilanthes argyrophylla (Sw.) Cordem.; Bull. Soc. Sci. Île Réunion 1890-91: 155 (1891)
= Cheilanthes farinosa Forssk.; Fl. Aegypt.-Arab. 187 (1775)
= Cheilanthes farinosa var. deltoidea Bonap.; Notes Ptéridol. 4: 77 (1917)
= Hemionitis aethiopica (Saiki) Christenh.; Global Fl. 4: 9 (2018)
= Hemionitis argyrophylla (Sw.) Christenh.; Global Fl. 4: 9 (2018)
= Hemionitis decursiva (Forssk.) Christenh.; Global Fl. 4: 12 (2018)
= Hemionitis farinosa (Forssk.) Christenh.; Global Fl. 4: 13 (2018)
= Hemionitis grandis Christenh.; Global Fl. 4: 14 (2018), nom. nov. for Aleuritopteris gigantea Saiki
= Hemionitis vandervekenii (Pic.Serm.) Christenh.; Global Fl. 4: 22 (2018)
= Pteris argyrophylla Sw.; Syn. Fil. (Kiliae) [O.Swartz] 105 (1806)
= Pteris decursiva Forssk.; Fl. Aegypt.-Arab. 186 (1775)
= Pteris farinosa Forssk.; Fl. Aegypt.-Arab. 187 (1775)
species :
Aleuritopteris formosana
(Hayata) Tagawa; Acta Phytotax. Geobot. 14: 191 (1952)
Distribution: China (Fujian, Guangdong, Guangxi, Guizhou, Sichuan, Yunnan); Tibet; Taiwan; Pakistan; Nepal; Bhutan; India (Jammu & Kashmir, Andhra Pradesh, Arunachal Pradesh, Assam State, Chhattisgarh, Himachal Pradesh, Karnataka, Jharkhand, Madhya Pradesh, Maharashtra, Manipur, Meghalaya, Mizoram, Nagaland, Odisha, Rajasthan, Sikkim, ?Tripura, Uttarakhand, Uttar Pradesh, West Bengal); Myanmar; N-Thailand; Vietnam; Philippines
36 CHI TIB 38 TAI 40 BHU IND NEP PAK 41 MYA THA VIE 42 PHI
Synonyms: hidden
= Aleuritopteris platychlamys Ching var. alpina Ching; Acta Phytotax. Sin. 19(1): 71 (1981) (fide Fraser-Jenkins 1997)
= Cheilanthes anceps var. brevifrondis Khullar & Mehra; (1972) (nom. nud.)
= Cheilanthes anceps var. brevifrons Khullar; Amer. Fern J. 66(1): 24 (1976) (fide Fraser-Jenkins 1997)
= Cheilanthes brevifrons (Khullar) Khullar; Indian Fern J. 1(1-2): 90 (1984 publ. 1985) (fide Fraser-Jenkins 1997)
= Cheilanthes farinosa H.Itô; Fil. Jap. Illustr., t. 64 (1944), non (Forssk.) Kaulf. 1824
= Cheilanthes formosana Hayata; Enum. Pl. Formosa 61 (1906)
= Hemionitis formosana (Hayata) Christenh.; Global Fl. 4: 14 (2018)
species :
Aleuritopteris fragilis
(Hook.) Windham & Schuettp.; Taxon 74(2), p. 12 of 32 (2025)
Distribution: China (S-Yunnan); Myanmar; N-Thailand; Laos
36 CHI 41 LAO MYA THA
Synonyms: hidden
= Cheilanthes fragilis Hook.; Fil. Exot., t. 96 (1859)
= Cheilosoria fragilis (Hook.) Ching & K.H.Shing; Glossary Lycopods Ferns 39 (1982)
= Hemionitis fragilis (Hook.) Christenh.; Global Fl. 4: 14 (2018)
= Oeosporangium fragile (Hook.) Fraser-Jenk.; Annot. Checkl. Ind. Pterid. 1: 265 (2016)
species :
Aleuritopteris gongshanensis
G.M.Zhang; Novon 14: 513 (2004)
Distribution: China (C-Yunnan, NW-Yunnan)
36 CHI
species :
Aleuritopteris grevilleoides
(Christ) G.M.Zhang & X.C.Zhang; Lycophytes Ferns China 231 (2012)
Distribution: China (N-Sichuan, W-Yunnan)
36 CHI
Conservation Status: IUCN R
Synonyms: hidden
= Cheilanthes grevilleoides Christ; Notul. Syst. (Paris) 1: 51, cum. fig. (1909)
= Hemionitis grevilleoides (Christ) Christenh.; Global Fl. 4: 14 (2018)
= Sinopteris grevilleoides (Christ) C.Chr. & Ching; Bull. Fan Mem. Inst. Biol., Bot. 4: 360 (1933)
species :
Aleuritopteris grisea
(Blanf.) Panigrahi; Bull. Bot. Surv. India 2: 321 (1961)
Distribution: China (Guangxi, Guizhou, Hebei, Sichuan, Yunnan); Tibet; Pakistan; Nepal; Bhutan; India (Jammu & Kashmir, Himachal Pradesh, Sikkim, Uttarakhand, ?West Bengal); Myanmar
36 CHI TIB 40 BHU IND NEP PAK 41 MYA
Synonyms: hidden
= Aleuritopteris farinosa var. grisea (Blanf.) Ching; Hong Kong Naturalist 10(3-4): 202 (1941)
= Aleuritopteris grisea var. alpina (Ching ex S.K.Wu) S.K.Wu; Fl. Reipubl. Popularis Sin. 3(1): 163 (1990) (fide Fraser-Jenkins 1997)
= Aleuritopteris platychlamys Ching; Fl. Tsinling. 2: 207 (1974)
= Aleuritopteris platychlamys var. alpina Ching ex S.K.Wu; Acta Phytotax. Sin. 19(1): 71 (1981)
= Cheilanthes farinosa f. minor C.B.Clarke & Baker; J. Linn. Soc., Bot. 25: 441 (1889)
= Cheilanthes farinosa var. grisea (Blanf.) Blanf.; J. Asiat. Soc. Bengal, Pt. 2, Nat. Hist. 57, 2(4): 250 (1881)
= Cheilanthes grisea Blanf.; J. Simla Naturalists´ Soc. 1(2): 21 (1886)
= Cheilanthes platychlamys (Ching) Fraser-Jenk.; New Sp. Syndr. Indian Pteridol. 75 (1997)
= Hemionitis grisea (Blanf.) Christenh.; Global Fl. 4: 14 (2018)
= Hemionitis platychlamys (Ching) Christenh.; Global Fl. 4: 19 (2018)
species :
Aleuritopteris guanchicus
(Bolle) Windham & Schuettp.; Taxon 74(2), p. 12 of 32 (2025)
Name: Guanchen-Schuppenfarn (DE); Helecho de roca, helecho chico (ES)
Distribution: Portugal; Spain; France; Corsica; Sardinia; Italy; Greece (only E-Peloponnisos); Morocco; Algeria; Tunisia; Madeira (Madeira Isl.); Canary Isl. (Fuerteventura, Gran Canaria, Tenerife, La Gomera, Hierro, La Palma Isl., Lanzarote); East Aegaean Isl. (Ikaria)
12 COR FRA POR SAR SPA 13 GRC ITA 20 ALG MOR TUN 21 CNY MDR 34 EAI
Synonyms: hidden
= Allosorus guanchicus (Bolle) Christenh.; Willdenowia 42(2): 284 (2012)
= Cheilanthes guanchica Bolle; Bonplandia (Hannover) 7: 107 (1859)
= Cheilanthes pteridioides subsp. guanchica (Bolle) O.Bolòs, Vigo, Masalles & Ninot; Fl. Man. Paisos Catalans 1213 (1990)
= Cheilanthes ×sventenii Benl; Nova Hedwigia 12: 147, t. 28 (1966)
= Hemionitis guanchica (Bolle) Christenh.; Global Fl. 4: 15 (2018)
= Oeosporangium guanchicum (Bolle) Fraser-Jenk. & Pariyar; Annot. Checkl. Ind. Pterid. 1: 265 (2016)
species :
Aleuritopteris hancockii
(Baker) Windham & Schuettp.; Taxon 74(2), p. 12 of 32 (2025)
Distribution: China (Gansu, Yunnan, W-Guizhou, Sichuan); Tibet; Bhutan
36 CHI TIB 40 BHU
Remarks: not in India
Synonyms: hidden
= Cheilanthes bonatiana Brause; Hedwigia 64: 203, t. 4f (1914)
= Cheilanthes hancockii Baker; Bull. Misc. Inform. Kew 1895: 54 (1895)
= Cheilanthes taliensis Christ; Mém. Soc. Bot. France 1 [Hist. Acrostich.]: 58 (1905)
= Cheilanthes wilsonii Christ; Bull. Acad. Int. Géogr. Bot., Sér. 3, 16: 132 (1906)
= Cheilosoria hancockii (Baker) Ching & K.H.Shing; Glossary Lycopods Ferns 39 (1982)
= Hemionitis hancockii (Baker) Christenh.; Global Fl. 4: 15 (2018)
= Oeosporangium hancockii (Baker) Fraser-Jenk.; Annot. Checkl. Ind. Pterid. 1: 257 (2016)
species :
Aleuritopteris hispanicus
(Mett.) Windham & Schuettp.; Taxon 74(2), p. 12 of 32 (2025)
Distribution: Portugal; Spain; France; Morocco; Algeria; ?Tunisia
12 FRA POR SPA 20 ALG MOR
Remarks: pteridioides agg.
