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Ver. 24.11
November 5th, 2024

044.0000 Family Thelypteridaceae Ching ex Pic.Serm.; Webbia 24: 711 (1970)
5-34 genera (depending on genus concept), 1100-1200 spp. New genus concept adopted here from Fawcett & Smith (2021), based on molecular results.
Species listed in database: 1180; hybrids: 38
044.2036 Genus Pronephrium C.Presl; Abh. Königl. Böhm. Ges. Wiss., ser. 5, 6: 258 (1851)
Distribution: trop. Asia, few Madagascar, Australia, Pacific
Species listed in database: 40; hybrids: 2
= Pronephrium subgen.Menisciopsis Holttum; Fl. Males. Ser. 2, Pterid. 1(5): 530 (1982)
= Thelypteris subgen.Dimorphopteris K.Iwats.; Mem. Coll. Sci. Univ. Kyoto, Ser. B, Biol. 31: 35 (1964)
= Dimorphopteris Tagawa & K.Iwats.; Acta Phytotax. Geobot. 19: 8 (1961)
= Thelypteris subgen.Haplodictyum K.Iwats.; Mem. Coll. Sci. Univ. Kyoto, Ser. B, Biol. 31: 32 (1964)
= Haplodictyum C.Presl; Abh. Königl. Böhm. Ges. Wiss., ser. 5, 6: 50 (1851)
= Nannothelypteris Holttum; Blumea 19: 38 (1971)

Search for: Pronephrium fidelei

Nothing found!
species :
Pronephrium affine (Blume) C.Presl; Abh. Königl. Böhm. Ges. Wiss., ser. 5, 6: 259 (1851)
Distribution: C-Sumatra; S-Sumatra; W-Java
42 JAW SUM
Synonyms: hidden
= Aspidium affine Blume; Enum. Pl. Javae 2: 148 (1828)
= Cyclosorus affinis (Blume) Mazumdar & Mukhop.; J. Bot. Soc. Bengal 67(1): 44 (2013)
= Dryopteris oxyotis Rosenst.; Meded. Rijks-Herb. no. 31: 5 (1917), nom. nov.
= Gymnogramma macrotis Kunze; Bot. Zeitung (Berlin) 114 (1848) (non Dryopteris macrotis (Hook.) Kuntze)
= Nephrodium auriculare C.Presl; Abh. Königl. Böhm. Ges. Wiss., ser. 5, 6 [Epim. Bot.]: 258 (1851)
= Sagenia menisciicarpa T.Moore; Index Fil. [T. Moore] 96 (1858)
= Thelypteris muminorum Christenh.; Global Fl. 1(4): 34 (2018), nom. nov. for Aspidium affine Blume
species :
Pronephrium amboinense (Willd.) Holttum; Blumea 20: 120 (1972)
Distribution: C-Sulawesi; SE-Sulawesi; Moluccas (Ambon)
42 MOL SUL
Synonyms: hidden
= Aspidium amboinense Willd.; Sp. Pl., ed. 4 [Willdenow] 5(1): 228 (1810)
= Aspidium canescens f. nephrodiiformis Christ; Ann. Jard. Bot. Buitenzorg 15: 131 (1898)
= Cyclosorus amboinensis (Willd.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 142 (1947)
= Dryopteris amboinensis (Willd.) Kuntze; Revis. Gen. Pl. 2: 812 (1891)
= Dryopteris subconformis C.Chr.; Bot. Jahrb. Syst. 66: 47 (1933)
= Nephrodium amboinense (Willd.) C.Presl; Tent. Pterid. 81 (1836)
= Nephrodium molle var. amboinense (Willd.) Bedd.; Ferns Brit. India 278 (1883)
= Pronephrium palopense Holttum; Blumea 20: 116 (1972)
= Thelypteris amboinensis (Willd.) C.F.Reed; Phytologia 17: 259 (1968)
species :
Pronephrium amphitrichum Holttum; Fl. Males. Ser. 2, Pterid. 1(5): 521 (1982)
Distribution: Philippines (Luzon)
42 PHI
Remarks: recently found again
Synonyms: hidden
= Thelypteris amphitricha (Holttum) Christenh.; Global Fl. 1(4): 27 (2018)
species :
