031.0000
Family
Dennstaedtiaceae Lotsy; Vortr. Bot. Stammesgesch. 2: 655 (1909), as ´Dennstaedtineaceae´
11 genera, ca. 230 spp. Phylogeny: Triana-Moreno et al. (2023)
Species listed in database: 237; hybrids: 9
= Hypolepidaceae Pic.Serm.; Webbia 24: 705 (1970)
= Pteridiaceae Ching; Acta Phytotax. Sin. 13: 96 (1975)
= Monachosoraceae Ching; Acta Phytotax. Sin. 14: 17 (1978)
031.2002
Genus
Hypolepis Bernh.; J. Bot. (Schrader) 1(2): 34 (1806)
Distribution: Worldwide tropical/subtropical
Species listed in database: 69; hybrids: 2
species :
Hypolepis acantha
Schwartsb.; Kew Bull. 67(4): 816 (2012)
Distribution: Brazil (Bahia, Espírito Santo, Paraná, Rio de Janeiro, Rio Grande do Sul, Santa Catarina, São Paulo)
84 BRA
species :
Hypolepis alpina
(Blume) Hook.; Sp. Fil. [W. J. Hooker] 2: 63 (1852)
Distribution: China (Guangdong, Guangxi, Hainan, Yunnan); S-Japan (Yakushima Isl.); Ryukyu Isl.; Bonin Isl.; Volcano Isl. (Iwojima); Taiwan; Borneo (Mt. Kinabalu, Sabah); Philippines; Java; New Guinea (Papua New Guinea)
36 CHI 38 JAP KZN NNS OGA TAI 42 BOR JAW NWG PHI
Remarks: not in India. The correct name is probably javensis
Synonyms: hidden
= Acrophorus javensis (Willd.) T.Moore; Index Fil. [T. Moore] 2 (1857)
= Aspidium javense Willd.; Sp. Pl., ed. 4 [Willdenow] 5(1): 284 (1810)
= Cheilanthes alpina Blume; Enum. Pl. Javae 2: 138 (1828)
= Cystopteris javensis (Willd.) Desv.; Mém. Soc. Linn. Paris 6(3): 265 (1827)
= Hypolepis alte-gracillima Hayata; Icon. Pl. Formosan. 6: 295, f. 118 (1915)
= Hypolepis robusta W.M.Chu; Acta Bot. Yunnan., Suppl. 5: 36 (1992)
species :
Hypolepis amaurorhachis
(Kunze) Hook.; Sp. Fil. [W. J. Hooker] 2: 62 (1852)
Name: Austral Ground fern
Distribution: Australia (SE-Victoria, Tasmania); Chatham Isl.; Auckland Isl.; Campbell Isl.
50 AUS 51 AUC CTM 91 CMP
Remarks: included by Schwartsb. & J. Prado (2013) in H. rugosula s. lat., but kept separate by Flora of Australia
Synonyms: hidden
= Cheilanthes amaurorhachis Kunze; Linnaea 28: 242, 306 (1850)
= Hypolepis australis Wakef.; Vict. Naturalist 72: 95 (1955)
= Hypolepis rugosula subsp. subantarctica (Brownsey & Chinnock) Schwartsb.; Acta Bot. Brasil. 28(2): 217 (2014)
= Hypolepis subantarctica Brownsey & Chinnock; New Zealand J. Bot. 22(1): 57 (1984)
species :
Hypolepis ambigua
(A.Rich.) Brownsey & Chinnock; New Zealand J. Bot. 22(1): 59 (1984)
Distribution: New Zealand (North Isl., South Isl., Three Kings Isl., Stewart Isl.); Chatham Isl.