Synonyms: hidden
= Allosorus hispanicus (Mett.) Christenh.; Willdenowia 42(2): 284 (2012)
= Cheilanthes hispanica Mett.; Abh. Senckenberg. Naturf. Ges. 3(1) [Aspl.]: 74 (1859)
= Hemionitis hispanica (Mett.) Christenh.; Global Fl. 4: 15 (2018)
= Oeosporangium hispanicum (Mett.) Fraser-Jenk. & Pariyar; Annot. Checkl. Ind. Pterid. 1: 265 (2016)
species :
Aleuritopteris insignis
(Ching) Windham & Schuettp.; Taxon 74(2), p. 13 of 32 (2025)
Distribution: China (N-Sichuan); E-Tibet
36 CHI TIB
Synonyms: hidden
= Oeosporangium insigne (Ching) Fraser-Jenk.; Annot. Checkl. Ind. Pterid. 1: 265 (2016)
= Cheilanthes insignis Ching; Fl. Tsinling. 2: 208 (1974)
= Cheilanthes tenuifolia C.Chr.; (1924) (non Sw. 1806)
= Cheilosoria insignis (Ching) Ching & K.H.Shing; Fl. Xizang. 1: 91 (1983)
= Hemionitis insignis (Ching) Christenh.; Global Fl. 4: 16 (2018)
species :
Aleuritopteris intramarginalis
(Kaulf. ex Link) Windham & Schuettp.; Taxon 74(2), p. 13 of 32 (2025)
Distribution: Mexico (Chiapas, Coahuila, Colima, Guanajuato, Guerrero, Hidalgo, Jalisco, Mexico State, Michoacan, Morelos, Nuevo Leon, Oaxaca, Puebla, Queretaro, San Luis Potosi, Sinaloa, Tamaulipas, Tlaxcala, Veracruz); Guatemala; El Salvador; Honduras; Nicaragua; Costa Rica; Panama; Hispaniola (Haiti); Ecuador; Galapagos
79 MEX 80 COS ELS GUA HON NIC PAN 81 HIS 83 ECU GAL
Synonyms: hidden
= Allosorus intramarginalis (Kaulf. ex Link) C.Presl; Tent. Pterid. 153 (1836)
= Cassebeera intramarginalis (Kaulf. ex Link) J.Sm.; J. Bot. [Hooker] 4: 159 (1841)
= Cheilanthes intramarginalis (Kaulf. ex Link) Hook.; Sp. Fil. [W.J.Hooker] 2: 112 (1852)
= Cheilanthes intramarginalis var. grosseserrata Hook.; Sec. Cent. Ferns, t. 72 (1861)
= Hemionitis intramarginalis (Kaulf. ex Link) Christenh.; Global Fl. 4: 16 (2018)
= Mildella intramarginalis (Kaulf. ex Link) Trevis.; Rendiconti Reale Ist. Lombardo Sci., Ser. 2, 9(20): 810 (1876)
= Pellaea intramarginalis (Kaulf. ex Link) J.Sm.; Cat. Ferns Kew 4 (1856)
= Platyloma intramarginale (Kaulf. ex Link) E.J.Lowe; Ferns 3: t. 31 (1857)
= Pteris intramarginalis Kaulf. ex Kunze; Analecta Pteridogr. 21, t. 17, f. 1 (1837)
= Pteris intramarginalis Kaulf. ex Link; Hort. Berol. [Link] 2: 34 (1833)
= Pteris intramarginalis Kaulf. ex Schltdl. & Cham.; Linnaea 5(4): 613 (1830) (nomen)
species :
Aleuritopteris khammouanensis
(Vongthavone & Phonep.) comb.ined.
Distribution: Laos
41 LAO
Synonyms: hidden
= Hemionitis khammouanensis Vongthavone & Phonep.; Kew Bull. 80(1): 5 of 12 (2025)
species :
Aleuritopteris krameri
(Franch. & Sav.) Ching; Hong Kong Naturalist 10: 202 (1941)
Distribution: Japan; Taiwan; Thailand (E-Thailand, N-Thailand); Sulawesi; Philippines; New Guinea (Irian Jaya, Papua New Guinea)
38 JAP TAI 41 THA 42 NWG PHI SUL
Conservation Status: EN in Japan
Synonyms: hidden
= Cheilanthes krameri Franch. & Sav.; Enum. Pl. Jap. 2: 212, 619 (1877)
= Hemionitis krameri (Franch. & Sav.) Christenh.; Global Fl. 4: 16 (2018)
species :
Aleuritopteris kuhnii
(Milde) Ching; Hong Kong Naturalist 10: 202 (1941)
Distribution: Russian Far East (Primorye); China (Gansu, Hebei, Henan, Jilin, Liaoning, Nei Mongol, Shaanxi, Shanxi, Sichuan, Yunnan); Tibet; North Korea; South Korea (incl. Cheju Isl.); Japan
31 RFE 36 CHI TIB 38 JAP KOR NKO
Remarks: not in India
Synonyms: hidden
= Aleuritopteris caesia (Christ) Ching; Hong Kong Naturalist 10: 202 (1941)
= Aleuritopteris caesia var. efarinosa H.S.Kung, Li Bing Zhang & X.S.Guo; Acta Bot. Yunnan. 17(4): 422 (1995)
= Aleuritopteris kuhnii f. efarinosa (Makino) Tagawa; Acta Phytotax. Geobot. 14(6): 191 (1952)
= Aleuritopteris kuhnii var. brandtii (Franch. & Sav.) Tagawa; Acta Phytotax. Geobot. 14(6): 191 (1952)
= Aleuritopteris tenella (Ching & S.K.Wu) Saiki; J. Phytogeogr. Taxon. 32(1): 11 (1984)
= Cheilanthes brandtii Franch. & Sav.; Enum. Pl. Jap. 2: 212, 620 (1877)
= Cheilanthes brandtii var. efarinosa Makino; J. Jap. Bot. 1(7): 26 (1917)
= Cheilanthes caesia Christ; Bull. Acad. Int. Géogr. Bot., Sér. 3, 16: 133 (1906)
= Cheilanthes farinosa var. brandtii (Franch. & Sav.) C.Chr.; Index Filic. 3: 172 (1905)
= Cheilanthes kuhnii f. efarinosa (Makino) Nakaike; J. Nippon Fernist Club 3(Suppl. 2): 40 (2004)
= Cheilanthes kuhnii Milde; Bot. Zeitung (Berlin) 25: 149 (1867)
= Cheilanthes kuhnii var. brandtii (Franch. & Sav.) Tagawa; J. Jap. Bot. 13(3): 184 (1934)
= Cheilanthes kuhnii var. caesia (Christ) C.Chr.; Acta Horti Gothob. 1: 90 (1924)
= Cheilanthes lanceolata C.Chr.; Bot. Gaz. 66: 334 (1913)
= Hemionitis brandtii (Franch. & Sav.) Christenh.; Global Fl. 4: 10 (2018)
= Hemionitis caesia (Christ) Christenh.; Global Fl. 4: 11 (2018)
= Hemionitis kuhnii (Milde) Christenh.; Global Fl. 4: 16 (2018)
= Leptolepidium caesium (Christ) K.H.Shing & S.K.Wu; Acta Bot. Yunnan. 1(1): 117 (1979)
= Leptolepidium kuhnii (Milde) K.H.Shing & S.K.Wu; Acta Bot. Yunnan. 1(1): 117 (1979)
= Leptolepidium kuhnii var. brandtii (Franch. & Savigny) K.H.Shing & S.K.Wu; Acta Bot. Yunnan. 1(1): 118 (1979)
= Leptolepidium tenellum Ching & S.K.Wu; Acta Bot. Yunnan. 1(1): 117 (1979)
= Oeosporangium kuhnii (Milde) Fraser-Jenk.; Annot. Checkl. Ind. Pterid. 1: 261 (2016)
species :
Aleuritopteris leonardii
(Maxon) Windham & Schuettp.; Taxon 74(2), p. 13 of 32 (2025)
Distribution: Hispaniola (Haiti)
81 HIS
Synonyms: hidden
= Cheilanthes leonardii Maxon; J. Wash. Acad. Sci. 14: 87 (1924)
= Hemionitis leonardii (Maxon) Christenh.; Global Fl. 4: 16 (2018)
= Mildella leonardii (Maxon) C.C.Hall & Lellinger; Amer. Fern J. 57: 126 (1967)
species :
Aleuritopteris leptolepis
(Fraser-Jenk.) Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 127 (2008)
Distribution: ?Yemen (?N-Yemen); China (SW-Sichuan, NW-Yunnan); Tibet; Pakistan (Gilgit, Baltistan, Murree, Hazara); Nepal; Bhutan; India (Jammu & Kashmir, Himachal Pradesh, Sikkim, Uttarakhand, West Bengal); Myanmar
36 CHI TIB 40 BHU IND NEP PAK 41 MYA
Synonyms: hidden
= Cheilanthes leptolepidium Fraser-Jenk.; (1992), nom. nud.