Pronephrium aoristisorum (Harr.) S.E.Fawc. & A.R.Sm.; Sida, Bot. Misc. 59: 72 (2021)
Distribution: Philippines (Panay, Luzon)
42 PHI
Synonyms: hidden
= Cyclosorus aoristisorus (Harr.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 142 (1947)
= Dryopteris aoristisora (Harr.) C.Chr.; Index Filic. 252 (1905)
= Nannothelypteris aoristisora (Harr.) Holttum; Blumea 19: 38 (1971)
= Nephrodium aoristisorum (Harr.) Copel.; Polypodiaceae Philipp. 28 (1905)
= Phegopteris aoristisora (Harr.) Alderw.; Malayan Ferns 501 (1908)
= Polypodium aoristisorum Harr.; J. Linn. Soc., Bot. 16: 30 (1877)
= Thelypteris aoristisora (Harr.) C.F.Reed; Phytologia 17: 260 (1968)
species :
Pronephrium aquatiloides (Copel.) Holttum; Blumea 20: 108 (1972)
Distribution: Borneo (W-Sarawak)
42 BOR
Synonyms: hidden
= Cyclosorus aquatiloides (Copel.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 242 (1941)
= Cyclosorus jacobsii Holttum; Blumea 11: 530 (1972)
= Dryopteris aquatiloides Copel.; Philipp. J. Sci. 7: 59 (1912)
= Thelypteris aquatiloides (Copel.) C.F.Reed; Phytologia 17: 260 (1968)
= Thelypteris jacobsii (Holttum) C.F.Reed; Phytologia 17: 285 (1968)
species :
Pronephrium borneense (Hook.) Holttum; Fl. Males. Ser. 2, Pterid. 1(5): 528 (1982)
Distribution: Borneo (Sarawak, Mt. Kinabalu, Sabah)
42 BOR
Synonyms: hidden
= Cyclosorus borneensis (Hook.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 142 (1947)
= Dryopteris labuanensis C.Chr.; Index Filic. 273 (1905), nom. nov., non D. borneensis (Hook.) Kuntze
= Phegopteris borneensis (Hook.) Alderw.; Malayan Ferns 500 (1908)
= Polypodium borneense Hook.; Sp. Fil. [W. J. Hooker] 5: 11 (1863)
= Thelypteris borneensis (Hook.) C.F.Reed; Phytologia 17: 264 (1968)
species :
Pronephrium camarinense (Holttum) S.E.Fawc. & A.R.Sm.; Sida, Bot. Misc. 59: 72 (2021)
Distribution: Philippines (Luzon)
42 PHI
Conservation Status: only known from type
Synonyms: hidden
= Nannothelypteris camarinensis Holttum; Kalikasan 5: 119 (1976)
= Thelypteris camarinensis (Holttum) Christenh.; Global Fl. 1(4): 28 (2018)
species :
Pronephrium celebicum (Baker) Holttum; Blumea 20: 119 (1972)
Distribution: Sulawesi; ?Lesser Sunda Isl. (?Sumba); Moluccas (Seram, Tanimbar); New Guinea (Irian Jaya)
42 MOL NWG SUL
Synonyms: hidden
= Acrostichum celebicum Baker; Bull. Misc. Inform. Kew 1901: 741 (1901)
= Aspidium canescens f. acrostichoides Christ; Ann. Jard. Bot. Buitenzorg 15: 132 (1898)
= Cyclosorus celebicus (Baker) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 142 (1947)
= Dryopteris acrostichoides (Christ) Alderw.; Malayan Ferns Fern Allies, Suppl. 1 (Corr.): 49 (1917), non Kuntze 1891
= Dryopteris celebica (Baker) Copel.; Philipp. J. Sci. 37: 410 (1928)
= Leptochilus celebicus (Baker) C.Chr.; Index Filic. 384 (1905)
= Meniscium hosei var. sumbensis Alderw.; Bull. Dépt. Agric. Indes Néerl. 21: 7 (1908)
= Phegopteris hosei var. sumbensis Alderw.; Bull. Dépt. Agric. Indes Néerl. 21: 7 (1908), nom. altern.
