034.0000
Family
Dryopteridaceae Herter; Revista Sudamer. Bot. 9: 15 (1949), nom. cons.
26 genera, ca. 2100 spp. Phylogeny: Zuo et al. (2024)
Species listed in database: 2174; hybrids: 201
= Aspidiaceae Burnett; Outlines Bot. 324, 1156 (1835), nom. illeg.
= Aspidiaceae Mett. ex A.B.Frank; Syn. Pflanzenk. [Leunis] ed. 2, 3: 1469 (1877), nom. illeg.
= Bolbitidaceae (Pic.Serm.) Ching; Acta Phytotax. Sin. 16: 15 (1978)
= Elaphoglossaceae Pic.Serm.; Webbia 23: 209 (1968)
= Filicaceae Juss.; Gen. Pl. [Jussieu] 14 (1789), as ´Filices´, nom. illeg., rej.
= Peranemataceae (C.Presl) Ching; Sunyatsenia 5: 246 (1940), nom. rej.
034.7005
Genus
Polystichum Roth; Tent. Fl. Germ. 3: 31, 69 (1799), nom. cons.
Distribution: Nearly worldwide, 55 in trop. America, more than half (224) in China. Many undescribed species remain (Le Pechon et al. 2016), but several species in China will probably still turn out to be synonyms.
Species listed in database: 415; hybrids: 70
= Acropelta Nakai; Bull. Natl. Sci. Mus., Tokyo, 33: 5 (1953)
= Adenoderris J.Sm.; Hist. Fil. 222 (1875)
= Aetopteron Ehrh.; Beitr. Naturk. [Ehrhart] 4: 148 (1789), ex House, Amer. Fern J. 10: 88 (1920)
= Amblia C.Presl; Tent. Pterid. 184 (1836)
= Cyrtogonellum Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 327 (1938)
= Cyrtomidictyum Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 182 (1940)
= Hemesteum H.Lév.; Fl. Kouy-Tchéou 496 (1915) (non Hemestheum Newman (1851))
= Hemigonum J.Sm.; Hist. Fil. 204 (1875)
= Hypopeltis Michx.; Fl. Bor.-Amer. (Michaux) 2: 226 (1803), in obs., ex Bory, Voy. Indes Or. [Bélanger] 2: 63 (1833)
= Papuapteris C.Chr.; Brittonia 2: 300 (1937)
= Plecosorus Fée; Mém. Foug. 5 (Gen. Filic.): 150 (1852)
= Polystichum sect.Sorolepidium (Christ) Tagawa; Acta Phytotax. Geobot. 9: 122 (1940)
= Porpaea C.Presl ms. ex Ettingsh.; Farnkr. Jetztw. [Ettingshausen]: 174 (1864)
= Sorolepidium Christ; Bot. Gaz. 51: 350 (1911)
species :
Polystichum acanthophyllum
(Franch.) Christ; Mém. Soc. Bot. France 1 [Hist. Acrostich.]: 30 (1905)
Distribution: China (Yunnan, W-Sichuan); Tibet; Taiwan
36 CHI TIB 38 TAI
Remarks: subgen. Polystichum, sect. Xiphopolystichum. Not in India
Synonyms: hidden
= Aspidium acanthophyllum Franch.; Bull. Soc. Bot. France 32: 23 (1885)
= Polystichum aculeatum var. acanthophyllum (Franch.) Bedd.; Ferns Brit. India Suppl. 43 (1892)
= Polystichum horridipinnum Hayata; Icon. Pl. Formosan. 4: 195, f. 132 (1914)
= Polystichum spinescens Tagawa; J. Jap. Bot. 14: 102 (1938)
species :
Polystichum acrostichoides
(Michx.) Schott; Gen. Fil. (Vindob.) t. 9 (1834)
Name: Christmas Fern
Distribution: Canada (New Brunswick, Nova Scotia, Ontario, Prince Edward Isl., Québec); USA (Alabama, Arkansas, Connecticut, District of Columbia, Delaware, Florida, Georgia, Iowa, Illinois, Indiana, Kansas, Kentucky, Louisiana, Massachusetts, Maryland, Maine, Michigan, Minnesota, Missouri, Mississippi, North Carolina, New Hampshire, New Jersey, New York, Ohio, Oklahoma, Pennsylvania, Rhode Island, South Carolina, Tennessee, Texas, Virginia, Vermont, Wisconsin, West Virginia); Mexico (Nuevo Leon, Tamaulipas)
72 CDN 74 USA 75 USA 77 USA 78 USA 79 MEX
Synonyms: hidden
= Aetopteron acrostichoides (Michx.) House; Amer. Fern J. 10: 88 (1920)
= Aspidium acrostichoides (Michx.) Sw.; Syn. Fil. [Swartz] 44 (1806)
= Aspidium acrostichoides var. incisum A.Gray; Ann. Lyceum Nat. Hist. New York 3: 238 (1835)
= Aspidium auriculatum Schkuhr; 24. Kl. Linn. Pfl.-Syst. 1: 31 (1806), nom. illeg.
= Aspidium schweinitzii L.C.Beck; Bot. North. Middle States [Beck] 449 (1833)
= Dryopteris acrostichoides (Michx.) Kuntze; Revis. Gen. Pl. 2: 812 (1891)
= Nephrodium acrostichoides Michx.; Fl. Bor.-Amer. (Michaux) 2: 267 (1803)
= Nephrodium acrostichoides var. crispum (Clute) Hopkins; Fern Bull. 15: 7 (1907)
= Nephrodium acrostichoides var. incisum (A.Gray) Hopkins; Fern Bull. 15: 7 (1907)
= Polypodium acrostichoides (Michx.) Poir.; Encycl. [J. Lamarck et al.] 5: 529 (1804), nom. illeg.
= Polystichum acrostichoides f. crispum Clute; Ferns in Their Haunts 317 (1901)
= Polystichum acrostichoides f. cristatum Clute; Fern Bull. 20: 81 (1912)
= Polystichum acrostichoides f. demittens C.F.Reed; Castanea 16: 65, fig. 1 (1951)
= Polystichum acrostichoides f. furcatum F.Seym.; Fl. Vermont, ed. 4: 28 (1969)
= Polystichum acrostichoides f. gymnosorum Neidorf; Amer. Fern J. 39: 103 (1949)
= Polystichum acrostichoides f. incisum (A.Gray) Gilbert; List N. Amer. Pterid.: 19 (1901)
= Polystichum acrostichoides f. lanceolatum Clute; Fern Bull. 20: 24 (1912)
= Polystichum acrostichoides f. ligulatum Gruber; Amer. Fern J. 30: 48 (1940)
= Polystichum acrostichoides f. multifidum Clute; Fern Bull. 15: 122 (1909)
= Polystichum acrostichoides f. orbiculatum Eames; Rhodora 33: 168 (1931)
= Polystichum acrostichoides f. recurvatum Clute; Fern Bull. 12: 79 (1904)
= Polystichum acrostichoides f. spathiforme C.F.Reed; Castanea 16: 69 (1951)
= Polystichum acrostichoides f. ventroperaferens Neidorf; Amer. Fern J. 39: 103, tab. 9 (1949)
= Polystichum acrostichoides var. crispum Clute; Ferns in Their Haunts 107 (1901)
= Polystichum acrostichoides var. incisum A.Gray; Manual (Gray) 632 (1848)
= Polystichum acrostichoides var. schweinitzii (Beck) Small; Bull. Torrey Bot. Club 20: 464 (1893)
= Thelypteris acrostichoides (Michx.) Nieuwl.; Amer. Midl. Naturalist 2: 277 (1912)
variant :
var. lonchitoides
M.G.Brooks; Castanea 12: 24 (1947)
Distribution: USA (West Virginia)
75 USA
species :
Polystichum aculeatum
(L.) Roth; Tent. Fl. Germ. 3: 79 (1799)
Name: Dorniger Schildfarn, Gelappter Schildfarn (DE); Portia (ES)
Distribution: England (England, Wales, Scotland, Outer Hebrides, Orkney Isl., Isle of Man); Ireland (Ireland, Northern Ireland); Denmark; Norway; Sweden; Netherlands; Belgium; Luxembourg; Germany (Brandenburg, +Berlin, Baden-Württemberg, Bayern, Hessen, +Mecklenburg-Vorpommern, Niedersachsen, Nordrhein-Westfalen, Rheinland-Pfalz, Schleswig-Holstein, Saarland, Sachsen, Sachsen-Anhalt, Thüringen); Switzerland; Liechtenstein; Austria; Poland; Czech Republic; Slovakia; Hungary; Spain; Andorra; France; Channel Isl. (Guernsey); Corsica; ?Sardinia; Italy; ?Sicily; San Marino; Slovenia; Croatia; Bosnia & Hercegovina; Montenegro; Serbia; Kosovo; North Macedonia; Albania; Romania; Bulgaria; ?European Turkey; Greece (widespread in mainland); +Estonia; Latvia; Lithuania; Belarus; W-European Russia; Moldova; Ukraine; Crimea; Morocco; Algeria; Canary Isl. (La Gomera, La Palma Isl., Tenerife); Siberia (Altai); Kyrgyzstan; Tajikistan; Northern Caucasus; Georgia [Caucasus]; Armenia; Azerbaijan (incl. Talysh); Turkey (Anatolia); Iraq; Iran; Lebanon; China (Xinjiang)
10 DEN GRB IRE NOR SWE 11 AUT BGM CZE GER HUN LIE LUX NET POL SLO SWI 12 ADO CIS COR FRA SPA 13 ALB BIH BUL CRO GRC ITA KOS MNT NMA ROM SLV SMA SRB 14 BLR EST KRY LAT LIT MLD UKR 20 ALG MOR 21 CNY 30 SIB 32 KGZ TZK 33 ARM AZE GEO NCS 34 IRN IRQ LEB TUR 36 CHI
Synonyms: hidden
= Aetopteron aculeatum (L.) House; Amer. Fern J. 10: 88 (1920)
= Aspidium aculeatum (L.) Sw.; J. Bot. (Schrader) 1800(2): 37 (1801)
= Aspidium aculeatum f. plukenetii (Loisel.) Döll; Fl. Baden [Döll] 1(1): 39 (1855)
= Aspidium aculeatum f. rotundatum Döll; Fl. Baden [Döll] 1(1): 38 (1855)
= Aspidium aculeatum f. simplicius Lindm.; Ark. Bot. 1: 224 (1903)
= Aspidium aculeatum f. umbraticum (Kuntze) Döll; Fl. Baden [Döll] 1(1): 38 (1855)
= Aspidium aculeatum subsp. lobatum (Huds.) Celak.; 10 (1867), isonym
= Aspidium aculeatum subsp. lobatum (Huds.) Milde; Sporenpfl. 63 (1865)
= Aspidium aculeatum subsp. lobatum (Huds.) Niessl; Verh. Naturf. Vereins Brünn 4: 305 (1866), isonym
= Aspidium aculeatum var. angulare A.Braun ex Döll; Rhein. Fl. 21 (1843)
= Aspidium aculeatum var. aristatum Christ; Ber. Schweiz. Bot. Ges. 1: 85 (1891)
= Aspidium aculeatum var. cambricum Gray; Nat. Arr. Brit. Pl. 2: 7 (1821 publ. 1822)
= Aspidium aculeatum var. intermedium Bellynck; Fl. Namur 316 (1855)
= Aspidium aculeatum var. lobatum (Huds.) Hampe; 63 (1836)
= Aspidium aculeatum var. lobatum (Huds.) Warnst.; Syn. Fl. Germ. Helv. 3: 2864 (1907), isonym
= Aspidium aculeatum var. longilobum Milde; Sporenpfl. 64 (1865)
= Aspidium aculeatum var. muscosum Gray; Nat. Arr. Brit. Pl. 2: 7 (1821 publ. 1822)
= Aspidium aculeatum var. plukenetii (Loisel.) Bellynck; Fl. Namur 316 (1855)
= Aspidium aculeatum var. plukenetii (Loisel.) Rouy; Fl. France [Rouy & Foucaud] 14: 418 (1913), isonym
= Aspidium aculeatum var. pseudolonchitis Bellynck; Fl. Namur 316 (1855)
= Aspidium aculeatum var. vulgare Döll; Rhein. Fl. 20 (1843)
= Aspidium angulare subsp. aculeatum (L.) C.Hartm.; 164 (1866), nom. illeg.
= Aspidium bosniacum Formánek; Oesterr. Bot. Z. 38: 243 (1888)
= Aspidium hastulatum subsp. lobatum (Huds.) Simonk.; 607 (1887)
= Aspidium intermedium Sadler; Descr. Pl. Epiph. Hung.: 16 (1820), nom. illeg.
= Aspidium lobatum (Huds.) Sw.; J. Bot. (Schrader) 1800(2): 37 (1801)
= Aspidium lobatum f. abnorme Zapal.; Conspic. Fl. Gallic. Crit. 1: 79 (1905)
= Aspidium lobatum f. plukenetii (Loisel.) Zapal.; Conspic. Fl. Gallic. Crit. 1: 79 (1905)
= Aspidium lobatum var. aristatum Christ; 85 (1891)
= Aspidium lobatum var. longilobum Milde; 186 (1858)
= Aspidium lobatum var. microlobum Milde; 187 (1858)
= Aspidium lobatum var. platylobum Milde; 187 (1858)
= Aspidium lobatum var. plukenetii (Loisel.) F.Hofer; 115 (1902)
= Aspidium lobatum var. ramosum Gray; Nat. Arr. Brit. Pl. 2: 8 (1821 publ. 1822)
= Aspidium lobatum var. rigidum Christ; Bull. Herb. Boissier, sér. 2, 4: 963 (1904)
= Aspidium lobatum var. subtripinnatum Milde; 186 (1858)
= Aspidium lobatum var. umbraticum Kunze; Flora 31: 357 (1848)
= Aspidium munitum Sadler; Fil. Ver. Hung. 34 (1830)
= Aspidium plukenetii Steud.; Nomencl. Bot. [Steudel] 2: 64 (1824)
= Aspidium pseudolonchitis Dumort.; Bull. Soc. Roy. Bot. Belgique 4: 346 (1865)
= Aspidium serrulatum Opiz; Kratos 2(1): 6 (1820)
= Aspidium sublobatum Blume; Enum. Pl. Javae 2: 166 (1828)
= Dryopteris aculeata (L.) Kuntze; Revis. Gen. Pl. 2: 812 (1891)
= Dryopteris aculeata subsp. lobata (Huds.) Briq.; Prodr. Fl. Corse 1: 14 (1910)
= Dryopteris lobata (Huds.) Beck; Oesterr. Bot. Z. 67: 113 (1918), isonym
= Dryopteris lobata (Huds.) Schinz & Thell.; Vierteljahrsschr. Naturf. Ges. Zürich 60: 340 (1915)
= Dryopteris setifera subsp. lobata (Huds.) Maire; Cat. Pl. Maroc [Jahandiez & Maire] 3 (1931)
= Hypopeltis aculeata (L.) Tod.; Giorn. Sci. Nat. Econ. Palermo 1: 238 (1866)
= Hypopeltis lobulata Bory; Exp. Sci. Morée, Bot. 3(2): 286 (1832)
= Nephrodium aculeatum (L.) Coss. & Germ.; Atlas Fl. Paris 673 (1845)
= Phymatodes plukenetii (Loisel.) C.Presl; Tent. Pterid. 196 (1836)
= Polypodium aculeatum L.; Sp. Pl. 2: 1090 (1753)
= Polypodium aculeatum var. lobatum (Huds.) C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 509 (1880)
= Polypodium aculeatum var. minus Lej.; Rev. Fl. Spa: 211 (1825)
= Polypodium appendiculatum Hoffm.; Deutschl. Fl. 2: 8 (1796)
= Polypodium lobatum Huds.; Fl. Angl. (Hudson) 459 (1762)
= Polypodium plukenetii Loisel.; Not. Fl. France: 146 (1810)
= Polystichum aculeatum (L.) Schott; Gen. Fil. (Vindob.) ad t. 9 (1834), isonym
= Polystichum aculeatum subsp. lobatum (Huds.) Vollm.; Fl. Bayern: 8 (1914)
= Polystichum aculeatum var. aristatum (Christ) Askerov; Zametki Sist. Geogr. Rast. 39: 6 (1983)
= Polystichum aculeatum var. aristatum (Christ) Vollm.; Fl. Bayern: 9 (1914)
= Polystichum aculeatum var. corymbiferum T.Moore ex J.Dix; Proc. Roy. Hort. Soc. London 1: 258 (1860)
= Polystichum aculeatum var. plukenetii (Loisel.) Vollm.; Fl. Bayern: 9 (1914)
= Polystichum lobatum (Huds.) Bastard; Essai Fl. Marne et Loire: 367 (1809)
= Polystichum lobatum (Huds.) C.Presl; Tent. Pterid. 83 (1836), isonym
= Polystichum lobatum f. bifidum Seeland; 78 (1938), nom. inval.
= Polystichum lobatum f. multifurcatum Seeland; 78 (1938), nom. inval.
= Polystichum lobatum f. pilosum Domin; 8 (1931)
= Polystichum lobatum var. aculeatum (L.) Syme & N.E.Br.; Engl. Bot., 3rd ed., vol. XII: 93 (1886), nom. illeg.
= Polystichum plukenetii (Loisel.) DC.; Fl. Franç. [Lam. & A. DC.], éd. 3, 6: 241 (1815)
= Polystichum pseudolochnitis Bellynck ex Dumort.; Bull. Soc. Roy. Bot. Belgique 4: 346 (1865)
= Polystichum stipitatum Chauvin; Mém. Soc. Linn. Calvados 2: 502 (1825)
= Tectaria aculeata (L.) Lag., Garcia & Clem.; Anales Ci. Nat. 5: 145 (1802)
species :
Polystichum acutidens
Christ; Bull. Acad. Int. Géogr. Bot. 259 (1902)
Distribution: China (Guangxi, Guizhou, Hubei, Hunan, Sichuan, Yunnan, Zhejiang); Tibet; ?Japan; Taiwan; Nepal; Bhutan; NE-India (Arunachal Pradesh, Manipur, Meghalaya, Nagaland, Sikkim, West Bengal); Myanmar; Thailand; Vietnam
36 CHI TIB 38 TAI 40 BHU IND NEP 41 MYA THA VIE
Remarks: subgen. Haplopolystichum, sect. Haplopolystichum
Synonyms: hidden
= Polypodium mucronatum Roxb.; Calcutta J. Nat. Hist. 4: 490 (1844), non (Sw.) Sw. 1806
= Polystichum deltodon var. acutidens (Christ) C.Chr.; Acta Horti Gothob. 1: 68 (1924)
= Polystichum subauriculatum Tagawa; Acta Phytotax. Geobot. 4: 139 (1935)
species :
Polystichum acutipinnulum
Ching & K.H.Shing; Wuyi Sci. J. 1: 9 (1981)
Distribution: China (Fujian, Guizhou, Henan, W-Hubei, Hunan, Sichuan, Yunnan)
36 CHI
Remarks: subgen. Polystichum, Sect. Hypopeltis
Synonyms: hidden
= Polystichum lobatopinnulum Ching; Boufford & K. H. Shing, J. Arnold Arbor. 64(1): 32 (1983) [not a synonym of P. makinoi fide Flora of China]
= Polystichum wuyishanense Ching & K.H.Shing; Wuyi Sci. J. 1: 9 (1981)
species :
Polystichum adungense
H.S.Kung & Li Bing Zhang ex Ching & Fraser-Jenk.; Acta Phytotax. Sin. 32(4): 370 (1994)
Distribution: China (Yunnan); NE-India (Arunachal Pradesh); N-Myanmar
36 CHI 40 IND 41 MYA
Conservation Status: IUCN CR for India (Fraser-Jenkins 2012)
species :
Polystichum albomarginatum
M.Kessler & A.R.Sm.; Brittonia 57(3): 218 (2005)
Distribution: Ecuador; Bolivia (La Paz)
83 BOL ECU
species :
Polystichum alcicorne
(Baker) Diels; Bot. Jahrb. Syst. 29: 194 (1900)
Distribution: China (Chongqing, NE-Guizhou, Sichuan)
36 CHI
Remarks: subgen. Haplopolystichum, sect. Sphaenopolystichum
Synonyms: hidden
= Aspidium alcicorne (Baker) Christ; Bull. Soc. Bot. Ital. 295 (1901)
= Polypodium alcicorne Baker; J. Bot. 26: 229 (1888)
species :
Polystichum alfaroi
(Christ) Barrington; Rhodora 94: 328 (1992)
Distribution: Mexico (Chiapas); Guatemala; El Salvador; Costa Rica; Panama
79 MEX 80 COS ELS GUA PAN
Remarks: subgen. Polystichum, sect. Polystichum
Synonyms: hidden
= Aspidium aculeatum var. alfaroi Christ; Bull. Herb. Boissier, sér. 2, 4: 963 (1904)
= Polystichum aculeatum var. flavidum Rosenst.; Repert. Spec. Nov. Regni Veg. 22: 9 (1925)
= Polystichum flavidum Rosenst.; Repert. Spec. Nov. Regni Veg. 22: 9 (1925)
species :
Polystichum alluvium
Li Bing Zhang, N.T.Lu & Yi F.Duan; Phytotaxa 323(1): 88 (2017)
Distribution: China (Guizhou)
36 CHI
species :
Polystichum alpinum
Rosenst.; Repert. Spec. Nov. Regni Veg. 12: 171 (1913)
Distribution: Moluccas (Seram); New Guinea (Irian Jaya, Papua New Guinea)
42 MOL NWG
Synonyms: hidden
= Polystichum habbemense Nakaike; Bull. Natl. Sci. Mus., Tokyo, B, 2(2): 51 (1976)
species :
Polystichum altum
Ching ex Li Bing Zhang & H.S.Kung; Acta Phytotax. Sin. 36(5): 465, pl. 1: 6-8 (1998)
Distribution: China (Chongqing, Sichuan, Yunnan)
36 CHI
species :
Polystichum ambiguum
Maxon; Contr. U. S. Natl. Herb. 16: 49, t. 27 (1912)
Distribution: Jamaica
81 JAM
Remarks: known only from type
species :
Polystichum amboroense
M.Kessler & A.R.Sm.; Brittonia 57(3): 224 (2005)
Distribution: Peru; Bolivia (Santa Cruz)
83 BOL PER
species :
Polystichum ammifolium
(Desv.) C.Chr.; Cat. Pl. Madag., Pterid. 31 (1932)
Distribution: Madagascar; Mauritius; La Réunion
29 MAU MDG REU
Remarks: subgen. Polystichum, Sect. Hypopeltis II
Synonyms: hidden
= Aspidium ammifolium (Poir.) Desv.; Mag. Neuesten Entdeck. Gesammten Naturk. Ges. Naturf. Freunde Berlin 5: 321 (1811)
= Aspidium schizolobium (Fée) Kuhn; Filic. Afr. 141 (1868)
= Aspidium stramineum Kaulf. ex Spreng.; Syst. Veg., ed. 16 [Sprengel] 4(1): 105 (1827)
= Aspidium vestitum Sieber; Flora Mauritiana, exsiccatae suppl. n. 48, nom. nud.
= Hypopeltis mauritiana Bory; Voy. Indes Or. [Bélanger] 2: 67 (1833)
= Hypopeltis straminea (Kaulf. ex Spreng.) Bory; Voy. Indes Or. [Bélanger] 2(1): 67 (1833)
= Phegopteris ammifolia (Desv.) Fée; Mém. Foug. 5 (Gen. Filic.): 243 (1852)
= Polypodium ammifolium (Desv.) Poir.; Encycl. [J. Lamarck et al.] 5: 554 (1863)
= Polystichum mauritianum (Bory) Fée; Mém. Foug. 5 (Gen. Filic.): 278 (1852)
= Polystichum schizolobium Fée; Mém. Foug. 8 (Icon. Esp. Nouv.): 99 (1857)
= Polystichum sieberianum C.Presl; Tent. Pterid. 83 (1836), nom. nov. for Aspidium vestitum Sieber, nom. nud.
= Polystichum stramineum (Kaulf. ex Spreng.) C.Presl; Tent. Pterid. 83 (1836)
species :
Polystichum andersonii
Hopkins; Amer. Fern J. 3: 116, pl. 9 (1913)
Name: Anderson´s hollyfern (EN)
Distribution: SE-Alaska; Canada (British Columbia); USA (Idaho, Montana, Oregon, Washington State)
70 ASK 71 CDN 73 USA
Remarks: subgen. Polystichum, basal in HYMASO superclade
Synonyms: hidden
= Polystichum braunii subsp. andersonii (Hopkins) Calder & Roy L.Taylor; Canad. J. Bot. 43: 1388 (1965)
= Polystichum braunii var. andersonii (Hopkins) Hultén; Ark. Bot., n. s. 7(1): 5 (1967 publ. 1968)
= Polystichum jenningsii Hopkins; Ann. Carnegie Mus. 11: 362, t. 37 (1917)
species :
Polystichum andinum
Phil.; Linnaea 29: 108 (1857)
Distribution: Argentina (Chubut, Neuquen, Rio Negro, Santa Cruz, Tierra del Fuego); Chile (Metropolitana de Santiago, O´Higgins, Maule, Ñuble, Biobío, Araucanía, Los Rios, Los Lagos, Aisén, Magallanes)
85 ARG CLE
species :
Polystichum annapurnicola
Fraser-Jenk.; N. Thapa, Pterid. Nepal: 128 (2002)
Distribution: C-Nepal
40 NEP
species :
Polystichum anomalum
(Hook. & Arn.) J.Sm.; Ferns Brit. Foreign 151 (1866)
subspecies :
subsp. anomalum
Distribution: Sri Lanka
40 SRL
Conservation Status: IUCN I (Sri Lanka); EN for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Aspidium anomalum (Hook. & Arn.) Hook.; Sp. Fil. [W. J. Hooker] 4: 27 (1862)
= Dryopteris anomala (Hook. & Arn.) Kuntze; Revis. Gen. Pl. 2: 812 (1891)
= Phegopteris anomala (Hook. & Arn.) Mett. ex Salomon; Nomencl. Gefäßkrypt. 273 (1883)
= Phegopteris eximia Mett.; Kuhn, Linnaea 36: 107 (1869)
= Polypodium anomalum Hook. & Arn.; Hooker´s J. Bot. Kew Gard Misc. 8: 360, t. 11 (1856)
= Polystichum eximium (Mett.) C.Chr.; Bull. Biol. Dept. Sci. Coll. Sun Yatsen Univ. 6: 8 (1933) (fide Fraser-Jenkins 2008)
subspecies :
subsp. travancoricum
(Bedd.) Fraser-Jenk.; Annot. Checkl. Ind. Pterid. 2: 311 (2018)
Distribution: S-India (Kerala, Tamil Nadu)
40 IND
Synonyms: hidden
= Polystichum aculeatum var. travancoricum Bedd.; Ferns Brit. India 209 (1883)
= Polystichum anomalum f. travancoricum (Bedd.) Fraser-Jenk.; Perspect. Pterid. Present & Future (eds. T. N. Bhardwaja & C. B. Gena) 265 (1991)
= Polystichum anomalum var. travancoricum (Bedd.) Sledge; Bull. Brit. Mus. (Nat. Hist.), Bot. 5: 33 (1973)
= Polystichum travancoricum (Bedd.) B.K.Nayar & Kaur; Companion Ferns Brit. India 49 (1974) (fide Fraser-Jenkins 1997)
species :
Polystichum anshunense
Li Bing Zhang; Phytotaxa 404(6): 209 (2019)
Distribution: China (Guizhou)
36 CHI
species :
Polystichum aquifolium
Underw. & Maxon; Bull. Torrey Bot. Club 29: 584 (1902), nom. nov.
Distribution: E-Cuba
81 CUB
Synonyms: hidden
= Aspidium triangulum var. ilicifolium (Fée) Kuhn; Bot. Jahrb. Syst. 24(1): 111 (1897)
= Polystichum ilicifolium Fée; Mém. Foug. 5 (Gen. Filic.): 279 (1852) (non T. Moore (1858))
species :
Polystichum archboldii
Copel.; Univ. Calif. Publ. Bot. 18: 221 (1942)
Distribution: New Guinea (Irian Jaya, Papua New Guinea)
42 NWG
species :
Polystichum articulatipilosum
H.G.Zhou & Hua Li; Guihaia 16(3): 205, f. 3 (1996)
Distribution: China (W-Guangxi); Vietnam
36 CHI 41 VIE
species :
Polystichum asiae-minoris
Tunckol & Li Bing Zhang; Phytotaxa 447(4): 296 (2020)
Distribution: Turkey
34 TUR
species :
Polystichum asperrimum
Li Bing Zhang; Phytotaxa 404(6): 212 (2019)
Distribution: China (Guizhou)
36 CHI
species :
Polystichum assurgentipinnum
W.M.Chu & B.Y.Zhang; Fl. Reipubl. Popularis Sin. [Fl. China Ed. Committee] 5(2): 224 (2001)
Distribution: China (Chongqing)
36 CHI
species :
Polystichum atkinsonii
Bedd.; Ferns Brit. India Suppl. 14, t 362 (1876)
Distribution: China (Chongqing, Guizhou, Hubei, Hunan, Shaanxi, Sichuan, Yunnan); Tibet; Japan; Taiwan; Nepal; Bhutan; India (Arunachal Pradesh, Sikkim, West Bengal); India (incl. Sikkim); Myanmar (Kachin, etc.)
36 CHI TIB 38 JAP TAI 40 BHU IND NEP 41 MYA
Remarks: subgen. Polystichum, Sect. Stenopolystichum
Conservation Status: EN in Japan
Synonyms: hidden
= Aspidium atkinsonii (Bedd.) C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 506 (1880)
= Polypodioides atkinsonii (Bedd.) Ching; (1978)
= Polystichum franchetii Christ; Mém. Soc. Bot. France 1 [Hist. Acrostich.]: 28 (1905)
= Polystichum gracilipes C.Chr.; Bot. Gaz. 66: 338 (1913)
= Polystichum gracilipes var. gemmiferum (Tagawa) Tagawa; Acta Phytotax. Geobot. 16: 75 (1956)
= Polystichum morii Hayata; Icon. Pl. Formosan. 7: 95 (1918) (fide Fraser-Jenkins)
species :
Polystichum auriculum
Ching; Bull. Fan Mem. Inst. Biol. Bot., ser. 2, 1: 309 (1949)
Distribution: China (SE-Yunnan); Vietnam
36 CHI 41 VIE
Remarks: subgen. Haplopolystichum, sect. Sphaenopolystichum
species :
Polystichum australiense
Tindale; S. Austral. Naturalist 28: 31, f. 1 (1954)
Distribution: Australia (Australian Capital Territory, E-New South Wales)
50 AUS
species :
Polystichum australium
Copel.; Occas. Pap. Bernice P. Bishop Mus. 14: 57, t. 8 (1938)
Distribution: Austral Isl. (Tubuai, Raivavae)
61 AUI
species :
Polystichum austropaleaceum
Fraser-Jenk.; Annot. Checkl. Ind. Pterid. 2: 312 (2018)
Distribution: S-India (Tamil Nadu); Sri Lanka
40 IND SRL
species :
Polystichum bachii
M.Kessler & A.R.Sm.; Brittonia 57(3): 214 (2005)
Distribution: Bolivia (Cochabamba)
83 BOL
Conservation Status: Known only from type
species :
Polystichum bakerianum
(Atk. ex C.B.Clarke) Diels; Nat. Pflanzenfam. [Engler & Prantl] I, 4: 191 (1899)
Distribution: China (W-Sichuan, NW-Yunnan); Tibet; Pakistan (Hazara); Nepal; Bhutan; India (Himachal Pradesh, Uttarakhand, Sikkim, Arunachal Pradesh, ?West Bengal, Jammu & Kashmir)
36 CHI TIB 40 BHU IND NEP PAK
Remarks: subgen. Polystichum
Synonyms: hidden
= Aspidium bakerianum (Atk. ex C.B.Clarke) Atk. ex Baker; Hooker´s Icon. Pl. 17: t. 1656 (1886)
= Aspidium prescottianum var. bakerianum Atk. ex C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 510 (1880)
= Polystichum castaneum var. bakerianum (Atk. ex C.B.Clarke) Bedd.; Ferns Brit. India 210 (1883)
= Polystichum prescottianum var. bakerianum (Atk. ex C.B.Clarke) Bedd.; Ferns Brit. India 210 (1883)
species :
Polystichum balansae
Christ; Trudy Imp. S.-Peterburgsk. Bot. Sada 28: 193 (1908)
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, E-Guizhou, Hainan, Hunan, Jiangxi, Zhejiang); Japan; NE-India (Arunachal Pradesh); Vietnam
36 CHI 38 JAP 40 IND 41 VIE
Remarks: subgen. Haplopolystichum, sect. Adenolepia
Conservation Status: IUCN CR for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Cyrtomium balansae (Christ) C.Chr.; Index Filic., Suppl. 2 (1906-1912): 23 (1913)
= Cyrtomium balansae f. edentatum Ching & K.H.Shing; Acta Phytotax. Sin., Addit. 1: 18 (1965)
= Cyrtomium kwantungense Ching; Sinensia 1: 44 (1930)
= Phanerophlebiopsis balansae (Christ) Fraser-Jenk. & Baishya; Bull. Nat. Mus. Nat. Sci., Ser. B, 38(4): 175 (2012)
= Polystichum anomophyllum var. miyajimense (Kodama) Nakai; Bot. Mag. (Tokyo) 39: 115 (1925)
= Polystichum miyasimense Kodama; Icon. Pl. Koisikav. 2: 51 (1914)
species :
Polystichum bamlerianum
Rosenst.; Repert. Spec. Nov. Regni Veg. 10: 330 (1912)
Distribution: New Guinea (Papua New Guinea)
42 NWG
Synonyms: hidden
= Polystichum pullenii Nakaike; Bull. Natl. Sci. Mus., Tokyo, B, 43(3): 129 (1978)
= Polystichum bamlerianum var. ferrugineopaleaceum Rosenst.
species :
Polystichum basipinnatum
(Baker) Diels; Nat. Pflanzenfam. [Engler & Prantl] I, 4: 189 (1899)
Distribution: China (Guangdong, Guangxi, Hongkong)
36 CHI
Synonyms: hidden
= Aspidium basipinnatum Baker; J. Bot. 27: 176 (1889)
= Cyrtomidictyum basipinnatum (Baker) Ching; Acta Phytotax. Sin. 6: 257, 262, t. 51 (1957)
species :
Polystichum biaristatum
(Blume) T.Moore; Index Fil. [T. Moore] 86 (1856)
Distribution: Taiwan; Sri Lanka; Bangladesh; Myanmar; Thailand (widespread); Vietnam; peninsular Malaysia; Sulawesi; Philippines; Java; Lesser Sunda Isl. (Lombok, Sumbawa, Timor)
38 TAI 40 BAN SRL 41 MYA THA VIE 42 JAW LSI MLY PHI SUL
Remarks: subgen. Polystichum, Sect. Macropolystichum
Synonyms: hidden
= Aspidium aculeatum var. biaristatum (Blume) C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 510 (1880)
= Aspidium biaristatum Blume; Enum. Pl. Javae 2: 164 (1828)
= Polystichum aculeatum var. biaristatum (Blume) Bedd.; Ferns Brit. India 209 (1883)
= Polystichum aculeatum var. durissimum Rosenthal; Hedwigia 56: 339 (1915)
= Polystichum aquilinum Copel.; Philipp. J. Sci. 81: 20 (1952)
species :
Polystichum bifidum
Ching; Icon. Filic. Sin. 5: t. 242 (1958)
Distribution: China (SE-Yunnan)
36 CHI
Remarks: subgen. Haplopolystichum, sect. Sphaenopolystichum
species :
Polystichum bigemmatum
Ching ex L.L.Xiang; Chenia 3-4: 133, f. 1-4 (1997)
Distribution: China (SW-Sichuan)
36 CHI
Remarks: possibly a synonym of P. lentum fide Fraser-Jenkins et al. (2015)
species :
Polystichum bissectum
C.Chr.; Index Filic., Suppl. 2 (1906-1912): 64 (1913)
Distribution: China (Chongqing, Sichuan)
36 CHI
Synonyms: hidden
= Polystichum omeiense Christ; Bull. Acad. Int. Géogr., sér. 3, 16: 114 (1906) (non C. Chr. (1905))
species :
Polystichum boboense
Hieron.; Hedwigia 46: 358, t. 8, f. 25, 26 (1906)
Distribution: Ecuador; Peru
83 ECU PER
Remarks: valid species separate from P. orbiculatum fide Moran et al. 2018
Synonyms: hidden
= Polystichum orbiculatum var. boboense (Hieron.) R.M.Tryon; Fieldiana, Bot., n. s. 27: 54 (1991)
= Polystichum boboense var. minus Hieron.; Hedwigia 46: 359, pl. 8, fig. 26 (1907)
species :
Polystichum bolanicum
Rosenst.; Repert. Spec. Nov. Regni Veg. 12: 170 (1913)
Distribution: New Guinea (Papua New Guinea)
42 NWG
Synonyms: hidden
= Polystichum bolanicum var. ovale Rosenst.