= Cheilanthes leptolepis Fraser-Jenk.; Bot. Helv. 102(2): 144 (1992)
= Hemionitis leptolepis (Fraser-Jenk.) Christenh.; Global Fl. 4: 16 (2018)
= Leptolepidium leptolepis (Fraser-Jenk.) Kholia & Punetha; J. Indian Bot. Soc. 184 (1995)
species :
Aleuritopteris mairei
(Brause) Windham & Schuettp.; Taxon 74(2), p. 13 of 32 (2025)
Distribution: China (Guizhou, Hunan, Shaanxi, SW-Sichuan, Yunnan)
36 CHI
Synonyms: hidden
= Cheilanthes brausei Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 122 (2008), nom. nov.
= Hemionitis mairei (Brause) Christenh.; Global Fl. 4: 17 (2018)
= Mildella mairei (Brause) C.C.Hall & Lellinger; Amer. Fern J. 57: 133 (1967)
= Oeosporangium mairei (Brause) Fraser-Jenk.; Annot. Checkl. Ind. Pterid. 1: 265 (2016)
= Pellaea mairei Brause; Hedwigia 64: 201, t. 4 C (1914), non Cheilanthes mairei Brause
species :
Aleuritopteris mexicana
Fée; Mém. Foug. 5 (Gen. Filic.) 154 (1852)
Distribution: Mexico (Aguascalientes, Chiapas, Chihuahua, Colima, Ciudad de Mexico, Durango, Guerrero, Hidalgo, Jalisco, Mexico State, Michoacan, Morelos, Nayarit, Oaxaca, Puebla, Queretaro, San Luis Potosi, Sinaloa, Tamaulipas, Veracruz, Zacatecas); Guatemala; Honduras; Costa Rica; Colombia (Cundinamarca); Venezuela (Tachira); Ecuador; Peru; Bolivia (Potosi)
79 MEX 80 COS GUA HON 82 CLM VEN 83 BOL ECU PER
Remarks: frequently included in A. farinosa s. lat.
Synonyms: hidden
= Aleuritopteris chihuahuaensis Saiki; J. Phytogeogr. Taxon. 32(2): 85 (1984)
= Aleuritopteris peruviana Saiki; J. Phytogeogr. Taxon. 32(2): 89 (1984)
= Cheilanthes chihuahuaensis (Saiki) Fraser-Jenk.; New Sp. Syndr. Indian Pteridol. 81 (1997)
= Cheilanthes mexicana (Fée) Punetha & Kholia; New Botanist, Int. Quart. J. Pl. Sci. Res. 16(1-4): 119 (1989) (non Davenp., nec sensu Punetha & Kholia)
= Cheilanthes pulveracea C.Presl; Reliq. Haenk. 1: 64 (1825), nom. illeg.
= Hemionitis chihuahuaensis (Saiki) Christenh.; Global Fl. 4: 11 (2018)
= Pellaea pulcherrima Rovirosa; Pteridogr. Sur Mexico t. 14 (1909), nom. nud.
species :
Aleuritopteris niphobola
(C.Chr.) Ching; Hong Kong Naturalist 10: 197 (1941)
Distribution: China (Gansu, Hebei, Nei Mongol, Ningxia, Shaanxi, Shanxi, Sichuan); Tibet
36 CHI TIB
Synonyms: hidden
= Aleuritopteris niphobola var. concolor Ching; Fl. Tsinling. 2: 207 (1974)
= Aleuritopteris niphobola var. pekingensis Ching & Y.P.Hsu; Acta Phytotax. Sin. 19: 64 (1981)
= Aleuritopteris pekingensis Ching; (1962)
= Cheilanthes niphobola C.Chr.; Acta Horti Gothob. 1: 88, t. 19 (1924)
= Hemionitis niphobola (C.Chr.) Christenh.; Global Fl. 4: 18 (2018)
species :
Aleuritopteris nitidula
(Wall. ex Hook.) Windham & Schuettp.; Taxon 74(2), p. 13 of 32 (2025)
Distribution: China (Fujian, Gansu, Guangdong, Guangxi, Guizhou, Hebei, Hunan, Jiangxi, Sichuan, Yunnan, Zhejiang); Tibet; Taiwan; Pakistan (Chitral, Swat, Hazara); Nepal; Bhutan; India (Jammu & Kashmir, Arunachal Pradesh, Himachal Pradesh, Sikkim, Uttarakhand); Vietnam
36 CHI TIB 38 TAI 40 BHU IND NEP PAK 41 VIE
Synonyms: hidden
= Aleuritopteris henryi (Christ) Fay W.Li; Taiwania 64(4): 378 (2019)
= Allosorus nitidulus C.Presl; Tent. Pterid. 152 (1836), nomen
= Cheilanthes henryi Christ; Bull. Acad. Int. Géogr. Bot., Sér. 3, 16: 133 (1906)
= Cheilanthes nitidula Hook.; Sp. Fil. [W.J.Hooker] 2: 112 (1852)
= Cheilanthes nitidula subsp. henryi (Christ) Fraser-Jenk.; Fern Gaz. 18(5): 219 (2009)
= Hemionitis nitidula (Hook.) Christenh.; Global Fl. 4: 18 (2018)
= Mildella henryi (Christ) C.C.Hall & Lellinger; Amer. Fern J. 57: 129 (1967)
= Mildella nitidula (Hook.) C.C.Hall & Lellinger; Amer. Fern J. 57: 127 (1967)
= Oeosporangium nitidulum (Hook.) Fraser-Jenk.; Annot. Checkl. Ind. Pterid. 1: 257 (2016)
= Pellaea henryi Christ; Bull. Herb. Boissier 7: 7 (1899)
= Pellaea nitidula (Hook.) Wall. ex Baker; Syn. Fil. [W.J.Hooker & Baker] 149 (1867)
= Pteris nitidula Wall.; Numer. List [Wallich] no. 89 (1829), nomen
species :
Aleuritopteris papuana
(C.Chr.) H.Schneid.; Guide Alpine Subalpine Fl. Mount Jaya [R.J.Johns et al.] 45 (2006)
Distribution: Moluccas (Seram); New Guinea (Papua New Guinea, Irian Jaya)
42 MOL NWG
Synonyms: hidden
= Cheilanthes papuana C.Chr.; Dansk Bot. Ark. 9(3): 57 (1937)
= Hemionitis papuana (C.Chr.) Christenh.; Global Fl. 4: 19 (2018)
species :
Aleuritopteris parishii
Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 129 (2008)
Distribution: Myanmar (SE-Myanmar, C-Myanmar)
41 MYA
species :
Aleuritopteris patula
(Baker) Windham & Schuettp.; Taxon 74(2), p. 14 of 32 (2025)
Distribution: China (Chongqing, Guangxi, Guizhou, W-Hubei)
36 CHI
Synonyms: hidden
= Oeosporangium patulum (Baker) Fraser-Jenk.; Annot. Checkl. Ind. Pterid. 1: 265 (2016)
= Cheilanthes patula Baker; J. Bot. 26: 225 (1888)
= Cheilosoria patula (Baker) P.S.Wang & X.Y.Wang; Pterid. Fl. Guizhou 188 (2001)
= Hemionitis patula (Baker) Christenh.; Global Fl. 4: 19 (2018)
= Pellaea patula (Baker) Ching; Acta Phytotax. Sin. 10: 302 (1965)
species :
Aleuritopteris paupercula
(Christ) Windham & Schuettp.; Taxon 74(2), p. 14 of 32 (2025)
Distribution: China (W-Sichuan)
36 CHI
Synonyms: hidden
= Cheilanthes christii Fraser-Jenk. & Yatsk.; Indian Fern J. 27(1-2): 213 (2011), nom. nov.