= Pronephrium celebicum var. sopuense Holttum; Blumea 33(2): 402 (1988)
= Thelypteris celebica (Baker) C.F.Reed; Phytologia 17: 267 (1968)
species :
Pronephrium clemensiae (Copel.) Holttum; Blumea 20: 118 (1972)
Distribution: Philippines (Luzon)
42 PHI
Remarks: recently found again
Synonyms: hidden
= Cyclosorus clemensiae (Copel.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 243 (1941)
= Dryopteris clemensiae Copel.; Philipp. J. Sci. 46: 213 (1931)
= Thelypteris clemensiae (Copel.) C.F.Reed; Phytologia 17: 268 (1968)
variant :
var. degener (Christ) Holttum; Fl. Males. Ser. 2, Pterid. 1(5): 522 (1982)
Distribution: Philippines (Luzon)
42 PHI
Synonyms: hidden
= Cyclosorus degener (Christ) Copel.; Fern Fl. Philipp. 1: 356 (1960)
= Dryopteris canescens var. degener Christ; Philipp. J. Sci. 2: 199 (1907)
= Phegopteris canescens var. degener (Christ) Alderw.; Malayan Ferns 507 (1908)
= Thelypteris degener (Christ) C.F.Reed; Phytologia 17: 271 (1968)
species :
Pronephrium euryphyllum (Rosenst.) Holttum; Blumea 20: 112 (1972)
Distribution: Sumatra
42 SUM
Remarks: status and genus uncertain
Conservation Status: only known from 2 sites
Synonyms: hidden
= Cyclosorus euryphyllus (Rosenst.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 142 (1947)
= Dryopteris euryphylla Rosenst.; Meded. Rijks-Herb. no. 31: 7 (1917)
= Dryopteris urophylla var. teysmannii Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 28: 24 (1918)
= Thelypteris euryphylla (Rosenst.) C.F.Reed; Phytologia 17: 276 (1968)
species :
Pronephrium exsculptum (Baker) Holttum; Blumea 20: 117 (1972)
Distribution: peninsular Thailand; Borneo (Sarawak); Brunei
41 THA 42 BOR BRN
Synonyms: hidden
= Acrostichum exsculptum Baker; J. Bot. 26: 326 (1888)
= Cyclosorus exsculptus (Baker) S.Linds.; Edinburgh J. Bot. 66(2): 359 (2009)
= Dryopteris brevipinna C.Chr.; Index Filic. 255 (1905)
= Dryopteris exsculpta (Baker) C.Chr. ex Copel.; Philipp. J. Sci. 37: 410 (1928)
= Leptochilus exsculptus (Baker) C.Chr.; Index Filic. 9 (1905), 385 (1906)
= Meniscium stenophyllum Baker; J. Bot. 29: 108 (1891) (non Nephrodium Sodiro (1883) nec Baker (1884) & (1886))
= Phegopteris stenophylla (Baker) Alderw.; Malayan Ferns 10 (1908)
= Thelypteris exsculpta (Baker) Christenh.; Global Fl. 1(4): 30 (2018), isonym
= Thelypteris exsculpta (Baker) K.Iwats.; Acta Phytotax. Geobot. 21: 170 (1965)
= Thelypteris stenophylla (Baker) C.F.Reed; Phytologia 17: 315 (1968)
species :
Pronephrium fideleae Rakotondr.; Adansonia, sér. 3, 34(2): 226 (2012)
Distribution: Madagascar
29 MDG
Remarks: genus wrong, probably in Pseudocyclosorus-Trigonospora clade
Synonyms: hidden
= Thelypteris fideleae (Rakotondr.) Christenh.; Global Fl. 1(4): 30 (2018)
species :
Pronephrium firmulum (Baker) Holttum; Blumea 20: 116 (1972)
Distribution: Borneo (Sarawak, Mt. Kinabalu, Sabah)
42 BOR
Synonyms: hidden
= Cyclosorus firmulus (Baker) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 142 (1947)
= Dryopteris firmula (Baker) C.Chr.; Index Filic. 266 (1905)
= Phegopteris firmula (Baker) Alderw.; Malayan Ferns 501 (1908)
= Polypodium firmulum Baker; Bull. Misc. Inform. Kew 1893: 211 (1893)
= Thelypteris firmula (Baker) K.Iwats.; Acta Phytotax. Geobot. 21: 170 (1965)
species :
Pronephrium giluwense Holttum; Fl. Males. Ser. 2, Pterid. 1(5): 520 (1982)
Distribution: New Guinea (Papua New Guinea)
42 NWG
Conservation Status: only known from type
Synonyms: hidden
= Thelypteris giluwensis (Holttum) Christenh.; Global Fl. 1(4): 30 (2018)
species :
Pronephrium granulosum (C.Presl) Holttum; Blumea 20: 119 (1972)
Distribution: Palawan; Philippines (Luzon, Balabac, Panay, Mindanao); Lesser Sunda Isl. (Bali, Timor)
42 LSI PHI PLW
Synonyms: hidden
= Cyclosorus chamaeotaria (Christ) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 142 (1947)
= Cyclosorus granulosus (C.Presl) Copel.; Fern Fl. Philipp. 2: 373 (1960)
= Cyclosorus subdimorphus Copel.; Philipp. J. Sci. 81: 38 (1951) (non Thelypteris subdimorphus (Copel.) C. F. Reed)
= Dryopteris chamaeotaria Christ; Philipp. J. Sci. 2: 203 (1907)
= Dryopteris granulosa (C.Presl) C.Chr.; Index Filic. 269 (1906)
= Dryopteris maquilingensis Copel.; Philipp. J. Sci. 56: 103 (1935)
= Goniopteris granulosa (C.Presl) C.Presl; Tent. Pterid. 183 (1836)
= Nephrodium granulosum (C.Presl) J.Sm.; Bot. Voy. Herald [Seemann] 428 (1857)
= Phegopteris chamaeotaria (Christ) Alderw.; Malayan Ferns 505 (1908)
= Phegopteris granulosa (C.Presl) Alderw.; Malayan Ferns 503 (1908) (p. p.)