= Polystichum bolanicum var. ovalifolium Rosenst.
species :
Polystichum bonapartii
Rosenst.; Repert. Spec. Nov. Regni Veg. 7: 297 (1909)
Distribution: Ecuador
83 ECU
species :
Polystichum bonseyi
W.H.Wagner & R.W.Hobdy; Contr. Univ. Michigan Herb. 19: 75 (1993)
Distribution: Hawaii (Maui, Hawaii Isl.)
63 HAW
Remarks: subgen. Polystichum, Sect. Hypopeltis
species :
Polystichum bradei
Rosenst.; Hedwigia 66: 365 (1915)
Distribution: Brazil (Minas Gerais, Rio de Janeiro)
84 BRA
species :
Polystichum brassii
Copel.; Philipp. J. Sci. 78: 394, t. 2 (1951)
Distribution: New Guinea (Irian Jaya, Papua New Guinea)
42 NWG
species :
Polystichum braunii
(Spenn.) Fée; Mém. Foug. 5 (Gen. Filic.): 278 (1852)
Name: Weicher Schildfarn, Zarter Schildfarn (DE), Braun´s hollyfern (EN)
Distribution: Norway; Sweden; Germany (Baden-Württemberg, Bayern, Hessen, +Rheinland-Pfalz, +Sachsen); Switzerland; ?Liechtenstein; Austria; Poland; Czech Republic; Slovakia; Hungary; France; Italy; Slovenia; Croatia; Montenegro; Serbia; Kosovo; Romania; Bulgaria; Latvia; Belarus; C-European Russia; W-European Russia; Moldova; Ukraine; Siberia (Altai, Krasnoyarsk, W-Siberia); Russian Far East (Kamchatka, Khabarovsk, Kuril Isl., Primorye, Sakhalin); Kazakhstan; Northern Caucasus; Georgia [Caucasus]; Armenia; Azerbaijan; Turkey (Anatolia); Iran; China (Anhui, S-Gansu, Hebei, Heilongjiang, W-Henan, NW-Hubei, Jilin, Liaoning, S-Gansu, Shaanxi, Shanxi, Sichuan, W-Henan, Xinjiang); Tibet; North Korea; South Korea (incl. Cheju Isl.); Japan; Alaska; Canada (British Columbia, Labrador, New Brunswick, Newfoundland, Nova Scotia, Ontario, Prince Edward Isl., Québec); St. Pierre et Miquelon; USA (Connecticut, Idaho, Massachusetts, Maine, Michigan, Minnesota, New Hampshire, New York, Pennsylvania, Vermont, Wisconsin)
10 NOR SWE 11 AUT CZE GER HUN POL SLO SWI 12 FRA 13 BUL CRO ITA KOS MNT ROM SLV SRB 14 BLR LAT MLD RUS UKR 30 SIB 31 RFE 32 KAZ 33 ARM AZE GEO NCS 34 IRN TUR 36 CHI TIB 38 JAP KOR NKO 70 ASK 71 CDN 72 CDN SPM 73 USA 74 USA 75 USA
Remarks: subgen. Polystichum, basal in HYMASO superclade
Synonyms: hidden
= Aetopteron braunii (Spenn.) House; Amer. Fern J. 10: 88 (1920)
= Aspidium aculeatum subsp. braunii (Spenn.) Arcang.; Comp. Fl. Ital. [Arcangeli] 810 (1882), isonym
= Aspidium aculeatum subsp. braunii (Spenn.) Celak.; 697 (1881)
= Aspidium aculeatum var. braunii (Spenn.) Döll; Rhein. Fl. 21 (1843)
= Aspidium aculeatum var. subtripinnatum Milde; Sporenpfl. 65 (1865)
= Aspidium braunii Spenn.; Fl. Friburg. 1: 9 (1825)
= Aspidium braunii var. subtripinnatum (Milde) Borbás; Math. Term. Közlem. 14: 447 (1877)
= Aspidium pilosum Schur; Verh. Mitth. Siebenbürg. Vereins Naturwiss. Hermannstadt 2: 168 (1851)
= Dryopteris aculeata var. braunii (Spenn.) Underw.; Native Ferns, ed. 4: 112 (1893)
= Dryopteris braunii (Spenn.) Underw.; Ill. Fl. N. U.S. (Britton & Brown), ed. 1: 15 (1896)
= Polystichum aculeatum var. braunii (Spenn.) Davenp.; Rhodora 4: 10 (1902)
= Polystichum braunii subsp. purshii (Fernald) Calder & Roy L.Taylor; Canad. J. Bot. 43: 1388 (1965)
= Polystichum braunii var. clarkei Christ; Bull. Herb. Boissier, sér. 1, 7: 25 (1899)
= Polystichum braunii var. purshii Fernald; Rhodora 30: 30 (1928)
= Polystichum shennongense Ching, Boufford & K.H.Shing; J. Arnold Arbor. 64: 34 (1983) [not a synonym of P. makinoi fide Flora of China]
species :
Polystichum bulbiferum
Barrington; Ann. Missouri Bot. Gard. 76(2): 371 (1989)
Distribution: Ecuador
83 ECU
species :
Polystichum calderonense
Proctor; Mem. New York Bot. Gard. 53: 273 (1989)
Distribution: Puerto Rico
81 PUE
Conservation Status: IUCN E (known only from two collections)
species :
Polystichum californicum
(D.C.Eaton) Diels; Nat. Pflanzenfam. [Engler & Prantl] I, 4: 191 (1899)
Name: California Sword Fern
Distribution: C-Canada (British Columbia, Vancouver Isl.); W-USA (Washington, Oregon, California)
71 CDN 73 USA 76 USA
Remarks: subgen. Polystichum, North American clade
Synonyms: hidden
= Aetopteron californicum (D.C.Eaton) House; Amer. Fern J. 10: 88 (1920)
= Aspidium aculeatum var. californicum (D.C.Eaton) D.C.Eaton; Rep. U.S. Geogr. Surv., Wheeler 6(Botany): 336 (1878)
= Aspidium californicum D.C.Eaton; Proc. Amer. Acad. Arts 6: 555 (1865)
= Dryopteris aculeata var. californica (D.C.Eaton) Underw.; Native Ferns, ed. 4 112 (1893)
= Dryopteris californica (D.C.Eaton) Kuntze; Revis. Gen. Pl. 2: 812 (1891)
= Polystichum aculeatum var. californicum (D.C.Eaton) Jeps.; Man. Fl. Pl. Calif. [Jepson] 36 (1923)
species :
Polystichum caruifolium
(Baker) Diels; Bot. Jahrb. Syst. 29: 194 (1900) [carvifolium]
Distribution: China (Chongqing, W-Guangxi, Guizhou, Sichuan, E-Yunnan)
36 CHI
Remarks: subgen. Haplopolystichum, sect. Sphaenopolystichum
Synonyms: hidden
= Acropelta caruifolia (Diels) Pic.Serm.; Webbia 31(1): 254 (1977)
= Aspidium caruifolium Baker; J. Bot. 26: 228 (1888) (non Kunze (1851))
= Polystichum faberi Christ; Notul. Syst. (Paris) 1: 37 (1909), nom. nov. superfl.
= Polystichum omeiense C.Chr.; Index Filic. 67 (1905), 585 (1906), nom. illeg. superfl., not Christ (1906)
species :
Polystichum caudatum
(Sw.) Fée; Mém. Foug. 5 (Gen. Filic.): 278 (1852)
Distribution: Brazil (Minas Gerais)
84 BRA
Synonyms: hidden
= Aspidium caudatum Sw.; Kongl. Vetensk. Acad. Handl. 1817: 62 (1817), nom. illeg., non Sw. 1806
= Aspidium freyreissii Wikstr.; Kongl. Vetensk. Acad. Handl. 1825: 443 (1825)
species :
Polystichum caudescens
Dutra; Anais Reunião Sul-Amer. Bot. 2: 49 (1938 publ. 1939)
Distribution: Brazil (Rio Grande do Sul, São Paulo)
84 BRA
species :
Polystichum cavernicola
Li Bing Zhang & H.He; Bot. Stud. (Taipei) 52(1): 122 (2011)
Distribution: China (S-Guizhou)
36 CHI
species :
Polystichum centronepalense
Fraser-Jenk. & Tamang; Taxon. Revis. Indian Subcontinental Pteridophytes 417 (2008)
Distribution: China (W-Sichuan); C-Nepal
36 CHI 40 NEP
Synonyms: hidden
= Polystichum kangdingense H.S.Kung & Li Bing Zhang; Act. Phytotax. Sin. 36: 254, t. 3, f. 1-4 (1998), nom. inval. without type, et Phytotaxa 60: 58 (2012)
species :
Polystichum chaparense
M.Kessler & A.R.Sm.; Brittonia 57(3): 225 (2005)
Distribution: Bolivia (Cochabamba)
83 BOL
species :
Polystichum cheilanthoides
Copel.; Univ. Calif. Publ. Bot. 18: 221 (1942)
Distribution: New Guinea (Papua New Guinea, Irian Jaya)
42 NWG
species :
Polystichum chilense
(Christ) Diels; Nat. Pflanzenfam. [Engler & Prantl] I, 4: 192 (1899)
Name: Pelomén-lahuén [Chile]
Distribution: Argentina (Chubut, Neuquen, Rio Negro, Santa Cruz, Tierra del Fuego); Chile (Maule, Ñuble, Biobío, Araucanía, Los Rios, Los Lagos, Aisén, Magallanes)
85 ARG CLE
Synonyms: hidden
= Aspidium bridgesii (Schott) J.W.Sturm; Abh. Naturhist. Ges. Nürnberg 2 [Enum. Pl. Vasc. Cr. Chil.]: 181 (1858)
= Aspidium paucicuspe J.W.Sturm; Abh. Naturhist. Ges. Nürnberg 2 [Enum. Pl. Vasc. Cr. Chil.]: ? (1858)
= Polystichum aculeatum var. chilense Christ; Ber. Schweiz. Bot. Ges. 3: 39 (1893)
= Polystichum bridgesii Schott; Gen. Fil. (Vindob.): t. 9 (1834)
variant :
var. dusenii
(C.Chr.) Looser ex R.Rodr.; Gayana, Bot. 44(1-4): 48 (1987 publ. 1988)
Distribution: Argentina (Chubut, Neuquen, Rio Negro, Santa Cruz, Tierra del Fuego); Chile (Los Lagos, Aisén, Magallanes)
85 ARG CLE
Synonyms: hidden
= Polystichum multifidum var. dusenii C.Chr.; Ark. Bot. 10(2): 19 (1910)
species :
Polystichum chingae
Ching; Icon. Filic. Sin. 4: t. 182 (1937)
Distribution: China (S-Yunnan); Vietnam
36 CHI 41 VIE
Remarks: subgen. Polystichum, Sect. Chingiarum
species :
Polystichum christianae
(Jenman) Underw. & Maxon; Maxon, Contr. U. S. Natl. Herb. 18: 30 t, 2 (1909)
Distribution: Jamaica
81 JAM
Synonyms: hidden
= Aspidium christianae Jenman; Bull. Bot. Dept., Jamaica 2: 285 (1895)
species :
Polystichum christii
Ching; Bull. Fan Mem. Inst. Biol. Bot. 2: 192, t. 7 (1931)
Distribution: China (Guizhou, SE-Yunnan); N-Vietnam
36 CHI 41 VIE
Remarks: subgen. Haplopolystichum, sect. Sphaenopolystichum
species :
Polystichum chunii
Ching; Sinensia 1: 2 (1929)
Distribution: China (Guangxi, Guizhou, SW-Hunan); Taiwan; Vietnam
36 CHI 38 TAI 41 VIE
Remarks: subgen. Polystichum, Sect. Macropolystichum. New record for Taiwan in Hsu (2014). Included in P. lentum by Fraser-Jenkins et al. (2018)
species :
Polystichum clarinervium
C.X.Li, N.R.Patel & Li Bing Zhang; Phytotaxa 280(3): 271-277 (2017)
Distribution: China (Sichuan: Emei Shan)
36 CHI
species :
Polystichum concinnum
Lellinger ex Barrington; Ann. Missouri Bot. Gard. 76: 367 (1989)
Distribution: Costa Rica; Panama
80 COS PAN
Remarks: subgen. Polystichum, sect. Polystichum
species :
Polystichum confusum
Li Bing Zhang; Phytotaxa 404(6): 215 (2019)
Distribution: China (Guizhou)
36 CHI
species :
Polystichum congestum
M.Kessler & A.R.Sm.; Brittonia 57(3): 212 (2005)
Distribution: Ecuador; Bolivia (La Paz)
83 BOL ECU
Remarks: to be expected in Peru
species :
Polystichum conjunctum
(Ching) Li Bing Zhang; Phytotaxa 60: 57 (2012)
Distribution: China (Jiangxi)
36 CHI
Synonyms: hidden
= Cyrtomidictyum conjunctum Ching; Acta Phytotax. Sin. 6: 263, t. 52 (1957)
species :
Polystichum consimile
Ching; Icon. Filic. Sin. 6: t. 237 (1958)
Distribution: +China (+Chongqing)
36 CHI
Remarks: a valid species fide Fraser-Jenkins et al. 2018
Conservation Status: probably extinct fide Flora of China
species :
Polystichum copelandii
Christ; Philipp. J. Sci. 2: 157 (1907)
Distribution: Philippines (Luzon)
42 PHI
species :
Polystichum costularisorum
Ching ex W.M.Chu & Z.R.He; Fl. Reipubl. Popularis Sin. [Fl. China Ed. Committee] 5(2): 224 (2001)
Distribution: China (W-Sichuan)
36 CHI
species :
Polystichum craspedosorum
(Maxim.) Diels; Nat. Pflanzenfam. [Engler & Prantl] I, 4: 189 (1899)
Distribution: Russian Far East (Khabarovsk, Primorye); China (Gansu, Guizhou, Hebei, Heilongjiang, Henan, Hubei, Hunan, Jilin, Liaoning, Ningxia, Shaanxi, Shandong, Shanxi, Sichuan, Zhejiang); North Korea; South Korea (incl. Cheju Isl.); Japan
31 RFE 36 CHI 38 JAP KOR NKO
Remarks: subgen. Haplopolystichum, sect. Mastigopteris
Synonyms: hidden
= Aspidium craspedosorum Maxim.; Bull. Acad. Imp. Sci. Saint-Pétersbourg 16: 231 (1871)
= Aspidium craspedosorum var. giraldii Christ; Nuovo Giorn. Bot. Ital., n. s. 4(1): 93 (1897)
= Aspidium craspedosorum var. japonicum Maxim.; Bull. Acad. Imp. Sci. Saint-Pétersbourg 15: 232 (1870)
= Aspidium craspedosorum var. mandshuricum Maxim.; Bull. Acad. Imp. Sci. Saint-Pétersbourg 15: 232 (1870)
= Dryopteris craspedosora (Maxim.) Kuntze; Revis. Gen. Pl. 2: 812 (1891)
= Hemesteum craspedosorum (Maxim.) H.Lév.; Fl. Kouy-Tcheou 497-498 (1915)
= Polystichum leucochlamys Christ; Bot. Gaz. 61: 351 (1911)
= Ptilopteris craspedosora (Maxim.) Hayata; Bot. Mag. (Tokyo) 41: 706 (1927)
species :
Polystichum crassinervium
Ching ex W.M.Chu & Z.R.He; Fl. Reipubl. Popularis Sin. [Fl. China Ed. Committee] 5(2): 224 (2001)
Distribution: China (N-Guangdong, Guangxi, S-Guizhou, S-Hunan)
36 CHI
species :
Polystichum crinigerum
(C.Chr.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 495 (1938)
Distribution: China (S-Yunnan)
36 CHI
Remarks: subgen. Polystichum, Sect. Crinigera
Synonyms: hidden
= Dryopteris crinigera C.Chr.; Index Filic., Suppl. 2 (1906-1912): 31 (1913), nom. nov.
= Polypodium crinitum Baker; Bull. Misc. Inform. Kew 1906: 12 (1906) (non Poir. (1804))
species :
Polystichum crinulosum
(Desv.) J.P.Roux; Novon 6(2): 202 (1996)
Distribution: +Mauritius
29 MAU
Synonyms: hidden
= Aspidium crinulosum Desv.; Mem. Soc. Linn. Paris 6(2): 249 (1827)
species :
Polystichum cuneatiforme
W.M.Chu & Z.R.He; Fl. Reipubl. Popularis Sin. [Fl. China Ed. Committee] 5(2): 177, pl. 50: 9-11 (2001), nom. nov.
Distribution: China (C-Guizhou)
36 CHI
Synonyms: hidden
= Polystichum cuneatum Ching ex P.S.Wang; Guizhou Sci. 12(2): 56.f. 2 (1994), not P. cuneatum Nakaike (1978)
species :
Polystichum cuneatum
Nakaike; Bull. Natl. Sci. Mus., Tokyo, B, 4(3): 128 (1978)
Distribution: New Guinea (Papua New Guinea)
42 NWG
species :
Polystichum cyclolobum
C.Chr.; H. Lév., Cat. Pl. Yun-Nan 111 (1916)
Distribution: China (S-Gansu, Guizhou, W-Sichuan, Yunnan); Tibet; Nepal; Bhutan; NE-India (Arunachal Pradesh, Nagaland, Sikkim, Uttarakhand)
36 CHI TIB 40 BHU IND NEP
Remarks: subgen. Polystichum, sect. Xiphopolystichum
Synonyms: hidden
= Polystichum delavayi (Christ) Ching ex Li Bing Zhang & H.S.Kung; Acta Phytotax. Sin. 36(5): 468 (1998)
= Polystichum ilicifolium var. delavayi Christ; Mém. Soc. Bot. France 1 [Hist. Acrostich.]: 31 (1905)
= Polystichum integrilimbum Ching & H.S.Kung; Acta Bot. Boreal.-Occid. Sin. 9(4): 275 (1989) (fide Fraser-Jenkins et al. 2018)
= Polystichum pseudorhomboideum H.S.Kung & Li Bing Zhang; Acta Phytotax. Sin. 36(5): 468 (1998), nom. nov. superfl. for P. rhomboideum Ching 1974
= Polystichum rhomboideum Ching; Fl. Tsinling. 2: 223 (1974), nom. illeg. (fide Fraser-Jenkins (1997) but kept separate by Flora of China)
= Polystichum stimulans var. delavayi (Christ) H.S.Kung; Acta Bot. Boreal.-Occid. Sin. 9(4): 268 (1989) (fide Fraser-Jenkins 1997)
species :
Polystichum cystostegia
(Hook.) Armstr.; Trans. & Proc. New Zealand Inst. 13: 364 (1881) (cystostigium)
Name: Alpine shield fern
Distribution: New Zealand (North Isl., South Isl., Stewart Isl.); Auckland Isl.; Campbell Isl.
51 AUC NZL 91 CMP
Remarks: subgen. Polystichum, Australasian clade
Synonyms: hidden
= Aspidium cystostegia Hook.; Sp. Fil. [W. J. Hooker] 4: 26, t. 227 (1862)
= Dryopteris cystostegia (Hook.) Kuntze; Revis. Gen. Pl. 2: 812 (1891)
= Polystichum cystostegia (Hook.) Cheeseman; (1925), isonym
species :
Polystichum daguanense
L.L.Xiang ex Ching; Acta Phytotax. Sin. 32(3): 263 (1994)
Distribution: China (NE-Yunnan)
36 CHI
species :
Polystichum dangii
P.S.Wang; Pterid. Fl. Guizhou 531.f. 133: 1-5 (2001)
Distribution: China (S-Guizhou, N-Guangxi); Vietnam
36 CHI 41 VIE
species :
Polystichum decoratum
Maxon; Contr. U. S. Natl. Herb. 13: 20, t. 3 (1909)
subspecies :
subsp. decoratum
Distribution: Cuba; ?Jamaica
81 CUB
subspecies :
subsp. habanense
Morejón & C.Sánchez; Willdenowia 42(2): 274 (2012)
Distribution: Cuba
81 CUB
species :
Polystichum deflexum
Ching ex W.M.Chu; Acta Bot. Yunnan., Suppl. 5: 47 (1992)
Distribution: China (Yunnan)
36 CHI
species :
Polystichum deltatum
Li Bing Zhang, M.Q.Han & Yan Liu; Phytotaxa 365(2): 146 (2018)
Distribution: China (Yunnan)
36 CHI
species :
Polystichum deltodon
(Baker) Diels; Nat. Pflanzenfam. [Engler & Prantl] I, 4: 191 (1899)
Distribution: China (Anhui, Guizhou, Hubei, Hunan, Sichuan, Yunnan, Zhejiang); Japan; Ryukyu Isl.; Myanmar; Vietnam; Philippines
36 CHI 38 JAP NNS 41 MYA VIE 42 PHI
Remarks: subgen. Haplopolystichum, sect. Haplopolystichum
Synonyms: hidden
= Arachniodes dimorphophylla (Hayata) Ching; Acta Bot. Sin. 10: 257 (1962) [fide Flora of China]
= Aspidium deltodon Baker; Gard. Chron., ser. 2, 14: 494 (1810)
= Aspidium tosaense Makino; Bot. Mag. (Tokyo) 6(60): 46 (1892)
= Hemesteum deltodon (Baker) H.Lév.; Fl. Kouy-Tcheou 497-498 (1915)
= Polystichum deltodon var. cultripinnum W.M.Chu & Z.R.He; Fl. Reipubl. Popularis Sin. [Fl. China Ed. Committee] 5(2): 225 (2001)
= Polystichum dimorphophyllum Hayata; J. Coll. Sci. Imp. Univ. Tokyo 30: 428 (1911) [fide Flora of China]
= Polystichum tosaense (Makino) Makino; Bot. Mag. (Tokyo) 23: 144 (1909)
= Rumohra dimorphophylla (Hayata) H.Itô; J. Jap. Bot. 11: 576, f. 5(3) (1935) [fide Flora of China]
species :
Polystichum deminuens
Maxon; Contr. U. S. Natl. Herb. 24: 53, t. 19 (1922)
Distribution: Cuba
81 CUB
Remarks: valid species according to Hernandez & Sánchez 2013, treated as synonym of P. decoratum and as ´P. diminuens´ in Mickel (1997)
species :
Polystichum diaphanum
(Zoll. & Moritzi) T.Moore; Index Fil. [T. Moore] 90 (1858)
Distribution: Java; Lesser Sunda Isl. (Bali, Sumbawa)
42 JAW LSI
Synonyms: hidden
= Aspidium diaphanum Zoll. & Moritzi; Natuur- Geneesk. Arch. Ned.-Indië 2: 203 (1845)
species :
Polystichum dielsii
Christ; Bull. Acad. Int. Géogr., sér. 3, 16: 238 (1906)
Distribution: China (W-Guangxi, Guizhou, SE-Yunnan); N-Vietnam
36 CHI 41 VIE
Synonyms: hidden
= Hemesteum dielsii (Christ) H.Lév.; Fl. Kouy-Tcheou 497-498 (1915)
= Hemesteum pinfaense (Christ) H.Lév.; Fl. Kouy-Tcheou 497-498 (1915)
= Polystichum craspedocarpium Ching & W.M.Chu ex L.L.Xiang; Acta Phytotax. Sin. 32: 259 (1994)
= Polystichum deltodon var. marginale (Christ) C.Chr.; Acta Horti Gothob. 1: 67 (1924)
= Polystichum hecatopterum var. marginale Christ; Bull. Acad. Int. Géogr. Bot. 114 (1904)
= Polystichum pinfaense Christ; Bull. Acad. Int. Géogr., sér. 3, 16: 110 (1906)
species :
Polystichum diffundens
H.S.Kung & Li Bing Zhang; Acta Bot. Yunnan. 17(1): 27 (1995)
Distribution: China (W-Sichuan)
36 CHI
species :
Polystichum discretum
(D.Don) J.Sm.; J. Bot. (Hooker) 3: 413 (1841)
Distribution: China (Yunnan); Tibet (Nyingchi, Medog); Japan; Pakistan (Swat, Hazara); Nepal; Bhutan; India (Jammu & Kashmir, Arunachal Pradesh, ?Assam State, Himachal Pradesh, Jharkhand, Madhya Pradesh, Meghalaya, Mizoram, Nagaland, Sikkim, Uttarakhand, West Bengal, Jammu & Kashmir); Myanmar (Kachin, Chin, etc.); Thailand
36 CHI TIB 38 JAP 40 BHU IND NEP PAK 41 MYA THA
Remarks: subgen. Polystichum, Sect. Hypopeltis III
Synonyms: hidden
= Aspidium aculeatum var. nigropaleaceum Christ; Ber. Schweiz. Bot. Ges. 3: 26, 38 (1893)
= Aspidium aculeatum var. yunnanense Christ; Bull. Herb. Boissier, sér. 1, 6(12): 964 (1898), non A. yunnanense Christ 1898
= Aspidium discretum D.Don; Prodr. Fl. Nepal. 4 (1825)
= Polystichum aculeatum var. angulare Bedd.; Ferns Brit. India 207 (1883) (p. p., non Aspidium angulare Wall.)
= Polystichum aculeatum var. nigropalaceum Christ; Ber. Schweiz. Bot. Ges. 3 (1893)
= Polystichum indicum Khullar & S.C.Gupta; Nova Hedwigia 32: 425 (1980), nom. nud. (fide Fraser-Jenkins 1997)
= Polystichum kathmanduense Nakaike; Y. Otani, Rep. Cryptogamic Study Nepal (Misc. Publ. Nat. Sci. Mus. Tokyo): 139 (1982) (fide Fraser-Jenkins 1997)
= Polystichum lobatum var. discretum (D.Don) C.Chr.; Index Filic. 10: 583 (1906)
= Polystichum nigropalaceum (Christ) Diels; Nat. Pflanzenfam. [Engler & Prantl] I, 4: 191 (1899)
= Polystichum setiferum var. nigropalaceum (Christ) Sledge; Bull. Brit. Mus. (Nat. Hist.), Bot. 5: 37 (1973)
= Polystichum wallichianum C.Presl; Tent. Pterid. 83 (1836), nom. nud.
species :
Polystichum disjunctum
Ching ex W.M.Chu & Z.R.He; Fl. Reipubl. Popularis Sin. [Fl. China Ed. Committee] 5(2): 226, pl. 3: 6-9 (2001)
Distribution: China (W-Guangxi, S-Yunnan); N-Vietnam
36 CHI 41 VIE
species :
Polystichum dissimulans
Maxon; Contr. U. S. Natl. Herb. 13, 31, t. 4 A (1909)
Distribution: Jamaica; Hispaniola (Dominican Republic)
81 HIS JAM
Remarks: recently recollected: Majure et al. 2020. The type is from Jamaica, the Hispaniola population might be a different taxon.
Synonyms: hidden
= Aspidium rhizophyllum var. sublobatum Christ; Bot. Jahrb. Syst. 24: 112 (1897)
species :
Polystichum distans
E.Fourn.; Mexic. Pl. 1: 91 (1872)
Distribution: Mexico (Chiapas, Colima, Guerrero, Hidalgo, Jalisco, Mexico State, Michoacan, Morelos, Oaxaca, Puebla, Queretaro, San Luis Potosi, Veracruz); Guatemala; El Salvador; Honduras; Colombia (Antioquia)
79 MEX 80 ELS GUA HON 82 CLM
Synonyms: hidden
= Polystichum pallidum E.Fourn.; Mexic. Pl. 1: 91 (1872) (non Gardner 1842)
species :
Polystichum dongchuanense
Z.L.Liang, Z.R.He & Li Bing Zhang; Phytotaxa 479(1): 114 (2021)
Distribution: China (Yunnan)
36 CHI
species :
Polystichum dracomontanum
Schelpe & N.C.Anthony; Contr. Bolus Herb. 10: 145 (1982)
Distribution: South Africa (KwaZulu-Natal, ?Free State); Lesotho
27 LES SAF
Remarks: subgen. Polystichum, Afra clade
species :
Polystichum drepanum
(Sw.) C.Presl; Tent. Pterid. 84 (1836)
Distribution: Madeira (Madeira Isl.)
21 MDR
Conservation Status: IUCN E
Synonyms: hidden
= Aspidium drepanum Sw.; J. Bot. (Schrader) 1800(2): 38 (1801)
= Phegopteris drepanum (Sw.) J.Sm.; Cat. Cult. Ferns: 16 (1857)
= Polypodium drepanum (Sw.) Lowe; Trans. Cambridge Philos. Soc. 6: 523 (1838)
species :
Polystichum dubium
(H.Karst.) Diels; Nat. Pflanzenfam. [Engler & Prantl] I, 4: 194 (1899)
Distribution: +Costa Rica; Panama; Colombia (Boyacá, Cundinamarca, Huila, Nariño, Quindío, Santander, Tolima, Valle); Venezuela (Tachira); Ecuador; Peru; Bolivia (Cochabamba, La Paz)
80 COS PAN 82 CLM VEN 83 BOL ECU PER
Synonyms: hidden
= Cyrtomium dubium (H.Karst.) R.M.Tryon & A.F.Tryon; Rhodora 83: 134 (1981)
= Phegopteris dictyophylla Mett.; Kuhn, Linnaea 36: 106 (1869)
= Phegopteris dubia H.Karst.; Fl. Columb. (H. Karst.) 1: 169, t. 84 (1861)
= Polypodium dubium (H.Karst.) Hook.; Sp. Fil. [W. J. Hooker] 5: 15 (1863)
= Polystichum dictyophyllum (Mett.) C.Chr.; Index Filic. 581 (1906)
species :
Polystichum dudleyi
Maxon; J. Wash. Acad. Sci. 8: 620 (1918)
Name: Dudley´s swordfern
Distribution: USA (W-California)
76 USA
Conservation Status: IUCN V
Synonyms: hidden
= Polystichum aculeatum var. dudleyi (Maxon) Jeps.; Man. Fl. Pl. Calif. [Jepson] 36 (1923)
species :
Polystichum duthiei
(C.Hope) C.Chr.; Index Filic. 72 (1905), 581 (1906)
Distribution: China (Gansu, Yunnan); Tibet; Nepal; ?Bhutan; India (Uttarakhand)
36 CHI TIB 40 IND NEP
Remarks: subgen. Polystichum, Sect. Sorolepidium
Conservation Status: IUCN CR for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Aspidium duthiei C.Hope; J. Bombay Nat. Hist. Soc. 12: 532 (1899)
= Sorolepidium duthiei (C.Hope) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 10 (1940)
species :
Polystichum duyunense
Li Bing Zhang, X.Y.Miao & Chun X.Li; Phytotaxa 292(3): 296 (2017)
Distribution: China (Guizhou)
36 CHI
species :
Polystichum echinatum
(J.F.Gmel.) C.Chr.; Index Filic. 83 (1905), 581 (1906)
Distribution: Guatemala; Cuba; Hispaniola; Puerto Rico
80 GUA 81 CUB HIS PUE
Remarks: subgen. Polystichum, sect. Polystichum
Synonyms: hidden
= Aspidium trapezoides Sw.; J. Bot. (Schrader) 1800(2): 31 (1801)
= Aspidium triangulum var. falcatum Kuhn; Bot. Jahrb. Syst. 24(1): 112 (1897)
= Aspidium triangulum var. trapezoides (Sw.) Griseb.; Fl. Brit. W. I. [Grisebach] 689 (1864)
= Aspidium triangulum var. trapezoides (Sw.) Kuhn; Bot. Jahrb. Syst. 24(1): 111 (1897), isonym
= Aspidium viviparum var. trapezoides (Sw.) Jenman; Ferns Brit. W. Ind. 199 (1907)
= Nephrodium sloanei C.Presl; Reliq. Haenk. 1: 33 (1825)
= Polypodium echinatum J.F.Gmel.; Syst. Nat. ed. 13[bis], 2(2): 1309 (1792)
= Polypodium muricatum Sw.; Prodr. Veg. Ind. Occ. 131 (1788) (non L. (1753))
= Polystichum cyphochlamys Fée; Mém. Foug. 5 (Gen. Filic.): 279 (1852)
= Polystichum sloanei C.Presl; Tent. Pterid. 83 (1836)
species :
Polystichum ekmanii
Maxon; Proc. Biol. Soc. Wash. 46: 108 (1933)
Distribution: Hispaniola (Dominican Republic, Haiti)
81 HIS
Remarks: subgen. Polystichum, sect. Polystichum. Treated as bona sp. in Mickel (1997)
species :
Polystichum elevatovenosum
Ching ex W.M.Chu & Z.R.He; Fl. Reipubl. Popularis Sin. [Fl. China Ed. Committee] 5(2): 226 (2001)
Distribution: China (Yunnan)
36 CHI
species :
Polystichum elmeri
Copel.; Philipp. J. Sci. 81: 21 (1952)
Distribution: Philippines (Mindanao)
42 PHI
species :
Polystichum eriorachis
Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 3, 5: 220 (1922)
Distribution: Sumatra
42 SUM
species :
Polystichum erosum
Ching & K.H.Shing; Acta Phytotax. Sin. 10: 303 (1965), nom. nov.