= Cheilanthes paupercula (Christ) Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 122 (2008)
= Hemionitis christii (Fraser-Jenk. & Yatsk.) Christenh.; Global Fl. 4: 11 (2018)
= Mildella paupercula (Christ) C.C.Hall & Lellinger; Amer. Fern J. 57: 131 (1967)
= Oeosporangium pauperculum (Christ) Fraser-Jenk. & Pariyar; Annot. Checkl. Ind. Pterid. 1: 265 (2016)
= Pellaea paupercula (Christ) Ching; Bull. Fan Mem. Inst. Biol., Bot. 2: 203 (1931)
= Pteris paupercula Christ; Bull. Acad. Int. Géogr. Bot., Sér. 3, 16: 131 (1906)
species :
Aleuritopteris pentagona
Saiki; J. Phytogeogr. Taxon. 32(1): 8 (1984)
Distribution: Myanmar; Thailand
41 MYA THA
Remarks: subdimorpha agg.
species :
Aleuritopteris persica
(Bory) Windham & Schuettp.; Taxon 74(2), p. 14 of 32 (2025)
Distribution: Italy; Croatia; Bosnia & Hercegovina; Montenegro; North Macedonia; Albania; Bulgaria; Greece (scattered mainland); Crete (rare); Crimea; ?Algeria; Kazakhstan; Turkmenistan; Uzbekistan; Kyrgyzstan; Tajikistan; Northern Caucasus; Georgia [Caucasus]; Armenia; Azerbaijan; East Aegaean Isl. (Lesvos, Chios, Ikaria, Samos, Simi); Turkey (Anatolia); NE-Iraq; Iran; Afghanistan; Pakistan (Chitral, Baluchistan, Murree, Gilgit, Baltistan); India (Himachal Pradesh, Jammu & Kashmir)
13 ALB BIH BUL CRO GRC ITA KRI MNT NMA 14 KRY 32 KAZ KGZ TKM TZK UZB 33 ARM AZE GEO NCS 34 AFG EAI IRN IRQ TUR 40 IND PAK
Remarks: pteridioides agg.
Synonyms: hidden
= Acrostichum microphyllum Bertol.; Propag. Agric. 6: 343 (1856)
= Allosorus persicus (Bory) Christenh.; Willdenowia 42(2): 284 (2012)
= Cheilanthes fimbriata Vis.; Fl. Dalmat. 1: 42, t. 1, f. 1 (1842)
= Cheilanthes persica (Bory) Mett.; Fil. Afr. [Kuhn]: 73 (1868)
= Cheilanthes szovitsii Fisch. & C.A.Mey.; Bull. Soc. Imp. Naturalistes Moscou 6: 260 (1833), nom. nud.
= Hemionitis persica (Bory) Christenh.; Global Fl. 4: 19 (2018)
= Myriopteris szovitzii (Fisch. & C.A.Mey.) J.Sm.; Hist. Fil. 281 (1875)
= Notholaena microphylla Bolle; Sitzungsber. Ges. Naturf. Freunde Berlin (1858)
= Notholaena persica (Bory) Bory; Voy. Indes Or. [Bélanger] 2(1): 23 (1833)
= Notholaena szovitsii (Fisch. & C.A.Mey.) Keyserl.; Polyp. Herb. Bunge 28 (1873)
= Oeosporangium fimbriatum (Vis.) Vis.; (1867)
= Oeosporangium persicum (Bory) Vis.; Mem. Reale Ist. Veneto Sci.16: 44 (1872)
= Oeosporangium szovitsii (Fisch. & C.A.Mey.) Vis.; Atti Reale Ist. Veneto Sci. Lett. Arti, Ser. 3, 12: 656 (1867)
species :
Aleuritopteris pseudoargentea
S.K.Wu; Mem. Coll. Sci. Kyoto Imp. Univ., Ser. B, Biol. 8(2): 155 (1983)
Distribution: Thailand
41 THA
Remarks: subdimorpha agg.
Synonyms: hidden
= Cheilanthes pseudoargentea (S.K.Wu) K.Iwats.; Fl. Thailand 3(4): 619 (1989)
species :
Aleuritopteris pteridioides
(Reichard) Windham & Schuettp.; Taxon 74(2), p. 14 of 32 (2025)
Name: Madeira-Schuppenfarn (DE); Culantrillo de risco (ES)
Distribution: Portugal; Spain; France; +Monaco; Corsica; Sardinia; Italy; Sicily; Pantelleria; Croatia; Bosnia & Hercegovina; Montenegro; North Macedonia; Albania; Bulgaria; Greece (Kiklades, E-Peloponnisos); Crete; Morocco; S-Algeria (Hoggar); Azores (Pico); Madeira (Madeira Isl.); Canary Isl. (Lanzarote, Fuerteventura, Gran Canaria, Tenerife, La Gomera, Hierro, La Palma Isl.); N-Chad (Tibesti); Ethiopia; Djibouti; Turkmenistan; Northern Caucasus; Armenia; Azerbaijan; East Aegaean Isl. (Karpathos, Dodecanese, Lesvos); Turkey (Anatolia); Iraq; Iran; Cyprus; Lebanon; Syria; Israel; Palestine; Sinai peninsula; Jordania; Saudi Arabia; United Arab Emirates; Oman; Yemen (N-Yemen)
12 COR FRA MCO POR SAR SPA 13 ALB BIH BUL CRO GRC ITA KRI MNT NMA PTL SIC 20 ALG MOR 21 AZO CNY MDR 24 CHA DJI ETH 32 TKM 33 ARM AZE NCS 34 CYP EAI IRN IRQ ISR JOR LEB PAL SIN SYR TUR 35 OMA SAU UAE YEM
Remarks: for a long time known as Cheilanthes maderensis, while Cheilanthes pteridioides in the previous sense is todays O. acrosticum
Synonyms: hidden
= Adiantum fragrans (L.) L.f.; Suppl. Pl. 447 (1782), nom. illeg.
= Adiantum odoratum Poir.; Encycl. [J.Lamarck et al.] Suppl. 1: 142 (1810)
= Adiantum pusillum All. ex Pollini; Fl. Veron. 3: 293 (1824)
= Adiantum pusillum Willd. ex Bernh.; Neues J. Bot. 1(2): 36 (1806)
= Adiantum suaveolens Poir.; Encycl. [J.Lamarck et al.] Suppl. 1: 142 (1810)
= Allosorus fragrans (L.f.) Farw.; Amer. Midl. Naturalist 12: 284 (1931)
= Allosorus pteridioides (Reichard) Christenh.; Willdenowia 42(2): 284 (2012)
= Cheilanthes fragrans (L.) Sw.; Syn. Fil. (Kiliae) [O.Swartz] 127 (1806) (nom. illeg.)
= Cheilanthes fragrans (L.) Webb ex Benth.; Hist. Nat. Iles Canaries (Phytogr.) 3: 452 (1847)
= Cheilanthes fragrans subsp. maderensis (Lowe) Benl; Mitt. Bot. Staatssamml. München 6: 33 (1966)
= Cheilanthes maderensis Lowe; J. Bot. [Hooker] 1: 26 (1834) (nomen), ex Trans. Cambridge Philos. Soc. 6: 528 (1838)
= Cheilanthes microphylla Buch; Abh. Königl. Akad. Wiss. Berlin 1816-1817: 360 (1819), nom. illeg.
= Cheilanthes odora proles maderensis (Lowe) Rouy; Fl. France [Rouy & Foucaud] 14: 389 (1913)
= Cheilanthes pteridiodes var. maderensis (Lowe) Trab.; Mém. Soc. Sci. Nat. Maroc. 8(1): 164 (1924)
= Cheilanthes pteridioides (Reichard) C.Chr.; Index Filic. 178 (1905)
= Cheilanthes pteridioides f. maderensis (Lowe) Maire & Weiller; Encycl. Biol. 33: 71 (1952)
= Cheilanthes pteridioides subsp. maderensis (Lowe) O.Bolòs, Vigo, Masalles & Ninot; Fl. Man. Paisos Catalans 1213 (1990)
= Cheilanthes pteridioides var. maderensis (Lowe) Trab.; Mém. Soc. Sci. Nat. Maroc 8(1): 164 (1924)
= Cheilanthes squamosa Heer & Regel; Linnaea 24: 174 (1851)
= Cheilanthes suaveolens Sw.; Syn. Fil. (Kiliae) [O.Swartz] 127 (1806), nom. superfl.
