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Ver. 24.11
November 5th, 2024

005.0000 Family Psilotaceae J.W.Griff. & Henfr.; Microgr. Dict.: 540 (1855) (Psiloteae)
2 genera, 18 spp.
Species listed in database: 19; hybrids: 1
= Tmesipteridaceae Nakai; Chosakuronbun Mokuroku [Ord. Fam. Trib. Nov.]: 206 (1943)
005.0001 Genus Psilotum Sw.; J. Bot. (Schrader) 1800(2): 8 (1802)
Distribution: Pantropic to warm-temperate, north to SE-USA, extreme SW-Europe, C-Japan and S-Korea. New taxonomy fide Tejero-Díez et al. 2024.
Species listed in database: 2; hybrids: 1
= Bernhardia Bernh.; Abh. Kurfürstl.-Mainz. Akad. Nützl. Wiss. Erfurt 2: 11 (1802)
= Hoffmannia Willd.; Bot. Mag. (Römer & Usteri) 2(6): 15, 17 (1789)
= Tristeca P.Beauv. ex Mirb.; Hist. Nat. Veg. [Lamarck & Mirbel] 3: 478 (1802), nom. superfl.
species :
Psilotum flaccidum Wall. ex Hook. & Grev.; Bot. Misc. 2(6): 363 (1831)
Distribution: Seychelles; Nicobar Isl.; peninsular Thailand; Vietnam; peninsular Malaysia (Perak, Selangor, Negeri Sembilan, Terengganu, Pahang, Johor); +Singapore; Sumatra; Borneo (Sabah, Sarawak, Kalimantan); Sulawesi; Philippines; Moluccas (Seram); New Guinea (Irian Jaya, Papua New Guinea, d´Entrecasteaux Arch.: Normanby Isl., Louisiade Arch.: Misima Isl., Woodlark Isl.); Bismarck Arch. (Admiralty Isl.); Solomon Isl. (Baga, Bougainville, Guadalcanal, Santa Ysabel, Kolombangara, Makira, Malaita, New Georgia, Rendova, Kirakira Isl., Ranongga, Tetepare, Vangunu, Vella Lavella); Australia (E-Queensland, NE-New South Wales); Santa Cruz Isl.; Tonga (Vava´u); Fiji (Viti Levu, Vanua Levu, Taveuni); Wallis & Futuna (Futuna); American Samoa (Tutuila, Ta´u); [Western] Samoa (Savai´i, ´Upolu); Tokelau (Swains Isl.); Cook Isl. (Rarotonga, Mauke); Society Isl. (Tahiti); Palau (Babeldaob, Malakal); Northern Marianas (Alamagan, Rota Isl.); Fed. States of Micronesia (Nukuoro, Pohnpei, Kosrae); Hawaii (Kauai, Oahu, Molokai, Lanai, Maui, Hawaii Isl.); Mexico (Chiapas, Chihuahua, Colima, Guerrero, Hidalgo, Jalisco, Mexico State, Morelos, Nayarit, Oaxaca, Puebla, Sinaloa, Sonora, Tamaulipas, Veracruz); Guatemala; El Salvador; Honduras; Nicaragua; Costa Rica; Cuba; Jamaica; Hispaniola (Haiti, Dominican Republic); Colombia (Cundinamarca); Peru
29 SEY 41 NCB THA VIE 42 BOR MLY MOL NWG PHI SNG SUL SUM 43 BIS SOL 50 AUS 60 ASA FIJ SAM SCZ TOK TON WAL 61 COO SCI 62 MIC MRN PLU 63 HAW 79 MEX 80 COS ELS GUA HON NIC 81 CUB HIS JAM 82 CLM 83 PER
Remarks: for a long time known as "P. complanatum Sw.", but this name belongs to the hybrid fide Tejero-Díez et al. 2024.
