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041.0000 Family Polypodiaceae J. & C.Presl; Delic. Prag. 159 (1822)
67 genera, ca. 1700 spp. Phylogeny: Wei & Zhang 2022
Species listed in database: 1739; hybrids: 48
= Platyceriaceae Ching; Acta Phytotax. Sin. 16(3): 18 (1978)
= Drynariaceae Ching; Acta Phytotax. Sin. 16: 1-37 (1978)
= Grammitidaceae Newman; Hist. Brit. Ferns 7 (1840) (Grammitaceae auct., missp.)
= Gymnogrammitidaceae Ching; Acta Phytotax. Sin. 11: 12 (1966)
= Loxogrammaceae Ching ex Pic.Serm.; Webbia 29: 11 (1975)
= Platyceriaceae Ching; Sunyatsenia 5: ? (1940)
= Pleurosoriopsidaceae Sa.Kurata & Ikebe ex Ching; Acta Phytotax. Sin. 16(4): 17 (1978)
041.6001 Genus Niphidium J.Sm.; Hist. Fil. 99 (1875)
Distribution: Tropical America. Sometimes included in Campyloneurum.
Species listed in database: 11
= Pessopteris Underw. ex Maxon; Contr. U.S. Natl. Herb. 10: 485 (1908)
= Anaxetum Schott; Gen. Fil. [Schott] t. 50 (1834) (nec Gaertner (1791))
= Pleuridium (C.Presl) Fée; Mém. Foug. 5 (Gen. Filic.) 273 (1852) (non Bridel 1819)
species :
Niphidium albopunctatissimum (J.Sm.) Lellinger; Amer. Fern J. 62: 109 (1972)
Distribution: Colombia (Antioquia, Cauca, Chocó, Cundinamarca, Meta, Nariño, Santander); Ecuador; Peru; Bolivia (Cochabamba, La Paz, Santa Cruz)
82 CLM 83 BOL ECU PER
Synonyms: hidden
= Campyloneurum albopunctatissimum (J.Sm.) Christenh.; Global Fl. 4: 40 (2018), as ´(Lellinger) Christenh.´
= Pessopteris albopunctatissima (J.Sm.) Pic.Serm.; Webbia 31: 248 (1977)
= Pessopteris crassifolia var. albopunctatissima (J.Sm.) Farw.; Amer. Midl. Naturalist 12: 298 (1931)
= Pleuridium albopunctatissimum J.Sm.; Ferns Brit. Foreign 95 (1866)
= Polypodium crassifolium var. longipes Rosenst.; Repert. Spec. Nov. Regni Veg. 11: 58 (1912)
species :
Niphidium anocarpos (Kunze) Lellinger; Amer. Fern J. 62: 120 (1972)
Distribution: Peru; Bolivia (Beni, Chuquisaca, Cochabamba, La Paz, Santa Cruz)
83 BOL PER
Synonyms: hidden
= Campyloneurum anocarpos (Kunze) Christenh.; Global Fl. 4: 40 (2018)
= Pessopteris anocarpos (Kunze) Pic.Serm.; Webbia 31(1): 248 (1977)
= Polypodium acrosorum Kunze; Linnaea 9: 39 (1843)
= Polypodium anocarpos Kunze; Linnaea 9: 40 (1843)
species :
Niphidium carinatum Lellinger; Amer. Fern J. 62: 113 (1972)
Distribution: Colombia (Nariño, Tolima); Ecuador; Peru; Bolivia (Beni, La Paz)
82 CLM 83 BOL ECU PER
Synonyms: hidden
= Campyloneurum carinatum (Lellinger) Christenh.; Global Fl. 4: 40 (2018)
= Pessopteris carinata (Lellinger) Pic.Serm.; Webbia 31(1): 248 (1977)
species :
Niphidium crassifolium (L.) Lellinger; Amer. Fern J. 62: 106 (1972)