= Hemionitis maderensis (Lowe) Christenh.; Global Fl. 4: 17 (2018)
= Hemionitis pteridioides (Reichard) Christenh.; Global Fl. 4: 19 (2018)
= Hemionitis quezelii (Tardieu) Christenh.; Global Fl. 4: 20 (2018)
= Negripteris quezelii Tardieu; Mission Bot. Tibesti 92, t. 1(1-5) (1958)
= Oeosporangium pteridioides (Reichard) Fraser-Jenk. & Pariyar; Annot. Checkl. Ind. Pterid. 1: 258 (2016)
= Polypodium fragrans L.; Mant. Pl. 2: 307 (1771) (non L. 1753)
= Polypodium pteridioides Reichard; Syst. Pl., ed. 4 [Willdenow] 4: 424 (1780), nom. nov. pro Polypodium fragrans L. (1771), non L. (1753), nec P. pterioides Lam. (1779)
= Polypodium pteryoides Bory; Ess. Isl. Fortunées 311 (1803)
= Pteris fragrans (L.) Lag.; Anales Ci. Nat. 5: 158 (1802)
species :
Aleuritopteris pulchella
(Bory ex Willd.) Windham & Schuettp.; Taxon 74(2), p. 14 of 32 (2025)
Name: Zierlicher Schuppenfarn (DE); Doradilla fina (ES)
Distribution: Madeira (Madeira Isl.); Canary Isl. (Gran Canaria, Tenerife, La Gomera, Hierro, La Palma Isl.)
21 CNY MDR
Synonyms: hidden
= Allosorus fragilis Christenh.; Willdenowia 42(2): 284 (2012), nom. nov.
= Allosorus pulchellus (Bory ex Willd.) C.Presl; Tent. Pterid. 152 (1836)
= Cheilanthes pulchella Bory ex Willd.; Sp. Pl., ed. 4 [Willdenow] 5(1): 456 (1810)
= Hemionitis pulchella (Bory ex Willd.) Christenh.; Global Fl. 4: 20 (2018)
= Oeosporangium pulchellum (Bory ex Willd.) Fraser-Jenk. & Pariyar; Annot. Checkl. Ind. Pterid. 1: 264 (2016)
species :
Aleuritopteris pygmaea
Ching; Acta Phytotax. Sin. 19(1): 61 (1981)
Distribution: Tibet
36 TIB
Synonyms: hidden
= Hemionitis pygmaea (Ching) Christenh.; Global Fl. 4: 20 (2018)
species :
Aleuritopteris rosulata
(C.Chr.) Ching; Hong Kong Naturalist 10: 197 (1941)
Distribution: China (W-Sichuan)
36 CHI
Synonyms: hidden
= Cheilanthes rosulata C.Chr.; Acta Horti Gothob. 1: 89, t. 20 (1924)
= Hemionitis rosulata (C.Chr.) Christenh.; Global Fl. 4: 20 (2018)
species :
Aleuritopteris rouxii
Fraser-Jenk. & E.Wollenw.; Ferns Fern-Allies Nepal 1: 244 (2015)
Distribution: Nigeria
22 NGA
Synonyms: hidden
= Hemionitis rouxii (Fraser-Jenk. & E.Wollenw.) Christenh.; Global Fl. 4: 20 (2018)
species :
Aleuritopteris rufa
(D.Don) Ching; Hong Kong Naturalist 10: 200 (1941)
Distribution: China (Guizhou, Yunnan, Guangxi); Nepal; India (Jammu & Kashmir, Arunachal Pradesh, Himachal Pradesh, Maharashtra, Madhya Pradesh, Manipur, Meghalaya, Sikkim, Uttarakhand, West Bengal); Myanmar; N-Thailand; N-Vietnam; ?peninsular Malaysia
36 CHI 40 IND NEP 41 MYA THA VIE
Synonyms: hidden
= Aleuritopteris intermedia Ching; Sporae Pterid. Sin. (1976), nom. nud.
= Cheilanthes rufa D.Don; Prodr. Fl. Nepal. 16 (1825)
= Hemionitis gmorgana Christenh.; Global Fl. 4: 14 (2018), nom. nov.
species :
Aleuritopteris scioana
(Chiov.) Fraser-Jenk. & Dulawat; Taxon. Revis. Indian Subcontinental Pteridophytes 133 (2008)
Distribution: Sudan; Ethiopia; Eritrea; Somalia; Soqotra; Kenya; Oman; Yemen (N-Yemen, S-Yemen); W-India (Rajasthan)
24 ERI ETH SOC SOM SUD 25 KEN 35 OMA YEM 40 IND
Remarks: A. tricholepifera is kept separate by Roux (2009)
Conservation Status: IUCN CR for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Cheilanthes scioana (Chiov.) J.P.Roux; Strelitzia 23: 185 (2009)
= Cheilanthes tricholepifera (Pic.Serm.) J.P.Roux; Strelitzia 23: 186 (2009) (fide Fraser-Jenkins 2008)
= Hemionitis scioana (Chiov.) Christenh.; Global Fl. 4: 21 (2018)
= Hemionitis tricholepifera (Pic.Serm.) Christenh.; Global Fl. 4: 22 (2018)
= Mohria scioana Chiov.; Atti Reale Accad. Italia, Mem. Cl. Sci. Fis. 11: 66 (1940)
= Negripteris scioana (Chiov.) Pic.Serm.; Nuovo Giorn. Bot. Ital., n. s. 53: 131, t. 14-16 (1946)
= Negripteris tricholepifera Pic.Serm.; Webbia 27: 411 (1972 publ. 1973)
species :
Aleuritopteris siamensis
S.K.Wu; Mem. Coll. Sci. Kyoto Imp. Univ., Ser. B, Biol. 8(2): 152 (1983)
Distribution: N-Thailand
41 THA
Conservation Status: DD in Thailand
Synonyms: hidden
= Cheilanthes siamensis (S.K.Wu) K.Iwats.; Fl. Thailand 3(4): 617 (1989)
= Hemionitis siamensis (S.K.Wu) Christenh.; Global Fl. 4: 21 (2018)
species :
Aleuritopteris sichouensis
Ching & S.K.Wu; Acta Phytotax. Sin. 19(1): 69 (1981)
Distribution: China (S-Yunnan, Guangxi, Guizhou); Vietnam
36 CHI 41 VIE
Synonyms: hidden
= Hemionitis sichouensis (Ching & S.K.Wu) Christenh.; Global Fl. 4: 21 (2018)
species :
Aleuritopteris smithii
(C.Chr.) Windham & Schuettp.; Taxon 74(2), p. 14 of 32 (2025)
Distribution: China (W-Sichuan, NW-Yunnan)
36 CHI
Synonyms: hidden
= Oeosporangium smithii (C.Chr.) Fraser-Jenk. & Pariyar; Annot. Checkl. Ind. Pterid. 1: 265 (2016)
= Cheilanthes smithii (C.Chr.) R.M.Tryon; Amer. Fern J. 76(4): 185 (1986)
= Hemionitis christensenii Christenh.; Global Fl. 4: 11 (2018), nom. nov. for Pellaea smithii C. Chr.
= Mildella smithii (C.Chr.) C.C.Hall & Lellinger; Amer. Fern J. 57: 132 (1967)
= Pellaea smithii C.Chr.; Acta Horti Gothob. 1: 84, t. 18 (1924)
species :
Aleuritopteris speciosa
Ching & S.K.Wu; Acta Phytotax. Sin. 19(1): 61 (1981)
Distribution: E-Tibet
36 TIB
Synonyms: hidden
= Hemionitis speciosa (Ching & S.K.Wu) Christenh.; Global Fl. 4: 21 (2018)
species :
Aleuritopteris squamosa
(C.Hope & C.H.Wright) Ching; Hong Kong Naturalist 10: 199 (1941)
Distribution: China (Yunnan, Hainan)
36 CHI
Synonyms: hidden
= Cheilanthes hopeana C.Chr.; Index Filic., Suppl. (1906-1912) 18 (1913)
= Doryopteris squamosa (C.Hope & C.H.Wright) C.Chr.; Index Filic. 245 (1905)
= Hemionitis hopeana (C.Chr.) Christenh.; Global Fl. 4: 15 (2018)
= Hemionitis squamosa (Gillies ex Hook. & Grev.) Christenh.; Global Fl. 4: 21 (2018)
= Pellaea squamosa C.Hope & C.H.Wright; J. Linn. Soc., Bot. 35: 518 (1903)
species :
Aleuritopteris stenochlamys
Ching; Acta Phytotax. Sin. 19(1): 72 (1981)
Distribution: Tibet; Nepal; Bhutan; India (Sikkim, Uttarakhand, Manipur); Thailand
36 TIB 40 BHU IND NEP 41 THA
Remarks: bona sp. and separate from A. grisea fide Fraser-Jenkins 2008 and 2015
Synonyms: hidden
= Cheilanthes stenochlamys (Ching) Khullar; Fern Fl. W. Himalaya 2: 179 (2024)
species :
Aleuritopteris straminea
(Ching) Windham & Schuettp.; Taxon 74(2), p. 14 of 32 (2025)
Distribution: China (Gansu, S-Qinghai, Xinjiang); Tibet; Bhutan; India (Sikkim)
36 CHI TIB 40 BHU IND
Synonyms: hidden
= Cheilanthes tibetica Fraser-Jenk. & Wangdi; Fern Gaz. 18(5): 218 (2009), nom. nov. for Pellaea straminea Ching
= Hemionitis straminea (Ching) Christenh.; Global Fl. 4: 21 (2018)
= Hemionitis tibetica (Fraser-Jenk. & Wangdi) Christenh.; Global Fl. 4: 22 (2018), nom. superfl.