Conservation Status: IUCN EN for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Bernhardia ramulosa Müll.Hal.; Bot. Zeitung (Berlin) 14: 222 (1856)
= Bernhardia schiedeana Müll.Hal.; Bot. Zeitung (Berlin) 14: 222, fig. 9-11 (1856)
= Psilotum complanatum var. flaccidum (Wall. ex Hook. & Grev.) Domin; Luerss., Biblioth. Bot. 85: 234 (1913)
= Psilotum complanatum var. latissimum Spring; Nouv. Mém. Acad. Roy. Sci. Bruxelles 24: 271 (1849)
= Psilotum flaccidum Wall.; Numer. List. [Wallich] no. 45 (1829), nomen
species :
Psilotum nudum (L.) P.Beauv.; Prodr. Aethéogam. 112 (1805)
Name: Whisk Fern (EN); Nacktfarn (DE)
Distribution: Spain (S-Spain); Canary Isl. [I] (Tenerife [I], Lanzarote [I]); Madeira [I] (Funchal [I]); Cape Verde Isl. (Santo Antao Isl., Ilha de Sao Nicolau, Ilha Brava); Senegal; Guinea; Liberia; Ghana; Benin; Nigeria; Cameroon; Bioko Isl. [Fernando Poo]; São Tomé; Annobon Isl.; Equatorial Guinea (Rio Muni); Gabon; Congo [Brazzaville]; D.R.Congo [Zaire]; Uganda; Kenya; Pemba Isl.; Tanzania; Angola; Zambia; Malawi; Zimbabwe; Mozambique; South Africa (Limpopo, Mpulamanga, Free State, KwaZulu-Natal, E-Cape Prov.); Lesotho; Eswatini [Swaziland]; Ascension Isl.; Madagascar; Comores (Mayotte); Mauritius; La Réunion; Rodrigues; Iles Glorieuses; Aldabra (Cosmoledo Isl.); Agalega Isl.; Chagos Arch. (Diego Garcia); Russian Far East (Kuril Isl.); Saudi Arabia; N-Yemen; China (Anhui, Chongqing, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hunan, Jiangsu, Jiangxi, Macau, Shaanxi, Sichuan, Yunnan, Zhejiang); South Korea (Cheju Isl.); Japan; Bonin Isl. (Chichijima, Minamijima, Anijima, Otohtojima, Hahajima, Meijima); Taiwan; Pakistan; Nepal; Bhutan; India (Andhra Pradesh, Arunachal Pradesh, Assam State, Bihar, Chhattisgarh, Himachal Pradesh, Karnataka, Kerala, Madhya Pradesh, Maharashtra, Meghalaya, Mizoram, Nagaland, Odisha, Punjab, Sikkim, Tamil Nadu, Uttarakhand, Uttar Pradesh, West Bengal); Lakshadweep Isl. [Laccadives]; Maldives; Sri Lanka; Bangladesh; Andaman Isl.; Nicobar Isl.; Myanmar; Thailand (widespread); Laos; Vietnam; peninsular Malaysia (widespread); Singapore; Borneo (Sabah, Sarawak); Sulawesi; Palawan; Philippines; Lesser Sunda Isl. (Bali); East Timor; Moluccas (Seram); Christmas Isl. [Austr.]; New Guinea (Irian Jaya, Papua New Guinea, d´Entrecasteaux Arch.: Normanby Isl., Louisiade Arch.: Sudest Isl.); Bismarck Arch. (incl. Admiralty Isl.); Solomon Isl. (Baga, Bougainville, Choiseul, Santa Ysabel, Kolombangara, Malaita, Mono, Tulagi, Guadalcanal, Ranongga, Rennell); Australia (Western Australia, Northern Territory, Queensland, South Australia, Victoria, New South Wales); Lord Howe Isl.; Norfolk Isl.; New Zealand (North Isl., Poor Knights Isl.); Kermadec Isl.; Santa Cruz Isl.; New Caledonia (incl. Loyalty Isl.); Rotuma Isl.; Tonga (Late Isl., Tongatapu, ´Eua, Kao, Niuatoputapu); Niue; Fiji (Viti Levu, Vanua Levu, Beqa Isl., Taveuni); Wallis & Futuna (Futuna Isl., Alofi, ´Uvea, Wallis); American Samoa (Ofu); [Western] Samoa (Savai´i, ´Upolu); Tuvalu (Nui Atoll, +Funafuti, Nanumea, Vaitupu, Nukulaelae, Niutao); Tokelau (Nukunonu, Fakaofo, Swains Isl.); Gilbert Isl. (Butaritari, Tarawa); Banaba [Ocean] Isl.; Nauru; N-Line Isl. (Palmyra, Washington, Fanning); S-Line Isl. (Caroline Atoll); Cook Isl. (Rarotonga, Mangaia, Penrhyn, Rakahanga, Manihiki, Pukapuka, Nassau, Suwarrow, Palmerston, Aitutaki, Manuae, Takutea, Mitiaro, Atiu, Mauke); Marquesas Isl. (Eiao, Nuku