Distribution: New Zealand [I] (North Isl. [I]); Mexico (Campeche, Chiapas, Guerrero, Hidalgo, Oaxaca, Puebla, Queretaro, Quintana Roo, San Luis Potosi, Veracruz); Guatemala; Belize; El Salvador; Honduras; Nicaragua; Costa Rica; Panama; Cuba; Jamaica; Hispaniola (Haiti, Dominican Republic); Puerto Rico; Saba; Montserrat; Guadeloupe; Dominica; Martinique; St. Lucia; St. Vincent; Barbados; Trinidad; Colombia (Amazonas, Antioquia, Bolívar, Boyacá, Caquetá, Cauca, Chocó, Cundinamarca, Huila, Meta, Putumayo, Quindío, Risaralda, Santander, Tolima, Valle, Vaupés); Venezuela (Amazonas, Apure, Aragua, Barinas, Bolívar, Carabobo, Venezuela Distrito Federal, Falcon, Lara, Merida, Miranda, Monagas, Portuguesa, Sucre, Tachira, Trujillo, Yaracuy, Zulia); Guyana; Surinam; French Guiana; Ecuador; Peru; Bolivia (Chuquisaca, Cochabamba, La Paz, Santa Cruz); Brazil (Acre, Alagoas, Amazonas, Bahia, Ceará, Brasilia Distrito Federal, Espírito Santo, Goiás, Maranhão, Minas Gerais, Mato Grosso do Sul, Mato Grosso, Pará, Pernambuco, Paraná, Rio de Janeiro, Roraima, Rio Grande do Sul, Santa Catarina, São Paulo, Tocantins); Paraguay (Caazapa); Argentina (Corrientes, Jujuy, Misiones, Salta, Tucuman)
51 nzl 79 MEX 80 BLZ COS ELS GUA HON NIC PAN 81 BBD CUB DOM GDL HIS JAM MAR MTS PUE SAB STL STV TRT 82 CLM FRG GUY SUR VEN 83 BOL ECU PER 84 BRA 85 ARG PAR
Remarks: what was called Serpocaulon giganteum is now Serpocaulon articulatum. Type of Polypodium giganteum belongs here fide Schwartsb. & A. R. Sm. 2013
Synonyms: hidden
= Anaxetum crassifolium (L.) Schott; Gen. Fil. [Schott] t. 1 (1834)
= Campyloneurum crassifolium (L.) Christenh.; Global Fl. 4: 40 (2018)
= Dipteris crassifolia (L.) J.Sm.; London J. Bot. 1: 196 (1842)
= Drynaria crassifolia (L.) J.Sm.; J. Bot. [Hooker] 4: 61 (1841)
= Pessopteris crassifolia (L.) Underw. & Maxon; Contr. U.S. Natl. Herb. 10: 485 (1908)
= Pessopteris crassifolia var. angusta (Fée) Farw.; Amer. Midl. Naturalist 12: 298 (1931)
= Phymatodes acrosora C.Presl; Tent. Pterid. 197 (1836)
= Phymatodes crassifolia (L.) C.Presl; Tent. Pterid. 197 (1836)
= Phymatodes porrecta (Willd.) C.Presl; Tent. Pterid. 197 (1836)
= Pleopeltis crassifolia (L.) T.Moore; Index Fil. [T. Moore] 78 (1857)
= Pleuridium angustum Fée; Mém. Foug. 8 (Icon. Esp. Nouv.): 98 (1857)
= Pleuridium crassifolium (L.) Fée; Mém. Foug. 5 (Gen. Filic.) 274 (1852)
= Polypodium brasiliense f. giganteum (Desv.) Domin; Rozpr. Kral. Ceske Spolecn. Nauk, Tr. Mat.-Prir. no. 2: 131 (1929)
= Polypodium coriaceum Raddi; Opusc. Sci. 3: 286 (1819)
= Polypodium crassifolium L.; Sp. Pl. [Linnaeus] 2: 1083 (1753)
= Polypodium giganteum Desv.; Mém. Soc. Linn. Paris 6(2): 236 (1827)
= Polypodium porrectum Willd.; Sp. Pl., ed. 4 [Willdenow] 5(1): 161 (1810)
= Serpocaulon giganteum (Desv.) A.R.Sm.; Taxon 55(4): 928 (2006)
species :
Niphidium longifolium (Cav.) C.V.Morton & Lellinger; Amer. Fern J. 61: 39 (1971)
Distribution: Colombia (Nariño); Ecuador
82 CLM 83 ECU
Synonyms: hidden
= Campyloneurum americanum (Hook.) Christenh.; Global Fl. 4: 40 (2018)
= Campyloneurum longifolium (Cav.) Christenh.; Global Fl. 4: 40 (2018)
= Cyclophorus americanus (Hook.) C.Chr.; Index Filic. 198 (1905)
= Cyclophorus longifolius (Cav.) C.Chr.; Dansk Bot. Ark. 9(3): 11 (1937), nom. illeg.
= Niphidium americanum (Hook.) J.Sm.; Hist. Fil. 99 (1875)
= Niphobolus americanus (Hook.) Diels; Nat. Pflanzenfam. [Engler & Prantl] I, 4: 325 (1899)