= Mildella straminea (Ching) C.C.Hall & Lellinger; Amer. Fern J. 57: 130 (1967)
= Oeosporangium stramineum (Ching) Fraser-Jenk.; Annot. Checkl. Ind. Pterid. 1: 260 (2016)
= Pellaea straminea Ching; Bull. Fan Mem. Inst. Biol., Bot. 2: 203, t. 17 (1931), non C. straminea Brause 1913
= Pellaea straminea var. tibetica Ching; Fl. Xizang. 1: 91 (1983)
species :
Aleuritopteris subargentea
Ching ex S.K.Wu; Acta Phytotax. Sin. 19(1): 65 (1981)
Distribution: China; Taiwan; Nepal
36 CHI 38 TAI 40 NEP
Remarks: accepted by several specialists including Knapp (Ferns of Taiwan, 2011 ff.) but Flora of China and Fraser-Jenkins et al. (2015) include it into A. argentea
Conservation Status: IUCN CR for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Cheilanthes subargentea (Ching) C.M.Kuo; Taiwania 30: 57 (1985), without full basionym
= Hemionitis subargentea (Ching ex S.K.Wu) Christenh.; Global Fl. 4: 21 (2018)
species :
Aleuritopteris subdimorpha
(C.B.Clarke & Baker) Fraser-Jenk.; Fern Gaz. 18(5): 223 (2009)
Distribution: SW-China (Himalayas); India (Arunachal Pradesh, Sikkim, Darjeeling, Uttarakhand, Meghalaya, Assam State, Manipur, Nagaland); Bangladesh; Myanmar; Thailand; Laos; Vietnam; Sumatra
36 CHI 40 BAN IND 41 LAO MYA THA VIE 42 SUM
Remarks: subdimorpha agg.
Conservation Status: DD in Thailand
Synonyms: hidden
= Aleuritopteris longipes Ching & S.K.Wu; Acta Bot. Yunnan. 5(2): 165 (1983) (fide Fraser-Jenkins 2008)
= Cheilanthes farinosa var. subdimorpha C.B.Clarke & Baker; J. Linn. Soc., Bot. 24: 411 (1888)
= Cheilanthes longipes (Ching & S.K.Wu) R.D.Dixit & Balkr.; Pl. Sci. Res. (India) 11: 10 (1989)
= Cheilanthes subdimorpha (C.B.Clarke & Baker) B.K.Nayar & Kaur; Companion Handb. Ferns Brit. India [Nayar & Kaur] 26 (1974)
= Hemionitis subdimorpha (C.B.Clarke & Baker) Christenh.; Global Fl. 4: 21 (2018)
species :
Aleuritopteris subvillosa
(Hook.) Ching; Hong Kong Naturalist 10: 203 (1941)
Distribution: China (Guizhou, Sichuan, Yunnan); Tibet; Pakistan (Hazara); Nepal; Bhutan; India (Jammu & Kashmir, Himachal Pradesh, Sikkim, Uttarakhand, West Bengal); Myanmar
36 CHI TIB 40 BHU IND NEP PAK 41 MYA
Synonyms: hidden
= Aleuritopteris subvillosa var. dilatata (Brause) H.S.Kung; Fl. Sichuan. 6: 256 (1988)
= Aleuritopteris subvillosa var. tibetica (Ching & S.K.Wu) H.S.Kung; Fl. Sichuan. 6: 258 (1988)
= Allosorus subvillosus (Hook.) Farw.; Amer. Midl. Naturalist 12: 284 (1931)
= Cheilanthes subvillosa Hook.; Sp. Fil. [W.J.Hooker] 2: 87, t. 98B (1852)
= Cheilanthes yunnanensis Brause; Hedwigia 54(2): 204 (1914)
= Cheilanthes yunnanensis var. dilatata Brause; Hedwigia 54(2): 205, t. 4, f. G (1913)
= Hemionitis subvillosa (Hook.) Christenh.; Global Fl. 4: 21 (2018)
= Leptolepidium subvillosum (Hook.) K.H.Shing & S.K.Wu; Acta Bot. Yunnan. 1(1): 116 (1979)
= Leptolepidium subvillosum var. dilatatum (Brause) K.H.Shing & S.K.Wu; Acta Bot. Yunnan. 1(1): 116 (1979)
= Leptolepidium subvillosum var. tibeticum Ching & S.K.Wu; Acta Bot. Yunnan. 1(1): 116 (1979)
= Oeosporangium subvillosum (Hook.) Fraser-Jenk. & Pariyar; Annot. Checkl. Ind. Pterid. 1: 260 (2016)
species :
Aleuritopteris tamburii
(Hook.) Ching; Hong Kong Naturalist 10: 198 (1941)
Distribution: China (S-Sichuan, NW-Yunnan); ?Tibet; Nepal; India (Arunachal Pradesh, Meghalaya, Sikkim, Uttarakhand); Myanmar
36 CHI 40 IND NEP 41 MYA
Conservation Status: IUCN EN for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Aleuritopteris punethae Kholia, Bhakuni & R.Punetha; Telopea 12(4): 486 (2010)
= Allosorus tamburii (Hook.) Kuntze; Revis. Gen. Pl. 2: 806 (1891)
= Cheilanthes argentea var. tamburii (Hook.) Bedd.; Ferns Brit. India 194 (1893)
= Cheilanthes tamburii (Hook.) T.Moore; Index Fil. [T. Moore] 254 (1861)
= Doryopteris tamburii (Hook.) C.Chr.; Acta Horti Gothob. 1: 86 (1924)
= Hemionitis tamburii (Hook.) Christenh.; Global Fl. 4: 22 (2018)
= Pellaea tamburii Hook.; Sp. Fil. [W.J.Hooker] 2: 134, t. 129 A (1858)
variant :
var. viridis
H.S.Kung; Acta Bot. Yunnan. 5(4): 356 (1983)
Distribution: China (SW-Sichuan)
36 CHI
species :
Aleuritopteris thwaitesii
(Mett.) Saiki; J. Phytogeogr. Taxon. 32(1): 11 (1984)
Distribution: S-India (Tamil Nadu); Sri Lanka
40 IND SRL
Conservation Status: IUCN CR for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Cheilanthes keralensis N.C.Nair & S.R.Ghosh; J. Indian Bot. Soc. 55 52 (1976)
= Cheilanthes laxa T.Moore; Index Fil. [T. Moore] 245 (1861), nomen
= Cheilanthes thwaitesii Mett.; Linnaea 36: 82 (1869)
= Hemionitis keralensis (N.C.Nair & S.R.Ghosh) Christenh.; Global Fl. 4: 16 (2018)
= Hemionitis thwaitesii (Mett.) Christenh.; Global Fl. 4: 22 (2018)
= Oeosporangium thwaitesii (Mett.) Fraser-Jenk.; Annot. Checkl. Ind. Pterid. 1: 262 (2016)
species :
Aleuritopteris tinaei
(Tod.) Windham & Schuettp.; Taxon 74(2), p. 14 of 32 (2025)
Name: Korsischer Schuppenfarn (DE); Helecho de risco (ES)
Distribution: Portugal; Spain; France; Corsica; Sardinia; Italy; Sicily; Morocco; Algeria; Tunisia; Madeira (Madeira Isl.); Canary Isl. (Tenerife); Turkey (Anatolia)
12 COR FRA POR SAR SPA 13 ITA SIC 20 ALG MOR TUN 21 CNY MDR 34 TUR
Synonyms: hidden
= Allosorus tinaei (Tod.) Christenh.; Willdenowia 42(2): 284 (2012)
= Cheilanthes corsica Reichst. & Vida; Candollea 28: 85 (1973)
= Cheilanthes pteridioides subsp. tinaei (Tod.) O.Bolòs, Vigo, Masalles & Ninot; Fl. Man. Paisos Catalans 1213 (1990)
= Cheilanthes tinaei Tod.; Giorn. Sci. Nat. Econ. Palermo 1: 217 (seors. 12) (1866)
= Hemionitis tinaei (Tod.) Christenh.; Global Fl. 4: 22 (2018)
= Oeosporangium tinaei (Tod.) Fraser-Jenk.; Annot. Checkl. Ind. Pterid. 1: 265 (2016)
species :
Aleuritopteris tirajanae
(T.S.Velázquez) Windham & Schuettp.; Taxon 74(2), p. 14 of 32 (2025)
Name: Tirajana-Schuppenfarn (DE)
Distribution: Canary Isl. (Gran Canaria)
21 CNY
Synonyms: hidden