Hiva, Ua Huka, Ua Pou, Hiva Oa, Tahuata, Fatu Hiva); Society Isl. (Maupiti, Tahiti, Tahaa, Moorea, Huahine, Tetiaroa Atoll, Tupai); Gambier Isl. (Mangareva); Tuamotu Arch. (Makatea, Rangiroa, Ahii, Takaroa, Manihi, Kaehi, Tahanea, Toau, Faite, Taenga, Makemo, Taiaro, Takapoto Atoll); Austral Isl. (Rapa Iti, Rurutu, Tubuai, Maria Rocks, Rimatara, Raivavae); Pitcairn Isl.; Henderson Isl.; +Easter Isl.; Palau (Babeldaob, Koror, Ngarakabesang, Malakal, Ulebsechel, Sonsorol); Northern Marianas (Pagan, Anatahan, Rota Isl., Saipan); Guam; Fed. States of Micronesia (Sonsorol, Yap, Truk, Moen, Fefan, Piis, Nukuoro, Kapingamarangi, Pohnpei, Kosrae); Marshall Isl.. (Enewetok, Kwajalein, Majuro, Arno, Jaluit); Hawaii (Niihau, Kauai, Oahu, Molokai, Lanai, Maui, Hawaii Isl., Midway Isl.); USA (Alabama, Arkansas, Arizona, Florida, Georgia, Louisiana, Mississippi, North Carolina, South Carolina, Texas); Bermuda; Mexico (Campeche, Chiapas, Colima, Ciudad de Mexico, Guerrero, Jalisco, Mexico State, Morelos, Oaxaca, Sonora, Tabasco, Tamaulipas, Veracruz, Yucatan); Revillagigedos Isl. (Isla Socorro); Guatemala; Belize; El Salvador; Honduras; Nicaragua; Costa Rica; Isla del Coco; Panama; Bahamas; Turks & Caicos Isl.; Cuba; Cayman Isl. (Grand Cayman); Jamaica; Hispaniola (Haiti, Dominican Republic); Puerto Rico; Virgin Isl. (Tortola, Virgin Gorda, St. John, St. Thomas); St. Martin/St. Maarten; St. Barthélémy; Saba; Nevis; St. Kitts; Redonda; Montserrat; Guadeloupe (incl. La Desirade); Marie-Galante; Dominica; Martinique; St. Lucia; St. Vincent; Grenada; Trinidad; Colombia (Antioquia, Caqueta, Chocó, Huila, Nariño, Valle); Venezuela (Amazonas, Anzoategui, Aragua, Carabobo, Cojedes, Venezuela Distrito Federal, Falcon, Guarico, Lara, Merida, Miranda, Monagas, Nueva Esparta, Sucre, Yaracuy); Surinam; French Guiana; Ecuador; Galapagos; Peru; Bolivia (Beni, Cochabamba, La Paz, Pando, Santa Cruz); Brazil (Acre, Amazonas, Bahia, Ceará, Brasilia Distrito Federal, Espírito Santo, Goiás, Maranhão, Minas Gerais, Mato Grosso do Sul, Mato Grosso, Pará, Paraíba, Pernambuco, Paraná, Rio Grande do Norte, Rio de Janeiro, Rondônia, Rio Grande do Sul, Santa Catarina, São Paulo, Sergipe, Tocantins); Ilha da Trindade; Paraguay (Alto Paraná, Amambay, Caaguazu, Canindeyu, Central, Concepcion, Cordillera, Paraguari, San Pedro); Uruguay (Rocha, Tacuarembo); Argentina (Corrientes, Misiones, Salta)
12 SPA 21 cny CVI mdr 22 BEN GHA GUI LBR NGA SEN 23 ANN BIO CMN CON EQG GAB SAO ZAI 25 KEN TAN UGA 26 ANG MLW MOZ ZAM ZIM 27 LES SAF SWZ 28 ASC 29 AGA ALD CGS COM IGL MAU MDG REU ROD 31 RFE 35 SAU YEM 36 CHI 38 JAP KOR OGA TAI 40 BAN BHU IND LDV MDV NEP PAK SRL 41 AND LAO MYA NCB THA VIE 42 BOR LSI MLY MOL NWG PHI PLW SNG SUL XMS 43 BIS SOL 50 AUS LHI NFK 51 KER NZL 60 ASA FIJ GIL NRU NUE NWC OCE ROT SAM SCZ TOK TON TUV WAL 61 AUI COO EAS GBI HEN LIN MRQ PIT SCI TUA 62 GUM MIC MRN PLU 63 HAW 76 USA 77 USA 78 USA 79 MEX RGI 80 BLZ COC COS ELS GUA HON NIC PAN 81 ABA BAH BER CAY CUB DOM GDL GRE HIS JAM MAR MTS PUE SAB SBA STK STL STM STV TCI TRT VIR 82 CLM FRG SUR VEN 83 BOL ECU GAL PER 84 BRA ITR 85 ARG PAR URU
Synonyms: hidden
= Bernhardia antillarum Müll.Hal.; Bot. Zeitung (Berlin) 14: 234 (1856)
= Bernhardia capensis Müll.Hal.; Bot. Zeitung (Berlin) 14: 239 (1856)
= Bernhardia deppeana Müll.Hal.; Bot. Zeitung (Berlin) 14: 235 (1856)
= Bernhardia dichotoma Willd.; Abh. Kurfürstl.-Mainz. Akad. Nützl. Wiss. Erfurt 2(6): 11 (1802), nom. superfl.