Distribution: China (Chongqing, Guizhou, Henan, Hubei, Hunan, Sichuan, NE-Yunnan); Taiwan
36 CHI 38 TAI
Remarks: subgen. Haplopolystichum, sect. Basigemmifera
Synonyms: hidden
= Polystichum lacerum Christ; Bot. Gaz. 61: 362 (1911) (non C. Presl 1851)
species :
Polystichum erythrosorum
A.R.Sm.; Proc. Calif. Acad. Sci. ser. 4, 40: 225 (1975)
Distribution: Mexico (Chiapas, Oaxaca); Guatemala; Honduras
79 MEX 80 GUA HON
species :
Polystichum exauriforme
H.S.Kung & Li Bing Zhang; Acta Bot. Yunnan. 17(1): 27 (1995)
Distribution: China (Sichuan)
36 CHI
species :
Polystichum excellens
Ching; Icon. Filic. Sin. 5: t. 234 (1958)
Distribution: China (Yunnan, Guizhou, Guangxi, Hunan); N-Vietnam
36 CHI 41 VIE
species :
Polystichum excelsius
Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(4): 16 (1984)
Distribution: China (Chongqing, Hubei, Hunan)
36 CHI
species :
Polystichum falcatilobum
Ching ex W.M.Chu & Z.R.He; Fl. Reipubl. Popularis Sin. [Fl. China Ed. Committee] 5(2): 227 (2001)
Distribution: China (Chongqing, N-Guizhou, S-Sichuan, NE-Yunnan)
36 CHI
Remarks: subgen. Haplopolystichum, sect. Adenolepia
Synonyms: hidden
= Cyrtomium uniseriale f. marginale Ching ex K.H.Shing
species :
Polystichum falcinellum
(Sw.) C.Presl; Tent. Pterid. 83 (1836)
Distribution: Madeira (Madeira Isl.)
21 MDR
Conservation Status: IUCN E
Synonyms: hidden
= Aspidium falcinellum Sw.; Syn. Fil. [Swartz] 46, 243 (1806)
= Dryopteris falcinella (Sw.) Kuntze; Revis. Gen. Pl. 2: 812 (1891)
species :
Polystichum fallax
Tindale; Proc. Linn. Soc. New South Wales 80: 54, t. 2 (1955)
Distribution: Australia (SE-Queensland, E-New South Wales)
50 AUS
Remarks: subgen. Polystichum, Australasian clade
species :
Polystichum faucicola
M.Kessler & Lehnert; Candollea 64(1): 124 (2009)
Distribution: Ecuador
83 ECU
species :
Polystichum fengshanense
Li Bing Zhang & H.He; Syst. Bot. 36(4): 859 (2012)
Distribution: China (N-Guangxi)
36 CHI
species :
Polystichum fibrillosopaleaceum
(Kodama) Tagawa; Acta Phytotax. Geobot. 14: 16 (1949)
Distribution: South Korea; Japan (Honshu, Shikoku)
38 JAP KOR
Synonyms: hidden
= Aetopteron aculeatum var. fibrillosopaleaceum (Kodama) Nakai; Bull. Natl. Sci. Mus., Tokyo, 33: 4 (1953)
= Polystichum aculeatum var. fibrillosopaleaceum Kodama; Bot. Mag. (Tokyo) 29(346): 327 (1915)
= Polystichum polyblepharon var. fibrillosopaleaceum (Kodama) Tagawa; J. Jap. Bot. 13: 186 (1937)
species :
Polystichum filiorum
Mickel; Holttum Mem. [R. J. Johns, ed.] 127, f. 4a (1997)
Distribution: Hispaniola (Dominican Republic)
81 HIS
species :
Polystichum fimbriatum
Christ; Bull. Acad. Int. Géogr., sér. 3, 16: 237 (1906)
Distribution: China (NW-Guangxi, S-Guizhou); N-Vietnam
36 CHI 41 VIE
Remarks: subgen. Polystichum, Sect. Fimbriata
species :
Polystichum flaccidum
Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 3, 6: 220 (1922)
Distribution: Sumatra
42 SUM
species :
Polystichum formosanum
Rosenst.; Hedwigia 66: 338 (1915)
Distribution: S-Japan; Ryukyu Isl.; Taiwan
38 JAP NNS TAI
Remarks: subgen. Haplopolystichum, sect. Adenolepia
Synonyms: hidden
= Polystichum iriomotense Tagawa; Acta Phytotax. Geobot. 4: 137 (1935)
= Polystichum obtusoauriculatum Hayata; Icon. Pl. Formosan. 5: 337 (1915)
species :
Polystichum formosum
Tindale; S. Austral. Naturalist 28: 32 (1954)
Distribution: Australia (SE-Queensland, E-New South Wales, E-Victoria)
50 AUS
species :
Polystichum fournieri
A.R.Sm.; Amer. Fern J. 70(1): 27 (1980), nom. nov.
Distribution: Mexico (Chiapas, Guerrero, Hidalgo, Mexico State, Michoacan, Morelos, Oaxaca, San Luis Potosi, Veracruz); Guatemala; El Salvador; Honduras; Costa Rica; Panama; Venezuela (Lara, Merida)
79 MEX 80 COS ELS GUA HON PAN 82 VEN
Synonyms: hidden
= Aspidium muelleri Mett.; non C. Chr.
= Polystichum muelleri Mett. ex E.Fourn.; Mexic. Pl. 1: 91 (1872) (non Schum. (1803))
species :
Polystichum fraxinellum
(Christ) Diels; Nat. Pflanzenfam. [Engler & Prantl] I, 4: 494 (1899)
Distribution: China (Guangxi, Guizhou, Hunan, Sichuan, Yunnan); Taiwan; N-Vietnam
36 CHI 38 TAI 41 VIE
Remarks: subgen. Haplopolystichum, sect. Cyrtogonellum
Synonyms: hidden
= Aspidium fraxinellum Christ; Bull. Herb. Boissier, sér. 1, 7: 15 (1899)
= Cyrtogonellum fraxinellum (Christ) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 329, t. 7(1) (1938)
= Cyrtogonellum omeiense Ching ex Y.T.Hsieh; Bull. Bot. Res., Harbin 9(3): 15 (1989)
= Cyrtomium fraxinellum (Christ) Christ; Bull. Acad. Int. Géogr. Bot. 264 (1902)
= Phanerophlebia fraxinella (Christ) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 111 (1947)
species :
Polystichum fuentesii
Espinosa; Revista Chilena Hist. Nat. 36: 239 (1932), et 38: 153, f. 38, 39 (1934)
Distribution: +Easter Isl.
61 EAS
Conservation Status: IUCN V
species :
Polystichum fugongense
Ching & W.M.Chu; Acta Bot. Yunnan., Suppl. 5: 49 (1992)
Distribution: China (Yunnan)
36 CHI
Remarks: included in P. longipaleatum by Fraser-Jenkins (1997) but kept separate by Flora of China
species :
Polystichum furfuraceum
A.R.Sm.; Proc. Calif. Acad. Sci. ser. 4, 40: 226 (1975)
Distribution: Mexico (Chiapas); Guatemala
79 MEX 80 GUA
species :
Polystichum fuscopaleaceum
Alston; Bol. Soc. Brot., ser. 2, 30: 22 (1956)
Distribution: Cameroon; Bioko Isl. [Fernando Poo]; D.R.Congo [Zaire]; Ethiopia; Somalia; Uganda; Kenya; Tanzania; Malawi; Zimbabwe; Comores (Grande Comore); La Réunion; N-Yemen
23 BIO CMN ZAI 24 ETH SOM 25 KEN TAN UGA 26 MLW ZIM 29 COM REU 35 YEM
Remarks: subgen. Polystichum, Sect. Hypopeltis. This species has been treated within P. sinense (wilsonii) for a long time. The status remains questionable.
Synonyms: hidden
= Polystichum aculeatum var. mildbraedii Brause; Bot. Jahrb. Syst. 53: 379, 380 (1915)
= Polystichum aculeatum var. rubescens Bonap.; Notes Ptéridol. 14: 214 (1923)
= Polystichum aculeatum var. stenophyllon Bonap.; Notes Ptéridol. 14: 215 (1923)
= Polystichum fuscopaleaceum var. ruwensoriense (Pirotta) Pic.Serm.; Webbia 32(1): 90 (1977)
= Polystichum lobatum var. ruwensoriense Pirotta; Savoia, Il Ruwensori, part. sci. 1: 478 (1909)
= Polystichum setiferum var. fuscopaleaceum (Alston) Schelpe; Bol. Soc. Brot., ser. 2, 41: 216 (1967)
species :
Polystichum fuscum
Copel.; Philipp. J. Sci. 81: 20 (1952)
Distribution: Philippines (Luzon)
42 PHI
species :
Polystichum gejiuense
Li Bing Zhang, M.Q.Han & Yan Liu; Phytotaxa 365(2): 148 (2018)
Distribution: China (Yunnan)
36 CHI
species :
Polystichum gelidum
(Kunze ex Klotzsch) Fée; Mém. Foug. 5 (Gen. Filic.): 278 (1852)
Distribution: Colombia; Venezuela (Lara, Merida); Ecuador; Peru
82 CLM VEN 83 ECU PER
Remarks: similar to P. lehmannii but kept separate by McHenry & Barrington (2014)
Synonyms: hidden
= Aspidium gelidum Kunze ex Klotzsch; Linnaea 20: 365 (1847)
species :
Polystichum glaciale
Christ; Mém. Soc. Bot. France 1 [Hist. Acrostich.]: 28 (1905)
Distribution: China (Gansu, Qinghai, Sichuan, Yunnan); Tibet; Taiwan; Nepal; Bhutan; India (Sikkim)
36 CHI TIB 38 TAI 40 BHU IND NEP
Remarks: subgen. Polystichum, Sect. Sorolepidium
Conservation Status: IUCN CR for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Polystichum duthiei var. subbipinnatum Tagawa; Acta Phytotax. Geobot. 9(3): 131 (1940)
= Sorolepidium glaciale (Christ) Christ; Bot. Gaz. 61: 350, f. 1 (1911)
= Sorolepidium ovale Y.T.Hsieh; Bull. Bot. Res., Harbin 11(2): 17 (1991)
species :
Polystichum glandulosum
C.Presl; Tent. Pterid. 82 (1836)
Distribution: Cuba; Jamaica
81 CUB JAM
Remarks: correct generic placement fide McHenry et al. 2013
Synonyms: hidden
= Adenoderris glandulosa (C.Presl) J.Sm.; Hist. Fil. 223 (1875)
= Adenoderris viscidula (Mett.) Maxon; Bot. Gaz. 39: 367, f. 1 (1905)
= Aspidium glandulosum (C.Presl) Hook. & Grev.; Icon. Filic. [W. J. Hooker & Greville] t. 140 (1829) (non Blume (1828))
= Aspidium viscidulum Mett.; Abh. Senckenberg. Naturf. Ges. 2(2) [Pheg. u. Asp.]: 322 (1858)
= Dryopteris viscidula (Mett.) Kuntze; Revis. Gen. Pl. 2: 811 (1891)
= Polystichum viscidulum (Mett.) C.Chr.; Index Filic. 75 (1905)
species :
Polystichum gonggashanense
H.S.Kung & Li Bing Zhang ex H.He & Li Bing Zhang; Phytotaxa 559(2): 207 (2022)
Distribution: China (Sichuan)
36 CHI
species :
Polystichum grandifrons
C.Chr.; Index Filic., Suppl. Tert.: 163 (1934)
Distribution: China (Guangxi, Guizhou, S-Yunnan); Japan (Kyushu); Taiwan; NE-India (Manipur, Nagaland); Vietnam; Philippines
36 CHI 38 JAP TAI 40 IND 41 VIE 42 PHI
Remarks: subgen. Polystichum, Sect. Neopolystichum
Conservation Status: IUCN CR for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Polystichum grande Ching; Bull. Fan Mem. Inst. Biol. Bot. 2: 189, t. 5 (1931) (non Fée (1857))
= Polystichum kiusiuense Tagawa; Acta Phytotax. Geobot. 16: 15 (1953)
species :
Polystichum guadalupense
Fée; Mém. Foug. 11 (Hist. Foug. Antil.): 74, t. 19, f.2 (1866)
Distribution: Guadeloupe
81 GDL
species :
Polystichum guajaibonense
Morejón & C.Sánchez; Willdenowia 42(2): 276 (2012)
Distribution: Cuba
81 CUB
species :
Polystichum guangxiense
W.M.Chu & H.G.Zhou; Guihaia 16(3): 205, f. 2 (1996)
Distribution: China (W-Guangxi)
36 CHI
species :
Polystichum gymnocarpium
Ching ex W.M.Chu & Z.R.He; Fl. Reipubl. Popularis Sin. [Fl. China Ed. Committee] 5(2): 227 (2001)
Distribution: China (N-Fujian, SW-Zhejiang)
36 CHI
species :
Polystichum hagiangense
N.T.Lu, Liang Zhang & L.-B.Zhang; Phytotaxa 175(5): 293-297 (2014)
Distribution: N-Vietnam
41 VIE
species :
Polystichum hainanicola
Li Bing Zhang, Liang Zhang & X.F.Gao; Phytotaxa 85(1): 10 (2013)
Distribution: China (Hainan)
36 CHI
species :
Polystichum haleakalense
Brack.; U.S. Expl. Exped., Filic. 16: 204, t. 28 (1854)
Distribution: Hawaii (Maui, Hawaii Isl.)
63 HAW
Remarks: subgen. Polystichum, Sect. Hypopeltis
Conservation Status: IUCN R
species :
Polystichum hancockii
(Hance) Diels; Nat. Pflanzenfam. [Engler & Prantl] I, 4: 191 (1899)
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Hunan, Jiangxi, Zhejiang); Ryukyu Isl.; Taiwan
36 CHI 38 NNS TAI
Remarks: subgen. Haplopolystichum, sect. Crucifilix
Synonyms: hidden
= Aspidium hancockii (Hance) Baker; Ann. Bot. (Oxford) 5: 314 (1891)
= Aspidium reductum Baker; J. Bot. 26: 105 (1888)
= Polystichum hancockii var. leptopteron (Hayata) Suzuki; Rep. (Annual) Taihoku Bot. Gard. 1: 111 (1931)
= Polystichum hancockii var. proliferum Seriz.; Shidekobushi 1(2): 92 (2009)
= Polystichum leptopteron Hayata; Icon. Pl. Formosan. 6: 340, f. 137 a, 141 (1915)
= Polystichum longistipes Hayata; Icon. Pl. Formosan. 6: 341, f. 137 b, 142 (1915)
= Polystichum parahancockii Ching; Bull. Bot. Res., Harbin 2(2): 68 (1982)
= Polystichum simplicipinnum Hayata; Icon. Pl. Formosan. 5: 343, f. 137 j. 146 (1915)
= Ptilopteris hancockii Hance; J. Bot. 22: 139 (1884)
species :
Polystichum hanmengqii
Li Bing Zhang & Yan Liu; Phytotaxa 365(2): 151 (2018)
Distribution: China (Guangdong)
36 CHI
species :
Polystichum harpophyllum
(Zenker ex Kunze) Sledge; J. Linn. Soc., Bot. 84(1): 7 (1982)
Distribution: S-India (Kerala, Tamil Nadu); Sri Lanka
40 IND SRL
Synonyms: hidden
= Aspidium auriculatum var. normale C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 507 (1880)
= Phegopteris nitens Fée; Mém. Foug. 5 (Gen. Filic.): 246 (1852)
= Polypodium harpophyllum Zenker ex Kunze; Linnaea 24: 256 (1851)
= Polystichum nitens (Fée) Fée; Mém. Foug. 8 (Icon. Esp. Nouv.): 127 (1857)
species :
Polystichum harrisii
Maxon; Contr. U. S. Natl. Herb. 13: 32, t. 4 B (1909)
Distribution: Jamaica
81 JAM
Synonyms: hidden
= Aspidium caudatum Jenman; J. Bot. 17: 260 (1879) (non Sw. (1806))
= Aspidium triangulum subsp. caudatum (Jenman) Jenman; Ferns Brit. W. Ind. 198 (1907)
= Aspidium viviparum subsp. caudatum (Jenman) Jenman; Bull. Bot. Dept., Jamaica 2: 268 (1895)
= Aspidium viviparum var. caudatum Jenman; Bull. Bot. Dept., Jamaica 2: 263 (1895)
species :
Polystichum hartwegii
(Klotzsch) Hieron.; Hedwigia 46: 355 (1906)
Distribution: Mexico (Chiapas, Guerrero, Hidalgo, Jalisco, Mexico State, Morelos, Nuevo Leon, Oaxaca, Puebla, Veracruz); Revillagigedos Isl. (Isla Socorro); Guatemala; El Salvador; Honduras; Nicaragua; Costa Rica; Panama; Colombia; Ecuador; Bolivia (La Paz)
79 MEX RGI 80 COS ELS GUA HON NIC PAN 82 CLM 83 BOL ECU
Synonyms: hidden
= Aspidium crenatum Mett.; Fil. Hort. Bot. Lips. 89 (1856), nom. illeg.
= Aspidium hartwegii Klotzsch; Linnaea 20: 366 (1847), as ´hartwigii´
= Aspidium lechleri Mett.; Sp. Fil. [W. J. Hooker] 4: 21 (1862)
= Aspidium trejoi Christ; Bull. Herb. Boissier, sér. 2, 5: 255 (1905)
= Polystichum drepanoides E.Fourn.; Mexic. Pl. 1: 92 (1872)
= Polystichum grande Fée; Mém. Foug. 8 (Icon. Esp. Nouv.): 98 (1857)
= Polystichum hartwigii (Klotzsch) Hieron.; Hedwigia 46: 365 (1906)
= Polystichum trejoi (Christ) Christ; Bull. Herb. Boissier, sér. 2, 5: 728 (1905)
species :
Polystichum hastipinnum
G.D.Tang & Li Bing Zhang; Phytotaxa 309(1): 66 (2017)
Distribution: China (Guangdong)
36 CHI
species :
Polystichum hecatopterum
Diels; Bot. Jahrb. Syst. 29: 193 (1900)
Distribution: China (Guangxi, Guizhou, Hubei, Hunan, Jiangxi, Sichuan, Yunnan, Zhejiang); Tibet; Taiwan; NE-India (Arunachal Pradesh)
36 CHI TIB 38 TAI 40 IND
Remarks: subgen. Polystichum, Sect. Hecatoptera
Conservation Status: IUCN CR for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Aspidium auriculatum var. stenophyllum Baker; J. Bot. 26(7): 227 (1888)
= Hemesteum hecatopterum (Diels) H.Lév.; Fl. Kouy-Tcheou 497-498 (1915)
= Polystichum auriculatum var. stenophyllum (Baker) C.G.Matthew; J. Linn. Soc., Bot. 39: 385 (1911)
= Polystichum pseudomaximowiczii Hayata; Icon. Pl. Formosan. 6: 334, f. 137 f-h., 139 (1915)
species :
Polystichum herbaceum
Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(4): 20 (1984)
Distribution: China (Chongqing, Guizhou, W-Hunan); Taiwan
36 CHI 38 TAI
Remarks: subgen. Polystichum, sect. Xiphopolystichum. Included by Fraser-Jenkins et al. (2018) in P. luctuosum
species :
Polystichum heterolepis
Fée; Mém. Foug. 5 (Gen. Filic.): 279 (1852)
Distribution: Cuba
81 CUB
Synonyms: hidden
= Aspidium viviparum (Fée) Mett.; Abh. Senckenberg. Naturf. Ges. 2(2) [Pheg. u. Asp.]: 328 (1858)
= Dryopteris vivipara (Fée) Kuntze; Revis. Gen. Pl. 2: 814 (1891)
= Polystichum viviparum Fée; Mém. Foug. 5 (Gen. Filic.): 280 (1852)
species :
Polystichum hillebrandii
Carruth.; Fl. Vit. [Seemann] 358 (1873)
Distribution: Hawaii (Maui, Hawaii Isl.)
63 HAW
Remarks: subgen. Polystichum, sect. Xiphopolystichum
Conservation Status: IUCN R
Synonyms: hidden
= Aspidium hillebrandii Hillebr.; Fl. Hawaiian Isl. [Hillebrand] 568 (1888)
species :
Polystichum holttumii
C.Chr.; Gard. Bull. Straits Settlem. 7: 256, t. 53 (1934)
Distribution: Borneo (Mt. Kinabalu, Sabah)
42 BOR
species :
Polystichum hookerianum
(C.Presl) C.Chr.; Index Filic. 67 (1905), 582 (1906)
Distribution: China (Guangxi, Guizhou, Hunan, Sichuan, Yunnan); Tibet; Japan; Nepal; Bhutan; India (Arunachal Pradesh, Manipur, Meghalaya, Nagaland, Sikkim, West Bengal); Myanmar (Kachin, etc.); N-Thailand; Vietnam
36 CHI TIB 38 JAP 40 BHU IND NEP 41 MYA THA VIE
Remarks: subgen. Haplopolystichum, sect. Adenolepia
Synonyms: hidden
= Aspidium caducum Wall. ex Hook. & Grev.; Icon. Filic. 2: t. 171 (1830), nom. illeg. (non Kunth (1815))
= Cyrtomium caducum (Wall. ex Hook. & Grev.) T.Moore; Index Fil. [T. Moore] 276 (1861), nom. nov. superfl.
= Cyrtomium hookerianum (C.Presl) C.Chr.; Index Filic., Suppl. 2 (1906-1912): 101 (1913)
= Cyrtomium tachiroanum (Luerss.) C.Chr.; Amer. Fern J. 20: 45 (1930)
= Dryopteris caduca (Wall. ex Hook. & Grev.) Kuntze; Revis. Gen. Pl. 2: 812 (1891)
= Lastrea hookeriana C.Presl; Tent. Pterid. 77 (1836), nom. nov. for A. caducum Wall.
= Phanerophlebia hookeriana (C.Presl) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 111 (1947)
= Phanerophlebia tachiroana (Luerss.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 111 (1947)
= Phanerophlebiopsis caduca (T.Moore) Fraser-Jenk.; New Species Syndr. Indian Pteridol. 190 (1997), nom. inval. (name rejected by the same author in the same publication)
= Phanerophlebiopsis hookeriana (C.Presl) Fraser-Jenk.; Himalayan Ferns (Guide to Polystichum) 42 (1997)
= Phanerophlebiopsis tachiroana (Luerss.) Fraser-Jenk.; Perspect. Pterid. Present & Future (eds. T. N. Bhardwaja & C. B. Gena) 280 (1991), et in Himalayan Ferns (Guide to Polystichum) 43 (1997), nom. inval.
= Polypodium tachiroanum Luerss.; Bot. Jahrb. Syst. 4: 362 (1883)
= Polystichum caducum (T.Moore) Diels; Nat. Pflanzenfam. [Engler & Prantl] I, 4: 191 (1900), nom. illeg.
= Polystichum tachiroanum (Luerss.) Tagawa; Mayebara, Fl. Austro-Higoensis 9 (1931)
species :
Polystichum houchangense
Ching ex P.S.Wang; Guizhou Sci. 12(2): 55, f. 1 (1994)
Distribution: China (C-Guizhou)
36 CHI
species :
Polystichum huashanicola
(W.M.Chu & Z.R.He) Li Bing Zhang; Phytotaxa 60: 58 (2012)
Distribution: China (NE-Yunnan)
36 CHI
Synonyms: hidden
= Polystichum daguanense var. huashanicola W.M.Chu & Z.R.He; Acta Bot. Yunnan. 23: 257 (2000) (huashanicolum)
species :
Polystichum hubeiense
Liang Zhang & Li Bing Zhang; Ann. Bot. Fenn. 50(1-2): 107 (2013)
Distribution: China (Hubei)
36 CHI
Remarks: subgen. Haplopolystichum, sect. Platylepia
species :
Polystichum ichangense
Christ; Mém. Soc. Bot. France 1 [Hist. Acrostich.]: 28 (1905)
Distribution: China (Chongqing, N-Guizhou, W-Hubei, N-Hunan)
36 CHI
species :
Polystichum igaense
Tagawa; Acta Phytotax. Geobot. 15: 15 (1953)
Distribution: Japan
38 JAP
Remarks: subgen. Polystichum, Sect. Hypopeltis III
species :
Polystichum imbricans
(D.C.Eaton) D.H.Wagner; Pteridologia 1: 50 (1979)
subspecies :
subsp. imbricans
Name: Narrowleaf swordfern
Distribution: C-Canada (S-British Columbia); W-USA (Washington, Oregon, California)
71 CDN 73 USA 76 USA
Remarks: subgen. Polystichum, North American clade
Synonyms: hidden
= Aspidium munitum var. imbricans D.C.Eaton; Ferns N. Amer. 1: 188 (1878)
= Polystichum munitum f. imbricans (D.C.Eaton) Clute ex M.Broun; Index No. Amer. Ferns: 149 (1938)
= Polystichum munitum subsp. imbricans (D.C.Eaton) Munz; Fl. S. Calif. (Munz) 21 (1974)
= Polystichum munitum var. imbricans (D.C.Eaton) Maxon; Fern Bull. 8: 30 (1900)
= Polystichum munitum var. nudatum (D.C.Eaton) Gilbert; List N. Amer. Pterid.: 20 (1901)
subspecies :
subsp. curtum
(Ewan) D.H.Wagner; Pteridologia 1: 51 (1979)
Distribution: USA (California)
76 USA
Synonyms: hidden
= Polystichum munitum subsp. curtum Ewan; Amer. Fern J. 32: 99 (1942)
species :
Polystichum incisopinnulum
H.S.Kung & Li Bing Zhang; Acta Bot. Yunnan. 17(1): 25 (1995)
Distribution: China (W-Sichuan)
36 CHI
species :
Polystichum incongruum
J.P.Roux; Bot. J. Linn. Soc. 125(1): 36 (1997)
Distribution: South Africa (W-Cape Prov., E-Cape Prov.)
27 SAF
Remarks: subgen. Polystichum, Afra clade
species :
Polystichum integrilobum
(Ching ex Y.T.Hsieh) W.M.Chu ex H.S.Kung; Acta Phytotax. Sin. 36(3): 244 (1998)
Distribution: China (NW-Yunnan)
36 CHI
Synonyms: hidden
= Sorolepidium integrilobum Ching ex Y.T.Hsieh; Bull. Bot. Res., Harbin 9(3): 48 (1989)
species :
Polystichum integripinnum
Hayata; Icon. Pl. Formosan. 4: 196, f. 133 (1914)
Distribution: Taiwan
38 TAI
Remarks: included in P. hookerianum by Flora of China but kept separate by Knapp (Ferns of Taiwan, 2011 ff.)
Synonyms: hidden
= Aspidium integripinnum Hayata; Icon. Pl. Formosan. 4: 196 (1914)
= Cyrtomium integripinnum (Hayata) Copel.; Philipp. J. Sci. 38: 136 (1929)
species :
Polystichum jamunae
Fraser-Jenk.; Aspects Pl. Sci. 13: 277 (1991), pro hybr.
Distribution: Nepal; India (Himachal Pradesh, Uttarakhand)
40 IND NEP
Remarks: Polystichum mehrae x squamosum
species :
Polystichum jinfoshanense
Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 3(4): 29 (1983)
Distribution: China (Chongqing, Guizhou, Sichuan, NE-Yunnan)
36 CHI
species :
Polystichum jinpingense
Z.L.Liang, Liang Zhang & Li Bing Zhang; Phytotaxa 539(1): 70 (2022)
Distribution: China (Yunnan)
36 CHI
species :
Polystichum jiucaipingense
P.S.Wang & Q.Luo; Bull. Bot. Res., Harbin 29(2): 134 (2009)
Distribution: China (NW-Guizhou)
36 CHI
Remarks: subgen. Polystichum, Sect. Micropolystichum
species :
Polystichum jiulaodongense
W.M.Chu & Z.R.He; Fl. Reipubl. Popularis Sin. [Fl. China Ed. Committee] 5(2): 227 (2001)
Distribution: China (Sichuan: Emei Shan)
36 CHI
Remarks: subgen. Haplopolystichum, sect. Basigemmifera
species :
Polystichum kadyrovii
Askerov & A.E.Bobrov; Bot. Zhurn. (Moscow & Leningrad) 57: 1297 (1972)
Distribution: Azerbaijan (Talysh)
33 AZE
species :
Polystichum kalambatitrense
Tardieu; Notul. Syst. (Paris) 15: 165, f. 1(1-4) (1956)
Distribution: Madagascar
29 MDG
species :
Polystichum kenwoodii
Lorence & W.L.Wagner; PhytoKeys 4: 22 (2011)
Distribution: Marquesas Isl. (Ua Huka, Hiva Oa, Tahuata)
61 MRQ
species :
Polystichum keysserianum
Rosenst.; Repert. Spec. Nov. Regni Veg. 10: 331 (1912)
Distribution: New Guinea (Irian Jaya, Papua New Guinea)
42 NWG
Synonyms: hidden
= Polystichum daymanense Nakaike; Bot. Exped. Papua New Guinea, Nation. Sci. Mus. (Tokyo) 102 (1975)
species :
Polystichum kilimanjaricum
Pic.Serm.; Webbia 27: 445 (1972 publ. 1973)
Distribution: Tanzania
25 TAN
species :
Polystichum kinabaluense
C.Chr.; Gard. Bull. Straits Settlem. 7: 255, t. 52 (1934)
Distribution: Borneo (Mt. Kinabalu, Sabah); Moluccas (Seram); New Guinea (Papua New Guinea); Bismarck Arch. (New Ireland)
42 BOR MOL NWG 43 BIS
Synonyms: hidden
= Polystichum clemensiae Copel.; Philipp. J. Sci. 56: 474, t. 4 (1935)
species :
Polystichum kropfii
Li Bing Zhang & Yi F.Duan; Phytotaxa 404(6): 216 (2019)
Distribution: China (Guizhou)
36 CHI
species :
Polystichum kruckebergii
W.H.Wagner; Amer. Fern J. 56: 4 (1966)
Name: Kruckeberg´s Hollyfern
Distribution: Alaska; Canada (British Columbia); USA (California, Idaho, Montana, Nevada, Oregon, Utah, Washington State)
70 ASK 71 CDN 73 USA 76 USA
species :
Polystichum kuankuoshuiense
Bo Wang, Q.W.Sun & Li Bing Zhang; Phytotaxa 592(2): 145 (2023)
Distribution: China (Guizhou)
36 CHI
species :
Polystichum kungianum
H.He & Li Bing Zhang; Bot. Stud. (Taipei) 51(3): 396 (2010)
Distribution: China (Chongqing, Hunan)
36 CHI
Remarks: subgen. Haplopolystichum, sect. Basigemmifera
species :
Polystichum kwakiutlii
D.H.Wagner; Amer. Fern J. 80: 50, fig. 1 (1990)
Name: Kwakiutl´s hollyfern
Distribution: Canada (C-British Columbia: Alice Arm)
Conservation Status: IUCN Ex/E (found only once)
species :
Polystichum kwangtungense
Ching; Icon. Filic. Sin. 5: t. 236 (1958)
Distribution: China (Guangdong)
36 CHI
species :
Polystichum lachenense
(Hook.) Bedd.; Ferns Brit. India t. 32 (1865)
Name: Aleutian Hollyfern
Distribution: China (Gansu, Sichuan, Xinjiang, Yunnan); Tibet; Japan; Taiwan; Pakistan (Swat, Gilgit, Hazara); Nepal; Bhutan; India (Jammu & Kashmir, Arunachal Pradesh, Sikkim, Uttarakhand, West Bengal); Myanmar; Aleutes (Adak Isl., Atka Isl.)