= Polypodium americanum Hook.; Sp. Fil. [W.J.Hooker] 5: 54 (1863)
= Polypodium longifolium Cav.; Descr. Pl. [Cavanilles] 245 (1801)
species :
Niphidium macbridei Lellinger; Amer. Fern J. 62: 117 (1972)
Distribution: Peru
83 PER
Synonyms: hidden
= Campyloneurum macbridei (Lellinger) Christenh.; Global Fl. 4: 40 (2018)
= Pessopteris macbridei (Lellinger) Pic.Serm.; Webbia 31(1): 248 (1977)
species :
Niphidium mortonianum Lellinger; Amer. Fern J. 62: 115 (1972)
Distribution: Colombia (Antioquia, Boyacá, Cauca, Cesar, Cundinamarca, La Guajira, Magdalena, Nariño, Norte de Santander); Venezuela (Anzoategui, Aragua, Venezuela Distrito Federal, Lara, Merida, Monagas, Tachira, Trujillo)
82 CLM VEN
Synonyms: hidden
= Campyloneurum mortonianum (Lellinger) Christenh.; Global Fl. 4: 40 (2018)
= Pessopteris mortoniana (Lellinger) Pic.Serm.; Webbia 31(1): 249 (1977)
species :
Niphidium nidulare (Rosenst.) Lellinger; Amer. Fern J. 62: 112 (1972)
Distribution: Costa Rica; Ecuador
80 COS 83 ECU
Synonyms: hidden
= Campyloneurum nidulare (Rosenst.) Christenh.; Global Fl. 4: 40 (2018)
= Pessopteris nidularis (Rosenst.) Pic.Serm.; Webbia 31(1): 249 (1977)
= Polypodium crassifolium var. nidulare Rosenst.; Repert. Spec. Nov. Regni Veg. 22: 18 (1925)
species :
Niphidium oblanceolatum A.Rojas; Brenesia 45-46: 28 (1997)
Distribution: Guatemala; Belize; ?Honduras; Nicaragua; Costa Rica; Panama; Colombia (Antioquia, Bolívar, Chocó); Ecuador; Peru
80 BLZ COS GUA NIC PAN 82 CLM 83 ECU PER
species :
Niphidium rufosquamatum Lellinger; Amer. Fern J. 62: 115 (1972)
Distribution: Bolivia (Beni, La Paz); Brazil (Paraná, Rio de Janeiro, Rio Grande do Sul, Santa Catarina, São Paulo); Argentina (Misiones)
83 BOL 84 BRA 85 ARG
Synonyms: hidden
= Campyloneurum rufosquamatum (Lellinger) Christenh.; Global Fl. 4: 40 (2018)
= Pessopteris rufosquamata (Lellinger) Pic.Serm.; Webbia 31(1): 249 (1977)
species :
Niphidium vittaria (Mett.) Lellinger; Amer. Fern J. 62: 119 (1972)
Distribution: Peru; Bolivia (La Paz, Santa Cruz)
83 BOL PER
Synonyms: hidden
= Campyloneurum vittaria (Mett.) Christenh.; Global Fl. 4: 40 (2018)
= Pessopteris vittaria (Mett.) Pic.Serm.; Webbia 31(1): 249 (1977)
= Polypodium vittaria Mett.; Fil. Lechl. 1: 8 (1856)
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A complete, synonymic checklist of the Ferns and Lycophytes of the World

Version 25.04 (April 17th, 2025)

Covering 13,178 accepted species,
819 hybrids, 950 accepted infraspecific taxa and 54,489 synonyms,
in total 69,436* names

[*about 1,200 dubious, "unplaced" names are omitted from the main database and listed under "Unplaced names"]

With complete, detailed distribution data (countries and regions) and literature references

Arranged in linear phylogenetic sequence reflecting true relationships

Fully searchable and continously updated

With 4,677 pictures of 1,435 taxa

Assembled through 40 years independently from other databases

In cooperation with Catalogue of Life