= Polypodium granulosum C.Presl; Reliq. Haenk. 1: 24, pl. 4, f. 2 (1825)
= Thelypteris chamaeotaria (Christ) C.F.Reed; Phytologia 17: 267 (1968)
= Thelypteris granulosa (C.Presl) C.F.Reed; Phytologia 17: 280 (1968)
= Thelypteris palawanensis C.F.Reed; Phytologia 17: 300 (1968), nom. nov. for Cyclosorus subdimorphus Copel.
species :
Pronephrium hewittii (Copel.) Holttum; Fl. Males. Ser. 2, Pterid. 1(5): 527 (1982)
Distribution: Borneo (W-Sarawak)
42 BOR
Synonyms: hidden
= Cyclosorus compactus (Copel.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 142 (1947)
= Cyclosorus hewittii (Copel.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 245 (1941)
= Dryopteris compacta Copel.; Philipp. J. Sci. 6: 137, pl. 18 (1911)
= Dryopteris hewittii Copel.; Philipp. J. Sci. 3: 344 (1909)
= Thelypteris compacta (Copel.) C.F.Reed; Phytologia 17: 268 (1968)
= Thelypteris hewittii (Copel.) C.F.Reed; Phytologia 17: 282 (1968)
species :
Pronephrium hosei (Baker) Holttum; Blumea 20: 120 (1972)
Distribution: Borneo (Sabah, Sarawak, Kalimantan); Brunei; Philippines (Zamboanga, Basilan)
42 BOR BRN PHI
Synonyms: hidden
= Cyclosorus hosei (Baker) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 142 (1947)
= Dryopteris hosei (Baker) C.Chr.; Index Filic. 271 (1905)
= Meniscium hosei Baker; J. Linn. Soc., Bot. 22: 142 (1886)
= Phegopteris hosei (Baker) Alderw.; Bull. Dépt. Agric. Indes Néerl. 21: 7 (1908)
= Thelypteris hosei (Baker) K.Iwats.; Acta Phytotax. Geobot. 21: 170 (1965)
species :
Pronephrium inaequilobatum (Holttum) S.E.Fawc. & A.R.Sm.; Sida, Bot. Misc. 59: 72 (2021)
Distribution: Philippines (Luzon)
42 PHI
Synonyms: hidden
= Nannothelypteris inaequilobata Holttum; Kalikasan 2: 67 (1973)
= Thelypteris inaequilobata (Holttum) Christenh.; Global Fl. 1(4): 32 (2018)
species :
Pronephrium kjellbergii Holttum; Blumea 20: 117 (1972)
Distribution: C-Sulawesi
42 SUL
Synonyms: hidden
= Pronephrium kjellbergii var. eglandulosum Holttum; Blumea 33(2): 402 (1988)
= Thelypteris kjellbergii (Holttum) Christenh.; Global Fl. 1(4): 32 (2018)
species :
Pronephrium lineatum (Blume) C.Presl; Abh. Königl. Böhm. Ges. Wiss., ser. 5, 6 [Epim. Bot.]: 259 (1851)
Distribution: Borneo (Sabah); Philippines (Tawi Tawi); C-Java; E-Java; Moluccas (Sula Arch.)
42 BOR JAW MOL PHI
Synonyms: hidden
= Abacopteris lineata (Blume) Fée ex Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 257 (1938)
= Alsophila fragilis Zoll. & Moritzi; Natuur- Geneesk. Arch. Ned.-Indië 1: 400 (1844)
= Aspidium lineatum Blume; Enum. Pl. Javae 2: 144 (1828)
= Cyclosorus lineatus (Blume) Tardieu & C.Chr.; Notul. Syst. (Paris) 7: 74 (1938)
= Cyclosorus lineatus var. uniformis Rosenst. ex Tardieu & C.Chr.; Notul. Syst. (Paris) 7: 74 (1938)
= Cyclosorus verruculosus (Alderw.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 250 (1941)
= Dryopteris lineata (Blume) C.Chr.; Index Filic. 276 (1906) (p. p.)