36 CHI TIB 38 JAP TAI 40 BHU IND NEP PAK 41 MYA 70 ASK
Remarks: subgen. Polystichum, Sect. Sorolepidium
Synonyms: hidden
= Aspidium lachenense Hook.; Sp. Fil. [W. J. Hooker] 4: 8, t. 212 (1862)
= Polystichum aleuticum C.Chr.; Hultén, Svensk Bot. Tidskr. 30: 515 (1936)
= Polystichum sinkiangense Ching ex Chang Y.Yang; Bull. Bot. Res., Harbin 8(2): 55 (1988)
= Polystichum tsuchuense Ching; Fl. Tsinling. 2: 222 (1974)
species :
Polystichum lanceolatum
(Baker) Diels; Bot. Jahrb. Syst. 29: 193 (1900)
Distribution: China (Guizhou, Hubei, ?Henan, Hunan, Jiangxi, Sichuan)
36 CHI
Synonyms: hidden
= Aspidium lanceolatum Baker; Gard. Chron., ser. 2, 14: 494 (1880), non Ettingsh. (1864)
= Hemesteum nanum (Christ) H.Lév.; Fl. Kouy-Tcheou 497-498 (1915)
= Hemesteum parvulum (Christ) H.Lév.; Fl. Kouy-Tcheou 497-498 (1915)
= Polystichum nanum Christ; Bull. Acad. Int. Géogr., sér. 3, 16: 238 (1906)
= Polystichum parvulum Christ; Bull. Acad. Int. Géogr. Bot. 114 (1904)
species :
Polystichum lancilobum
C.Chr.; Kongl. Svenska Vetensk. Acad. Handl., ser. 3, 16(2): 40, t. 9(1-4) (1937)
Distribution: Hispaniola (Dominican Republic)
81 HIS
Remarks: recently recollected (Majure et al. 2020), close to P. dissimulans and P. heterolepis
species :
Polystichum langchungense
Ching ex H.S.Kung; Chin. J. Appl. Environ. Biol. 3(2): 134, pl. 1: 1 (1997)
Distribution: China (Guizhou, Sichuan, Yunnan)
36 CHI
Synonyms: hidden
= Polystichum fengjieense Ching ex S.F.Wu; W. T. Wang, Keys Vasc. Pl. Wuling Mts. 58 (1995), nom. nud.
species :
Polystichum laniceps
Rosenst.; Hedwigia 46: 112 (1906)
Distribution: Brazil (Rio Grande do Sul)
84 BRA
species :
Polystichum lehmannii
Hieron.; Bot. Jahrb. Syst. 34: 452 (1904)
Distribution: Colombia (Antioquia, Boyacá, Caldas, Cauca, Cundinamarca); Ecuador; Peru; Bolivia (Cochabamba, La Paz)
82 CLM 83 BOL ECU PER
Synonyms: hidden
= Polystichum yungense Rosenst.; Repert. Spec. Nov. Regni Veg. 11: 55 (1912)
species :
Polystichum lemmonii
Underw.; Native Ferns, ed. 6: 116 (1900)
Name: Shasta Fern, Lemmon´s hollyfern
Distribution: SW-Canada (S-British Columbia); NW-USA (N-Washington, NE-Oregon, N-California)
71 CDN 73 USA 76 USA
Synonyms: hidden
= Aetopteron lemmonii (Underw.) House; Amer. Fern J. 10: 88 (1920)
= Polystichum mohrioides var. lemmonii (Underw.) Fernald; Rhodora 26: 92 (1924)
species :
Polystichum lentum
(D.Don) T.Moore; Index Fil. [T. Moore] 86 (1858)
Distribution: Tibet (Medog); Pakistan (Swat); Nepal; Bhutan; India (Jammu & Kashmir, Himachal Pradesh, Uttarakhand, Sikkim, Darjeeling, Meghalaya, Nagaland, Manipur, Arunachal Pradesh, Assam State, Mizoram, Tripura, West Bengal); Myanmar (Kachin, Chin, etc.); New Zealand [I]
36 TIB 40 BHU IND NEP PAK 41 MYA 51 nzl
Remarks: subgen. Polystichum, Sect. Macropolystichum
Synonyms: hidden
= Aspidium auriculatum var. lentum (D.Don) C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 507 (1880)
= Aspidium auriculatum var. subbipinnatum Hook.; Sp. Fil. [W. J. Hooker] 4: 11 (1862)
= Aspidium lentum D.Don; Prodr. Fl. Nepal. 4 (1825)
= Aspidium ocellatum Wall.; Numer. List. [Wallich] no. 360 (1829), nomen
= Aspidium ocellatum Wall. ex Mett.; Abh. Senckenberg. Naturf. Ges. 2(2) [Pheg. u. Asp.]: 43 (1858)
= Polystichum auriculatum var. lentum (D.Don) Bedd.; Ferns Brit. India 204 (1883)
= Polystichum auriculatum var. subbipinnatum (Hook.) Bedd.; Ferns Brit. India t. 136 (1866) non P. nepalense var. subbipinnatum C. Chr. 1931
= Polystichum lentum var. bifurcatum M.C.Biswas & S.K.Basu; J. Econ. Taxon. Bot. 3(3): 961 (1982 publ. 1983) (fide Fraser-Jenkins 1997)
= Polystichum longipes Ching & S.K.Wu; Fl. Xizangica 1: 213 (1983), not P. longipes Maxon (1909)
= Polystichum medogense Ching & S.K.Wu; Fl. Xizangica 1: 214 (1983)
= Polystichum ocellatum (Wall.) Schott; Gen. Fil. (Vindob.) ad t. 9 (1834)
= Polystichum panigrahianum Sarn.Singh; Ferns Fern Allies Arunachal Pradesh 1: 301 (2005)
= Polystichum sublentum Sarn.Singh & Panigrahi; Ferns Fern Allies Arunachal Pradesh 1: 307 (2005) (fide Fraser-Jenkins 2008)
species :
Polystichum lepidocaulon
(Hook.) J.Sm.; Ferns Brit. Foreign 286 (1866)
Distribution: China (S-Anhui, Fujian, Guangxi, Hunan, S-Jiangsu, Jiangxi, Zhejiang); South Korea (incl. Cheju Isl.); Japan; Taiwan
36 CHI 38 JAP KOR TAI
Remarks: subgen. Haplopolystichum, sect. Cyrtomiopsis
Synonyms: hidden
= Aspidium lepidocaulon Hook.; Sp. Fil. [W. J. Hooker] 4: 12, t. 217 (1862)
= Cyrtomidictyum lepidocaulon (Hook.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 182 (1940)
= Dryopteris lepidocaulis (Hook.) Kuntze; Revis. Gen. Pl. 2: 813 (1891)
= Polystichum lepidocaulon var. appendiculatum Nakai; Bot. Mag. (Tokyo) 46(550): 632 (1932)
species :
Polystichum lepidotum
Sundue & M.Kessler; Brittonia 57(3): 210 (2005)
Distribution: Bolivia (La Paz)
83 BOL
Remarks: subgen. Polystichum, sect. Polystichum
Conservation Status: Known only from type collection
species :
Polystichum leveillei
C.Chr.; Bull. Acad. Int. Géogr. Bot. 143 (1918)
Distribution: China (Guizhou)
36 CHI
Synonyms: hidden
= Hemesteum leveillei (C.Chr.) H.Lév.; Fl. Kouy-Tcheou 497-498 (1915)
species :
Polystichum levingei
C.Hope ex Christ; Bull. Acad. Int. Géogr., sér. 3, 11: 260 (1902)
Distribution: China (S-Gansu, SW-Henan, W-Hubei, W-Sichuan, N-Yunnan); Tibet; Taiwan; Nepal; Bhutan; India (Arunachal Pradesh, Sikkim); Myanmar
36 CHI TIB 38 TAI 40 BHU IND NEP 41 MYA
Remarks: subgen. Polystichum, Sect. Stenopolystichum
Synonyms: hidden
= Polystichum caespitosum var. stenophyllum Franch.; Nouv. Arch. Mus. Hist. Nat., sér. 2, 10: 117 (1887)
= Polystichum conaense Ching & S.K.Wu; Fl. Xizangica 1: 217 (1983)
= Polystichum deversum Christ; Bot. Gaz. 51(5): 353-354 (1911)
= Polystichum levingei N.C.Nair; Bull. Bot. Surv. India 11: 192 (1969 publ. 1971), non C. Hope ex Christ 1902
= Polystichum stenophyllum (Franch.) Christ; Mém. Soc. Bot. France 1 [Hist. Acrostich.]: 27 (1905)
= Polystichum stenophyllum var. abbreviatum Tagawa; Acta Phytotax. Geobot. 3(2): 93 (1934)
= Polystichum stenophyllum var. conaense (Ching & S.K.Wu) W.M.Chu & Z.R.He; Fl. Reipubl. Popularis Sin. [Fl. China Ed. Committee] 5(2): 8 (2001)
species :
Polystichum libingii
Yi F.Duan & Kropf; Phytotaxa 404(6): 216 (2019)
Distribution: China (Guizhou)
36 CHI
species :
Polystichum liboense
P.S.Wang & X.Y.Wang; Guizhou Sci. 21(1-2): 109 (2003)
Distribution: China (S-Guizhou); Vietnam
36 CHI 41 VIE
species :
Polystichum lilianae
Barrington; Brittonia 55(4): 321 (2003)
Distribution: Costa Rica
80 COS
species :
Polystichum lindsaeifolium
Scort.; J. Malayan Branch Roy. Asiat. Soc. 4: 61 (1926)
Distribution: Thailand; peninsular Malaysia (Kelantan, Pahang, Selangor)
41 THA 42 MLY
species :
Polystichum lineare
(C.Chr.) Copel.; Philipp. J. Sci. 78: 397 (1951)
Distribution: New Guinea (Papua New Guinea)
42 NWG
Synonyms: hidden
= Papuapteris linearis C.Chr.; Brittonia 2: 300, f. 2 (1937)
species :
Polystichum lionelii
O.G.Martínez & M.A.Hern.; Phytotaxa 619(2): 190 (2023)
Distribution: Argentina (Salta)
85 ARG
species :
Polystichum liui
Ching; Bull. Bot. Res., Harbin 3(4): 28 (1983)
Distribution: China (Chongqing, ?Guangxi, Guizhou, Hunan)
36 CHI
species :
Polystichum lonchitis
(L.) Roth; Tent. Fl. Germ. 3: 71 (1799)
Name: Lanzen-Schildfarn (DE), Northern Hollyfern (EN)
Distribution: Iceland; Faeroe Isl.; England (N-England, N-Wales, Scotland, Isle of Man); Ireland (W- & N-Ireland); Norway; Sweden; Finland; Belgium; Luxembourg; Germany (Brandenburg [I], Berlin [I], Baden-Württemberg [I], Bayern, Hessen [I], Mecklenburg-Vorpommern [I], Niedersachsen, Nordrhein-Westfalen, Rheinland-Pfalz [I], Saarland, Sachsen [I], Sachsen-Anhalt, Thüringen); Switzerland; Liechtenstein; Austria; Poland; Czech Republic; Slovakia; Hungary; Spain; Andorra; France; Corsica; Italy; Slovenia; Croatia; Bosnia & Hercegovina; Montenegro; Serbia; Kosovo; North Macedonia; Albania; Romania; Bulgaria; Greece (mountains in mainland); Crete (Lefka Ori); Estonia; Latvia; N-European Russia; W-European Russia; Ukraine; Crimea; Morocco; Siberia (Altai, Buryatia, Irkutsk, Krasnoyarsk, W-Siberia); Russian Far East (Kamchatka, Kuril Isl., Sakhalin); Kazakhstan; Uzbekistan; Kyrgyzstan; Tajikistan; Northern Caucasus; Georgia [Caucasus]; Armenia; Azerbaijan; Turkey (Anatolia); E-Afghanistan; China (Xinjiang); Japan; Pakistan (Gilgit, Chitral, Swat, Hazara); NW-India (Jammu & Kashmir); Alaska; Greenland; Canada (Alberta, British Columbia, Newfoundland, Nova Scotia, Northern Territories, Ontario, Québec, Yukon); USA (Arizona, California, Colorado, Idaho, Michigan, Minnesota, Montana, Nevada, New York, Oregon, South Dakota, Utah, Washington State, Wyoming)
10 FIN FOR GRB ICE IRE NOR SWE 11 AUT BGM CZE GER HUN LIE LUX POL SLO SWI 12 ADO COR FRA SPA 13 ALB BIH BUL CRO GRC ITA KOS KRI MNT NMA ROM SLV SRB 14 EST KRY LAT RUS UKR 20 MOR 30 SIB 31 RFE 32 KAZ KGZ TZK UZB 33 ARM AZE GEO NCS 34 AFG TUR 36 CHI 38 JAP 40 IND PAK 70 ASK CDN GNL 71 CDN 72 CDN 73 USA 74 USA 75 USA 76 USA
Remarks: subgen. Polystichum, sect. Polystichum
Synonyms: hidden
= Aetopteron lonchitis (L.) House; Amer. Fern J. 10: 88 (1920)
= Aspidium aculeatum subsp. lonchitis (L.) Bonnier & Layens; Tabl. Syn. Pl. Vasc. France: 378 (1894)
= Aspidium asperum Gray; Nat. Arr. Brit. Pl. 2 (1821)
= Aspidium lonchitis (L.) Sw.; J. Bot. (Schrader) 1800(2): 30 (1802)
= Dryopteris lonchitis (L.) Kuntze; Revis. Gen. Pl. 2: 813 (1891)
= Hypopeltis lonchitis (L.) Tod.; Giorn. Sci. Nat. Econ. Palermo 1: 237 (1866)
= Polypodium lonchitis L.; Sp. Pl. 2: 1088 (1753)
= Polystichum asperum (Gray) Bubani; Fl. Pyren. (Bubani) 4: 435 (1901)
= Polystichum lonchitis f. marginale McColl; Amer. Fern J. 14: 107 (1924)
= Polystichum lonchitis var. japonicum Nakai & H.Itô; Iconogr. Pl. Asiae Orient. 1: t. 28 (1936)
= Polystichum marginale (McColl) Cody; Naturaliste Canad. 93: 273 (1966)
species :
Polystichum longifrons
Sa.Kurata; Sci. Rep. Yokosuka City Mus. 10: 31 (1964)
Distribution: South Korea; Japan
38 JAP KOR
Remarks: subgen. Polystichum, Sect. Hypopeltis III
species :
Polystichum longipaleatum
Christ; Notul. Syst. (Paris) 1: 35 (1909)
Distribution: China (Guangxi, Guizhou, Hunan, Sichuan, Yunnan); Tibet; Nepal; Bhutan; India (Uttarakhand, Sikkim, Darjeeling, Arunachal Pradesh, Nagaland, Meghalaya, Manipur, West Bengal); Myanmar (Kachin, etc.)
36 CHI TIB 40 BHU IND NEP 41 MYA
Synonyms: hidden
= Aspidium aculeatum var. setosum Wall. ex C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 510 (1880)
= Aspidium setosum Wall.; Numer. List. [Wallich] no. 371 (1829), nomen (non Sw. 1801)
= Aspidium wallichianum Kunze; Linnaea 24: 291 (1851)
= Hypopeltis setosa (Wall. ex C.B.Clarke) Bory; Voy. Indes Or. [Bélanger] 2: 68 (1833)
= Lastrea setosa Wall. ex T.Moore; Index Fil. [T. Moore] 103 (1858)
= Polystichum aculeatum var. setosum (C.B.Clarke) Bedd.; Ferns Brit. India 209 (1883)
= Polystichum aculeatum var. setulosum Rosenst.; Repert. Spec. Nov. Regni Veg. 13(355-358): 129-130 (1914)
= Polystichum nudisorum Ching; Bull. Fan Mem. Inst. Biol. Bot. 11: 71 (1941) (fide Fraser-Jenkins et al. 2018)
= Polystichum pianmaense W.M.Chu; Acta Bot. Yunnan., Suppl. 5: 51 (1992) (fide Fraser-Jenkins et al. 2018)
= Polystichum setosum (Wall. ex C.B.Clarke) Khullar & Gupta; Nova Hedwigia 32: 420 (1980), isonym
= Polystichum setosum (Wall. ex C.B.Clarke) Schott ex Diels; Nat. Pflanzenfam. [Engler & Prantl] 1(4): 191 (1899), non (Thunb.) C. Presl 1836
= Polystichum setosum (Wall.) Schott; Gen. Fil. (Vindob.): t. 9 (1834), nom. nud.
species :
Polystichum longispinosum
Ching ex Li Bing Zhang & H.S.Kung; Acta Phytotax. Sin. 36(5): 467, pl. 1: 1 (1998)
Distribution: China (Guizhou, Sichuan, Yunnan)
36 CHI
Remarks: subgen. Polystichum, Sect. Neopolystichum
species :
Polystichum longissimum
Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(4): 16 (1984)
Distribution: China (Chongqing, Guizhou)
36 CHI
species :
Polystichum loratum
H.He & Li Bing Zhang; Novon 22(2): 166 (2012)
Distribution: China (N-Guizhou)
36 CHI
species :
Polystichum luctuosum
(Kunze) T.Moore; Index Fil. [T. Moore] 95 (1858)
Name: Korean rockfern
Distribution: E-Zimbabwe; South Africa (Limpopo, Mpulamanga, Free State, KwaZulu-Natal, E-Cape Prov.); Lesotho; Eswatini [Swaziland]; Madagascar; La Réunion; China (Anhui, Fujian, S-Gansu, Guangxi, Guizhou, Henan, Hubei, W-Hunan, Jiangxi, Jilin, S-Shaanxi, Shandong, Sichuan, Yunnan, Zhejiang); Tibet; South Korea (Cheju Isl.); Japan; Taiwan; W-Nepal; India (Himachal Pradesh, Uttarakhand, Arunachal Pradesh); Vietnam; USA [I] (Florida [I], Louisiana [I])
26 ZIM 27 LES SAF SWZ 29 MDG REU 36 CHI TIB 38 JAP KOR TAI 40 IND NEP 41 VIE 78 usa
Remarks: subgen. Polystichum, sect. Xiphopolystichum. Inclusion of P. tsus-simense fide Fraser-Jenkins et al. (2015)
Synonyms: hidden
= Aspidium aculeatum var. pallescens Franch.; Nouv. Arch. Mus. Hist. Nat., sér. 2, 10: 117 (1887)
= Aspidium luctuosum Kunze; Linnaea 10: 548 (1836)
= Aspidium tsus-simense Hook.; Sp. Fil. [W. J. Hooker] 4: 16, t. 220 (1862)
= Polystichum falcilobum Ching; Fl. Xizangica 1: 236 (1983)
= Polystichum lobatum var. luctuosum (Kunze) Christ; Ber. Schweiz. Bot. Ges. 3: 34 (1893)
= Polystichum lobatum var. tsus-simense (Hook.) C.Chr.; Index Filic. 588 (1906)
= Polystichum polyblepharum Nakai; Bot. Mag. (Tokyo) 39: 117 (1925), nom. illeg. (non C. Presl (1849))
= Polystichum tsus-simense (Hook.) J.Sm.; Hist. Fil. 219 (1875)
= Polystichum tsus-simense var. parvipinnulum W.M.Chu; Acta Bot. Yunnan., Suppl. 5: 52 (1992)
species :
Polystichum luteoviride
Li Bing Zhang, Yi F.Duan, N.T.Lu & Liang Zhang; Phytotaxa 313(3): 296 (2017)
Distribution: China (Guizhou)
36 CHI
species :
Polystichum macleae
(Baker) Diels; Nat. Pflanzenfam. [Engler & Prantl] I, 4: 190 (1899)
Distribution: South Africa (Limpopo, Mpulamanga); Eswatini [Swaziland]
27 SAF SWZ
Remarks: subgen. Polystichum, Afra clade
Synonyms: hidden
= Aspidium macleae Baker; Hooker´s Icon. Pl. t. 1654 (1886)
species :
Polystichum macrodon
Li Bing Zhang; Phytotaxa 404(6): 222 (2019)
Distribution: China (Guizhou)
36 CHI
species :
Polystichum maevaranense
Tardieu; Notul. Syst. (Paris) 15: 166, f. l(5-9) (1956)
Distribution: Madagascar
29 MDG
Remarks: subgen. Polystichum, Sect. Hypopeltis II
species :
Polystichum magnificum
F.Ballard; Kew Bull. 12: 48b, f. 1 (1957)
Distribution: S-Ethiopia; Uganda; Kenya; Tanzania
24 ETH 25 KEN TAN UGA
species :
Polystichum makinoi
(Tagawa) Tagawa; Acta Phytotax. Geobot. 6: 258 (1936)
Distribution: China (Anhui, Fujian, S-Gansu, Guangxi, Guizhou, Hebei, Henan, W-Hubei, Hunan, Jiangsu, Jiangxi, N-Shaanxi, Sichuan, Yunnan, Zhejiang); Tibet; Japan; Bhutan
36 CHI TIB 38 JAP 40 BHU
Remarks: subgen. Polystichum, Sect. Hypopeltis. Not in India
Synonyms: hidden
= Polystichum aculeatum var. makinoi Tagawa; Acta Phytotax. Geobot. 1: 88 (1932)
= Polystichum aculeatum var. nigropaleaceum Makino; J. Jap. Bot. 2: 21 (1922), nom. illeg.
= Polystichum aculeatum var. veitchii Christ; Bull. Herb. Boissier, sér. 2, 3: 513 (1903)
= Polystichum longiaristatum Ching, Boufford & K.H.Shing; J. Arnold Arbor. 64: 33 (1983)
species :
Polystichum malipoense
Li Bing Zhang, M.Q.Han & Yan Liu; Phytotaxa 365(2): 154 (2018)
Distribution: China (Yunnan)
36 CHI
species :
Polystichum manickamianum
Benniamin, Fraser-Jenk. & Irudayaraj; Novon 18(3): 289 (2008)
Distribution: S-India (Tamil Nadu: Agasthiar Hills)
40 IND
Conservation Status: IUCN CR for India (Fraser-Jenkins 2012)
species :
Polystichum manmeiense
(Christ) Nakaike; Y. Otani, Rep. Cryptogamic Study Nepal (Misc. Publ. Nat. Sci. Mus. Tokyo): 141 (1982)
Distribution: China (Guizhou, Yunnan); Tibet; Taiwan; Nepal; Bhutan; India (Himachal Pradesh, Uttarakhand, Sikkim, Darjeeling, Arunachal Pradesh, West Bengal); Myanmar (Kachin, Chin, etc.); Philippines
36 CHI TIB 38 TAI 40 BHU IND NEP 41 MYA 42 PHI
Remarks: subgen. Polystichum, Sect. Achroloma
Synonyms: hidden
= Aspidium manmeiense Christ; Bull. Herb. Boissier, sér. 1, 6: 965 (1898)
= Polystichum falcatipinnum Hayata; Icon. Pl. Formosan. 4: 192, f. 130 (1914)
= Polystichum nepalense f. subbipinnatum (C.Chr.) Ching; Icon. Filic. Sin. 5: t. 243, f. 4 (1958)
= Polystichum nepalense var. subbipinnatum C.Chr.; Contr. U. S. Natl. Herb. 26: 284 (1931) (fide Fraser-Jenkins 2008)
species :
Polystichum marionense
Alston & Schelpe; J. S. African Bot. 23: 106. i. l(a), t. 34 (1957)
Distribution: Marion Isl.; Prince Edward Isl.; Iles Crozet; Ile Amsterdam
91 ASP CRZ MPE
species :
Polystichum marquesense
E.D.Br.; Bull. Bernice P. Bishop Mus. 89 [Fl. S. E. Polynesia II]: 40, t. 6 (1931)
Distribution: Marquesas Isl. (Nuku Hiva, Ua Huka, Ua Pou, Hiva Oa, Tahuata, Fatu Hiva)
61 MRQ
species :
Polystichum martini
Christ; Bull. Acad. Int. Géogr., sér. 3, 11: 263 (1902)
Distribution: China (C-Guizhou)
36 CHI
species :
Polystichum maximum
M.Kessler & A.R.Sm.; Brittonia 59(2): 197 (2007)
Distribution: Bolivia (La Paz)
83 BOL
Remarks: subgen. Polystichum, sect. Polystichum
Synonyms: hidden
= Polystichum giganteum M.Kessler & A.R.Sm.; Brittonia 57(3): 215 (2005), nom. illeg., non Fée 1869
species :
Polystichum mayebarae
Tagawa; Acta Phytotax. Geobot. 3: 91 (1934)
Distribution: China (Gansu, Guizhou, Henan, Hubei, Sichuan, Yunnan, Zhejiang); South Korea (Cheju Isl.); Japan; Taiwan
36 CHI 38 JAP KOR TAI
Remarks: subgen. Polystichum, sect. Xiphopolystichum
Synonyms: hidden
= Polystichum tsus-simense var. mayebarae (Tagawa) Sa.Kurata; Hokuriku J. Bot. 6: 9 (1957)
species :
Polystichum meeboldii
Rosenst.; Repert. Spec. Nov. Regni Veg. 12: 246 (1913)
Distribution: China (Guangxi, Guizhou, Yunnan); Taiwan; Nepal; Bhutan; India (Arunachal Pradesh, ?Assam State, Manipur, Meghalaya, Nagaland, Sikkim); Myanmar; Thailand (widespread); Laos; Vietnam
36 CHI 38 TAI 40 BHU IND NEP 41 LAO MYA THA VIE
Remarks: subgen. Polystichum, Sect. Macropolystichum. Valid name disputed due to missing type of P. meeboldii, possibly P. attenuatum is valid.
Synonyms: hidden
= Polystichum attenuatum Tagawa & K.Iwats.; Acta Phytotax. Geobot. 23: 113, f. 9 (1968)
= Polystichum attenuatum var. subattenuatum (Ching & W.M.Chu) W.M.Chu & Z.R.He; Fl. Reipubl. Popularis Sin. [Fl. China Ed. Committee] 5(2): 17 (2001)
= Polystichum mannii C.Hope ex Fraser-Jenk.; Aspects Pl. Sci. 13: 263 (1991)
= Polystichum subattenuatum Ching & W.M.Chu; Acta Bot. Yunnan., Suppl. 5: 51 (1992)
species :
Polystichum mehrae
Fraser-Jenk. & Khullar; Indian Fern J. 2(1-2): 10 (1985) (nom. inval., lectotype missing, et Indian Fern J. 3(1-2): 121 (1986)
Distribution: China (Yunnan, W-Sichuan); E-Tibet; Pakistan (Hazara); Nepal; Bhutan; India (Jammu & Kashmir, Himachal Pradesh, Arunachal Pradesh, Uttarakhand); ?Myanmar
36 CHI TIB 40 BHU IND NEP PAK
Synonyms: hidden
= Polystichum acanthophyllum var. indicum Christ; Mém. Soc. Bot. France 1 [Hist. Acrostich.]: 31 (1905)
= Polystichum aculeatum var. contortum C.Hope ex E.W.Trotter; Ferns Punjab 13 (1889)
= Polystichum mehrae f. latifundum H.S.Kung & Li Bing Zhang; Li Bing Zhang, H. S. Kung & X. S. Guo, Acta Bot. Yunnan. 17(1): 25 (1995)
species :
Polystichum menglaense
Z.L.Liang & Li Bing Zhang; Phytotaxa 479: 117 (2021)
Distribution: China (Yunnan)
36 CHI
species :
Polystichum mengziense
Li Bing Zhang; Phytotaxa 60: 58 (2012), nom. nov.
Distribution: China (SE-Yunnan)
36 CHI
Synonyms: hidden
= Polystichum deltodon var. henryi Christ; Mém. Soc. Bot. France 1 [Hist. Acrostich.]: 27 (1905)
species :
Polystichum mexiae
Copel.; Univ. Calif. Publ. Bot. 19: 299, t. 55 (1941)
Distribution: Ecuador; ?Peru; Bolivia (Cochabamba, La Paz, Santa Cruz)
83 BOL ECU
species :
Polystichum mickelii
A.R.Sm.; Amer. Fern J. 70: 25 (1980)
Distribution: Mexico (Chiapas, Oaxaca, Puebla, Veracruz); Guatemala; El Salvador; Honduras
79 MEX 80 ELS GUA HON
species :
Polystichum microchlamys
(Christ) Matsum.; Index Pl. Jap. 1: 343 (1904)
Name: Attu Hollyfern
Distribution: Russian Far East (Kamchatka, Kuril Isl.); Japan; Aleutes (Attu Isl.)
31 RFE 38 JAP 70 ASK
Remarks: subgen. Polystichum, North American clade
Synonyms: hidden
= Aspidium microchlamys Christ; Bull. Herb. Boissier, sér. 1, 7: 820 (1899)
= Polystichum azumiense (Seriz.) Seriz.; Shidekobushi 1(2): 93 (2009)
= Polystichum braunii subsp. kamtschaticum (C.Chr. & Hultén) Á.Löve & D.Löve; Taxon 26(2-3): 326 (1977)
= Polystichum braunii var. kamtschaticum C.Chr. & Hultén; Kungl. Svenska Vetenskapsakad. Handl. 3, 5: 38, t. 2 (1927)
= Polystichum kamtschaticum (C.Chr. & Hultén) Fomin; Fl. Sibir. Orient. Extremi 5: 94 (1930)
= Polystichum microchlamys (Christ) Kodama; Matsum., Icon. Pl. Koisikav. 3: t. 147 (1915)
= Polystichum microchlamys var. azumiense Seriz.; J. Jap. Bot. 45(9): 262 (1970)
species :
Polystichum minimum
(Y.T.Hsieh) Li Bing Zhang; Phytotaxa 60: 58 (2012)
Distribution: China (Chongqing, NW-Guangxi, Guizhou); Vietnam
36 CHI 41 VIE
Remarks: subgen. Haplopolystichum, sect. Cyrtogonellum
Synonyms: hidden
= Cyrtogonellum inaequale (Christ) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 331, t. 7 (4) (1938)
= Cyrtogonellum minimum Y.T.Hsieh; Bull. Bot. Res., Harbin 9(3): 17 (1989)
= Cyrtomium fraxinellum var. inaequale Christ; Bull. Acad. Int. Géogr., sér. 3, 11: 264 (1902)
species :
Polystichum minutissimum
Li Bing Zhang & H.He; Bot. Stud. (Taipei) 50: 353 (2009)
Distribution: China (S-Guizhou)
36 CHI
species :
Polystichum mohrioides
(Bory) C.Presl; Tent. Pterid. 83 (1836)
Distribution: Falkland Isl.; South Georgia Isl.
91 FAL SGE
Remarks: subgen. Polystichum, North American clade. Possibly a synonym of P. plicatum
Synonyms: hidden
= Aspidium mohrioides Bory; Mém. Soc. Linn. Paris 4: 597 (1824)
= Dryopteris mohrioides (Bory) Kuntze; Revis. Gen. Pl. 2: 813 (1891)
= Nephrodium mohrioides (Bory) Desv.; Mem. Soc. Linn. Paris 6(2): 261 (1827)
= Polystichum mohrioides var. plicatum C.Chr.; Ark. Bot. 10(2): 17 (1910)
species :
Polystichum moluccense
(Blume) T.Moore; Index Fil. [T. Moore] 84 (1858)
Distribution: Borneo (Mt. Kinabalu, Sabah); Sulawesi; Philippines; Moluccas (Ternate, Seram, etc.); New Guinea (Papua New Guinea); Bismarck Arch. (New Ireland); Solomon Isl. (Bougainville, Guadalcanal, Kolombangara, Makira, Malaita, Rendova, Nggatokae, Santa Ysabel, Vella Lavella); ?Fiji
42 BOR MOL NWG PHI SUL 43 BIS SOL
Synonyms: hidden
= Aspidium aculeatum var. batjanense Christ; Farnkr. Erde 238 (1897)
= Aspidium batjanense (Christ) Christ; Bull. Herb. Boissier, sér. 2, 6: 996 (1906)
= Aspidium moluccense Blume; Enum. Pl. Javae 2: 168 (1828)
= Polystichum aculeatum var. batjanense (Christ) C.Chr.; Farnkr. Erde 128 (1913)
= Polystichum acutifolium C.Presl; Abh. Königl. Böhm. Ges. Wiss., ser. 5, 6 [Epim. Bot.]: 259 (1851)
= Polystichum dendrophilum Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 3, 5: 221 (1922) (fide Fraser-Jenkins et al. 2018)
= Polystichum horizontale C.Presl; Abh. Königl. Böhm. Ges. Wiss., ser. 5, 6 [Epim. Bot.]: 57 (1851)
= Polystichum nudum Copel.; Philipp. J. Sci. 1, Suppl. II: 145 (1906) (fide Fraser-Jenkins et al. 2018)
= Polystichum oppositum Copel.; Philipp. J. Sci. 56: 474, t. 5 (1935)
= Polystichum papyrifolium Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 28: 41 (1918) (fide Fraser-Jenkins et al. 2018)
species :
Polystichum montevidense
(Spreng.) Rosenst.; Hedwigia 46: 111 (1906)
Distribution: Costa Rica; Colombia (Antioquia, Boyacá, Caldas, Cauca, Cundinamarca, Huila, Nariño, Santander, Valle); Venezuela (Merida); Ecuador; Peru; Bolivia (Chuquisaca, Cochabamba, La Paz, Santa Cruz, Tarija); Brazil (Minas Gerais, São Paulo, Rio de Janeiro, Rio Grande do Sul, Santa Catarina); Paraguay (Paraguari); Uruguay (Maldonado); Argentina (Buenos Aires, Cordoba, Jujuy, La Rioja, Misiones, Salta, San Luis, Tucuman)
80 COS 82 CLM VEN 83 BOL ECU PER 84 BRA 85 ARG PAR URU
Remarks: subgen. Polystichum, sect. Polystichum
Synonyms: hidden
= Aspidium aculeatum var. montevidense (Spreng.) Stuck.; Anales Mus. Nac. Buenos Aires ser. 3, 1: 296 (1902)
= Aspidium macrolepidum Desv.; Mem. Soc. Linn. Paris 6(2): 248 (1827)
= Aspidium montevidense (Spreng.) Hieron.; Bot. Jahrb. Syst. 22: 364 (1896)
= Aspidium montevidense f. imbricatum Hieron.; Bot. Jahrb. Syst. 22: 366 (1896)
= Aspidium montevidense var. subtripinnatum Rosenst.; Hedwigia 43: 223 (1904)
= Aspidium muricatum var. montevidense (Spreng.) Kuhn; Abh. Naturf. Ges. Halle 11: 35 (1870)
= Phegopteris montevidensis (Spreng.) Mett. ex Salomon; Nomencl. Gefäßkrypt. 276 (1883)
= Polypodium montevidense Spreng.; Syst. Veg., ed. 16 [Sprengel] 4(1): 59 (1827)
= Polystichum aculeatum var. microsorium (Fée) Farw.; Amer. Midl. Naturalist 12: 260 (1931)
= Polystichum aculeatum var. montevidense (Spreng.) Luetzelb.; Estud. Bot. Nordéste 3: 244 (1923)
= Polystichum microsorium Fée; Crypt. Vasc. Brésil 1: 125, t. 40, f. 1 (1869)
= Polystichum montevidense f. imbricatum (Hieron.) Hicken; Revista Mus. La Plata 15: 236 (1908)
= Polystichum sellowianum C.Presl; Tent. Pterid. 83 (1836)
variant :
var. squamulosum
(Hieron.) Hieron.; Hedwigia 46: 356 (1907)
Distribution: Argentina (Cordoba, Jujuy, Salta, Tucuman)
85 ARG
Synonyms: hidden
= Aspidium montevidense f. squamulosum Hieron.; Bot. Jahrb. Syst. 22(3): 366 (1896)
= Polystichum montevidense f. squamulosum (Hieron.) Hicken; Revista Mus. La Plata 15: 236 (1908)
species :
Polystichum monticola
N.C.Anthony & Schelpe; Bothalia 15(3-4): 554 (1985)
Distribution: W-Zimbabwe; South Africa (Free State, KwaZulu-Natal, E-Cape Prov., N-Cape Prov., W-Cape Prov.); Lesotho
26 ZIM 27 LES SAF
species :
Polystichum moorei
Christ; Ark. Bot. 4(12): 3 (1905) ex W. R. B. Oliv.
Distribution: Lord Howe Isl.
50 LHI
Remarks: subgen. Polystichum, Australasian clade
Conservation Status: IUCN R
Synonyms: hidden
= Polystichum kingii Watts; Proc. Linn. Soc. New South Wales 87: 401 (1912)
species :
Polystichum mucronifolium
(Blume) C.Presl; Abh. Königl. Böhm. Ges. Wiss., ser. 5, 6: 55 (1851)
Distribution: China (Yunnan); Taiwan; Nepal; Bhutan; India (Uttarakhand, Manipur, Meghalaya, Nagaland, Sikkim, Tamil Nadu, West Bengal); Sri Lanka; Myanmar; N-Thailand; Vietnam; Sumatra; Sulawesi; Java; Lesser Sunda Isl. (Bali, Lombok, Flores, Alor)
36 CHI 38 TAI 40 BHU IND NEP SRL 41 MYA THA VIE 42 JAW LSI SUL SUM
Remarks: subgen. Polystichum, Sect. Macropolystichum. It is assumed that records of "P. aculeatum" from the Lesser Sunda Isl. belong here.