= Cheilanthes tirajanae T.S.Velázquez; Bot. Macaronés. 28: 22 (2013)
species :
Aleuritopteris trichophylla
(Baker) Windham & Schuettp.; Taxon 74(2), p. 14 of 32 (2025)
Distribution: China (W-Yunnan, W-Sichuan); E-Tibet; ?India (?Sikkim)
36 CHI TIB
Synonyms: hidden
= Aleuritopteris delavayi (Baker) B.K.Nayar & Kaur; Companion Handb. Ferns Brit. India [Nayar & Kaur] 26 (1974)
= Cheilanthes delavayi Baker; Ann. Bot. (Oxford) 5: 211 (1891) (non Gymnogramma delavayi Baker)
= Cheilanthes trichophylla Baker; Ann. Bot. (Oxford) 5: 211 (1891)
= Cheilanthes undulata C.Hope & C.H.Wright; Gard. Chron., Ser. 3, 34: 397 (1903)
= Hemionitis trichophylla (Baker) Christenh.; Global Fl. 4: 22 (2018)
= Oeosporangium trichophyllum (Baker) Fraser-Jenk.; Annot. Checkl. Ind. Pterid. 1: 262 (2016)
= Pellaea trichophylla (Baker) Ching; Acta Phytotax. Sin. 10: 302 (1965)
species :
Aleuritopteris veitchii
(Christ) Ching; Hong Kong Naturalist 10: 200 (1941)
Distribution: China (Guangxi, S-Sichuan, NW-Yunnan)
36 CHI
Synonyms: hidden
= Aleuritopteris cremea Ching; Acta Phytotax. Sin. 19(1): 65 (1981)
= Aleuritopteris likiangensis Ching; Acta Phytotax. Sin. 19(1): 64 (1981) (fide Fraser-Jenkins et al. 2018)
= Cheilanthes veitchii (Christ) Ching; Index Filic., Suppl. Tert. 55 (1934)
= Doryopteris veitchii Christ; Bull. Acad. Int. Géogr. Bot. 134 (1904)
= Hemionitis cremea (Ching) Christenh.; Global Fl. 4: 12 (2018)
= Hemionitis likiangensis (Ching) Christenh.; Global Fl. 4: 17 (2018)
= Hemionitis veitchii (Christ) Christenh.; Global Fl. 4: 22 (2018)
species :
Aleuritopteris velutina
(Tardieu & C.Chr.) Windham & Schuettp.; Taxon 74(2), p. 14 of 32 (2025)
Distribution: SW-Thailand; Vietnam
41 THA VIE
Synonyms: hidden
= Cheilanthes velutina (Tardieu & C.Chr.) Fraser-Jenk.; New Sp. Syndr. Indian Pteridol. 84 (1997)
= Notholaena velutina Tardieu & C.Chr.; Notul. Syst. (Paris) 6: 167, t. 1(5-7) (1938)
= Hemionitis velutina (Tardieu & C.Chr.) Christenh.; Global Fl. 4: 22 (2018)
species :
Aleuritopteris welwitschii
(Hook. ex Baker) Ching; Hong Kong Naturalist 10: 202 (1941)
Distribution: Tanzania; Angola; Malawi; Zimbabwe; Mozambique
25 TAN 26 ANG MLW MOZ ZIM
Synonyms: hidden
= Cheilanthes mossambicensis Schelpe; J. S. African Bot. 30: 183, f. 1b (1964)
= Cheilanthes welwitschii Hook.; Syn. Fil. [W.J.Hooker & Baker] 142 (1867)
= Hemionitis welwitschii (Hook. ex Baker) Christenh.; Global Fl. 4: 23 (2018)
species :
Aleuritopteris wollenweberi
Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 127 (2008)
Distribution: S-India (Kerala, Tamil Nadu); Sri Lanka
40 IND SRL
species :
Aleuritopteris yalungensis
H.S.Kung; Acta Bot. Yunnan. 5(4): 356 (1983)
Distribution: China (SW-Sichuan)
36 CHI
Synonyms: hidden
= Hemionitis yalungensis (H.S.Kung) Christenh.; Global Fl. 4: 23 (2018)
species :
Aleuritopteris yunnanensis
(Ching) Windham & Schuettp.; Taxon 74(2), p. 15 of 32 (2025)
Distribution: China (Sichuan, Yunnan); Bhutan; India (?Sikkim)
36 CHI 40 BHU IND
Conservation Status: IUCN CR for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Cheilanthes bhutanica Fraser-Jenk. & Wangdi; Fern Gaz. 18(5): 220 (2009), nom. nov.
= Hemionitis bhutanica (Fraser-Jenk. & Wangdi) Christenh.; Global Fl. 4: 10 (2018), nom. superfl.
= Hemionitis yunnanensis (Ching) Christenh.; Global Fl. 4: 23 (2018)
= Oeosporangium yunnanense (Ching) Fraser-Jenk. & Pariyar; Annot. Checkl. Ind. Pterid. 1: 264 (2016)
= Pellaea yunnanensis Ching; Acta Phytotax. Sin. 20(2): 235 (1982), non Brause (1913)
species :
Aleuritopteris ×confundans
Fraser-Jenk.; Ferns Fern-Allies Nepal 1: 252 (2015), nom. inval., et Annot. Checkl. Indian Pteridophytes 2: 421 (2018)
Distribution: Nepal; India (Uttarakhand)
40 IND NEP
Remarks: Aleuritopteris anceps x formosana
species :
Aleuritopteris ×duriensis
(Mendonça & Vasc.) Windham & Schuettp.; Taxon 74(2), p. 12 of 32 (2025)
Distribution: Portugal; Spain
12 POR SPA
Remarks: Aleuritopteris hispanicus x pteridioides
Synonyms: hidden
= Allosorus ×prototinaei (Rasbach, Reichst. & Schneller) Rumsey; Fern Gaz. 19(7): 277 (2014)
= Cheilanthes ×duriensis Mendonça & Vasc.; Anais Inst. Vinho Porto 15(4): 47 (1956)
= Cheilanthes ×prototinaei Rasbach, Reichst. & Schneller; Webbia 37(1): 45 (1983)
= Hemionitis ×prototinaei (Rasbach, Reichst. & Schneller) F.M.Vázquez; Folia Bot. Extremadur. 15: 149 (2021)
= Oeosporangium ×prototinaei (Rasbach, Reichst. & Schneller) Arana & Mor.-Saiz; Phytotaxa 433(3): 192 (2020)
species :
Aleuritopteris ×fraser-jenkinsii
(Thapa) Fraser-Jenk. & Khullar; Taxon. Revis. Indian Subcontinental Pteridophytes 131 (2008)
Distribution: Nepal
40 NEP
Remarks: Aleuritopteris dealbatus x formosanus
Synonyms: hidden
= Cheilanthes ×fraser-jenkinsii Thapa; Pterid. Nepal 66 (2002)
species :
Aleuritopteris ×gardneri
Fraser-Jenk.; Ferns Fern-Allies Nepal 1: 252 (2015)
Distribution: Nepal
40 NEP
Remarks: Aleuritopteris dealbatus x rufus
species :
Aleuritopteris ×godavariensis
Fraser-Jenk. & Khullar; Annot. Checkl. Ind. Pterid. 1: 230 (2016)
Distribution: Nepal; India (Uttarakhand)
40 IND NEP
Remarks: Aleuritopteris anceps x bicolor
species :
Aleuritopteris ×hamiltonii
Fraser-Jenk.; Ferns Fern-Allies Nepal 1: 253 (2015)
Distribution: Nepal; India (Sikkim)
40 IND NEP
Remarks: Aleuritopteris dubius x formosanus
species :
Aleuritopteris ×iberica
(Rasbach & Reichst.) Windham & Schuettp.; Taxon 74(2), p. 13 of 32 (2025)
Distribution: Spain
12 SPA
Remarks: Aleuritopteris hispanicus x tinaei
Synonyms: hidden
= Allosorus ×ibericus (Rasbach & Reichst.) Rumsey; Fern Gaz. 19(7): 277 (2014)
= Cheilanthes ×iberica Rasbach & Reichst.; Webbia 35(2): 264 (1982)
= Hemionitis ×iberica (Rasbach & Reichst.) F.M.Vázquez; Folia Bot. Extremadur. 15: 149 (2021)
= Oeosporangium ×ibericum (Rasbach & Reichst.) Arana & Mor.-Saiz; Phytotaxa 433(3): 191 (2020)