= Dryopteris verruculosa Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 11: 12 (1913)
= Meniscium fragile (Zoll. & Moritzi) Kunze; Bot. Zeitung (Berlin) 6: 259 (1848)
= Meniscium lineatum (Blume) Kunze; Bot. Zeitung (Berlin) 6: 259 (1848)
= Nephrodium lineatum (Blume) C.Presl; Abh. Königl. Böhm. Ges. Wiss., ser. 5, 6: 48 (1851)
= Thelypteris lineata (Blume) K.Iwats.; Mem. Coll. Sci. Univ. Kyoto, Ser. B, Biol. 31: 34 (1964)
= Thelypteris verruculosa (Alderw.) C.F.Reed; Phytologia 17: 323 (1968)
species :
Pronephrium manuselense M.Kato; Acta Phytotax. Geobot. 48(1): 48 (1997)
Distribution: Moluccas (Seram)
42 MOL
Remarks: genus probably incorrect, probably in Menisciopsis
species :
Pronephrium marojejyensis Rakotondr.; Adansonia, sér. 3, 34(2): 230 (2012)
Distribution: Madagascar
29 MDG
Remarks: genus wrong, probably in Pronephrium or Sphaerostephanos
Synonyms: hidden
= Thelypteris marojejyensis (Rakotondr.) Christenh.; Global Fl. 1(4): 33 (2018)
species :
Pronephrium menisciicarpon (Blume) Holttum; Blumea 20: 111 (1972)
Distribution: ?Nicobar Isl.; peninsular Thailand; SW-Thailand; Vietnam; peninsular Malaysia (Kedah, Kelantan, Negeri Sembilan, Pahang, Perak, Selangor, Terengganu); Singapore; Sumatra; Borneo (Mt. Kinabalu, Sabah, Sarawak, Kalimantan); Sulawesi; Philippines (Samar, Leyte); W-Java; Moluccas (Seram); New Guinea (Irian Jaya, Papua New Guinea); Bismarck Arch. (New Britain)
41 THA VIE 42 BOR JAW MLY MOL NWG PHI SNG SUL SUM 43 BIS
Conservation Status: endangered (Singapore)
Synonyms: hidden
= Abacopteris holophylla (Baker) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 241 (1938)
= Abacopteris menisciicarpos (Blume) Holttum; Rev. Fl. Malaya 2: 290 (1955)
= Abacopteris truncata Fée; Mém. Foug. 5 (Gen. Filic.): 310 (1852)
= Aspidium menisciicarpon Blume; Enum. Pl. Javae 2: 142 (1828)
= Cyclosorus holophyllus (Baker) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 142 (1947)
= Cyclosorus latifolius (C.Presl) Copel.; Fern Fl. Philipp. 1: 370 (1960)
= Cyclosorus menisciicarpos (Blume) Holttum; Dansk Bot. Ark. 25: 39 (1967)
= Cyclosorus menisciicarpos (Blume) Panigrahi; Res. J. Pl. Environ. 9: 66 (1993), isonym
= Drynaria menisciicarpos (Blume) J.Sm.; J. Bot. (Hooker) 3: 421 (1841)
= Dryopteris cordifolia Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 11: 19, pl. 5 (1913)
= Dryopteris holophylla C.Chr.; Index Filic. 271 (1905), nom. nov. for Polypodium holophyllum Baker (1888), non Baker (1879)
= Dryopteris korthalsii Rosenst.; Meded. Rijks-Herb. no. 31: 5 (1917)
= Dryopteris menisciicarpos (Blume) Rosenst.; Varenfl. Jav. 61 (1939)
= Dryopteris mirabilis Copel.; Philipp. J. Sci. 6: 137, t. 19 (1911)
= Dryopteris urophylla var. peraspera Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 3, 5: 303 (1922)
= Dryopteris verruculosa var. sumatrana Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 3, 2: 151 (1920)
= Nephrodium latifolium C.Presl; Abh. Königl. Böhm. Ges. Wiss., ser. 5, 6: 45 (1851) (p. p.)
= Phegopteris cordifolia Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 11: 19, t. 5 (1913)
= Phegopteris holophylla (Baker) Alderw.; Malayan Ferns 500 (1909)
= Polypodium holophyllum Baker; J. Bot. 26: 325 (1888), nom. illeg., non Baker 1879
= Polypodium menisciicarpon (Blume) Hook.; Sp. Fil. [W. J. Hooker] 5: 86 (1863) (p. p.)