Synonyms: hidden
= Aspidium aculeatum var. mucronifolium (Blume) C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 509 (1880)
= Aspidium amblyotus Kunze; Bot. Zeitung (Berlin) 283 (1848)
= Aspidium congener Blume; Enum. Pl. Javae 2: 165 (1828)
= Aspidium microphyllum Blume; Enum. Pl. Javae 2: 163 (1828)
= Aspidium mucronifolium Blume; Enum. Pl. Javae 2: 164 (1828)
= Aspidium tacticopterum Kunze; Linnaea 24: 210 (1851)
= Polystichum aculeatum var. formosanum Kodama; Bot. Mag. (Tokyo) 29: 328 (1915)
= Polystichum aculeatum var. mucronifolium (Blume) Bedd.; Ferns Brit. India 210 (1883)
= Polystichum amblyotus (Kunze) C.Presl; Abh. Königl. Böhm. Ges. Wiss., ser. 5, 6 [Epim. Bot.]: 55 (1851)
= Polystichum heteropaleaceum N.C.Nair & Nag; J. Bombay Nat. Hist. Soc. 75(1): 153 (1978)
= Polystichum kodamae Tagawa; J. Jap. Bot. 13: 186 (1937) (fide Fraser-Jenkins 1997)
= Polystichum microphyllum (Blume) C.Presl; Abh. Königl. Böhm. Ges. Wiss., ser. 5, 6 [Epim. Bot.]: 55 (1851)
= Polystichum mucronifolium (Blume) B.K.Nayar & Kaur; Companion Ferns Brit. India 49 (1974), isonym
= Polystichum tacticopterum (Kunze) T.Moore; Index Fil. [T. Moore] 105 (1858)
= Polystichum tenggerense Rosenst.; Repert. Spec. Nov. Regni Veg. 8: 164 (1910) (fide Fraser-Jenkins 1997)
species :
Polystichum muenchii
(Christ) C.Chr.; Index Filic. 585 (1906), as ´münchii´
Distribution: Mexico (Chiapas); Guatemala
79 MEX 80 GUA
Conservation Status: IUCN R
Synonyms: hidden
= Aspidium muenchii Christ; Bull. Herb. Boissier, sér. 2, 1: 1120 (1901), as ´münchii´
species :
Polystichum multifidum
(Mett.) T.Moore; Index Fil. [T. Moore] 84 (1857)
Distribution: Argentina (Chubut, Neuquen, Rio Negro, Santa Cruz, Tierra del Fuego); Chile (Los Rios, Los Lagos, Aisen, Magallanes)
85 ARG CLE
Synonyms: hidden
= Aspidium multifidum Mett.; Fil. Lechl. 1: 20, t. 3, f. 14-17 (1856)
= Aspidium pearcei (Phil.) F.Phil.; Anales Univ. Chile 385 (1881)
= Dicksonia andina Phil.; Anales Univ. Chile 94: 309 (1896)
= Dryopteris metteniana Kuntze; Revis. Gen. Pl. 2: 811 (1891)
= Polystichum multifidum f. autranii Hicken; Anales Soc. Ci. Argent. 62: 172 (1906)
= Polystichum multifidum var. pearcei (Phil.) R.A.Rodr.; Gayana, Bot. 44: 51 (1987)
= Polystichum nahuelhuapiense Diem; Darwiniana 12: 67, f. 1 (1960)
= Polystichum pearcei Phil.; Linnaea 33: 305 (1865)
species :
Polystichum mulunense
X.L.Shen & R.H.Jiang; Pl. Diversity & Resources 37(6): 737 (2015)
Distribution: China (Guangxi)
36 CHI
species :
Polystichum munitum
(Kaulf.) C.Presl; Tent. Pterid. 83 (1836)
Name: Schwert-Schildfarn, Schwertfarn (DE), Common Sword Fern (EN)
Distribution: Germany [I]; Alaska; Canada (British Columbia); USA (California, Idaho, Montana, Oregon, Pennsylvania [I], South Dakota, Washington State); Mexico (Baja California Norte); Isla Guadalupe
11 ger 70 ASK 71 CDN 73 USA 74 USA 75 USA 76 USA 79 GLP MEX
Remarks: subgen. Polystichum, North American clade
Synonyms: hidden
= Aetopteron munitum (Kaulf.) House; Amer. Fern J. 10: 88 (1920)
= Aspidium munitum Kaulf.; Enum. Filic. [Kaulfuss] 236 (1824)
= Aspidium munitum var. incisoserratum D.C.Eaton; Ferns N. Amer. 1: 188 (1878)
= Aspidium munitum var. nudatum D.C.Eaton; Rep. U.S. Geogr. Surv., Wheeler 6(Botany): 335 (1878)
= Dryopteris munita (Kaulf.) Kuntze; Revis. Gen. Pl. 2: 813 (1891)
= Nephrodium plumula C.Presl; Reliq. Haenk. 1: 33 (1825)
= Polypodium munitum (Kaulf.) C.Presl; C. Presl, Tent. Pterid. 180 (1836)
= Polystichum munitum f. flabellatum A.A.Eaton; Fern Bull. 9: 8 (1901)
= Polystichum munitum f. incisoserratum (D.C.Eaton) Clute & H.St.John; Proc. Biol. Soc. Wash. 41: 191 (1928)
= Polystichum munitum f. nudatum (D.C.Eaton) M.Broun; Index No. Amer. Ferns: 148 (1938)
= Polystichum munitum subsp. inciso-serratum (D.C.Eaton) Piper & Beattie; Fl. N. W. Coast [Piper & Beattie] 7 (1915)
= Polystichum munitum subsp. nudatum (D.C.Eaton) Ewan; Amer. Fern J. 32: 100 (1942)
= Polystichum munitum subsp. solitarium Maxon; Fern Bull. 11: 39 (1903)
= Polystichum munitum var. incisoserratum (D.C.Eaton) Underw.; Native Ferns, ed. 6: 116 (1900)
= Polystichum plumula (C.Presl) C.Presl; Tent. Pterid. 83 (1836)
= Polystichum solitarium (Maxon) Underw.; Maxon, Contr. U. S. Natl. Herb. 10: 493 (1908)
species :
Polystichum muricatum
(L.) Fée; Mém. Foug. 5 (Gen. Filic.): 278 (1852)
Distribution: Mexico (Chiapas, Colima, Guerrero, Oaxaca, Veracruz); Revillagigedos Isl. (Isla Socorro); El Salvador; Honduras; Nicaragua; Costa Rica; Panama; Jamaica; Hispaniola (Haiti, Dominican Republic); Colombia (Antioquia, Boyacá, Caldas, Cauca, Cundinamarca, Huila, Magdalena, Meta, Nariño, Norte de Santander); Venezuela (Aragua, Venezuela Distrito Federal, Lara, Merida, Miranda, Sucre, Yaracuy); Ecuador; Galapagos
79 MEX RGI 80 COS ELS HON NIC PAN 81 HIS JAM 82 CLM VEN 83 ECU GAL
Remarks: subgen. Polystichum, sect. Polystichum
Synonyms: hidden
= Aspidium aculeatum subsp. moritzianum (Klotzsch) Jenman; Bull. Bot. Dept., Jamaica n. s. 2: 270 (1895)
= Aspidium aculeatum var. moritzianum (Klotzsch) Griseb.; Fl. Brit. W. I. [Grisebach] 689 (1864)
= Aspidium moritzianum Klotzsch; Linnaea 20: 367 (1847)
= Aspidium muricatum (L.) Willd.; Sp. Pl., ed. 4 [Willdenow] 5(1): 264 (1810)
= Aspidium muricatum var. incisum Kuhn; Abh. Naturf. Ges. Halle 11: 35 (1870)
= Aspidium muricatum var. muelleri Mett. ex Kuhn; Abh. Naturf. Ges. Halle 11: 35 (1870)
= Dryopteris aculeata var. muricata (L.) Kuntze; Revis. Gen. Pl. 3(3): 378 (1898)
= Polypodium muricatum L.; Sp. Pl. 2: 1093 (1753)
= Polystichum aculeatum var. moritzianum (Klotzsch) Hook.
= Polystichum aculeatum var. muricatum (L.) C.Chr.
= Polystichum moritzianum (Klotzsch) Hieron.; Hedwigia 46: 354, t. 8, f. 20 (1906)
= Polystichum moritzianum var. setosodentatum Hieron.; Hedwigia 46: 354 (1907)
species :
Polystichum muscicola
Ching ex W.M.Chu & Z.R.He; Fl. Reipubl. Popularis Sin. [Fl. China Ed. Committee] 5(2): 228 (2001), nom. inval. (holotype location not cited), et Fl. China 2-3: 706 (2013)
Distribution: China (W-Hubei, W-Sichuan)
36 CHI
species :
Polystichum muticum
Copel.; Univ. Calif. Publ. Bot. 18: 221 (1942)
Distribution: New Guinea (Irian Jaya, Papua New Guinea)
42 NWG
species :
Polystichum myer-dreesii
Copel.; Univ. Calif. Publ. Bot. 18: 221 (1942) (´meijerdreesii´ auct.)
Distribution: New Guinea (Papua New Guinea, Irian Jaya)
42 NWG
species :
Polystichum nanchuanicum
Ching; Bull. Fan Mem. Inst. Biol. Bot., ser. 2, 1: 310 (1949)
Distribution: China (Chongqing)
36 CHI
species :
Polystichum napoense
C.Xiong & R.H.Jiang; Phytotaxa 601(1): 43 (2023)
Distribution: China (Guangxi)
36 CHI
species :
Polystichum nayongense
P.S.Wang & X.Y.Wang; Acta Bot. Yunnan. 19(1): 41, f. 1 (1997)
Distribution: China (W-Guizhou)
36 CHI
species :
Polystichum neoacutidens
Li Bing Zhang, Pollawatn & Liang Zhang; Phytotaxa 450(1): 115 (2020)
Distribution: Thailand
41 THA
species :
Polystichum neocavernicola
Y.Y.Liang & Li Bing Zhang; Phytotaxa 559(1): 95-97 (2022)
Distribution: China (Guangxi)
36 CHI
species :
Polystichum neolobatum
Nakai; Bot. Mag. (Tokyo) 39: 118 (1925), nom. nov. for P. lobatum var. chinense Christ
Distribution: China (Anhui, Chongqing, Gansu, Guizhou, Henan, Hubei, Hunan, Jiangxi, ?Ningxia, Shaanxi, Sichuan, Yunnan, Zhejiang); Tibet; Japan; Taiwan; Nepal; Bhutan; India (Himachal Pradesh, Uttarakhand, Sikkim, Darjeeling, Nagaland, Manipur, Arunachal Pradesh); Myanmar
36 CHI TIB 38 JAP TAI 40 BHU IND NEP 41 MYA
Remarks: subgen. Polystichum, sect. Xiphopolystichum
Synonyms: hidden
= Polystichum assurgens Ching & S.K.Wu; Fl. Xizangica 1: 207 (1983)
= Polystichum baoxingense Ching & H.S.Kung; Acta Bot. Boreal.-Occid. Sin. 9(4): 271 (1989) (fide Fraser-Jenkins et al. 2018)
= Polystichum garhwalicum N.C.Nair & Nag; J. Jap. Bot. 51: 138, t. 1 (1976) (fide Fraser-Jenkins et al. 2018)
= Polystichum latilepis Ching & H.S.Kung; Acta Bot. Boreal.-Occid. Sin. 9(4): 273-277 (1989) (fide Fraser-Jenkins et al. 2018)
= Polystichum lobatum var. chinense Christ; Nuovo Giorn. Bot. Ital., n. s. 4: 92 (1897)
= Polystichum neolobatum var. brevipinnum Tagawa; Act. Phytotax. Geobot. 9(2): 92-93 (1940)
= Polystichum rhombiforme Ching & S.K.Wu; Fl. Xizangica 1: 209 (1983) (fide Fraser-Jenkins et al. 2018)
= Polystichum squarrosum var. chinense (Christ) C.Chr.; Acta Horti Gothob. 1(2): 69-70 (1924)
species :
Polystichum neozelandicum
Fée; Mém. Foug. 8 (Icon. Esp. Nouv.): 99 (1857)
subspecies :
subsp. neozelandicum
Name: Common shield fern, Pikopiko, Tutoke (Maori)
Distribution: New Zealand; Chatham Isl.
51 CTM NZL
Remarks: see also P. wawranum and P. oculatum
Synonyms: hidden
= Aspidium richardii Hook.; Sp. Fil. [W. J. Hooker] 4: 23, t. 222 (1862)
= Dryopteris richardii (Hook.) Kuntze; Revis. Gen. Pl. 2: 813 (1891)
= Polystichum richardii (Hook.) J.Sm.; Hist. Fil. 220 (1875)
subspecies :
subsp. zerophyllum
(Colenso) Perrie; New Zealand J. Bot. 41(2): 209 (2003)
Distribution: New Zealand; Chatham Isl.
51 CTM NZL
Synonyms: hidden
= Aspidium zerophyllum Colenso; Trans. & Proc. New Zealand Inst. 29: 418 (1897)
= Polystichum zerophyllum (Colenso) C.Chr.; Index Filic. 98 (1905), 589 (1906)
species :
Polystichum nepalense
(Spreng.) C.Chr.; Index Filic. 84 (1905), 585 (1906)
Distribution: China (Yunnan, Sichuan); Tibet; Taiwan; Nepal; Bhutan; India (Arunachal Pradesh, Himachal Pradesh, Sikkim, Uttarakhand, West Bengal); Myanmar; Vietnam; Philippines
36 CHI TIB 38 TAI 40 BHU IND NEP 41 MYA VIE 42 PHI
Remarks: subgen. Polystichum, Sect. Achroloma
Synonyms: hidden
= Aspidium auriculatum D.Don; Prodr. Fl. Nepal. 3 (1824), nom. illeg.
= Aspidium auriculatum var. marginatum C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 126 (1880)
= Aspidium marginatum Wall. ex Mett.; Abh. Senckenberg. Naturf. Ges. 4: 39 (1858) non (Wall.) C. B. Clarke ex Christ 1905
= Aspidium nepalense Spreng.; Syst. Veg., ed. 16 [Sprengel] 4(1): 97 (1827), nom. nov. for Polypodium auriculatum D. Don
= Polystichum atroviridissimum Hayata; Icon. Pl. Formosan. 4: 190 t 128 (1914)
= Polystichum auriculatum D.Don; Prodr. Fl. Nepal. 3 (1824), non (L.) Sw. 1802
= Polystichum auriculatum var. marginatum Bedd.; Ferns Brit. India 201 (1883)
= Polystichum marginatum (Wall.) Schott; Gen. Fil. (Vindob.) ad t. 9 (1834)
species :
Polystichum ningshenense
Ching & Y.P.Hsu; Fl. Tsinling. 2: 223 (1974)
Distribution: China (Shaanxi)
36 CHI
species :
Polystichum normale
Ching ex P.S.Wang & Li Bing Zhang; Novon 22(2): 250 (2012)
Distribution: China (S-Chongqing, E-Guizhou, NW-Hunan)
36 CHI
species :
Polystichum nudicaule
Rosenst.; Repert. Spec. Nov. Regni Veg. 11: 56 (1912)
Distribution: Ecuador; Peru; Bolivia (Chuquisaca, Cochabamba, La Paz, Potosí, Tarija)
83 BOL ECU PER
Remarks: if synonymy of P. wolfii is confirmed, it would have priority
Synonyms: hidden
= Polystichum montevidense var. nudicaule (Rosenst.) R.M.Tryon; Fieldiana, Bot., n. s. 27: 52 (1991)
= Polystichum nudicaule f. majus Rosenst.; Repert. Spec. Nov. Regni Veg. 11: 56 (1912)
= Polystichum wolfii Hieron.; Hedwigia 46: 356 (1907)
= Polystichum wolfii var. huilensis Hieron.; Hedwigia 46: 357 (1907)
species :
Polystichum obae
Tagawa; Acta Phytotax. Geobot. 2: 194 (1933)
Distribution: Ryukyu Isl. (Amami Oshima)
38 NNS
Remarks: subgen. Polystichum, Sect. Hypopeltis III
Conservation Status: CR in Japan. IUCN E
species :
Polystichum oblanceolatum
H.He & Li Bing Zhang; Novon 22(2): 160, figs. 1-2 (2012)
Distribution: China (N-Guangxi)
36 CHI
species :
Polystichum obliquum
(D.Don) T.Moore; Index Fil. [T. Moore] 87 (1858)
Distribution: China (W-Guangxi, W-Guizhou, W-Sichuan, W-Yunnan); Taiwan; Nepal; Bhutan; India (Jammu & Kashmir, Arunachal Pradesh, Himachal Pradesh, Meghalaya, Sikkim, Uttarakhand, West Bengal); Myanmar; Philippines (Luzon)
36 CHI 38 TAI 40 BHU IND NEP 41 MYA 42 PHI
Synonyms: hidden
= Aspidium auriculatum var. caespitosum (Wall. ex Mett.) C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 507 (1880), as ´caespitosa´
= Aspidium auriculatum var. obliquum (D.Don) Baker; Syn. Fil. [W. J. Hooker & Baker], ed. 2: 493 (1874)
= Aspidium caespitosum Wall.; Numer. List. [Wallich] no. 367 (1829), nomen
= Aspidium caespitosum Wall. ex Mett.; Abh. Senckenberg. Naturf. Ges. 2(2) [Pheg. u. Asp.]: 232 (1858)
= Aspidium obliquum D.Don; Prodr. Fl. Nepal. 3 (1825)
= Hemesteum obliquum (D.Don) H.Lév.; Fl. Kouy-Tcheou 497-498 (1915)
= Polystichum auriculatum var. caespitosum (Wall. ex Mett.) Bedd.; Ferns Brit. India 204 (1883)
= Polystichum caespitosum (Wall. ex Mett.) Bedd.; Ferns Brit. India t. 33 (1865)
species :
Polystichum oblongipinnarum
Li Bing Zhang, M.Q.Han & Yan Liu; Phytotaxa 365(2): 156 (2018)
Distribution: China (Yunnan)
36 CHI
species :
Polystichum oblongum
Ching ex W.M.Chu & Z.R.He; Fl. Reipubl. Popularis Sin. [Fl. China Ed. Committee] 5(2): 228 (2001)
Distribution: China (SW-Yunnan)
36 CHI
species :
Polystichum obtusum
J.Sm. ex C.Presl; Abh. Königl. Böhm. Ges. Wiss. 5(6): 413 (1849)
Distribution: Borneo (Mt. Kinabalu, Sabah); Philippines (Luzon)
42 BOR PHI
Synonyms: hidden
= Aspidium obtusum (J.Sm. ex C.Presl) Kunze ex Mett.; Abh. Senckenberg. Naturf. Ges. 2(2) [Pheg. u. Asp.]: 335 (1858), non (Spreng.) Sw.
= Dryopteris obtusa (J.Sm. ex C.Presl) Kuntze; Revis. Gen. Pl. 2: 813 (1891)
= Polystichum edanoi Copel.; Philipp. J. Sci. 83: 97, t. 2 (1954)
= Polystichum obtusum J.Sm.; J. Bot. (Hooker) 3: 412 (1841), nomen
= Polystichum obtusum var. densum Alderw.; (1911)
species :
Polystichum oculatum
Armstr.; Trans. & Proc. New Zealand Inst. 13: 364 (1881)
Name: Common shield fern, Pikopiko, Tutoke (Maori)
Distribution: New Zealand (North Isl., South Isl., Stewart Isl., Three Kings Isl.); Chatham Isl.
51 CTM NZL
Remarks: subgen. Polystichum, Australasian clade
Synonyms: hidden
= Aspidium coriaceum var. acutidentatum A.Rich.; (1832)
= Aspidium oculatum Hook.; Sp. Fil. [W. J. Hooker] 4: 24, t. 228 (1862)
= Dryopteris oculata (Hook.) Kuntze; Revis. Gen. Pl. 2: 813 (1891)
= Polystichum aristatum Hook.f.; Bot. Antarct. Voy. II (Fl. Nov.-Zel.) 1: t. 78 (1854) (non C. Presl (1836))
species :
Polystichum ohmurae
Sa.Kurata; Hokuriku J. Bot. 5: 79 (1956)
Distribution: Japan
38 JAP
Conservation Status: IUCN R
variant :
var. fujipedis
Nameg. & Sa.Kurata; Sci. Rep. Yokosuka City Mus. 15: 46 (1969)
Distribution: Japan
38 JAP
species :
Polystichum orbiculatum
(Desv.) J.Rémy & Fée; Fl. Chil. [Gay] 6: 515 (1853)
Distribution: Costa Rica; Colombia; Ecuador; Peru; Bolivia (Cochabamba, La Paz)
80 COS 82 CLM 83 BOL ECU PER
Remarks: subgen. Polystichum, sect. Polystichum. P. polyphyllum, P. sodiroi and P. pycnolepis have been recently separated. Synonymy and distribution uncertain.
Synonyms: hidden
= Aspidium orbiculatum Desv.; Mag. Neuesten Entdeck. Gesammten Naturk. Ges. Naturf. Freunde Berlin 5: 321 (1811)
= Phegopteris saxatilis (Klotzsch) Mett. ex Salomon; Nomencl. Gefäßkrypt. 278 (1883)
= Polypodium saxatile Klotzsch; Linnaea 2: 383 (1847)
= Polystichum orbiculare (Desv.) Christ; Ark. Bot. 4(12): 3 (1905)
= Polystichum orbiculatum (Desv.) Gay; Fl. Chil. [Gay] 6: 515 (1853), isonym
= Polystichum orbiculatum var. saxatile (Klotzsch) Hieron.; Hedwigia 46: 358 (1907)
species :
Polystichum ordinatum
(Kunze) Liebm.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd. 5, 1: 276 (1849)
Distribution: Mexico (Chiapas, Hidalgo, Oaxaca, Puebla, Veracruz); Guatemala; Honduras
79 MEX 80 GUA HON
Synonyms: hidden
= Aspidium ordinatum Kunze; Linnaea 18: 347 (1844)
= Polystichum bicolor A.R.Sm.; Proc. Calif. Acad. Sci. ser. 4, 40: 223 (1975)
species :
Polystichum oreodoxa
Ching ex H.S.Kung & Li Bing Zhang; Acta Phytotax. Sin. 33: 309 (1995)
Distribution: China (W-Yunnan)
36 CHI
species :
Polystichum otomasui
Sa.Kurata; J. Geobot. 12: 67 (1963)
Distribution: China (Fujian, Jiangxi); Japan (Kyushu)
36 CHI 38 JAP
Remarks: subgen. Polystichum, Sect. Hypopeltis
Conservation Status: IUCN R
species :
Polystichum otophorum
(Franch.) Bedd.; Suppl. Ferns Brit. Ind. 42 (1892)
Distribution: China (W-Sichuan); E-Tibet
36 CHI TIB
Remarks: subgen. Polystichum, sect. Xiphopolystichum
Synonyms: hidden
= Aspidium otophorum Franch.; Nouv. Arch. Mus. Hist. Nat., sér. 2, 10: 116 (1887)
species :
Polystichum ovatopaleaceum
(Kodama) Sa.Kurata; Sci. Rep. Yokosuka City Mus. 10: 35 (1964)
Distribution: China (Anhui); South Korea (Cheju Isl.); Japan
36 CHI 38 JAP KOR
Synonyms: hidden
= Polystichum aculeatum var. ovatopaleaceum Kodama; Bot. Mag. (Tokyo) 29: 323 (1915)
= Polystichum retrosopaleaceum var. ovatopaleaceum (Kodama) Tagawa; J. Jap. Bot. 13(3): 187 (1937)
variant :
var. coraiense
(Christ) Sa.Kurata; Sci. Rep. Yokosuka City Mus. 10: 35 (1964)
Distribution: China; South Korea (incl. Cheju Isl.); Japan
36 CHI 38 JAP KOR
Synonyms: hidden
= Aetopteron aculeatum var. coraiense (Christ) Nakai; Bull. Natl. Sci. Mus., Tokyo, 31: 4 (1952)
= Polystichum aculeatum var. coraiense Christ; Repert. Spec. Nov. Regni Veg. 5: 11 (1908)
= Polystichum retrosopaleaceum var. coraiense (Christ) Tagawa; J. Jap. Bot. 13: 187 (1937)
species :
Polystichum paleatum
Copel.; Occas. Pap. Bernice P. Bishop Mus. 14: 58, t. 9 (1938)
Distribution: Society Isl. (Tahiti)
61 SCI
species :
Polystichum palniense
Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 333 (2008)
Distribution: S-India (Kerala, Tamil Nadu: Palni Hills)
40 IND
Conservation Status: IUCN NT for India (Fraser-Jenkins 2012)
species :
Polystichum papuanum
C.Chr.; Brittonia 2: 299 (1937)
Distribution: New Guinea (Irian Jaya, Papua New Guinea)
42 NWG
species :
Polystichum paradeltodon
L.L.Xiang; Acta Phytotax. Sin. 32(3): 265 (1994)
Distribution: China (S-Yunnan)
36 CHI
species :
Polystichum paramartini
R.H.Jiang & X.C.Zhang; Turczaninowia 25(2): 87-88 (2022)
Distribution: China (Guizhou)
36 CHI
species :
Polystichum paramicola
M.Kessler & A.R.Sm.; Brittonia 57(3): 222 (2005)
Distribution: Ecuador; Bolivia (Cochabamba)
83 BOL ECU
Remarks: to be expected in Peru
species :
Polystichum paramoupinense
Ching; Fl. Xizangica 1: 226 (1983)
Distribution: China (W-Sichuan); Tibet (Bomi)
36 CHI TIB
Remarks: included in P. prescottianum by Fraser-Jenkins but kept separate by Flora of China
species :
Polystichum parvifoliolatum
W.M.Chu; Acta Bot. Yunnan., Suppl. 5: 49 (1992)
Distribution: China (Yunnan)
36 CHI
species :
Polystichum parvipinnulum
Tagawa; Acta Phytotax. Geobot. 6: 160 (1937)
Distribution: Taiwan
38 TAI
Remarks: subgen. Polystichum, Sect. Hypopeltis III
Synonyms: hidden
= Polystichum sozanense H.S.Kung & Li Bing Zhang ex Ching; Acta Phytotax. Sin. 33(3): 309 (1995)
species :
Polystichum pauciaculeatum
Bonap.; Notes Ptéridol. 7: 206 (1918)
Distribution: Madagascar
29 MDG
Synonyms: hidden
= Polystichum coursii Tardieu; Mém. Inst. Sci. Madagascar, Sér. B, Biol. Vég. 7: 42 (1956)
species :
Polystichum peishanii
Li Bing Zhang & H.He; Bot. Stud. (Taipei) 50(1): 102 (2009)
Distribution: China (S-Guizhou)
36 CHI
species :
Polystichum perditum
Li Bing Zhang; Phytotaxa 404(6): 222 (2019)
Distribution: China (Guizhou)
36 CHI
species :
Polystichum perpusillum
Li Bing Zhang & H.He; Ann. Bot. Fenn. 49: 68 (2012)
Distribution: China (S-Guizhou)
36 CHI
species :
Polystichum piceopaleaceum
Tagawa; Acta Phytotax. Geobot. 5: 255 (1936)
Distribution: China (Gansu, N-Guizhou, S-Shaanxi, Sichuan, Yunnan); Tibet; Japan; Taiwan; Pakistan (Chitral, Dir, Swat, Pakistani Punjab, Hazara); Nepal; Bhutan; India (Jammu & Kashmir, Arunachal Pradesh, Himachal Pradesh, Manipur, Meghalaya, Nagaland, Sikkim, Uttarakhand, West Bengal); Sri Lanka; Myanmar; Philippines
36 CHI TIB 38 JAP TAI 40 BHU IND NEP PAK SRL 41 MYA 42 PHI
Remarks: subgen. Polystichum, Sect. Hypopeltis
Synonyms: hidden
= Polystichum aculeatum var. fargesii Christ; Mém. Soc. Bot. France 1 [Hist. Acrostich.]: 29 (1905)
= Polystichum bicolor Ching & S.K.Wu; Fl. Xizangica 1: 240 (1983), non A. R. Sm. (1975)
= Polystichum doianum Tagawa; Acta Phytotax. Geobot. 5: 256 (1936)
= Polystichum makinoi var. chuanzangense Ching & S.K.Wu; Fl. Xizangica 1: 239 (1983
= Polystichum orientalitibeticum Ching; Fl. Xizangica 1: 240 (1983) (fide Fraser-Jenkins 1997 et al.)
= Polystichum pulogense Copel.; Philipp. J. Sci. 81: 19 (1952)
= Polystichum setiferum var. fargesii (Christ) C.Chr.; Acta Horti Gothob. 1: 71 (1924)
= Polystichum xizangicum Panigrahi; Pl. Sci. Res. (India) 20(1-2): 43 (1998)
= Polystichum yunnanense var. fargesii (Christ) C.Chr.; Contr. U. S. Natl. Herb. 26: 287 (1931)
species :
Polystichum pilosum
Copel.; J. Arnold Arbor. 30: 439 (1949)
Distribution: Fiji (Viti Levu, Vanua Levu)
60 FIJ
species :
Polystichum pingbianense
Li Bing Zhang, M.Q.Han & Yan Liu; Phytotaxa 365(2): 158 (2018)
Distribution: China (Yunnan)
36 CHI
species :
Polystichum plaschnickianum
(Kunze) T.Moore; Index Fil. [T. Moore] 100 (1858)
Distribution: Jamaica; Hispaniola (Haiti, Dominican Republic)
81 HIS JAM
Synonyms: hidden
= Aspidium plaschnickianum Kunze; Linnaea 28: 229, 302 (1850)
= Dryopteris plaschnickiana (Kunze) Kuntze; Revis. Gen. Pl. 2: 813 (1891)
species :
Polystichum platylepis
Fée; Crypt. Vasc. Brésil 1: 129, t. 41, f. 1 (1869)
Distribution: Brazil (Bahia, Espírito Santo, Minas Gerais, Mato Grosso, Paraná, Rio de Janeiro, Rio Grande do Sul, Santa Catarina, São Paulo)
84 BRA
Synonyms: hidden
= Polystichum aculeolatum Fée; Crypt. Vasc. Brésil 1: 128, t. 38, f. 2 (1869)
= Polystichum giganteum Fée; Crypt. Vasc. Brésil 1: 127, t. 39, f. 2 (1869)
= Polystichum longecuspis Fée; Crypt. Vasc. Brésil 1: 129, t. 41, f. 2 (1869)
= Polystichum tijucense Fée; Crypt. Vasc. Brésil 1: 127, t. 38, f. 1 (1869)
species :
Polystichum platyphyllum
(Willd.) C.Presl; Tent. Pterid. 84 (1836)
Distribution: Mexico (Chiapas, Oaxaca, Veracruz); Honduras; Nicaragua; Panama; Cuba; Jamaica; Hispaniola (Haiti, Dominican Republic); Puerto Rico; Colombia (Antioquia, Boyacá, Cauca, Cesar, Chocó, Cundinamarca, Huila, Magdalena, Meta, Nariño, Putumayo, Quindío, Risaralda, Tolima, Valle); Venezuela (Amazonas, Aragua, Barinas, Cojedes, Venezuela Distrito Federal, Falcon, Lara, Merida, Miranda, Portuguesa, Tachira, Trujillo, Yaracuy); Ecuador; Peru; Bolivia (Beni, Chuquisaca, Cochabamba, La Paz, Santa Cruz, Tarija); Brazil (Espírito Santo, Goiás, Minas Gerais, Mato Grosso do Sul, Mato Grosso, Rio de Janeiro, São Paulo); Paraguay (Guaira, San Pedro); Argentina (Jujuy, Misiones, Salta, Tucuman)
79 MEX 80 HON NIC PAN 81 CUB HIS JAM PUE 82 CLM VEN 83 BOL ECU PER 84 BRA 85 ARG PAR
Remarks: subgen. Polystichum, sect. Polystichum
Synonyms: hidden
= Aspidium aculeatum var. phegopteroideum Baker; Fl. Bras. [Martius] 1(2): 462, tab. 28, fig. 9-12 (1870)
= Aspidium aculeatum var. platyphyllum (Willd.) Griseb.; Fl. Brit. W. I. [Grisebach] 689 (1864)
= Aspidium platyphyllum Willd.; Sp. Pl., ed. 4 [Willdenow] 5(1): 255 (1810)
= Nephrodium platyphyllum (Willd.) Desv.; Mem. Soc. Linn. Paris 6(2): 261 (1827)
= Phegopteris platyphylla (Willd.) Mett.; Abh. Senckenberg. Naturf. Ges. 2(2): [Pheg. u. Asp. 12, no. 9] (1858)
= Phegopteris platyphylla var. phegopteroidea (Baker ex Mart.) Farw.
= Phegopteris polystichiformis Fée; Mém. Foug. 5 (Gen. Filic.): 243 (1852)
= Phegopteris polystichops Wawra; Maxim. Reise 201, t. 102 (1866)
= Polypodium platyphyllum (Willd.) Hook.; Sp. Fil. [W. J. Hooker] 4: 248 (1862) (non Sw. (1806))
= Polypodium polystichoides Klotzsch; Linnaea 20: 383 (1847)
= Polystichum aculeatum var. platyphyllum (Willd.) C.Chr.