species :
Aleuritopteris ×insularis
(Rasbach & Reichst.) Windham & Schuettp.; Taxon 74(2), p. 13 of 32 (2025)
Distribution: Spain; Corsica; Sardinia; Canary Isl. (Tenerife)
12 COR SAR SPA 21 CNY
Remarks: Aleuritopteris guanchicus x tinaei
Synonyms: hidden
= Allosorus ×insularis (Rasbach & Reichst.) Rumsey; Fern Gaz. 19(7): 277 (2014)
= Cheilanthes ×insularis Rasbach & Reichst.; Webbia 35(2): 266 (1982)
= Hemionitis ×insularis (Rasbach & Reichst.) F.M.Vázquez; Folia Bot. Extremadur. 15: 149 (2021)
= Oeosporangium ×insulare (Rasbach & Reichst.) Arana & Mor.-Saiz; Phytotaxa 433(3): 191 (2020)
species :
Aleuritopteris ×khasiana
Fraser-Jenk.; Annot. Checkl. Ind. Pterid. 1: 231 (2016)
Distribution: India (Meghalaya)
40 IND
Remarks: Aleuritopteris formosanus x subdimorphus
species :
Aleuritopteris ×kochiana
(Rasbach, Reichst. & Schneller) Windham & Schuettp.; Taxon 74(2), p. 13 of 32 (2025)
Distribution: Spain
12 SPA
Remarks: Aleuritopteris pteridioides x tinaei
Synonyms: hidden
= Allosorus ×kochianus (Rasbach, Reichst. & Schneller) Rumsey; Fern Gaz. 19(7): 277 (2014)
= Allosurus ×kochianus (Rasbach, Reichst. & Schneller) L.Sáez & Aymerich; Orsis 29: 26 (2015), isonym
= Cheilanthes ×kochiana Rasbach, Reichst. & Schneller; Webbia 37(1): 48 (1983)
= Hemionitis ×kochiana (Rasbach, Reichst. & Schneller) F.M.Vázquez; Folia Bot. Extremadur. 15: 149 (2021)
= Oeosporangium ×kochianum (Rasbach, Reichst. & Schneller) L.Sáez & Aymerich; Orsis 31: 33 (2017), as ´kochianus´
species :
Aleuritopteris ×kurdica
(Rasbach & Reichst.) Windham & Schuettp.; Taxon 74(2), p. 13 of 32 (2025)
Distribution: NE-Iraq; E-Afghanistan
34 AFG IRQ
Remarks: Aleuritopteris acrosticus x persicus
Synonyms: hidden
= Allosorus ×kurdicus (Rasbach & Reichst.) Rumsey; Fern Gaz. 19(7): 277 (2014)
= Cheilanthes ×kurdica Rasbach & Reichst.; Webbia 37(1): 58 (1983)
= Oeosporangium ×kurdicum (Rasbach & Reichst.) Fraser-Jenk. & Pariyar; Annot. Checkl. Ind. Pterid. 1: 266 (2016)
species :
Aleuritopteris ×malacitensis
(Rasbach & Reichst.) Windham & Schuettp.; Taxon 74(2), p. 13 of 32 (2025)
Distribution: Spain
12 SPA
Remarks: Aleuritopteris acrostichus x guanchicus
Synonyms: hidden
= Cheilanthes ×malacitensis Rasbach & Reichst.; Webbia 35(2): 269 (1982)
= Oeosporangium ×malacitense (Rasbach & Reichst.) Arana & Mor.-Saiz; Phytotaxa 433(3): 191 (2020)
= Allosorus ×malacitensis (Rasbach & Reichst.) Rumsey; Fern Gaz. 19(7): 277 (2014)
= Hemionitis ×malacitensis (Rasbach & Reichst.) F.M.Vázquez; Folia Bot. Extremadur. 15: 149 (2021)
species :
Aleuritopteris ×marchettiana
(Rasbach, Reichst. & Schneller) Windham & Schuettp.; Taxon 74(2), p. 13 of 32 (2025)
Distribution: Italy
13 ITA
Remarks: Aleuritopteris acrosticus x pteridioides
Synonyms: hidden
= Allosorus ×marchettianus (Rasbach, Reichst. & Schneller) Rumsey; Fern Gaz. 19(7): 277 (2014)
= Cheilanthes ×marchettiana Rasbach, Reichst. & Schneller; Webbia 37(1): 55 (1983)
= Hemionitis ×marchettiana (Rasbach, Reichst. & Schneller) F.M.Vázquez; Folia Bot. Extremadur. 15: 149 (2021)
= Oeosporangium pteridioides subsp. reichsteinii Fraser-Jenk. & Pariyar; Annot. Checkl. Ind. Pterid. 1: 266 (2016) [nothosubsp.]
= Oeosporangium ×marchettianum (Rasbach, Reichst. & Schneller) Arana & Mor.-Saiz; Phytotaxa 433(3): 191 (2020)
species :
Aleuritopteris ×meridionalis
(F.M.Vázquez) Windham & Schuettp.; Taxon 74(2), p. 13 of 32 (2025)
Distribution: Spain
12 SPA
Remarks: Aleuritopteris acrostichus x tinaei
Synonyms: hidden
= Hemionitis ×meridionalis F.M.Vázquez; Folia Bot. Extremadur. 15: 150 (2021)
species :
Aleuritopteris ×nepalensis
Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 132 (2008)
Distribution: Nepal; India (Meghalaya)
40 IND NEP
Remarks: Aleuritopteris rufus x formosanus
species :
Aleuritopteris ×pangteyi
Fraser-Jenk. & E.Wollenw.; Taxon. Revis. Indian Subcontinental Pteridophytes 130 (2008)
Distribution: Nepal; India (Jammu & Kashmir, Himachal Pradesh, Uttarakhand)
40 IND NEP
Remarks: Aleuritopteris bicolor x formosanus
species :
Aleuritopteris ×teneriffae
(Rasbach & Reichst.) Windham & Schuettp.; Taxon 74(2), p. 14 of 32 (2025)
Distribution: Canary Isl. (Tenerife)
21 CNY
Remarks: Aleuritopteris pteridioides x pulchellus
Synonyms: hidden
= Allosorus ×teneriffae (Rasbach & Reichst.) Rumsey; Fern Gaz. 19(7): 277 (2014)
= Cheilanthes ×teneriffae Rasbach & Reichst.; Webbia 35(2): 262 (1982)
= Oeosporangium ×teneriffae (Rasbach & Reichst.) Arana & Mor.-Saiz; Phytotaxa 433(3): 192 (2020)
species :
Aleuritopteris ×tolocensis
(Rasbach, Reichst. & Schneller) Windham & Schuettp.; Taxon 74(2), p. 14 of 32 (2025)
Distribution: Spain
12 SPA
Remarks: Aleuritopteris guanchicus x pteridioides
Synonyms: hidden
= Allosorus ×tolocensis (Rasbach, Reichst. & Schneller) Rumsey; Fern Gaz. 19(7): 277 (2014)
= Cheilanthes ×tolocensis Rasbach, Reichst. & Schneller; Webbia 37(1): 52 (1983)
= Hemionitis ×tolocensis (Rasbach, Reichst. & Schneller) F.M.Vázquez; Folia Bot. Extremadur. 15: 150 (2021)
= Oeosporangium ×tolocense (Rasbach, Reichst. & Schneller) Arana & Mor.-Saiz; Phytotaxa 433(3): 192 (2020)
species :
Aleuritopteris ×unicolor
Fraser-Jenk. & Khullar; Taxon. Revis. Indian Subcontinental Pteridophytes 131 (2008)
Distribution: Nepal
40 NEP
Remarks: Aleuritopteris rufus x bicolor
species :
Aleuritopteris ×vermae
(Fraser-Jenk. & Viane) Fraser-Jenk. & Khullar; Taxon. Revis. Indian Subcontinental Pteridophytes 130 (2008)
Distribution: India (Jammu & Kashmir, Himachal Pradesh, Uttarakhand)
40 IND
Remarks: Aleuritopteris albomarginatus x bicolor
Synonyms: hidden
= Cheilanthes ×vermae Fraser-Jenk. & Viane; Bot. Helv. 102(2): 154 (1992)
species :
Aleuritopteris ×wallichiana
Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 132 (2008)
Distribution: Nepal
40 NEP
Remarks: Aleuritopteris bicolor x dealbatus