= Thelypteris latifolia (C.Presl) C.F.Reed; Phytologia 17: 287 (1968)
= Thelypteris menisciicarpos (Blume) K.Iwats.; Acta Phytotax. Geobot. 21: 171 (1965)
species :
Pronephrium merrillii (Christ) Holttum; Blumea 20: 117 (1972)
Distribution: Palawan; ?Philippines (?Balabac)
42 PLW
Synonyms: hidden
= Cyclosorus merrillii (Christ) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 246 (1941)
= Dryopteris merrillii Christ; Philipp. J. Sci. 2: 201 (1907)
= Phegopteris merrillii (Christ) Alderw.; Malayan Ferns Fern Allies 505 (1917)
= Pronephrium balabacense P.M.Zamora & Co; Nat. Appl. Sci. Bull. Univ. Philipp. 32(1-4): 45 (1980)
= Thelypteris balabacensis (P.M.Zamora & Co) Christenh.; Global Fl. 1(4): 27 (2018)
= Thelypteris merrillii (Christ) C.F.Reed; Phytologia 17: 292 (1968)
species :
Pronephrium millarae Holttum; Blumea 20: 115 (1972)
Distribution: Moluccas (Seram); New Guinea (Papua New Guinea, Goodenough Isl.)
42 MOL NWG
Synonyms: hidden
= Thelypteris millarae (Holttum) Christenh.; Global Fl. 1(4): 33 (2018)
species :
Pronephrium minahassae Holttum; Blumea 20: 119 (1972)
Distribution: N-Sulawesi
42 SUL
Conservation Status: only known from type
Synonyms: hidden
= Thelypteris minahassae (Holttum) Christenh.; Global Fl. 1(4): 33 (2018)
species :
Pronephrium moniliforme (Tagawa & K.Iwats.) Holttum; Blumea 19: 36 (1971)
Distribution: Moluccas (Halmaheira, Bacan, Seram)
42 MOL
Synonyms: hidden
= Dimorphopteris moniliformis Tagawa & K.Iwats.; Acta Phytotax. Geobot. 19: 8, f. 14 (1961)
= Thelypteris moniliformis (Tagawa & K.Iwats.) K.Iwats.; Mem. Coll. Sci. Univ. Kyoto, Ser. B, Biol. 31: 36 (1964)
species :
Pronephrium murkelense (M.Kato) S.E.Fawc. & A.R.Sm.; Sida, Bot. Misc. 59: 72 (2021)
Distribution: Moluccas (Seram)
42 MOL
Synonyms: hidden
= Nannothelypteris murkelensis M.Kato; Acta Phytotax. Geobot. 48(1): 50 (1997)
species :
Pronephrium nervosum (Fee) S.E.Fawc. & A.R.Sm.; Sida, Bot. Misc. 59: 72 (2021)
Distribution: Philippines (Mindanao, Samar)
42 PHI
Synonyms: hidden
= Lastrea nervosa (Fée) Copel.; Philipp. J. Sci. 81: 32 (1952), et Fern Fl. Philipp. 2: 323 (1960)
= Nannothelypteris nervosa (Fée) Holttum; Kalikasan 2: 66 (1973)
= Phegopteris nervosa Fée; Mém. Foug. 5 (Gen. Filic.): 244 (1852)
= Polystichum auriculatum var. nervosum Christ; Bull. Herb. Boissier, sér. 1, 6: 192 (1898)
= Thelypteris grima Christenh.; Global Fl. 1(4): 31 (2018), nom. nov. for Phegopteris nervosa Fée
species :
Pronephrium palauense (Hosok.) Holttum; Blumea 20: 121 (1972)
Distribution: Palau (Babeldoap); Fed. States of Micronesia (Yap)
62 MIC PLU
Synonyms: hidden
= Cyclosorus palauensis (Hosok.) Lorence; Micronesica 43(1): 91 (2012)
= Glaphyropteris palauensis (Hosok.) Hosok.; Trans. Nat. Hist. Soc. Formosa 32: 285 (1942), nom. illeg.