= Polystichum hottense C.Chr.; Kongl. Svenska Vetensk. Acad. Handl., ser. 3, 16(2): 41 (1936)
= Polystichum lanosum Fée; Crypt. Vasc. Brésil 1: 126 t. 40 f. 2 (1869)
= Polystichum laxum Wawra; Oesterr. Bot. Z. 13: 143 (1863)
= Polystichum opacum Rosenst.; Hedwigia 46: 112 (1906)
= Polystichum pallidum Gardner; London J. Bot. 1: 547 (1842)
= Polystichum platyphyllum f. mettenii Rosenst.; Hedwigia 46: 112 (1906)
= Polystichum platyphyllum f. portoalegrense Rosenst.; Hedwigia 46: 112 (1906)
= Polystichum polystichiformis (Fée) Maxon; Contr. U. S. Natl. Herb. 18: 35 (1909) [fide Hernandez & Sánchez 2013]
= Polystichum tenue Gilbert; Fern Bull. 8: 63 (1900)
variant :
var. klotzschii
Rosenst. ex Hicken; Revista Mus. La Plata 15: 235 (1908)
Distribution: Brazil (Rio Grande do Sul, Santa Catarina); Uruguay (Tacuarembo); Argentina (Misiones)
84 BRA 85 ARG URU
variant :
var. kurtziana
Hicken; Anales Soc. Ci. Argent. 62: 169 (1906)
Distribution: Argentina (Corrientes)
85 ARG
species :
Polystichum plicatum
(Poepp. ex Kunze) Hicken ex Hosseus; Trab. Inst. Bot. Farmacol. 33: 9 (1915)
Distribution: Ecuador; Argentina (Buenos Aires, Chubut, Cordoba, Mendoza, Neuquen, Rio Negro, Santa Cruz, Tierra del Fuego); Chile (Coquimbo, Valparaíso, Metropolitana de Santiago, O´Higgins, Maule, Ñuble, Biobío, Araucanía, Los Rios, Los Lagos, Aisén, Magallanes)
83 ECU 85 ARG CLE
Synonyms: hidden
= Aspidium elegans (J.Rémy) J.W.Sturm; Enum. Pl. Vasc. Cr. Chil. 32 (1858) (non Mett. (1858))
= Aspidium fuscatum Willd.; Sp. Pl., ed. 4 [Willdenow] 5(1): 250 (1810)
= Aspidium plicatum Poepp. ex Kunze; Linnaea 9: 94 (1834)
= Aspidium plicatum var. laxum Kunze; Linnaea 9: 94 (1834 (1835))
= Aspidium plicatum var. rigidum Kunze; Linnaea 9: 94 (1834 (1835))
= Polystichum elegans J.Rémy ex Fée; Mém. Foug. 5 (Gen. Filic.): 278 (1852)
= Polystichum mohrioides f. latifolium Hicken; Anales Soc. Ci. Argent. 62: 171 (1906)
= Polystichum mohrioides var. elegans (J.Rémy) C.Chr.; Ark. Bot. 10, no. 2: 17 (1910)
= Polystichum mohrioides var. plicatum (Poepp. ex Kunze) C.Chr.; Ark. Bot. 10, no. 2: 17 (1910)
species :
Polystichum polyblepharon
(Roem. ex Kunze) C.Presl; Abh. Königl. Böhm. Ges. Wiss., ser. 5, 6 [Epim. Bot.]: 56 (1851)
Distribution: China (Jiangsu, Zhejiang); South Korea (incl. Cheju Isl.); Japan
36 CHI 38 JAP KOR
Remarks: subgen. Polystichum, Sect. Hypopeltis III
Synonyms: hidden
= Aetopteron polyblepharon (Roem. ex Kunze) Nakai; Bull. Natl. Sci. Mus., Tokyo, 33: 4 (1953)
= Aspidium aculeatum var. japonicum Franch. & Sav.; Enum. Pl. Jap. 2: 231 (1877)
= Aspidium polyblepharon Roem. ex Kunze; Bot. Zeitung (Berlin) 572 (1848)
= Polystichum aculeatum var. japonicum (Franch. & Sav.) Diels; Nat. Pflanzenfam. [Engler & Prantl] I, 4: 191 (1899)
= Polystichum polyblepharon f. scabiosum (Sa.Kurata) Seriz.; J. Phytogeogr. Taxon. 29: 24 (1981)
= Polystichum polyblepharon var. scabiosum Sa.Kurata; J. Geobot. 11: 38 (1962)
species :
Polystichum polyodon
Ching; Bull. Chin. Bot. Soc. 2: 93 (1936)
Distribution: ?SW-China; NE-India (Arunachal Pradesh, Manipur, Meghalaya, Mizoram); ?Myanmar
40 IND
Remarks: generic placement fide Fraser-Jenkins et al. 2018
Conservation Status: IUCN NT for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Arachniodes polyodon (Ching) Mazumdar; Phytotaxa 159(1): 26 (2014)
= Phanerophlebiopsis polyodon (Ching) Fraser-Jenk.; Bull. Nat. Mus. Nat. Sci., Ser. B, 38(3), 93-119 (2012)
species :
Polystichum polyphyllum
(C.Presl) C.Presl; Tent. Pterid. 83 (1836)
Distribution: Mexico (Chiapas); Guatemala; Costa Rica; Colombia (Antioquia, Arauca, Boyacá, Caldas, Cauca, Cesar, Cundinamarca, Magdalena, Meta, Nariño, Norte de Santander, Risaralda, Santander, Tolima, Valle); Ecuador; Peru; Bolivia (Cochabamba, La Paz)
79 MEX 80 COS GUA 82 CLM 83 BOL ECU PER
Remarks: subgen. Polystichum, sect. Polystichum. Valid species separate from P. orbiculatum fide McHenry & Barrington 2014. Synonymy provisional - some synonyms might belong to P. orbiculatum
Synonyms: hidden
= Aspidium polyphyllum (C.Presl) Mett.; Fil. Lechl. 1: 20 (1856)
= Aspidium trapezoides (C.Presl) Kunze; Linnaea 9: 93 (1834)
= Jamesonia paleacea Kunze; Bot. Zeitung (Berlin) 739 (1844)
= Nephrodium polyphyllum C.Presl; Reliq. Haenk. 1: 37 (1825)
= Nephrodium trapezoides C.Presl; Reliq. Haenk. 1: 37 (1825)
= Nephrodium trapezoides var. minus C.Presl; Reliq. Haenk. 1: 37 (1825)
= Phegopteris rigida Mett.; Abh. Senckenberg. Naturf. Ges. 2(2): [Pheg. u. Asp. 10, no. 5] (1858)
= Polypodium crenato-dentatum Klotzsch; Linnaea 20: 384 (1847)
= Polypodium rigidum Hook. & Grev.; Icon. Filic. 2: t. 163 (1830), nom. illeg.
= Polypodium rigidum var. majus Hook.; Sp. Fil. [W. J. Hooker] 4: 247 (1862)
= Polypodium rigidum var. polyphyllum (C.Presl) Hook.; Sp. Fil. [W. J. Hooker] 4: 247 (1862)
= Polypodium rigidum var. spruceanum Hook.; Sp. Fil. [W. J. Hooker] 4: 247 (1862)
= Polypodium rigidum var. vulgare Hook.; Sp. Fil. [W. J. Hooker] 4: 246 (1862)
= Polystichum haenkeanum C.Presl; Tent. Pterid. 83 (1836)
= Polystichum orbiculatum var. crenato-dentatum (Klotzsch) Hieron.; Hedwigia 46: 357 (1907)
= Polystichum orbiculatum var. trapezoides (C.Presl) Hieron.; Hedwigia 46: 357 (1907)
= Polystichum pygmaeum Copel.; Univ. Calif. Publ. Bot. 19: 300, t. 56 (1941)
= Polystichum rigidum (Mett.) C.Chr.; Ark. Bot. 20 A: 13 (1926)
species :
Polystichum prescottianum
(Wall. ex Mett.) T.Moore; Index Fil. [T. Moore] 101 (1858)
Distribution: E-Afghanistan; China (S-Gansu, Hubei, Shaanxi, W-Sichuan, NW-Yunnan); Tibet; Taiwan; Pakistan (Kurram, Swat, Chitral, Hazara, Baltistan); Nepal; Bhutan; India (Jammu & Kashmir, Garhwal, Himachal Pradesh, Uttarakhand, Sikkim, Darjeeling); Myanmar (Kachin, etc.)
34 AFG 36 CHI TIB 38 TAI 40 BHU IND NEP PAK 41 MYA
Remarks: subgen. Polystichum, Sect. Sorolepidium. Taxonomy following Fraser-Jenkins et al. (2018)
Synonyms: hidden
= Aspidium moupinense Franch.; Nouv. Arch. Mus. Hist. Nat., sér. 2, 10: 115 (1887)
= Aspidium prescottianum var. castaneum C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 509 (1880)
= Aspidium prescottianum Wall.; Numer. List. [Wallich] no. 363 (1829), nomen
= Aspidium prescottianum Wall. ex Mett.; Abh. Senckenberg. Naturf. Ges. 2(2) [Pheg. u. Asp.]: 48 (1858)
= Dryopteris prescottiana (Wall. ex Mett.) Kuntze; Revis. Gen. Pl. 2: 813 (1891)
= Polystichum aculeatum var. castaneum (C.B.Clarke) Bedd.; (1883)
= Polystichum bomiense Ching & S.K.Wu; Fl. Xizangica 1: 223 (1983) (fide Fraser-Jenkins et al. 2018)
= Polystichum brunneum Ching & S.K.Wu; Fl. Xizangica 1: 224 (1983) (fide Fraser-Jenkins et al. 2018)
= Polystichum castaneum (C.B.Clarke) B.K.Nayar & Kaur; Companion Ferns Brit. India 50 (1883)
= Polystichum erinaceum Ching & S.K.Wu; Fl. Xizangica 1: 221 (1983)
= Polystichum moupinense (Franch.) Bedd.; Suppl. Ferns Brit. Ind. 42 (1892)
= Polystichum moupinense (Franch.) Christ; Mém. Soc. Bot. France 1 [Hist. Acrostich.]: 30 (1905)
= Polystichum prescottianum f. castaneum (C.B.Clarke) Kholia; Ferns Fern Allies Sikkim 2: 78 (2014)
= Polystichum prescottianum var. castaneum (C.B.Clarke) Bedd.; Ferns Brit. India 210 (1883)
= Polystichum prescottianum var. moupinense (Franch.) C.Chr.; Acta Horti Gothob. 1: 70, 169 (1924)
= Polystichum pseudocastaneum Ching & S.K.Wu; Fl. Xizangica 1: 222 (1983) (fide Fraser-Jenkins et al. 2018)
= Polystichum salwinense Ching & H.S.Kung; Bull. Bot. Res., Harbin 9(3): 7 (1989) (fide Fraser-Jenkins et al. 2018)
= Polystichum yadongense Ching & S.K.Wu; Fl. Xizangica 1: 220 (1983) (fide Fraser-Jenkins et al. 2018)
species :
Polystichum prionolepis
Hayata; Icon. Pl. Formosan. 4: 197, f. 134 (1914)
Distribution: China (SC-Yunnan); Taiwan; Vietnam
36 CHI 38 TAI 41 VIE
Synonyms: hidden
= Polystichum rectipinnum Hayata; Icon. Pl. Formosan. 4: 199 t 136 (1914)
= Polystichum lentum var. gelidum Rosenst.; Hedwigia 56(5): 339 (1915)
species :
Polystichum proliferum
(R.Br.) C.Presl; Tent. Pterid. 83 (1836)
Name: Mother shield fern
Distribution: Australia (E-New South Wales, Australian Capital Territory, S-Victoria, Tasmania); New Zealand [I]
50 AUS 51 nzl
Remarks: subgen. Polystichum, Australasian clade
Synonyms: hidden
= Aspidium proliferum R.Br.; Prodr. Fl. Nov. Holland. 147 (1810)
= Aspidium radicans (C.Presl) Sieber; Exsicc. (Fl. Mixta) no. 104 (1827)
= Hypopeltis prolifera (R.Br.) Bory; Voy. Indes Or. [Bélanger] 2: 67 (1833)
= Polystichum aculeatum var. proliferum (R.Br.) Domin; Biblioth. Bot. 20: 54 (1914)
= Polystichum radicans C.Presl; Tent. Pterid. 83 (1836)
species :
Polystichum pseudoacutidens
Ching ex W.M.Chu & Z.R.He; Acta Bot. Yunnan. 22(3): 257, f. 2: 3, 4 (2000)
Distribution: China (NW-Yunnan)
36 CHI
species :
Polystichum pseudodangii
Li Bing Zhang, N.T.Lu & Liang Zhang; Phytotaxa 607(1): 86 (2023)
Distribution: Vietnam
41 VIE
species :
Polystichum pseudodeltodon
Tagawa; Acta Phytotax. Geobot. 6: 159 (1937)
Distribution: Taiwan
38 TAI
Remarks: separate from P. deltodon fide Wan et al. 2023
Synonyms: hidden
= Polystichum deltodon var. pseudodeltodon (Tagawa) Tagawa; Acta Phytotax. Geobot. 9(3): 136 (1940)
species :
Polystichum pseudolanceolatum
Ching ex P.S.Wang; Pterid. Fl. Guizhou 548.f. 130: 3, 4 (2001)
Distribution: China (C-Guizhou)
36 CHI
species :
Polystichum pseudomakinoi
Tagawa; Acta Phytotax. Geobot. 5: 257 (1936)
Distribution: China (Anhui, Guangxi, Guizhou, Henan, Hunan, Jiangsu, Jiangxi, Sichuan, Zhejiang); South Korea; Japan
36 CHI 38 JAP KOR
Remarks: subgen. Polystichum, Sect. Hypopeltis II
species :
Polystichum pseudomicrophyllum
Nakaike; Bull. Natl. Sci. Mus., Tokyo, B, 9(3): 103 (1983)
Distribution: New Guinea (Papua New Guinea)
42 NWG
species :
Polystichum pseudosetosum
Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(4): 19 (1984)
Distribution: China (Chongqing)
36 CHI
species :
Polystichum pseudostenophyllum
Tagawa; Acta Phytotax. Geobot. 3: 92 (1934)
Distribution: Taiwan
38 TAI
Remarks: included by Flora of China and Fraser-Jenkins et al. (2018) in P. stenophyllum but kept separate by Knapp (Ferns of Taiwan, 2011 ff.) and Wan et al. 2023
Synonyms: hidden
= Polystichum niitakayamense Hayata; J. Coll. Sci. Imp. Univ. Tokyo 22: 243 (1906), nomen, et Bot. Mag. (Tokyo) 21: 14 (1907)
species :
Polystichum pseudotsus-simense
Ching; Bull. Fan Mem. Inst. Biol. Bot. 2: 190 (1931)
Distribution: SW-China; Nepal; Bhutan; NE-India (Arunachal Pradesh, Assam State, Manipur, Meghalaya, Mizoram, Nagaland, Sikkim, West Bengal); Myanmar (Chin, etc.); SW-Thailand; Vietnam
36 CHI 40 BHU IND NEP 41 MYA THA VIE
Synonyms: hidden
= Aspidium affine Wall.; Numer. List. [Wallich] no. 370 (1829), nomen
= Aspidium affine Wall. ex Mett.; Abh. Senckenberg. Naturf. Ges. 2(2) [Pheg. u. Asp.]: 104 (1857), nom. illeg.
= Polystichum affine Wall. ex C.Presl; Tent. Pterid. 83 (1836), nom. nud., non Ledeb. 1853
species :
Polystichum punctiferum
C.Chr.; Contr. U. S. Natl. Herb. 26: 288, t. 17 (1931)
Distribution: China (W-Yunnan); Tibet; Nepal; Bhutan; NE-India (Arunachal Pradesh, Sikkim, West Bengal); Myanmar
36 CHI TIB 40 BHU IND NEP 41 MYA
Synonyms: hidden
= Polystichum glingense Ching & Y.X.Lin; Acta Phytotax. Sin. 22(5): 397 (1984)
= Polystichum virescens Ching & S.K.Wu; Fl. Xizangica 1: 243 (1983)
species :
Polystichum pungens
(Kaulf.) C.Presl; Tent. Pterid. 83 (1836)
Distribution: South Africa (Limpopo, Mpulamanga, Free State, KwaZulu-Natal, W-Cape Prov., E-Cape Prov.); Lesotho; Eswatini [Swaziland]
27 LES SAF SWZ
Synonyms: hidden
= Aspidium capense Desv.; Mem. Soc. Linn. Paris 6(2): 260 (1827), nom. illeg.
= Aspidium pungens Kaulf.; Enum. Filic. [Kaulfuss] 242 (1824)
= Dryopteris pungens (Kaulf.) Kuntze; Revis. Gen. Pl. 2: 813 (1891)
species :
Polystichum puteicola
Li Bing Zhang, H.He & Q.Luo; Bot. Stud. (Taipei) 51(1): 128 (2010)
Distribution: China (NW-Guizhou)
36 CHI
species :
Polystichum putuoense
Li Bing Zhang; Phytotaxa 60: 58 (2012), nom. nov.
Distribution: China (Zhejiang)
36 CHI
Remarks: subgen. Haplopolystichum, sect. Cyrtomiopsis
Synonyms: hidden
= Cyrtomidictyum faberi (Baker) Ching; Acta Phytotax. Sin. 6: 259, 265, t. 54 (1957)
= Dryopteris faberi (Baker) C.Chr.; Index Filic. 264 (1905)
= Nephrodium faberi Baker; Ann. Bot. (Oxford) 5: 316 (1891)
species :
Polystichum pycnolepis
(Kunze ex Klotzsch) T.Moore; Index Fil. [T. Moore] 92 (1858)
Distribution: Colombia; Venezuela (Aragua, Cojedes, Venezuela Distrito Federal, Lara, Merida); Ecuador; Galapagos; Peru; Bolivia (Chuquisaca, Cochabamba, La Paz)
82 CLM VEN 83 BOL ECU GAL PER
Remarks: separate species from P. orbiculatum/polyphyllum fide Morero et al. (2016). Priority of name P. pycnolepis over P. cochleatum see Kessler et al. (2023)
Synonyms: hidden
= Aspidium pycnolepis Kunze ex Klotzsch; Linnaea 20: 365 (1847)
= Aspidium robustum Kunze ex Mett.; Fil. Lechl. 1: 20 (1856)
= Phegopteris cochleata (Klotzsch) Mett.; Abh. Senckenberg. Naturf. Ges. 2(2) [Pheg. u. Asp.]: 295 (1858)
= Phegopteris platyphylla var. cochleata (Klotzsch) Farw.; Amer. Midl. Naturalist 12: 293 (1931)
= Phegopteris pycnolepis (Kunze ex Klotzsch) Mett.; Abh. Senckenberg. Naturf. Ges. 2(2): [Pheg. u. Asp. 11, no. 7] (1858)
= Polypodium cochleatum Klotzsch; Linnaea 20: 388 (1847)
= Polypodium pycnolepis (Kunze ex Klotzsch) Hook.; Sp. Fil. [W. J. Hooker] 4: 247 (1862)
= Polypodium pycnolepis var. elatum Hook.; Sp. Fil. [W. J. Hooker] 4: 248 (1862)
= Polystichum cochleatum (Klotzsch) Hieron.; Bot. Jahrb. Syst. 34: 452 (1904)
= Polystichum pflanzii Hieron.; Bot. Jahrb. Syst. 49: 179 (1912)
= Polystichum pycnolepis var. gehrigeri Vareschi; Fl. Venez. 1(1): 403 (1969)
species :
Polystichum pycnopterum
(Christ) Ching ex W.M.Chu & Z.R.He; Fl. Reipubl. Popularis Sin. [Fl. China Ed. Committee] 5(2): 17 (2001)
Distribution: China (SE-Yunnan)
36 CHI
Remarks: subgen. Polystichum, Sect. Macropolystichum
Synonyms: hidden
= Aspidium aculeatum var. pycnopterum Christ; Bull. Herb. Boissier, sér. 1, 7: 14 (1899)
species :
Polystichum quangbinhense
Li Bing Zhang, N.T.Lu & Liang Zhang; Phytotaxa 283(2): 295 (2016)
Distribution: Vietnam
41 VIE
species :
Polystichum rachichlaena
Fée; Mém. Foug. 8 (Icon. Esp. Nouv.): 100 (1857)
Distribution: Mexico (Chiapas, Ciudad de Mexico, Guanajuato, Guerrero, Jalisco, Mexico State, Michoacan, Morelos, Nayarit, Oaxaca, Sinaloa); Guatemala; Honduras
79 MEX 80 GUA HON
Synonyms: hidden
= Polystichum confluens E.Fourn.; Mexic. Pl. 1: 92 (1872)
species :
Polystichum rapense
E.D.Br.; Bull. Bernice P. Bishop Mus. 89 [Fl. S. E. Polynesia II]: 39, t. 4 A (1931)
Distribution: Cook Isl. (Rarotonga); Austral Isl. (Rapa Iti)
61 AUI COO
species :
Polystichum recavum
H.J.Wei & Li Bing Zhang; Phytotaxa 374(2): 167 (2018)
Distribution: China (Guangxi)
36 CHI
species :
Polystichum rectum
Li Bing Zhang, M.Q.Han & Yan Liu; Phytotaxa 365(2): 161 (2018)
Distribution: China (Yunnan)
36 CHI
species :
Polystichum retrosopaleaceum
(Kodama) Tagawa; J. Jap. Bot. 13: 187 (1937)
Distribution: China (Anhui, Hubei, Jiangxi, Zhejiang); South Korea (incl. Ulleung Isl.); Japan
36 CHI 38 JAP KOR
Remarks: subgen. Polystichum, Sect. Hypopeltis III
Synonyms: hidden
= Aetopteron retrosopaleaceum (Kodama) Nakai; Bull. Natl. Sci. Mus., Tokyo, 33: 5 (1953)
= Polystichum aculeatum var. retrosopaleaceum Kodama; Bot. Mag. (Tokyo) 29: 330 (1915)
= Polystichum tsingkanshanense Ching ex K.H.Shing & J.F.Cheng; Jiangxi Sci. 8(3): 46 (1990)
species :
Polystichum revolutum
P.S.Wang; Pterid. Fl. Guizhou 549.f. 133: 6-12 (2001)
Distribution: China (N-Guizhou, Sichuan)
36 CHI
Remarks: subgen. Polystichum, sect. Xiphopolystichum
species :
Polystichum rhizophorum
(Jenman) Maxon; Contr. U. S. Natl. Herb. 18: 36, t. 7 (1909)
Distribution: ?Cuba; Jamaica; Hispaniola (Haiti)
81 HIS JAM
Synonyms: hidden
= Aspidium triangulum subsp. rhizophorum (Jenman) Jenman; Ferns Brit. W. Ind. 198 (1908)
= Aspidium viviparum subsp. rhizophorum Jenman; Bull. Bot. Dept., Jamaica 2: 268 (1895)
species :
Polystichum rhizophyllum
(Sw.) C.Presl; Tent. Pterid. 82 (1836)
subspecies :
subsp. rhizophyllum
Distribution: Jamaica; Puerto Rico; Guadeloupe
81 GDL JAM PUE
Synonyms: hidden
= Aspidium rhizophyllum (Sw.) Sw.; J. Bot. (Schrader) 1800(2): 31 (1801)
= Aspidium rhizophyllum var. plaschnickianum Krug; Bot. Jahrb. Syst. 24: 112 (1897)
= Dryopteris rhizophylla (Sw.) Kuntze; Revis. Gen. Pl. 2: 813 (1891)
= Nephrodium rhizophyllum (Sw.) C.Presl; Reliq. Haenk. 1: 31 (1825)
= Polypodium rhizophyllum Sw.; Prodr. Veg. Ind. Occ. 132 (1788)
= Polystichum krugii Maxon; Proc. Biol. Soc. Wash. 18: 215 (1905)
subspecies :
subsp. cubense
(Mickel) Morejón & C.Sánchez; Willdenowia 43(2): 328 (2013)
Distribution: Cuba
81 CUB
Synonyms: hidden
= Polystichum rhizophyllum var. cubense Mickel; Holttum Mem. [R. J. Johns, ed.] 137 (1997)
species :
Polystichum rigens
Tagawa; Acta Phytotax. Geobot. 6: 91 (1937)
Distribution: China (Chongqing, Gansu, Hubei, Shaanxi); Japan
36 CHI 38 JAP
Remarks: subgen. Polystichum, sect. Xiphopolystichum
Synonyms: hidden
= Polystichum platychlamys Ching; Fl. Tsinling. 2: 224 (1974)
species :
Polystichum robustum
Ching ex Li Bing Zhang & H.S.Kung; Acta Phytotax. Sin. 36(5): 468, pl. 1: 2-5 (1998)
Distribution: China (NE-Yunnan)
36 CHI
species :
Polystichum rochaleanum
Glaz. ex Fée; Crypt. Vasc. Brésil 2: 69, t. 99, f. 2 (1873)
Distribution: Brazil (Espírito Santo, Minas Gerais, Rio de Janeiro)
84 BRA
Synonyms: hidden
= Aspidium rochaleanum (Glaz. ex Fée) Christ; Bull. Herb. Boissier, sér. 2, 2: 558 (1902)
= Polystichum aculeatum var. rochaleanum (Glaz. ex Fée) Luetzelb.; Estud. Bot. Nordéste 3: 244 (1923)
species :
Polystichum rufopaleaceum
Ching ex Li Bing Zhang & H.S.Kung; Acta Phytotax. Sin. 36(3): 246, pl. 1: 1-4 (1998)
Distribution: China (W-Sichuan, W-Yunnan); Tibet
36 CHI TIB
species :
Polystichum rufum
M.Kessler & A.R.Sm.; Brittonia 57(3): 217 (2005)
Distribution: Peru; Bolivia (Cochabamba, La Paz, Santa Cruz)
83 BOL PER
species :
Polystichum rupicola
Ching ex W.M.Chu; Acta Bot. Yunnan., Suppl. 5: 51 (1992)
Distribution: China (NW-Yunnan); ?N-Myanmar
36 CHI
species :
Polystichum sanchezii
Morejón; Willdenowia 42(2): 278 (2012)
Distribution: Cuba
81 CUB
species :
Polystichum saxicola
Ching ex H.S.Kung & Li Bing Zhang; Acta Phytotax. Sin. 36(3): 250, pl. 2: 1-4 (1998)
Distribution: China (Sichuan: Emei Shan)
36 CHI
species :
Polystichum scariosum
(Roxb.) C.V.Morton; Contr. U. S. Natl. Herb. 38: 359 (1974)
Distribution: China (Guangxi, Guizhou, Hainan, Hongkong, S-Hunan, Jiangxi, S-Sichuan, Yunnan, Zhejiang); Tibet; S-Japan (Yakushima Isl.); Nepal; Bhutan; India (Arunachal Pradesh, Manipur, Meghalaya, Mizoram, Nagaland, Sikkim, West Bengal); Myanmar (Kachin, etc.); peninsular Thailand; Vietnam; peninsular Malaysia; Sumatra; Borneo (Mt. Kinabalu, Sabah); Moluccas (Ambon)
36 CHI TIB 38 JAP NNS 40 BHU IND NEP 41 MYA THA VIE 42 BOR MLY MOL SUM
Remarks: subgen. Polystichum, Sect. Macropolystichum
Synonyms: hidden
= Aspidium scariosum Roxb.; Griffith, Calcutta J. Nat. Hist. 4: 494 (1844)
= Polypodium scariosum Roxb.; Calcutta J. Nat. Hist. 4: 494 (1844)
= Polystichum aculeatum var. variiforme Hayata; Icon. Pl. Formosan. 6: 332 (1915)
= Polystichum birii Jamir & R.R.Rao; J. Jap. Bot. 58(12): 376 (1983) (fide Fraser-Jenkins 1997)
= Polystichum eximium var. minus Tagawa; Acta Phytotax. Geobot. 10(4): 292 (1941)
= Polystichum fibrillosum Ching; Bull. Fan Mem. Inst. Biol. Bot. 2: 189, t. 4 (1931) (fide Fraser-Jenkins 2008)
= Polystichum gemmiferum Tagawa; Acta Phytotax. Geobot. 3: 31 (1934)
= Polystichum gemmiparum C.Chr.; Gard. Bull. Straits Settlem. 7: 256, t. 54 (1934) (fide Fraser-Jenkins et al. 208)
= Polystichum indochinense Tardieu & C.Chr.; Notul. Syst. (Paris) 7: 82 (1938)
= Polystichum prolificans Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 3, 2: 170 (1920) (fide Fraser-Jenkins 2018)
= Polystichum setiferum var. crenatum N.C.Nair; Amer. Fern J. 64: 15 (1974) (fide Fraser-Jenkins 1997)
= Polystichum tialooshanense Ching; Acta Phytotax. Sin. 9: 366 (1964)
= Polystichum variiforme (Hayata) Tagawa; J. Jap. Bot. 13: 186 (1937)
species :
Polystichum schizophyllum
Lorea-Hern. & A.R.Sm.; Amer. Fern J. 89: 184 (1999)
Distribution: Mexico (Guerrero)
79 MEX
species :
Polystichum scopulinum
(D.C.Eaton) Maxon; Fern Bull. 8: 29 (1900)
Name: Mountain hollyfern
Distribution: Canada (British Columbia, Newfoundland, Québec); USA (Arizona, California, Colorado, Idaho, Montana, Nevada, Oregon, Utah, Washington State, Wyoming)
71 CDN 72 CDN 73 USA 76 USA
Synonyms: hidden
= Aetopteron scopulinum (D.C.Eaton) House; Amer. Fern J. 10: 88 (1920)
= Aspidium aculeatum var. scopulinum D.C.Eaton; Ferns N. Amer. 2: 125 (1880)
= Polystichum aculeatum var. scopulinum (D.C.Eaton) Gilbert; List N. Amer. Pterid.: 20 (1901)
= Polystichum lonchitis var. scopulinum (D.C.Eaton) M.E.Jones; Bull. Montana State Univ., Biol. Ser. 15: 7 (1910)
= Polystichum mohrioides var. scopulinum (D.C.Eaton) Fernald; Rhodora 26: 89 (1924)
species :
Polystichum semifertile
(C.B.Clarke) Ching; Lingnan Sci. J. 15: 398 (1938)
Distribution: China (Sichuan); Tibet; Nepal; Bhutan; India (Arunachal Pradesh, Manipur, Meghalaya, Mizoram, Nagaland, Sikkim, West Bengal); Myanmar (Kachin, Chin, etc.); N-Thailand; Vietnam
36 CHI TIB 40 BHU IND NEP 41 MYA THA VIE
Remarks: subgen. Polystichum, Sect. Macropolystichum
Synonyms: hidden
= Aspidium aculeatum var. semifertile C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 509 (1880)
= Polystichum aculeatum var. semifertile (C.B.Clarke) Bedd.; Ferns Brit. India 209 (1883)
= Polystichum longipinnulum N.C.Nair; Amer. Fern J. 64: 15 (1974) (fide Fraser-Jenkins (1997), kept separate by Flora of China)
= Polystichum nyalamense Ching; Fl. Xizangica 1: 241 (1983)
= Polystichum yunnanense var. submuticum C.Chr.
species :
Polystichum setiferum
(Forssk.) T.Moore ex Woyn.; Mitt. Naturwiss. Vereines Steiermark 49: 181 (1913)
Name: Borstiger Schildfarn (DE); Penco (ES)
Distribution: England (England, Wales, Scotland, Outer Hebrides, Isle of Man); Ireland (Ireland, Northern Ireland); Luxembourg; Germany (Baden-Württemberg, +Bayern, Hessen, Nordrhein-Westfalen, Rheinland-Pfalz, Saarland); Switzerland; Austria; Poland; Hungary; Portugal; Spain; Baleares; France; Channel Isl. (Alderney, Guernsey, Jersey); Corsica; Sardinia; Italy; Sicily; San Marino; Slovenia; Croatia; Montenegro; Serbia; Kosovo; North Macedonia; Albania; Bulgaria; European Turkey; Greece (widespread in mainland, incl. Kiklades, Samothraki); Crete (Lefka Ori); Crimea; Morocco; Algeria; Tunisia; Azores (Santa Maria Isl., Sao Miguel Isl., Terceira, Graciosa, Sao Jorge, Pico, Faial, Flores Isl., Corvo Isl.); Madeira (Madeira Isl.); Canary Isl. (Tenerife, La Gomera, La Palma Isl., Hierro); Northern Caucasus (Daghestan); Georgia [Caucasus]; Azerbaijan (incl. Nachichevan); East Aegaean Isl. (Lesvos, Ikaria); Turkey (Anatolia); ?Iraq; Iran; New Zealand [I]
10 GRB IRE 11 AUT GER HUN LUX POL SWI 12 BAL CIS COR FRA POR SAR SPA 13 ALB BUL CRO GRC ITA KOS KRI MNT NMA SIC SLV SMA SRB TUE 14 KRY 20 ALG MOR TUN 21 AZO CNY MDR 33 AZE GEO NCS 34 EAI IRN TUR 51 nzl
Remarks: subgen. Polystichum. Not in India
Synonyms: hidden
= Aspidium aculeatum subsp. angulare (Kit. ex Willd.) Arcang.; Comp. Fl. Ital. [Arcangeli] 810 (1882)
= Aspidium aculeatum subsp. angulare (Willd.) Hook.f.; 465 (1870)
= Aspidium aculeatum subsp. swartzianum (W.D.J.Koch) Celak.; 72 (1887)
= Aspidium aculeatum var. hastulatum (Ten.) Christ; Bot. Jahrb. Syst. 24(1): 113 (1897), isonym
= Aspidium aculeatum var. hastulatum (Ten.) Kunze; Flora 31: 366 (1848)
= Aspidium aculeatum var. microlobum Christ; Farnkr. Schweiz 1: 123 (1900)
= Aspidium aculeatum var. swartzianum W.D.J.Koch; Syn. Fl. Germ. Helv., ed. 2, 3: 976 (1845)
= Aspidium angulare Kit. ex Willd.; Sp. Pl., ed. 4 [Willdenow] 5(1): 257 (1810) (p. p.)
= Aspidium hastulatum Ten.; Atti Real Ist. Incoragg. Sci. Nat. Napoli 5: 149, tab. 4, fig. 7 (1830)
= Aspidium lobatum var. angulare (Kit. ex Willd.) Mett.; Fil. Hort. Bot. Lips. 88 (1856)
= Dryopteris aculeata subsp. angularis (Kit. ex Willd.) Schinz & R.Keller; Fl. Suisse 7 (1908)
= Dryopteris aculeata subsp. angularis (Kit. ex Willd.) Schinz & Thell.; Fl. Schweiz (Schinz), ed. 3, 2: 3 (1914), isonym
= Dryopteris setifera (Forssk.) Woyn. ex Schinz & Thell.; Vierteljahrsschr. Naturf. Ges. Zürich 60: 340 (1915)
= Dryopteris setifera subsp. angularis (Kit. ex Willd.) Maire; Cat. Pl. Maroc [Jahandiez & Maire] 3 (1931)
= Hypopeltis hastulata (Ten.) Tod.; Giorn. Sci. Nat. Econ. Palermo 1: 237 (1866)
= Polypodium angulare (Kit. ex Willd.) Fr.; Novit. Fl. Suec. Mant. 1(1-3): 20 (1832)
= Polypodium setiferum Forssk.; Fl. Aegypt.-Arab. 185 (1775)
= Polystichum aculeatum subsp. angulare (Kit. ex Willd.) Vollm.; Fl. Bayern: 9 (1914)
= Polystichum aculeatum subsp. setiferum (Forssk.) Nordh.; 12 (1940)
= Polystichum aculeatum var. angulare (Kit. ex Willd.) Rupr.; Distr. Crypt. Vasc. Ross. 38 (1845)
= Polystichum aculeatum var. angulare C.Presl ex Bedd.; Ferns S. India t. 121 (1865)
= Polystichum angulare (Kit. ex Willd.) C.Presl; Tent. Pterid. 83 (1836)
= Polystichum angulare var. lineare J.Dix; Proc. Roy. Hort. Soc. London 1: 293 (1860)
= Polystichum setiferum (Forssk.) Rosend.; Ark. Bot. 14(14) 1 c. tab. (1916)
= Polystichum setiferum var. hastulatum (Ten.) Hayek; Repert. Spec. Nov. Regni Veg. Beih. 30(1): 29 (1927)
species :
Polystichum setigerum
(C.Presl) C.Presl; Tent. Pterid. 83 (1836)
Name: Alaska Sword Fern
Distribution: Aleutes (Attu Isl.); S-Alaska; C-Canada (British Columbia, Vancouver Isl.)