= Bernhardia novae-hollandiae Müll.Hal.; Bot. Zeitung (Berlin) 14: 237 (1856)
= Bernhardia oahuensis Müll.Hal.; Bot. Zeitung (Berlin) 14: 235 (1856)
= Hoffmannia aphylla Willd.; Bot. Mag. (Römer & Usteri) 2(6): 17 (1789) (nom. illeg superfl.)
= Hoffmannia uxpanapense Cast.-Campos & Baut.-Bello; Acta Bot. Mex. 119: 152 (2017) (tentative synonym)
= Lycopodium nudum L.; Sp. Pl. 2: 1100 (1753)
= Psilotum domingense Gand.; Bull. Soc. Bot. France 66: 306 (1919)
= Psilotum flabellatum Gand.; Bull. Soc. Bot. France 66: 306 (1919)
= Psilotum floridanum Michx.; Fl. Bor.-Amer. (Michaux) 2: 281 (1803)
= Psilotum heterocarpum Colenso; Trans. New. Zealand Inst. 20: 237 (1888)
= Psilotum neocaledonicum Gand.; Bull. Soc. Bot. France 66: 306 (1919)
= Psilotum novae-zelandiae Gand.; Bull. Soc. Bot. France 66: 306 (1919)
= Psilotum nudum (L.) Griseb.; Syst. Unters. 130 (1857), isonym
= Psilotum nudum var. fallicinum (Domin) Domin; Biblioth. Bot. 20: 232 (1915)
= Psilotum nudum var. molesworthii Iranzo, Prada & Salvo; Fl. Pterid. Andalucia: 117 (1982) (fide Fraser-Jenkins 1997)
= Psilotum triquetrum f. insulare Brade; Bradea 1(1): 9, tab. 5 (1969)
= Psilotum triquetrum Sw.; J. Bot. (Schrader) 1800(2): 109 (1801) (nom. illeg. superfl.)
= Psilotum triquetrum var. fallacinum Domin; Repert. Spec. Nov. Regni Veg. 9: 551 (1911)
= Psilotum triquetrum var. gracile Grev. & Hook.; Bot. Misc. 2: 362 (1831)
= Psilotum triquetrum var. latum Desv.; Mem. Soc. Linn. Paris 6(2): 192 (1827)
= Tristeca nuda (L.) P.Beauv. ex J.St.-Hil.; Expos. Fam. Nat. 1: 39 (1805)
species :
Psilotum ×complanatum Sw.; J. Bot. (Schrader) 1800(2): 110 (1801), pro spec.
Distribution: Hawaii (Kauai, Oahu, Lanai, Maui); Ecuador; El Salvador; Guadeloupe; Guatemala; Hispaniola (Haiti); Honduras; Jamaica; Mexico (Chihuahua, Ciudad de México, Guanajuato, Guerrero, Jalisco, Morelos, Nayarit, Puebla, Sinaloa, Sonora, Veracruz, Zacatecas); Montserrat; Puerto Rico; Venezuela (Caracas)
63 HAW 79 MEX 80 ELS GUA HON 81 GDL HIS JAM MTS PUE 82 VEN 83 ECU
Remarks: Psilotum flaccidum x nudum
Synonyms: hidden
= Bernhardia complanata (Sw.) Willd.; Sp. Pl., ed. 4 [Willdenow] 5(1): 57 (1810)
= Psilotum complanatum var. mexicanum Domin; Biblioth. Bot. 20: 234 (1915)
= Psilotum triquetrum var. complanatum (Sw.) Christ; (1897)
= Psilotum ×intermedium W.H.Wagner; Contr. Univ. Michigan Herb. 19: 79 (1993)
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A complete, synonymic checklist of the Ferns and Lycophytes of the World

CAUTION: Because two families were added, the families Desmophlebiaceae and Hemidictyaceae (with together 3 species) are not displayed in "World Ferns" any more. We are asap fixing this. In the meantime consult the main list if you need those families.

Version 24.11 (November 5th, 2024)

Covering 13,146 accepted species,
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in total 69,233* names

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