= Meniscium palauense Hosok.; Trans. Nat. Hist. Soc. Formosa 28: 148 (1938)
= Thelypteris palauensis (Hosok.) C.F.Reed; Phytologia 17: 300 (1968)
= Thelypteris peliliuensis Fosberg; Smithsonian Contr. Bot. 45: 4 (1980) (nom. nov. for Glaphyropteris palauensis)
species :
Pronephrium peltatum (Alderw.) Holttum; Fl. Males. Ser. 2, Pterid. 1(5): 529 (1982)
Distribution: C-Sumatra; S-Sumatra; W-Java
42 JAW SUM
Synonyms: hidden
= Cyclosorus peltatus (Alderw.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 143 (1947)
= Dryopteris lineata var. subacrostichoides Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 28: 24 (1918)
= Dryopteris peltata Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 16: 12 (1914)
= Dryopteris zippelii Rosenst.; Meded. Rijks-Herb. no. 31: 6 (1917)
= Thelypteris peltata (Alderw.) C.F.Reed; Phytologia 17: 303 (1968)
= Thelypteris peltata (Alderw.) Christenh.; Global Fl. 1(4): 35 (2018), isonym
variant :
var. aberrans Holttum; Fl. Males. Ser. 2, Pterid. 1(5): 530 (1982)
Distribution: Borneo (Mt. Kinabalu, Sabah)
42 BOR
variant :
var. peninsulare Holttum; Fl. Males. Ser. 2, Pterid. 1(5): 530 (1982)
Distribution: peninsular Malaysia (Main Range and Taiping Hills)
42 MLY
variant :
var. persetiferum Holttum; Fl. Males. Ser. 2, Pterid. 1(5): 530 (1982)
Distribution: Borneo (Sabah, Sarawak, Kalimantan); Moluccas (Seram)
42 BOR MOL
variant :
var. tenompokense (C.Chr.) Holttum; Fl. Males. Ser. 2, Pterid. 1(5): 530 (1982)
Distribution: Borneo (Sabah, Mt. Kinabalu)
42 BOR
Synonyms: hidden
= Dryopteris tenompokensis C.Chr.; Gard. Bull. Straits Settlem. 7: 248 (1934)
species :
Pronephrium peramalense Holttum; Blumea 20: 115 (1972)
Distribution: New Guinea (Irian Jaya)
42 NWG
Conservation Status: only known from type
Synonyms: hidden
= Cyclosorus peramalensis (Holttum) Mazumdar & R.Mukhop.; Bionature 33(1): 28 (2013)
= Thelypteris peramelensis (Holttum) Christenh.; Global Fl. 1(4): 35 (2018)
species :
Pronephrium philippinum (C.Presl) S.E.Fawc. & A.R.Sm.; Sida, Bot. Misc. 59: 72 (2021)
Distribution: Philippines (Luzon)
42 PHI
Synonyms: hidden
= Aspidium exiguum Kunze ex Mett.; Abh. Senckenberg. Naturf. Ges. 2(2) [Pheg. u. Asp.]: 360 (1858) (p. p.)
= Dryopteris confusa Copel.; Philipp. J. Sci. 8: 146 (1911)
= Dryopteris exigua (Kunze ex Mett.) Kuntze; Revis. Gen. Pl. 2: 812 (1891)
= Dryopteris philippina (C.Presl) C.Chr.; Index Filic. 284 (1905)
= Haplodictyum exiguum (Kunze ex Mett.) Fée; Mém. Foug. 5 (Gen. Filic.): 309 (1852)
= Lastrea confusa (Copel.) Copel.; Fern Fl. Philipp. 2: 324 (1960)
= Lastrea exigua (Kunze ex Mett.) J.Sm.; J. Bot. (Hooker) 3: 412 (1841), nomen
= Nannothelypteris philippina (C.Presl) Holttum; Kalikasan 2: 66 (1973)
= Nephrodium exiguum (Kunze ex Mett.) Hook.; Sp. Fil. [W. J. Hooker] 4: 92 (1862)
= Physematium philippinum C.Presl; Abh. Königl. Böhm. Ges. Wiss., ser. 5, 6 [Epim. Bot.]: 34 (1851)
= Thelypteris confusa (Copel.) C.F.Reed; Phytologia 17: 268 (1968)
= Thelypteris exigua (Kunze ex Mett.) C.F.Reed; Phytologia 17: 276 (1968)
= Thelypteris philippina (C.Presl) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 253 (1941)
species :
Pronephrium rhombeum (Christ) Holttum; Blumea 20: 120 (1972)
Distribution: Borneo (Sabah); Philippines (Luzon, Mindanao)
42 BOR PHI
Synonyms: hidden
= Cyclosorus lanceola (Christ) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 142 (1947)
= Cyclosorus rhombeus (Christ) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 192 (1938)
= Dryopteris acrostichoides var. rhombea (Christ) Alderw.; Malayan Ferns Fern Allies (1917) Corr. 49
= Dryopteris diversiloba subvar. lanceola Christ; Philipp. J. Sci. 2: 200 (1907)
= Dryopteris diversiloba subvar. rhombea Christ; Philipp. J. Sci. 2: 200 (1907)
= Dryopteris lanceola (Christ) Copel.; Philipp. J. Sci. 56: 102, pl. 7 (1935)
= Dryopteris rhombea (Christ) Copel.; Philipp. J. Sci. 56: 102 (1935)