70 ASK 71 CDN
Conservation Status: IUCN E (Alaska), IUCN V (B. C.)
Synonyms: hidden
= Nephrodium setigerum C.Presl; Reliq. Haenk. 1: 37 (1825)
= Polystichum alaskense Maxon; Amer. Fern J. 8: 35 (1918)
= Polystichum braunii subsp. alaskense (Maxon) Calder & Roy L.Taylor; Canad. J. Bot. 43: 1388 (1965)
= Polystichum braunii var. alaskense (Maxon) Hultén; Acta Univ. Lund., ser. 2, 37(1): 31 (1941)
species :
Polystichum setillosum
Ching; Bull. Fan Mem. Inst. Biol. Bot. 2: 188, t. 3 (1931) (setulosum?)
Distribution: China (W-Sichuan)
36 CHI
species :
Polystichum shandongense
J.X.Li & Y.Wei; Acta Phytotax. Sin. 22(2): 164 (1984)
Distribution: China (Shandong)
36 CHI
Remarks: possibly conspecific with P. craspedosorum
species :
Polystichum shensiense
Christ; Bull. Acad. Int. Géogr., sér. 3, 16: 113 (1906)
Distribution: China (Chongqing, S-Gansu, Shaanxi, W-Sichuan, W-Yunnan); Tibet; Pakistan (Kurram, Hazara); Nepal; Bhutan; India (Jammu & Kashmir, Arunachal Pradesh, Sikkim, Himachal Pradesh, Uttarakhand)
36 CHI TIB 40 BHU IND NEP PAK
Synonyms: hidden
= Dryopteris lichiangensis (C.H.Wright) C.Chr.; Index Filic., Suppl. 2 (1906-1912): 34 (1913)
= Nephrodium lichiangense C.H.Wright; Bull. Misc. Inform. Kew 1909: 267 (1909)
= Polystichum ellipticum Ching & S.K.Wu; Fl. Xizangica 1: 233 (1983)
= Polystichum frigidicola H.S.Kung & Li Bing Zhang; Acta Bot. Yunnan. 17(1): 28 (1995) (fide Fraser-Jenkins et al. 2018)
= Polystichum habaense Ching & H.S.Kung; Bull. Bot. Res., Harbin 9(3): 4 (1989) (fide Fraser-Jenkins et al. 2018)
= Polystichum lichiangense (C.H.Wright) Ching ex H.S.Kung; Bull. Bot. Res., Harbin 9(3): 6 (1989)
= Polystichum macrochlaenum Ching & S.K.Wu; Fl. Xizangica 1: 227 (1983) (fide Fraser-Jenkins et al. 2018)
= Polystichum melanostipes Ching & H.S.Kung; Bull. Bot. Res., Harbin 9(3)- 10 (1989) (fide Fraser-Jenkins et al. 2018)
= Polystichum nigrum Ching & H.S.Kung; Bull. Bot. Res., Harbin 9(3): 8 (1989) (fide Fraser-Jenkins et al. 2018)
= Polystichum obtusipinnum Ching & H.S.Kung; Bull. Bot. Res., Harbin 9(3): 6 (1989) (fide Fraser-Jenkins et al. 2018)
= Polystichum prescottianum var. shensiense (Christ) C.Chr.; Acta Horti Gothob. 1: 70 (1924)
= Polystichum qamdoense Ching & S.K.Wu; Fl. Xizangica 1: 227 (1983) (fide Fraser-Jenkins et al. 2018)
= Polystichum qamdoense var. elongatum Ching & S.K.Wu; Fl. Xizangica 1: 228 (1983)
= Polystichum tibeticum Ching; Fl. Xizangica 1: 219 (1983) (fide Fraser-Jenkins et al. 2018)
= Polystichum tumbatzense Ching; Fl. Xizangica 1: 232 (1983) (fide Fraser-Jenkins et al. 2018)
species :
Polystichum shimurae
Sa.Kurata ex Seriz.; J. Jap. Bot. 54(5): 139 (1979)
Distribution: China (Zhejiang); Japan
36 CHI 38 JAP
Conservation Status: IUCN R
species :
Polystichum silvaticum
(Colenso) Diels; Nat. Pflanzenfam. [Engler & Prantl] I, 4: 192 (1899)
Distribution: New Zealand (North Isl., South Isl.)
51 NZL
Synonyms: hidden
= Polypodium sylvaticum Colenso; Tasmanian J. Nat. Sci. 2: 163 (1844)
= Polystichum aculeatum var. sylvaticum Cheeseman; (1906)
species :
Polystichum simile
(Ching ex Y.T.Hsieh) Li Bing Zhang; Phytotaxa 60: 59 (2012)
Distribution: China (Guizhou, Yunnan)
36 CHI
Remarks: possibly a synonym of P. tenuius
Synonyms: hidden
= Cyrtogonellum simile Ching ex Y.T.Hsieh; Bull. Bot. Res., Harbin 9(3): 16 (1989)
species :
Polystichum sinense
(Christ) Christ; Mém. Soc. Bot. France 1 [Hist. Acrostich.]: 30 (1905)
Distribution: ?Rwanda; South Africa (Mpulamanga, Free State, KwaZulu-Natal, W-Cape Prov., E-Cape Prov.); Lesotho; China (Gansu, Hebei, Nei Mongol, Qinghai, Shaanxi, Shanxi, W-Sichuan, Xinjiang, NW-Yunnan); Tibet; Taiwan; Pakistan (Swat, Hazara); Nepal; Bhutan; India (Jammu & Kashmir, Himachal Pradesh, Uttarakhand, Sikkim, Darjeeling)
27 LES SAF 36 CHI TIB 38 TAI 40 BHU IND NEP PAK
Remarks: subgen. Polystichum, Sect. Hypopeltis. Nomenclature with priority of P. sinense over P. wilsonii as well as synonymy following Fraser-Jenkins (2008, 2018). All African records except South Africa are doubtful (Fraser-Jenkins 2018) and need to be reassessed.
Synonyms: hidden
= Aspidium prescottianum var. sinense Christ; Bull. Soc. Bot. Ital. 10: 289 (1901)
= Polystichum alatawshanicum C.Y.Yang; Fl. Xinjiangensis 1: 302 (1992 publ. 1993)
= Polystichum alticola Schelpe & N.C.Anthony; Contr. Bolus Herb. 10: 144 (1982) (fide Fraser-Jenkins 2008)
= Polystichum atuntzeense Ching; Fl. Xizangica 1: 231 (1983) (fide Fraser-Jenkins 2018)
= Polystichum austrotibeticum Ching & S.K.Wu; Fl. Xizangica 1: 233 (1983)
= Polystichum decorum Ching & S.K.Wu; Fl. Xizangica 1: 229 (1983) (fide Fraser-Jenkins 2018)
= Polystichum fukuyamae Tagawa; Acta Phytotax. Geobot. 6: 159 (1937)
= Polystichum gongboense Ching & S.K.Wu; Fl. Xizangica 1: 228 (1983) (fide Fraser-Jenkins 2018)
= Polystichum lhasaense Ching; Fl. Xizangica 1: 230 (1983) (fide Fraser-Jenkins 2018)
= Polystichum mediocre Ching & S.K.Wu; Fl. Xizangica 1: 235, t. 57, f. 1-4 (1983) (fide Fraser-Jenkins et al. 2018)
= Polystichum mollissimum Ching; Fl. Xizangica 1: 232 (1983) (fide Fraser-Jenkins et al. 2018)
= Polystichum mollissimum var. laciniatum H.S.Kung & Li Bing Zhang; Acta Phytotax. Sin. 36(3): 250 (1998)
= Polystichum parasinense Chang Y.Yang; Fl. Xinjiangensis 1: 302 (1993)
= Polystichum prescottianum var. sinense Christ; Bull. Soc. Bot. Ital. 1901: 289 (1901)
= Polystichum prescottianum var. wilsonii (Christ) C.Chr.; Acta Horti Gothob.1: 71 (1924)
= Polystichum rarum Ching & S.K.Wu; Fl. Xizangica 1: 230, t. 56, f. 5-8 (1983) (fide Fraser-Jenkins 2018)
= Polystichum sinense var. lobatum H.S.Kung; Acta Phytotax. Sin. 36(3): 249 (1998)
= Polystichum wilsonii Christ; Bot. Gaz. 61: 353 (1911)
species :
Polystichum sinotsus-simense
Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(4): 18 (1984)
Distribution: China (Chongqing, C-Guizhou, W-Hunan)
36 CHI
Remarks: subgen. Polystichum, sect. Xiphopolystichum
species :
Polystichum sodiroi
Christ; Ark. Bot. 4(12): 4 (1905)
Distribution: Ecuador; ?Bolivia; etc.
83 ECU
Remarks: subgen. Polystichum, sect. Polystichum. Valid species separate from P. orbiculatum / polyphyllum fide McHenry & Barrington (2014)
species :
Polystichum solomonii
M.Kessler & A.R.Sm.; Brittonia 57(3): 220 (2005)
Distribution: Venezuela; Peru; Bolivia (Cochabamba, La Paz, Santa Cruz)
82 VEN 83 BOL PER
species :
Polystichum speciosissimum
(A.Braun ex Kunze) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 109 (1947)
Distribution: Mexico (Chiapas, Colima, Ciudad de Mexico, Guanajuato, Guerrero, Hidalgo, Jalisco, Mexico State, Michoacan, Morelos, Oaxaca, Puebla, Queretaro, San Luis Potosi, Tlaxcala, Veracruz); Guatemala; Costa Rica; Panama
79 MEX 80 COS GUA PAN
Remarks: subgen. Polystichum, sect. Polystichum
Synonyms: hidden
= Cheilanthes scariosa M.Martens & Galeotti; Nouv. Mém. Acad. Roy. Sci. Bruxelles 15 (Mém. Lycopod.): 74 (1842)
= Cheilanthes speciosissima A.Braun ex Kunze; Analecta Pteridogr. 35 (1837)
= Cheilanthes speciosissima var. elata Liebm.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 5, 1: 106 [reprint page] (1849)
= Phegopteris speciosissima (A.Braun ex Kunze) Mett.; Abh. Senckenberg. Naturf. Ges. 3(1) [Aspl.]: 4 in nota (1859)
= Plecosorus mexicanus Fée; Mém. Foug. 5 (Gen. Filic.): 151, t. 13 (1852)
= Plecosorus speciosissimus (A.Braun ex Kunze) T.Moore; Index Fil. [T. Moore] 40 (1857)
= Polystichum speciosissimum (A.Braun ex Kunze) R.M.Tryon & A.F.Tryon; Rhodora 83(833): 136 (1981)
species :
Polystichum speluncicola
Li Bing Zhang & H.He; Syst. Bot. 35(1): 13 (2010)
Distribution: China (S-Guizhou)
36 CHI
species :
Polystichum spongiosum
Maxon; Amer. Fern J. 19: 47 (1929)
Distribution: Hispaniola (Haiti)
81 HIS
species :
Polystichum squarrosum
(D.Don) Fée; Mém. Foug. 5 (Gen. Filic.): 278 (1852)
Distribution: China (SW-Sichuan); Tibet; Pakistan (Pakistani Punjab, Hazara); Nepal; Bhutan; India (Jammu & Kashmir, Arunachal Pradesh, Assam State, Himachal Pradesh, Karnataka, Madhya Pradesh, Maharashtra, Nagaland, Sikkim, Tamil Nadu, Uttar Pradesh, Uttarakhand, West Bengal); Myanmar (Chin, etc.)
36 CHI TIB 40 BHU IND NEP PAK 41 MYA
Remarks: mostly treated as P. neolobatum in Fl. China fide Fraser-Jenkins (2008)
Synonyms: hidden
= Aspidium aculeatum var. rufibarbatum Wall. ex C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 509 (1880)
= Aspidium brachypterum Kunze; Linnaea 24: 288 (1851)
= Aspidium rufobarbatum Wall.; Numer. List. [Wallich] no. 369 (1829), nomen
= Aspidium squarrosum D.Don; Prodr. Fl. Nepal. 4 (1825)
= Hypopeltis squarrosa (D.Don) Bory; Voy. Indes Or. [Bélanger] 2: 67 (1833)
= Polypodium squarrosum (D.Don) Roxb.; Calcutta J. Nat. Hist. 4: 494 (1844)
= Polystichum aculeatum var. rufibarbatum (Wall. ex C.B.Clarke) Bedd.; Ferns Brit. India 207 (1883)
= Polystichum apicisterile Ching & S.K.Wu; Fl. Xizangica 1: 212 (1983)
= Polystichum brachypterum (Kunze) Ching; Fl. Xizangica 1: 209 (1983)
= Polystichum integripinnulum Ching; Fl. Xizangica 1: 210 (1983)
= Polystichum lobatum var. squarrosum (D.Don) C.Chr.; Index Filic. 583 (1906)
= Polystichum meiguense Ching & H.S.Kung; Acta Bot. Boreal.-Occid. Sin. 9(4): 269 (1989) (fide Fraser-Jenkins et al. 2018)
= Polystichum rufobarbatum (Wall. ex C.B.Clarke) Schott; Gen. Fil. (Vindob.) ad t. 9 (1834)
= Polystichum rufobarbatum (Wall. ex C.B.Clarke) Schott ex Diels; Nat. Pflanzenfam. [Engler & Prantl] I, 4: 191 (1902)
= Polystichum yigongense Ching & S.K.Wu; Fl. Xizangica 1: 210 (1983) (fide Fraser-Jenkins et al. 2018)
species :
Polystichum stimulans
(Kunze ex Mett.) Bedd.; Ferns Brit. India t. 31 (1865)
Distribution: China (Yunnan, Sichuan); Tibet; Nepal; Bhutan; India (Arunachal Pradesh, Himachal Pradesh, Sikkim, Uttarakhand, ?West Bengal)
36 CHI TIB 40 BHU IND NEP
Synonyms: hidden
= Aspidium ilicifolium D.Don; Prodr. Fl. Nepal. 3 (1825)
= Aspidium pungens Wall.; Numer. List. [Wallich] no. 368 (1829), nomen
= Aspidium stimulans Kunze ex Mett.; Abh. Senckenberg. Naturf. Ges. 2(2) [Pheg. u. Asp.]: 327 (1858)
= Dryopteris ilicifolia (D.Don) Kuntze; Revis. Gen. Pl. 2: 813 (1891)
= Polystichum ilicifolium (D.Don) T.Moore; Index Fil. [T. Moore] 94 (1858) (non Fée (1850)
= Polystichum stimulans (Kunze) C.Presl; Tent. Pterid. 83 (1836)
species :
Polystichum stokesii
E.D.Br.; Bull. Bernice P. Bishop Mus. 89 [Fl. S. E. Polynesia II]: 40, t. 5 (1931)
Distribution: Austral Isl. (Rapa Iti)
61 AUI
species :
Polystichum stuebelii
Hieron.; Hedwigia 46: 355, t. 8, f. 21 (1906)
Distribution: Colombia; Ecuador; Peru; Bolivia (Cochabamba, La Paz)
82 CLM 83 BOL ECU PER
species :
Polystichum subacutidens
Ching ex L.L.Xiang; Acta Phytotax. Sin. 32: 261 (1994)
Distribution: China (S-Guizhou, NW-Guangxi, SE-Yunnan); N-Vietnam
36 CHI 41 VIE
Synonyms: hidden
= Polystichum deltodon var. cultratum Christ
species :
Polystichum subapiciflorum
Hayata; Icon. Pl. Formosan. 6: 335, f. 140 (1915)
Distribution: Taiwan
38 TAI
Remarks: subgen. Polystichum, Sect. Macropolystichum. Separate from P. biaristatum fide Wan et al. 2023
species :
Polystichum subdeltodon
Ching; Bull. Fan Mem. Inst. Biol. Bot., ser. 2, 1: 308 (1949)
Distribution: China (Chongqing)
36 CHI
Synonyms: hidden
= Polystichum grossidentatum Ching; Icon. Filic. Sin. 6: t. 239 (1958)
species :
Polystichum subfimbriatum
W.M.Chu & Z.R.He; Acta Phytotax. Sin. 36(4): 382, pl. 1: 1-6 (1998)
Distribution: China (NE-Yunnan)
36 CHI
Remarks: subgen. Polystichum, Sect. Subfimbriata
species :
Polystichum subinerme
(Kunze) Fraser-Jenk.; Bhardwaja & Gena (eds.), Perspect. Pterid. Present & Future: 265 (1991), et in Himalayan Ferns (Guide to Polystichum) 23 (1997)
Distribution: S-India (Kerala, Tamil Nadu, Karnataka)
40 IND
Conservation Status: IUCN EN for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Aspidium subinerme Kunze; Linnaea 24: 290 (1851)
= Polystichum kunthianum B.K.Nayar & Geev.; Fern Fl. Malabar: 214 (1993)
species :
Polystichum subintegerrimum
(Hook. & Arn.) R.Rodr.; Gayana, Bot. 44(1-4): 48 (1987 publ. 1988)
Distribution: Chile (Ñuble, Biobío, Araucanía, Los Rios, Los Lagos, Aisén, Magallanes)
85 CLE
Synonyms: hidden
= Aspidium brongniartianum (J.Rémy) J.W.Sturm; Abh. Naturhist. Ges. Nürnberg 2 [Enum. Pl. Vasc. Cr. Chil.]: 181 (1858)
= Aspidium subintegerrimum Hook. & Arn.; Bot. Beechey Voy. 52 (1832)
= Polystichum aculeatum var. brongniartianum (J.Rémy) C.Chr.; Index Fil. [T. Moore] 576 (1905 publ. 1906)
= Polystichum aculeatum var. subintegerrimum (Hook. & Arn.) C.Chr.; Index Fil. [T. Moore] 576 (1905 publ. 1906)
= Polystichum brongniartianum J.Rémy ex Gay; Fl. Chil. [Gay] 6: 518 (1853)
= Polystichum subintegerrimum (Hook. & Arn.) Barrington; Ann. Missouri Bot. Gard. 76(2): 373 (1989), isonym
species :
Polystichum sublanceolatum
R.H.Jiang & X.C.Zhang; Turczaninowia 25(2): 87-88 (2022)
Distribution: China (Guangxi)
36 CHI
species :
Polystichum submarginale
(Baker) L.L.Xiang ex Ching; Acta Phytotax. Sin. 32(3): 263 (1994)
Distribution: China (Sichuan)
36 CHI
Remarks: subgen. Haplopolystichum, sect. Haplopolystichum
Synonyms: hidden
= Aspidium auriculatum var. submarginale Baker; J. Bot. 26(7): 227 (1888)
= Polystichum deltodon var. submarginale (Baker) C.Chr.; Acta Horti Gothob. 1: 67 (1924)
species :
Polystichum submite
(Christ) Diels; Bot. Jahrb. Syst. 29: 192 (1900)
Distribution: China (Gansu, Henan, Shaanxi, Sichuan)
36 CHI
Synonyms: hidden
= Aspidium submite Christ; Nuovo Giorn. Bot. Ital., n. s. 4: 93 (1897)
= Polystichum paradoxum Ching & Y.P.Hsu; Fl. Tsinling. 2: 224 (1974)
species :
Polystichum submucronatum
(Christ) Morejón & C.Sánchez; Willdenowia 43(2): 328 (2013)
Distribution: Cuba; Jamaica
81 CUB JAM
Synonyms: hidden
= Aspidium triangulum var. submucronatum Christ; Bot. Jahrb. Syst. 24: 111 (1897)
= Polystichum woodsioides Mickel; Holttum Mem. [R. J. Johns, ed.] 142 (1997), nom. illeg., non Christ 1911
species :
Polystichum subobliquum
Tagawa; Acta Phytotax. Geobot. 4: 138 (1935)
Distribution: Taiwan
38 TAI
Remarks: separate from P. obliquum fide Wan et al. 2023
species :
Polystichum subulatum
Ching ex Li Bing Zhang; Acta Bot. Yunnan. 16(2): 133 (1994)
Distribution: China (Sichuan, Guizhou)
36 CHI
species :
Polystichum suiyangense
Bo Wang, Q.W.Sun & Li Bing Zhang; Phytotaxa 567(1): 94 (2022)
Distribution: China (Guizhou)
36 CHI
species :
Polystichum sunhangii
Liang Zhang, Yong L.Qiu & Li Bing Zhang; Phytotaxa 618(1): 21 (2023)
Distribution: Tibet
36 TIB
species :
Polystichum superum
Li Bing Zhang, M.Q.Han & Yan Liu; Phytotaxa 365(2): 164 (2018)
Distribution: China (Yunnan)
36 CHI
species :
Polystichum tagawanum
Sa.Kurata; J. Geobot. 7: 41 (1958)
Distribution: South Korea (incl. Cheju Isl.); Japan
38 JAP KOR
Remarks: subgen. Polystichum, Sect. Hypopeltis II
Synonyms: hidden
= Polystichum pseudomakinoi var. ambiguum Tagawa; Acta Phytotax. Geobot. 5: 258 (1936)
species :
Polystichum taizhongense
H.S.Kung; Acta Phytotax. Sin. 35(5): 461, f. 1 (1997)
Distribution: Taiwan
38 TAI
species :
Polystichum takakii
Nakaike; Bot. Exped. Papua New Guinea, Nation. Sci. Mus. (Tokyo) 111 (1975)
Distribution: New Guinea (Papua New Guinea)
42 NWG
species :
Polystichum talamancanum
Barrington; Ann. Missouri Bot. Gard. 76: 369 (1989)
Distribution: Costa Rica; Panama
80 COS PAN
Remarks: subgen. Polystichum, sect. Polystichum
species :
Polystichum tangmaiense
H.S.Kung & Tateishi; Acta Phytotax. Sin. 32(3): 268 (1994)
Distribution: SW-China (Yunnan); Tibet (Bomi); India (Manipur, Arunachal Pradesh, Nagaland, Mizoram)
36 CHI TIB 40 IND
Conservation Status: IUCN VU for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Polystichum oligocarpum Ching ex H.S.Kung & Li Bing Zhang; Acta Phytotax. Sin. 33(3): 308 (1995) (fide Fraser-Jenkins 2008, 2018)
species :
Polystichum tenuius
(Ching) Li Bing Zhang; Phytotaxa 60: 59 (2012)
Distribution: China (Chongqing, Guangxi, Guizhou, Hunan, Sichuan, Yunnan); Taiwan; N-Vietnam
36 CHI 38 TAI 41 VIE
Remarks: subgen. Haplopolystichum, sect. Cyrtogonellum. New record for Taiwan see Hsu (2014)
Synonyms: hidden
= Cyrtogonellum caducum Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 330, t. 7(3) (1938)
= Cyrtogonellum falcilobum Ching ex Y.T.Hsieh; Bull. Bot. Res., Harbin 9(3): 17 (1989)
= Cyrtogonellum salicifolium Ching & Y.T.Hsieh; Bull. Bot. Res., Harbin 9(3): 16 (1989)
= Cyrtogonellum tenuius Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 329, t. 7(2) (1938)
species :
Polystichum tetragonum
Fée; Mém. Foug. 8 (Icon. Esp. Nouv.): 99 (1857)
Distribution: Juan Fdz. Isl.
85 JNF
Conservation Status: IUCN V
Synonyms: hidden
= Aspidium tetragonum (Fée) J.W.Sturm; Abh. Naturhist. Ges. Nürnberg 2 [Enum. Pl. Vasc. Cr. Chil.]: ? (1858), nom. illeg.
species :
Polystichum thomsonii
(Hook.f.) Bedd.; Ferns Brit. India 1: t. 126 (1865)
Distribution: E-Afghanistan; China (Chongqing, Guizhou, Hubei, Yunnan, Sichuan); E-Tibet; Japan (Honshu); Taiwan; Pakistan (Swat, Hazara); Nepal; Bhutan; India (Jammu & Kashmir, Arunachal Pradesh, Himachal Pradesh, Sikkim, Uttarakhand, West Bengal); Myanmar
34 AFG 36 CHI TIB 38 JAP TAI 40 BHU IND NEP PAK 41 MYA
Remarks: subgen. Polystichum, Sect. Micropolystichum. P. capillipes is sometimes kept separate (in Taiwan, for example)
Synonyms: hidden
= Aspidium capillipes Baker; J. Bot. 26: 228 (1888)
= Aspidium thomsonii Hook.f.; Hook., Sec. Cent. Ferns, t. 25 (1860)
= Aspidium thomsonii var. gracile C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 508 (1880)
= Dryopteris gracilis (T.Moore ex Bedd.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 455 (1938) (non (Heward) Domin, nec Copel.)
= Dryopteris inaensis Tagawa; Acta Phytotax. Geobot. 3: 90 (1934)
= Dryopteris thomsonii (Hook.f.) Kuntze; Revis. Gen. Pl. 2: 813 (1891)
= Hemesteum michelii (Christ) H.Lév.; Fl. Kouy-Tcheou 497-498 (1915)
= Lastrea gracilis T.Moore ex Bedd.; Ferns Brit. India t. 198 (1866)
= Nephrodium sparsum var. gracile (T.Moore ex Bedd.) Baker; Syn. Fil. [W. J. Hooker & Baker], ed. 2: 498 (1874)
= Polystichum capillipes (Baker) Diels; Nat. Pflanzenfam. [Engler & Prantl] I, 4: 191 (1899)
= Polystichum inaense (Tagawa) Tagawa; J. Jap. Bot. 14: 709 (1938)
= Polystichum longidens Ching & S.K.Wu; Fl. Xizangica 1: 217 (1983) (fide Fraser-Jenkins 1997)
= Polystichum michelii Christ; Bull. Acad. Int. Géogr. Bot. 16 (1910)
= Polystichum minusculum Christ; Mém. Soc. Bot. France 1 [Hist. Acrostich.]: 31 (1905)
= Polystichum molliculum Christ; Bot. Gaz. 61: 354 (1911)
= Polystichum silaense Ching; Bull. Fan Mem. Inst. Biol. Bot. 11: 71 (1941)
= Polystichum thomsonii var. gracile Bedd.; Ferns Brit. India 207 (1883)
species :
Polystichum tiandengense
H.He & Li Bing Zhang; Novon 25(2): 153 (2017)
Distribution: China (Guangxi)
36 CHI
species :
Polystichum tiankengicola
Li Bing Zhang, Q.Luo & P.S.Wang; Novon 22(2): 186 (2012)
Distribution: China (NW-Guizhou)
36 CHI
species :
Polystichum tonkinense
(Christ) W.M.Chu & Z.R.He; Fl. Reipubl. Popularis Sin. [Fl. China Ed. Committee] 5(2): 169, pl. 48: 1, 2 (2001)
Distribution: China (Guangxi, Guizhou, Yunnan); N-Vietnam
36 CHI 41 VIE
Remarks: subgen. Haplopolystichum, sect. Sphaenopolystichum
Synonyms: hidden
= Aspidium aculeatum var. tonkinense Christ; Bull. Sci. France Belgique 28: 268, pl. 12: 6 (1898)
species :
Polystichum tortuosum
Nakaike; Bull. Natl. Sci. Mus., Tokyo, B, 2(3): 102 (1976)
Distribution: New Guinea (Papua New Guinea)
42 NWG
species :
Polystichum transkeiense
W.Jacobsen; J. S. African Bot. 44(2): 169 (1978)
Distribution: South Africa (Limpopo, Mpulamanga, KwaZulu-Natal, E-Cape Prov.)
27 SAF
Remarks: subgen. Polystichum, Afra clade
species :
Polystichum transvaalense
N.C.Anthony; Contr. Bolus Herb. 10: 146 (1982)
Distribution: Cameroon; Bioko Isl. [Fernando Poo]; D.R.Congo [Zaire]; Rwanda; South Sudan; Ethiopia; Eritrea; Uganda; Kenya; Tanzania; Zambia; Malawi; E-Zimbabwe; Mozambique; South Africa (Limpopo, Mpulamanga, Free State, KwaZulu-Natal, E-Cape Prov.); Lesotho
23 BIO CMN RWA ZAI 24 ERI ETH SSU 25 KEN TAN UGA 26 MLW MOZ ZAM ZIM 27 LES SAF
species :
Polystichum trapezoides
(Sw.) C.Presl; Tent. Pterid. 83 (1836)
Distribution: Cuba; Jamaica; Hispaniola (Haiti, Dominican Republic)
81 CUB HIS JAM
Synonyms: hidden
= Polypodium trapezoides (C.Presl) Sw.; Fl. Ind. Occid. 3: 1647 (1806)
= Polystichum falcatum Fée; Mém. Foug. 5 (Gen. Filic.): 279 (1852)
= Polystichum killipii Maxon; Contr. U. S. Natl. Herb. 24: 53, t. 20 (1922)
species :
Polystichum trapezoideum
(Ching & K.H.Shing ex K.H.Shing) Li Bing Zhang; Phytotaxa 60: 59 (2012)
Distribution: China (Sichuan, Guangdong)
36 CHI
Synonyms: hidden
= Cyrtomium trapezoideum Ching & K.H.Shing; Acta Phytotax. Sin., Addit. 1: 15 (1965)
species :
Polystichum triangulum
(L.) Fée; Mém. Foug. 5 (Gen. Filic.): 279 (1852)
subspecies :
subsp. triangulum
Distribution: Cuba; Hispaniola (Dominican Republic, Haiti)
81 CUB HIS
Synonyms: hidden
= Aspidium triangulare (J.F.Gmel.) Desv.; Mem. Soc. Linn. Paris 6(2): 247 (1827)
= Aspidium triangulum (L.) Sw.; J. Bot. (Schrader) 1800(2): 31 (1801)
= Aspidium triangulum f. proliferum Christ; Bot. Jahrb. Syst. 24(1): 112 (1897)
= Aspidium triangulum var. laxum Hook.; Fil. Exot.: t. 33 (1858) et Sp. Fil. [W. J. Hooker] 4: 14 (1862)
= Dryopteris triangula (L.) Kuntze; Revis. Gen. Pl. 2: 814 (1891)
= Polypodium triangulare J.F.Gmel.; Syst. Nat., ed. 13[bis] 2(2): 1309 (1791)
= Polypodium triangulum L.; Sp. Pl. 2: 1088 (1753), et Syst. Nat. ed. XIII: 787 (1774)
= Polystichum triangulum var. laxum R.Sim ex J.Dix; Proc. Roy. Hort. Soc. London 1: 244 (1860)
subspecies :
subsp. mucronatum
(Sw.) Morejón & C.Sánchez; Willdenowia 43(2): 326 (2013)
Distribution: Jamaica
81 JAM
Synonyms: hidden
= Aspidium mucronatum Sw.; J. Bot. (Schrader) 1800(2): 30 (1801)
= Aspidium mucronatum var. pinnatifidum Jenman; Bull. Bot. Dept., Jamaica 2: 266 (1895)
= Aspidium triangulum var. bipinnatum (Jenman) Jenman; Ferns Brit. W. Ind.: 198 (1907)
= Aspidium viviparum var. bipinnatum (Jenman) Jenman; Bull. Bot. Dept., Jamaica 2: 268 (1895)
= Dryopteris mucronata (Sw.) Kuntze; Revis. Gen. Pl. 2: 813 (1891)
= Polypodium mucronatum (Sw.) Sw.; Fl. Ind. Occid. 3: 1649 (1806)
= Polystichum mucronatum (Sw.) C.Presl; Tent. Pterid. 83 (1936)
= Polystichum struthionis Maxon; Contr. U. S. Natl. Herb. 13: 37, t. 8 (1909)
species :
Polystichum tridens
(Hook.) Fée; Mém. Foug. 11 (Hist. Foug. Antil.): 74 (1866)
Distribution: Jamaica; Hispaniola (Dominican Republic)
81 HIS JAM
Synonyms: hidden
= Aspidium triangulum var. tridens (T.Moore ex Hook.) Griseb.; Fl. Brit. W. I. [Grisebach] 689 (1864)
= Aspidium tridens T.Moore ex Hook.; Sp. Fil. [W. J. Hooker] 4: 15 (1862)
= Dryopteris tridens (Hook.) Kuntze; Revis. Gen. Pl. 2: 814 (1891)
species :
Polystichum tripteron
(Kunze) C.Presl; Abh. Königl. Böhm. Ges. Wiss., ser. 5, 6: 55 (1851)
Distribution: Russian Far East (Kuril Isl., Primorye, Sakhalin); China (Anhui, Beijing, Fujian, Gansu, Guangdong, Guangxi, Guizhou, Hebei, Heilongjiang, Henan, Hubei, Hunan, Jiangsu, Jiangxi, Jilin, Liaoning, Shaanxi, Shandong, Sichuan, Zhejiang); North Korea; South Korea (incl. Cheju Isl.); Japan (Tsushima Isl.)
31 RFE 36 CHI 38 JAP KOR NKO
Remarks: subgen. Haplopolystichum, sect. Crucifilix
Synonyms: hidden
= Aspidium tripteris D.C.Eaton; Narr. Exped. China Japan [Perry] 2: 330 (1856)
= Aspidium tripteron Kunze; Bot. Zeitung (Berlin) 569 (1848)
= Dryopteris tripteros (Kunze) Kuntze; Revis. Gen. Pl. 2: 814 (1891)
= Polystichum subbipinnatum Kodama ex Nakai; Rep. Veg. Ooryongto 14 (1919), nom. nud.