= Thelypteris lanceola (Christ) C.F.Reed; Phytologia 17: 286 (1968)
= Thelypteris rhombea (Christ) C.F.Reed; Phytologia 17: 309 (1968)
species :
Pronephrium samarense (Copel.) Holttum; Blumea 20: 119 (1972)
Distribution: Philippines (Samar)
42 PHI
Synonyms: hidden
= Cyclosorus samarensis Copel.; Philipp. J. Sci. 81: 35 (1952)
= Thelypteris samarensis (Copel.) C.F.Reed; Phytologia 17: 311 (1968)
species :
Pronephrium simillimum (C.Chr.) Holttum; Blumea 20: 116 (1972)
Distribution: Borneo; SW-Sulawesi; Moluccas (Seram)
42 BOR MOL SUL
Synonyms: hidden
= Cyclosorus simillimus (C.Chr.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 249 (1941)
= Dryopteris simillima C.Chr.; Index Filic. 292 (1905)
= Nephrodium simulans Baker; J. Bot. 26: 325 (1888) (non (1874))
= Thelypteris simillima (C.Chr.) K.Iwats.; Acta Phytotax. Geobot. 21: 169 (1965)
species :
Pronephrium solsonicum Holttum; Kalikasan 3: 197 (1974)
Distribution: Philippines (Luzon)
42 PHI
Conservation Status: only known from type
Synonyms: hidden
= Thelypteris solsonica (Holttum) Christenh.; Global Fl. 1(4): 38 (2018)
species :
Pronephrium thysanoides Holttum; Fl. Males. Ser. 2, Pterid. 1(5): 525 (1982)
Distribution: Borneo (Sarawak, Mt. Mulu)
42 BOR
Conservation Status: only known from type
Synonyms: hidden
= Thelypteris thysanoides (Holttum) Christenh.; Global Fl. 1(4): 39 (2018)
species :
Pronephrium trachyphyllum Holttum; Fl. Males. Ser. 2, Pterid. 1(5): 525 (1982)
Distribution: Philippines (Mindanao)
42 PHI
Conservation Status: only known from type
Synonyms: hidden
= Thelypteris trachyphylla (Holttum) Christenh.; Global Fl. 1(4): 39 (2018)
species :
Pronephrium xiphioides (Christ) Holttum; Blumea 20: 119 (1972)
Distribution: Philippines
42 PHI
Synonyms: hidden
= Cyclosorus edanyoi Copel.; Philipp. J. Sci. 81: 37 (1952)
= Cyclosorus xiphioides (Christ) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 250 (1941)
= Cyclosorus xiphioides (Christ) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 143 (1947), isonym
= Dryopteris xiphioides Christ; Philipp. J. Sci. 2: 201 (1907)
= Phegopteris xiphioides (Christ) Alderw.; Malayan Ferns 501 (1909)
= Thelypteris edanyoi (Copel.) C.F.Reed; Phytologia 17: 274 (1968)
= Thelypteris xiphioides (Christ) C.F.Reed; Phytologia 17: 324 (1968)
species :
Pronephrium ×interruptum Holttum; Blumea 33(2): 403 (1988)
Distribution: Sulawesi
42 SUL
Remarks: Pronephrium kjellbergii x ?
Synonyms: hidden
= Thelypteris ×silmarillion Christenh.; Global Fl. 1(4): 38 (2018), nom. nov.
species :
Pronephrium ×occultum (C.Hope) R.Kr.Singh & V.K.Rawat; J. Biodivers. Conservation 8(4): 40 (2024)
Distribution: India (Uttarakhand)
40 IND
Remarks: a hybrid involving P. penangianum and maybe Glaphyropteridopsis erubescens fide Fraser-Jenkins et al. (2017)
Synonyms: hidden
= Cyclosorus ×occultus (C.Hope) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 228 (1938)
= Dryopteris ×occulta (C.Hope) C.Chr.; Index Filic. 280 (1905)
= Nephrodium ×occultum C.Hope; J. Bombay Nat. Hist. Soc. 12: 627, t. 13 (1899)
= Thelypteris ×occulta (C.Hope) C.F.Reed; Phytologia 17: 297 (1968)
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A complete, synonymic checklist of the Higher Plants of the World


Version 24.11 (November 5th, 2024)

Covering 355,576 accepted species,
8,715 hybrids, 47,863 accepted infraspecific taxa and 1,000,581 synonyms,
in total 1,412,785* names

[*about 25,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus" are deliberately omitted]


With complete, detailed distribution data (countries and regions) and literature references

Arranged in linear phylogenetic sequence reflecting true relationships

Fully searchable and continously updated

With 38,209 pictures of 15,787 species

Assembled through 40 years independently from other databases

In cooperation with Catalogue of Life