= Polystichum subtripteron Tzvelev; Novosti Sist. Vyssh. Rast. 26: 7 (1989)
= Polystichum tripteron f. subbipinnatum H.Itô; J. Jap. Bot. 11: 428 (1935)
= Polystichum tripteron var. subpinnatum (H.Itô) Honda; Nom. Pl. Jap., emend.: 15 (1957)
= Ptilopteris tripteros (Kunze) Hayata; Bot. Mag. (Tokyo) 41: 706 (1927)
= Ptilopteris tripteros var. subpinnata (Kodama) Nakai; Bull. Natl. Sci. Mus., Tokyo, 31 (1952)
species :
Polystichum tsaratananense
Tardieu; Mém. Inst. Sci. Madagascar, Sér. B, Biol. Vég. 7: 44 (1956)
Distribution: Madagascar
29 MDG
species :
Polystichum turrialbae
Christ; Bull. Herb. Boissier, sér. 2, 6: 163 (1906)
Distribution: Mexico (Chiapas, Ciudad de Mexico, Hidalgo, Mexico State, Michoacan, Oaxaca, Veracruz); El Salvador; Costa Rica; Panama; Bolivia (Cochabamba, La Paz)
79 MEX 80 COS ELS PAN 83 BOL
Synonyms: hidden
= Polystichum smithii Mickel & Beitel; Mem. New York Bot. Gard. 46 [Pterid. Fl. Oaxaca]: 315 (1988)
species :
Polystichum uahukaense
Lorence & W.L.Wagner; PhytoKeys 4: 25 (2011)
Distribution: Marquesas Isl. (Ua Huka)
61 MRQ
species :
Polystichum underwoodii
Maxon; Contr. U. S. Natl. Herb. 13: 38, t. 9 (1909), nom. nov.
Distribution: Jamaica
81 JAM
Synonyms: hidden
= Aspidium triangulum subsp. latipinnum (Jenman) Jenman; Ferns Brit. W. Ind. 198 (1907)
= Aspidium triangulum var. latipinnum Jenman; J. Bot. 17: 260 (1879)
= Aspidium viviparum subsp. latipinnum (Jenman) Jenman; Bull. Bot. Dept., Jamaica 2: 269 (1895)
species :
Polystichum unicum
Li Bing Zhang, N.T.Lu & Liang Zhang; Phytotaxa 607(1): 87 (2023)
Distribution: Vietnam
41 VIE
species :
Polystichum uniseriale
(Ching ex K.H.Shing) Li Bing Zhang; Phytotaxa 60: 59 (2012)
Distribution: China (Chongqing, Sichuan)
36 CHI
Synonyms: hidden
= Cyrtomium uniseriale Ching; Acta Phytotax. Sin., Addit. 1: 15 (1965)
= Cyrtomium uniseriale f. marginale Ching; Acta Phytotax. Sin., Addit. 1: 16 (1965)
species :
Polystichum venosum
R.H.Jiang & X.C.Zhang; Turczaninowia 25(2): 89-90 (2022)
Distribution: China (Guangxi)
36 CHI
species :
Polystichum vestitum
(G.Forst.) C.Presl; Tent. Pterid. 83 (1836)
Name: Prickly shield fern, Punui (Maori)
Distribution: New Zealand (North Isl., South Isl., Stewart Isl.); Snares Isl.; Chatham Isl.; Antipodes Isl.; Auckland Isl.; Macquarie Isl.; Campbell Isl.
51 ATP AUC CTM NZL 91 CMP MAQ
Remarks: subgen. Polystichum, Australasian clade
Synonyms: hidden
= Aspidium aculeatum var. vestitum (Sw.) Hook.f.; Handb. N. Zeal. Fl. 2: 375 (1867)
= Aspidium asterolepis Trevis.; Hedwigia 13: 145 (1874)
= Aspidium perelegans Colenso; Trans. & Proc. New Zealand Inst. 29: 416 (1897)
= Aspidium pulcherrimum Colenso; Tasmanian J. Nat. Sci. 2: 168 (1845)
= Aspidium tasmaniae Mett.; Fil. Hort. Bot. Lips. 88 (1856)
= Aspidium venustum Hook.f.; Fl. Antarct. 1: 106 (1844)
= Aspidium vestitum (G.Forst.) Sw.; J. Bot. (Schrader) 1800(2): 37 (1801)
= Aspidium waikarense Colenso; Tasmanian J. Nat. Sci. 2: 168 (1845)
= Polypodium vestitum G.Forst.; Fl. Ins. Austr.: 82 (1786)
= Polystichum aculeatum var. vestitum (G.Forst.) Rupr.; Distr. Crypt. Vasc. Ross. 39 (1945)
= Polystichum perelegans (Colenso) C.Chr.; Index Filic. 87 (1905), 586 (1906)
= Polystichum venustum (Hook.) Hombr. & Jacquinot; Voy. au Pole sud, t. 5 (1852)
= Polystichum vestitum var. venustum J.Dix; Proc. Roy. Hort. Soc. London 1: 568 (1861)
species :
Polystichum volkensii
(Hieron.) C.Chr.; Index Filic. 97 (1905), 586 (1906)
Distribution: Kenya; Tanzania
25 KEN TAN
Synonyms: hidden
= Aspidium volkensii Hieron.; Pflanzenw. Ost-Afrikas [Engler] C: 86 (1895)
= Polystichum barbatum C.Chr.; Notizbl. Königl. Bot. Gart. Berlin 9: 178 (1924)
species :
Polystichum vulcanicum
(Blume) T.Moore; Index Fil. [T. Moore] 108 (1858)
Distribution: Java
42 JAW
Synonyms: hidden
= Aspidium vulcanicum Blume; Enum. Pl. Javae 2: 163 (1828)
species :
Polystichum walkerae
(Hook.) Sledge; Bull. Brit. Mus. (Nat. Hist.), Bot. 5: 29 (1973)
Distribution: Sri Lanka
40 SRL
Synonyms: hidden
= Arachniodes bipinnata (Bedd.) B.K.Nayar & Kaur; Companion Ferns Brit. India 56 (1974) (fide Fraser-Jenkins 1997)
= Arachniodes macrocarpa (Bedd.) B.K.Nayar & Kaur; Companion Ferns Brit. India 56 (1974) (fide Fraser-Jenkins 1997)
= Arachniodes pinnatifida (Bedd.) B.K.Nayar & Kaur; Companion Handb. Ferns Brit. India56 (1974) (fide Fraser-Jenkins 1997)
= Arachniodes walkerae (Hook.) Ching; Acta Bot. Sin. 10: 260 (1962)
= Dryopteris walkerae (Hook.) Kuntze; Revis. Gen. Pl. 2: 814 (1891)
= Lastrea walkerae (Hook.) Bedd.; Ferns Brit. India Correct. 11 (1870)
= Lastrea walkerae var. bipinnata Bedd.; Ferns Brit. India 234 (1883)
= Lastrea walkerae var. macrocarpa Bedd.; Ferns Brit. India 233 (1883)
= Lastrea walkerae var. pinnatifida Bedd.; Ferns Brit. India 233 (1883)
= Nephrodium walkerae (Hook.) Baker; Syn. Fil. [W. J. Hooker & Baker], ed. 2: 493 (1874)
= Phegopteris walkerae (Hook.) J.Sm.; Ferns Brit. Foreign 169 (1866)
= Polypodium walkerae Hook.; Sp. Fil. [W. J. Hooker] 4: 233 (1862)
= Polypodium walkerae var. bipinnatum Bedd.; Ferns S. India 78, t. 235 (1864)
= Polypodium walkerae var. macrocarpum Bedd.; Ferns S. India 78, t. 235 (1864)
= Polypodium walkerae var. pinnatifidum Bedd.; Ferns S. India 78, t. 235 (1864)
= Rumohra walkerae (Hook.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 496 (1938)
species :
Polystichum wattii
(Bedd.) C.Chr.; Index Filic. 98 (1905), 589 (1906)
Distribution: China (Yunnan); Tibet; NE-India (Manipur, Arunachal Pradesh); Myanmar; Vietnam
36 CHI TIB 40 IND 41 MYA VIE
Conservation Status: IUCN VU for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Aspidium wattii Bedd.; J. Bot. 26: 234 (1888)
= Lastreopsis wattii (Bedd.) Tagawa; J. Jap. Bot. 22: 163 (1949)
= Lastrea wattii (Bedd.) Bedd.; Ferns Brit. India Suppl. 49 (1892)
species :
Polystichum wawranum
(Szyszyl.) Perrie; New Zealand J. Bot. 41(2): 204 (2003)
Name: Black Shield Fern, Shore Shield Fern
Distribution: New Zealand; Chatham Isl.
51 CTM NZL
Remarks: subgen. Polystichum, Australasian clade
Synonyms: hidden
= Aspidium wawranum Szyszyl.; It. Princip. Coburg. 126 (1888), as ´wawraeanum´
species :
Polystichum weimingii
Li Bing Zhang & H.He; Syst. Bot. 33(1): 13 (-16, f.1-2) (2009)
Distribution: China (S-Yunnan); ?Myanmar; ?Laos; ?N-Vietnam
36 CHI
Remarks: subgen. Polystichum, Sect. Hypopeltis III
species :
Polystichum whiteleggei
Watts; Proc. Linn. Soc. New South Wales 39: 257-262 (1914)
Distribution: Lord Howe Isl.
50 LHI
Conservation Status: IUCN R
species :
Polystichum wilesianum
Proctor; Brit. Fern Gaz. 10: 24, t. 3 (1968)
Distribution: Jamaica; Hispaniola
81 HIS JAM
species :
Polystichum woodsioides
Christ; Bot. Gaz. 61: 354 (1911)
Distribution: China (Sichuan); Tibet; Nepal; Bhutan; India (Arunachal Pradesh, Sikkim, ?Uttarakhand); Myanmar (Kachin, etc.)
36 CHI TIB 40 BHU IND NEP 41 MYA
Remarks: bona sp. fide Fraser-Jenkins (2008 and 2015), not a synonym of P. prescottianum. Called P. moupinense in Fl. of China
species :
Polystichum woronowii
Fomin; Moniteur Jard. Bot. Tiflis, livr. 18: 21 (1911)
Distribution: Georgia [Caucasus]; Azerbaijan (Talysh); Turkey (NE-Anatolia); NW-Iran
33 AZE GEO 34 IRN TUR
species :
Polystichum wrightii
(Baker) C.Chr. ex Maxon; Contr. U. S. Natl. Herb. 16: 50 (1912)
Distribution: Cuba
81 CUB
Remarks: known only from two collections
Synonyms: hidden
= Dryopteris sauvallei C.Chr.; Index Filic. 291 (1905)
= Nephrodium wrightii (Baker) Diels; Nat. Pflanzenfam. [Engler & Prantl] I, 4: 168 (1899)
= Polypodium wrightii Baker; Syn. Fil. [W. J. Hooker & Baker] 304 (1867)
= Polystichum longipes Maxon; Contr. U. S. Natl. Herb. 18: 34, t. 6 (1909)
species :
Polystichum wulingshanense
S.F.Wu; Keys Vasc. Pl. Wuling Mts. 568 (1995)
Distribution: China (NW-Hunan, W-Hubei)
36 CHI
species :
Polystichum wusugongii
Liang Zhang, X.M.Zhou & Li Bing Zhang; Phytotaxa 618(1): 22 (2023)
Distribution: Tibet
36 TIB
species :
Polystichum xichouense
(S.K.Wu & Mitsuta) Li Bing Zhang; Phytotaxa 60: 59 (2012)
Distribution: China (SE-Yunnan); Vietnam
36 CHI 41 VIE
Remarks: subgen. Haplopolystichum, sect. Cyrtogonellum
Synonyms: hidden
= Cyrtogonellum xichouense S.K.Wu & Mitsuta; Acta Phytotax. Geobot. 36(1-3): 25 (1985)
species :
Polystichum xinfeniae
Liang Zhang, W.B.Ju & Li Bing Zhang; Phytotaxa 618(1): 24 (2023)
Distribution: Tibet
36 TIB
species :
Polystichum xiphophyllum
(Baker) Diels; Nat. Pflanzenfam. [Engler & Prantl] I, 4: 189 (1906)
Distribution: China (Chongqing, Guangdong, Guizhou, Hunan, Jiangxi, Sichuan, Yunnan); Taiwan; peninsular Malaysia
36 CHI 38 TAI 42 MLY
Remarks: subgen. Polystichum, sect. Xiphopolystichum
Synonyms: hidden
= Aspidium monotis Christ; Bull. Soc. Bot. Ital. 1901: 294 (1901)
= Aspidium xiphophyllum Baker; J. Bot. 26: 227 (1888)
= Polystichum gladiipinnum Tagawa; Acta Phytotax. Geobot. 9: 91 (1940)
= Polystichum monotis (Christ) C.Chr.; Index Filic. 83 (1905), 584 (1906)
= Polystichum praelongum Christ; Bull. Acad. Int. Géogr. Bot. 260 (1902)
= Polystichum pseudoxiphophyllum Ching ex H.S.Kung; Chin. J. Appl. Environ. Biol. 3(2): 135, pl. 1: 2, 3 (1997) (fide Knapp (Ferns of Taiwan), kept separate by Flora of China)
= Polystichum xiphophyllum f. bipinnatum Ching; Icon. Filic. Sin. 5: 244: 7 (1959)
species :
Polystichum xuansonense
Li Bing Zhang, N.T.Lu & Liang Zhang; Phytotaxa 607(1): 89 (2023)
Distribution: Vietnam
41 VIE
species :
Polystichum yaanense
Liang Zhang & Li Bing Zhang; Novon 22(2): 244 (2012)
Distribution: China (Sichuan)
36 CHI
Remarks: subgen. Haplopolystichum, sect. Platylepia
species :
Polystichum yaeyamense
(Makino) Makino; Bot. Mag. (Tokyo) 23: 144 (1909)
Distribution: South Korea (Cheju Isl.); Ryukyu Isl. (Okinawa Isl., Iriomote)
38 KOR NNS
Remarks: subgen. Haplopolystichum, sect. Crucifilix
Conservation Status: IUCN R
Synonyms: hidden
= Aspidium yaeyamense Makino; Bot. Mag. (Tokyo) 11: 18 (1897)
= Dryopteris yaeyamensis (Makino) C.Chr.; Index Filic. 301 (1905)
species :
Polystichum yifanii
Li Bing Zhang & Kropf; Phytotaxa 404(6): 227 (2019)
Distribution: China (Guizhou)
36 CHI
species :
Polystichum yuanum
Ching; Icon. Filic. Sin. 5: t. 235 (1958)
Distribution: China (NW-Yunnan)
36 CHI
Remarks: subgen. Haplopolystichum, sect. Haplopolystichum
species :
Polystichum yunnanense
Christ; Notul. Syst. (Paris) 1: 34 (1909)
Distribution: Afghanistan; China (W-Guizhou, SW-Sichuan, Yunnan); Tibet; Pakistan (Swat, Chitral, Pakistani Punjab, Hazara); Nepal; Bhutan; N-India (Jammu & Kashmir, Arunachal Pradesh, Himachal Pradesh, Manipur, Meghalaya, Nagaland, Sikkim, Uttarakhand, West Bengal); Myanmar
34 AFG 36 CHI TIB 40 BHU IND NEP PAK 41 MYA
Synonyms: hidden
= Polystichum gyirongense Ching; Fl. Xizangica 1: 238 (1983) (fide Fraser-Jenkins 1997)
= Polystichum huae H.S.Kung & Li Bing Zhang; Acta Phytotax. Sin. 33(3): 311 (1995)
= Polystichum jizhushanense Ching; Fl. Xizangica 1: 238 (1983) (fide Fraser-Jenkins 1997)
species :
Polystichum zambesiacum
Schelpe; Bol. Soc. Brot., ser. 2, 41: 215 (1967)
Distribution: Tanzania; Malawi; Zimbabwe; Mozambique
25 TAN 26 MLW MOZ ZIM
species :
Polystichum zayuense
W.M.Chu & Z.R.He; Fl. Reipubl. Popularis Sin. [Fl. China Ed. Committee] 5(2): 228, pl. 45: 5, 6 (2001)
Distribution: Tibet (Zayu); Nepal
36 TIB 40 NEP
Synonyms: hidden
= Polystichum thomsonii f. himalaicum Kholia; Ferns Fern Allies Sikkim 2: 85 (2014)
species :
Polystichum zhijinense
Li Bing Zhang, Yi F.Duan & Kropf; Phytotaxa 331(1): 124 (2017)
Distribution: China (Guizhou)
36 CHI
species :
Polystichum ×amboversum
Sa.Kurata; Sci. Rep. Yokosuka City Mus. 10: 36 (1964)
Distribution: Japan
38 JAP
Remarks: Polystichum ovatopaleaceum x retrorsopaleaceum
species :
Polystichum ×anceps
Sa.Kurata; Sci. Rep. Yokosuka City Mus. 10: 32 (1964)
Distribution: Japan
38 JAP
Remarks: Polystichum longifrons x polyblepharum
species :
Polystichum ×arendsii
(F.Wirtg. ex Christ) R.Thiemann ex S.Piller; Amer. Fern J. 110(1): 48 (2020)
Distribution: Germany [c]
11 ger
Remarks: P. aculeatum x P. munitum; formed spontaneously in cultivation in Germany
Synonyms: hidden
= Aspidium ×arendsii F.Wirtg. ex Christ; Allg. Bot. Z. Syst. 12: 4 (1906)
species :
Polystichum ×bicknellii
(Christ) Hahne; Allg. Bot. Z. Syst. 10: 103 (1904)
Distribution: England (England, Wales, Scotland); Ireland (Ireland, Northern Ireland); Netherlands; Luxembourg; Germany (Baden-Württemberg, Bayern, Rheinland-Pfalz, Saarland); Channel Isl. (Guernsey); Spain; France; Romania; European Russia; etc.
10 GRB IRE 11 GER LUX NET 12 CIS FRA SPA 13 ROM 14 RUS
Remarks: Polystichum aculeatum x setiferum
Synonyms: hidden
= Aspidium ×bicknellii Christ; Burnat, Mater. Fl. Alpes Marit.: 20 (1900)
= Polystichum stansfieldii Druce; nom. nud.
species :
Polystichum ×dycei
Sleep; Pteridologist 3(3): 52 (1998)
Distribution: Great Britain
10 GRB
Remarks: Polystichum braunii x P. proliferum
species :
Polystichum ×flemingii
Fraser-Jenk.; Aspects Pl. Sci. 13: 277 (1991)
Distribution: Nepal; India (Jammu & Kashmir, Uttarakhand, West Bengal)
40 IND NEP
Remarks: Polystichum discretum x yunnanense
species :
Polystichum ×fominii
Askerov & A.E.Bobrov; Bot. Zhurn. (Moscow & Leningrad) 57: 1298 (1972)
Distribution: Northern Caucasus
33 NCS
Remarks: Polystichum aculeatum x woronowii
species :
Polystichum ×fujisanense
Seriz.; J. Jap. Bot. 45: 263 (1970)
Distribution: Japan
38 JAP
Remarks: Polystichum braunii x ohmurae
species :
Polystichum ×gemmilachenense
Miyam. & T.Nakam.; J. Jap. Bot. 58(5): 149 (1983)
Distribution: Taiwan
38 TAI
Remarks: subgen. Polystichum, Sect. Sorolepidium. Polystichum lachenense x pseudostenophyllum
species :
Polystichum ×hagenahii
Cody; Amer. Fern J. 58: 30 (1968)
Distribution: Canada (Ontario); USA (New York, Pennsylvania)
72 CDN 75 USA
Remarks: Polystichum acrostichoides x Polystichum lonchitis
Synonyms: hidden
= Polystichum ×marginale (V.R.McColl) Cody
species :
Polystichum ×hakonense
Sa.Kurata; J. Geobot. 9: 99 (1961)
Distribution: Japan (Honshu)
38 JAP
Remarks: Polystichum longifrons x P. pseudomakinoi
species :
Polystichum ×hitoyoshiense
Sa.Kurata; Sci. Rep. Yokosuka City Mus. 15: 47 (1969)
Distribution: Japan
38 JAP
Remarks: Polystichum otomasui x pseudomakinoi
species :
Polystichum ×hokurikuense
Sa.Kurata; Sci. Rep. Yokosuka City Mus. 10: 39 (1964)
Distribution: Japan
38 JAP
Remarks: Polystichum longifrons x retrosopaleaceum
species :
Polystichum ×ida-klodeae
R.Thiemann; Pteridologist 7(1): 33 (2020), no type cited
Distribution: E-Asia
Remarks: Polystichum atkinsonii x P. proliferum
species :
Polystichum ×iidanum
Sa.Kurata; Sci. Rep. Yokosuka City Mus. 10: 33 (1964)
Distribution: Japan
38 JAP
Remarks: Polystichum longifrons x makinoi
species :
Polystichum ×illyricum
(Borbás) Hahne; Allg. Bot. Z. Syst. 10: 103 (1904)
Distribution: England (Scotland); Sweden; Germany (Baden-Württemberg, Bayern); Austria; Andorra; Slovakia; Croatia; Montenegro; European Russia
10 GRB SWE 11 AUT GER SLO 12 ADO 13 CRO MNT 14 RUS
Remarks: Polystichum aculeatum x lonchitis
Synonyms: hidden
= Aspidium murbeckii Raimann; Wiener Ill. Gart.-Zeitung 16: 417 (1891)
= Aspidium ×illyricum Borbás; Oesterr. Bot. Z. 41: 354 (1891)
= Dryopteris illyrica (Borbás) Beck; Oesterr. Bot. Z. 67: 113 (1918)
= Polystichum ×eberlei D.E.Mey.; Nova Hedwigia 1: 107 (1959)
species :
Polystichum ×inadae
Sa.Kurata; J. Geobot. 11: 6 (1962)
Distribution: Japan
38 JAP
Remarks: Polystichum polyblepharon x P. retrosopaleaceum
species :
Polystichum ×inayatii
Fraser-Jenk.; Aspects Pl. Sci. 13: 277 (1991)
Distribution: Pakistan (Hazara); Nepal
40 NEP PAK
Remarks: Polystichum piceopaleaceum x yunnanense
species :
Polystichum ×izuense
Sa.Kurata; J. Geobot. 7: 42 (1958)
Distribution: Japan
38 JAP
Remarks: Polystichum makinoi x tagawanum
species :
Polystichum ×jitaroi
Sa.Kurata; Sci. Rep. Yokosuka City Mus. 10: 33 (1964)
Distribution: Japan
38 JAP
Remarks: Polystichum fibrillosopaleaceum x P. pseudomakinoi
species :
Polystichum ×kai-montanum
Seriz.; J. Jap. Bot. 54(5): 143 (1979)
Distribution: Japan
38 JAP
Remarks: Polystichum ohmurae x ovatopaleaceum var. coraiense
species :
Polystichum ×kasayamense
Sa.Kurata; Sci. Rep. Yokosuka City Mus. 15: 47 (1969)
Distribution: Japan
38 JAP
Remarks: Polystichum braunii x polyblepharum
species :
Polystichum ×kiyozumianum
Sa.Kurata; J. Geobot. 9: 99 (1961)
Distribution: Japan (Honshu)
38 JAP
Remarks: Polystichum pseudomakinoi x tagawanum
species :
Polystichum ×kumamontanum
Sa.Kurata; J. Geobot. 12: 68 (1963)
Distribution: Japan
38 JAP
Remarks: Polystichum otomasui x polyblepharum
species :
Polystichum ×kunioi
Sa.Kurata; Sci. Rep. Yokosuka City Mus. 15: 48 (1969)
Distribution: Japan
38 JAP
Remarks: Polystichum braunii x makinoi
species :
Polystichum ×kuratae
Seriz.; J. Jap. Bot. 54(5): 143 (1979)
Distribution: Japan
38 JAP
Remarks: Polystichum polyblepharum x pseudomakinoi
species :
Polystichum ×kurokawae
Tagawa; J. Jap. Bot. 26: 20 (1951)
Distribution: Japan
38 JAP
Remarks: Polystichum makinoi x P. ovatopaleaceum
species :
Polystichum ×lesliei
Rumsey & Acock; Fern Gaz. 16(5): 247 (2001 publ. 2002)
Distribution: England (Surrey, Cornwall); etc.
10 GRB
Remarks: Polystichum munitum x setiferum
species :
Polystichum ×lonchitiforme
(Halácsy) Bech.; Boll. Soc. Ticinese Sci. Nat. 36: 17 (1941)
Distribution: Ireland; Spain; etc.
10 IRE 12 SPA
Remarks: Polystichum lonchitis x setiferum
Synonyms: hidden
= Aspidium ×lonchitiforme Halácsy; Verh. K. K. Zool.-Bot. Ges. Wien 54: 120-129 (1904)
species :
Polystichum ×luerssenii
(Dörfl.) Hahne; Allg. Bot. Z. Syst. 10: 103 (1904)
Distribution: Germany (Baden-Württemberg, Hessen); Czech Republic; Slovakia; European Russia; Northern Caucasus
11 CZE GER SLO 14 RUS 33 NCS
Remarks: Polystichum aculeatum x braunii
Synonyms: hidden
= Aspidium lobatiforme Waisb.; Oesterr. Bot. Z. 49: 65 (1899)
= Aspidium luerssenii Dörfl.; Oesterr. Bot. Z. 40: 227 (1890)
species :
Polystichum ×machaerophyllum
Sloss.; Bull. Torrey Bot. Club 40: 688, t. 26, f. 4-5 (1913)
Distribution: Cuba
81 CUB
Remarks: Polystichum echinatum x P. ilicifolium
species :
Polystichum ×maderense
Johnson; Ann. Mag. Nat. Hist. III(17): 287 (1866)
Distribution: Madeira (Madeira Isl.)
21 MDR
Remarks: Polystichum falcinellum x Polystichum setiferum
species :
Polystichum ×mashikoi
Sa.Kurata; J. Geobot. 7: 116 (1958)
Distribution: Japan
38 JAP
Remarks: Polystichum polyblepharon x P. tagawanum
species :
Polystichum ×meyeri
Sleep & Reichst.; Bauhinia 3: 303, f. 7, 13, 15, 17 (1967)
Distribution: Europe; Austria; European Russia
11 AUT 14 RUS
Remarks: Polystichum braunii x lonchitis
species :
Polystichum ×microlepis
Sa.Kurata; Sci. Rep. Yokosuka City Mus. 10: 37 (1964)
Distribution: Japan
38 JAP
Remarks: Polystichum makinoi x retrosopaleaceum
species :
Polystichum ×midoriense
t.Oka & Ohtani; Sci. Rep. Yokosuka City Mus. 27: 57 (1980)
Distribution: Japan
38 JAP
Remarks: Polystichum fibrillosopaleaceum x ovatopaleaceum
species :
Polystichum ×minamitanii
Sa.Kurata ex Seriz.; J. Jap. Bot. 54(5): 141 (1979)
Distribution: Japan
38 JAP
Remarks: Polystichum makinoi x otomasui
species :
Polystichum ×miuranum
Sa.Kurata; Sci. Rep. Yokosuka City Mus. 15: 46 (1969)
Distribution: Japan
38 JAP
Remarks: Polystichum fimbrillosopaleaceum x polyblepharum
species :
Polystichum ×mucrolentum
Fraser-Jenk.; Annot. Checkl. Ind. Pterid. 2: 352 (2018)
Distribution: India (Meghalaya)
40 IND
Remarks: Polystichum lentum x mucronifolium
species :
Polystichum ×namegatae
Sa.Kurata; Hokuriku J. Bot. 6: 113 (1957)
Distribution: Japan
38 JAP
Remarks: Polystichum makinoi x pseudomakinoi
species :
Polystichum ×ohtanii
Sa.Kurata; Sci. Rep. Yokosuka City Mus. 15: 46 (1969)
Distribution: Japan
38 JAP
Remarks: Polystichum fimbrillosopaleaceum x longifrons
species :
Polystichum ×okanum
Sa.Kurata; J. Geobot. 8: 16 (1959)
Distribution: Japan
38 JAP
Remarks: Polystichum retrosopaleaceum x tagawanum
species :
Polystichum ×ongataense
Sa.Kurata; Hokuriku J. Bot. 6: 114 (1957)
Distribution: Japan
38 JAP
Remarks: Polystichum ovatopaleaceum x P. pseudomakinoi
species :
Polystichum ×phulchowkiense
Fraser-Jenk.; Annot. Checkl. Ind. Pterid. 2: 352 (2018)
Distribution: Nepal
40 NEP
Remarks: Polystichum mucronifolium x yunnanense
species :
Polystichum ×potteri
Barrington; Rhodora 88(855): 298 (1986)
Distribution: Canada (New Brunswick, Nova Scotia, Québec); USA (Massachusetts, Maine, New Hampshire, New York, Pennsylvania, Vermont)
72 CDN 75 USA
Remarks: Polystichum acrostichoides x Polystichum braunii
species :
Polystichum ×pseudobraunii
Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 337 (2008)
Distribution: Nepal
40 NEP
Remarks: Polystichum piceopaleaceum x sinense
species :
Polystichum ×pseudocraspedosorum
Seriz.; Shidekobushi 1(2): 94 (2009)
Distribution: Japan (Honshu)
38 JAP
Remarks: Polystichum capillipes x P. craspedosorum
species :
Polystichum ×pseudolentum
Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 420 (2008)
Distribution: India (Meghalaya)
40 IND
Remarks: Polystichum lentum x pseudotsus-simense
species :
Polystichum ×pseudoovatopaleaceum
Akasawa; Bull. Kochi Women´s Univ., Ser. Nat. Sci. 27: 2 (1979)
Distribution: Japan
38 JAP
Remarks: P. ovatopaleaceum x P. tagawanum
species :
Polystichum ×pseudoparvipinnulum
Miyam. & T.Nakam.; J. Jap. Bot. 58(5): 146 (1983)
Distribution: Taiwan
38 TAI
Remarks: Polystichum parvipinnulum x piceopaleaceum
species :
Polystichum ×pseudosemifertile
Nakaike & Gurung; Bull. Natl. Sci. Mus., Tokyo, B, 14(3): 81 (1988)
Distribution: Nepal; India (West Bengal)
40 IND NEP
Remarks: Polystichum lentum x semifertile
species :
Polystichum ×rupestre
P.S.Wang & Li Bing Zhang; Phytotaxa 60: 58 (2012)
Distribution: China (C-Guizhou)
36 CHI
Synonyms: hidden
= Cyrtogonellum ×rupicola P.S.Wang & X.Y.Wang; Pterid. Fl. Guizhou 242, f. 61: 2 (2001)
species :
Polystichum ×safarovii
Askerov & A.E.Bobrov; Bot. Zhurn. (Moscow & Leningrad) 57: 1298 (1972)
Distribution: Northern Caucasus
33 NCS
Remarks: Polystichum braunii x woronowii (or P. setiferum)
species :
Polystichum ×saltum
J.P.Roux; Bot. J. Linn. Soc. 124(4): 376 (1997)
Distribution: South Africa (KwaZulu-Natal)
27 SAF
Remarks: Polystichum sinense x transvaalense
species :
Polystichum ×sarukurense
Seriz.; J. Jap. Bot. 54(5): 144 (1979)
Distribution: Japan
38 JAP
Remarks: Polystichum braunii x microchlamys var. azumiense
species :
Polystichum ×semiyunnanense
Fraser-Jenk. & Kandel; Ferns Fern Allies Nepal 2: 323 (2019)
Distribution: Nepal
40 NEP
Remarks: Polystichum semifertile x P. yunnanense
species :
Polystichum ×shin-tashiroi
Sa.Kurata; J. Geobot. 11: 38 (1962)
Distribution: Japan
38 JAP
Remarks: Polystichum microchlamys x retrosopaleaceum
species :
Polystichum ×shizuokaense
Nakaike; New Fl. Japan, Pterid. 842 (1999), nom. nov.
Distribution: Japan (Honshu)
38 JAP
Remarks: Polystichum fibrillosopaleaceum x P. igaense
Conservation Status: CR in Japan
Synonyms: hidden
= Polystichum fibrillosopaleaceum var. marginale Seriz.; (1971)
species :
Polystichum ×silviamontanum
Miyam. & T.Nakam.; J. Jap. Bot. 58(5): 147 (1983)
Distribution: Taiwan
38 TAI
Remarks: Polystichum piceopaleaceum x wilsonii
species :
Polystichum ×stewartii
Fraser-Jenk.; Aspects Pl. Sci. 13: 276 (1991)
Distribution: Pakistan (Swat)
40 PAK
Remarks: Polystichum lonchitis x yunnanense
species :
Polystichum ×suginoi
Sa.Kurata; J. Geobot. 12: 68 (1963)
Distribution: Japan
38 JAP
Remarks: Polystichum otomasui x tagawanum
species :
Polystichum ×suyamanum
Sa.Kurata ex Seriz.; J. Jap. Bot. 54(5): 142 (1979)
Distribution: Japan
38 JAP
Remarks: Polystichum igaense x longifrons
species :
Polystichum ×takaosanense
Sa.Kurata; J. Geobot. 8: 17 (1959)
Distribution: Japan
38 JAP
Remarks: Polystichum longifrons x P. ovatopaleaceum
species :
Polystichum ×tare-bhirense
Nakaike & Gurung; Bull. Natl. Sci. Mus., Tokyo, B, 14(3): 83 (1988)
Distribution: Nepal; NW-India (Jammu & Kashmir)
40 IND NEP
Remarks: Polystichum squarrosum x yunnanense
species :
Polystichum ×tetsuyamense
Sa.Kurata ex Seriz.; J. Jap. Bot. 54(5): 141 (1979)
Distribution: Japan
38 JAP
Remarks: Polystichum otomasui x ovato-paleaceum
species :
Polystichum ×titibuense
Sa.Kurata; J. Geobot. 12: 68 (1963)
Distribution: Japan
38 JAP
Remarks: Polystichum braunii x P. ovatopaleaceum
species :
Polystichum ×tokyoense
Sa.Kurata ex Seriz.; J. Jap. Bot. 54(5): 140 (1979)
Distribution: Japan
38 JAP
Remarks: Polystichum ovatopaleaceum x shimurae
species :
Polystichum ×utsumii
(Sa.Kurata) Sa.Kurata; Sci. Rep. Yokosuka City Mus. 10: 37 (1964)
Distribution: Japan
38 JAP
Remarks: Polystichum pseudomakinoi x retrosopaleaceum
Synonyms: hidden
= Polystichum ×ongataense var. utsumii Sa.Kurata
species :
Polystichum ×wirtgenii
Hahne; Allg. Bot. Z. Syst. 10: 103 (1904)
Distribution: C-Europe; Croatia; European Russia; Northern Caucasus; etc.
13 CRO 14 RUS 33 NCS
Remarks: Polystichum braunii x setiferum
species :
Polystichum ×yokohamense
T.Oka & Ohtani; Sci. Rep. Yokosuka City Mus. 27: 57 (1980)
Distribution: Japan
38 JAP
Remarks: Polystichum fibrillosopaleaceum x tagawanum