51 CTM NZL
Synonyms: hidden
= Cheilanthes ambigua A.Rich.; Voy. Astrolabe 1: 84 (1832)
= Hypolepis petrieana Carse; Trans. New Zealand Inst. 50: 64 (1918)
= Polypodium amplum Colenso; Trans. New Zealand Inst. 24: 396 (1892), nom. illeg., non Willd. 1810
species :
Hypolepis aspidioides
Christ; Bot. Jahrb. Syst. 23: 343 (1896)
Distribution: [Western] Samoa (Savai´i, ´Upolu); ?American Samoa
60 SAM
species :
Hypolepis bamleriana
Rosenst.; Repert. Spec. Nov. Regni Veg. 10: 325 (1912)
Distribution: Borneo (Mt. Kinabalu, Sabah); Sulawesi; Philippines; Moluccas (Seram); New Guinea (Papua New Guinea, Irian Jaya); Fed. States of Micronesia (Pohnpei)
42 BOR MOL NWG PHI SUL 62 MIC
Synonyms: hidden
= Hypolepis punctata var. obscura Brause; Bot. Jahrb. Syst. 56: 161 (1921)
species :
Hypolepis barringtonii
Schwartsb.; Amer. Fern J. 108(4): 155 (2018)
Name: Creeping Bramble Fern
Distribution: USA (S- & C-Florida)
78 USA
Remarks: previously treated under H. repens
species :
Hypolepis blepharochlaena
Mickel & Beitel; Mem. New York Bot. Gard. 46 [Pterid. Fl. Oaxaca]: 219 (1988)
Distribution: Mexico (Chiapas, Hidalgo, Oaxaca, Puebla, Veracruz); Guatemala; El Salvador; Honduras; Costa Rica
79 MEX 80 COS ELS GUA HON
species :
Hypolepis bogotensis
H.Karst. ex Mett.; Ann. Sci. Nat., Bot., sér. 5, 2: 238 (1864), nomen, et ex Karst, Fl. Columb. 2: 91, t. 147 (1864)
Distribution: Colombia (Cundinamarca, Distrito Capital)
82 CLM
species :
Hypolepis crassa
Maxon; Contr. Gray Herb. 165: 69, t. 4 (1947)
Distribution: Ecuador (Napo, Tungurahua)
83 ECU
species :
Hypolepis cubensis
Schwartsb.; Amer. Fern J. 108(4): 156 (2018)
Distribution: Cuba
81 CUB
Remarks: previously treated in Cuba as ´H. repens´
species :
Hypolepis dicksonioides
(Bory) Hook.; Sp. Fil. [W. J. Hooker] 2: 61 (1852)
Distribution: Australia [I] (South Australia [I]); Norfolk Isl.; New Zealand (North Isl., South Isl.); Kermadec Isl.; Chatham Isl.; ?[Western] Samoa; Cook Isl. (Rarotonga); Marquesas Isl. (Nuku Hiva, Ua Pou, Hiva Oa)
50 aus NFK 51 CTM KER NZL 61 COO MRQ
Synonyms: hidden
= Cheilanthes dicksonioides Bory ex Endl.; Prodr. Fl. Norfolk. [Endlicher] 15 (1833)
= Cheilanthes pellucida Colenso; Tasmanian J. Nat. Sci. 2: 173 (1844)
= Hypolepis endlicheriana C.Presl; Tent. Pterid. 162 (1832)
species :
Hypolepis ditrichomatis
R.C.Moran; Ann. Missouri Bot. Gard. 77: 850 (1990)
Distribution: Costa Rica; Panama
80 COS PAN
species :
Hypolepis ekmanii
Maxon; Proc. Biol. Soc. Wash. 43: 84 (1930)
Distribution: Hispaniola (Haiti)
81 HIS
Remarks: nuda agg., bona sp. fide Schwartsb. & J. Prado (2015)
species :
Hypolepis elegans
Carruth.; Fl. Vit. [Seemann] 347 (1873)
subspecies :
subsp. elegans
Distribution: New Guinea (Papua New Guinea); Bismarck Arch. (New Ireland); Solomon Isl. (Bougainville, Guadalcanal, Kolombangara, Malaita, Ranongga, Rendova); Australia (NE-Queensland, NE-New South Wales, S-Victoria); Lord Howe Isl.; New Caledonia; Vanuatu; Fiji (Viti Levu, Vanua Levu, Taveuni); American Samoa (Tutuila, Ta´u); [Western] Samoa; Austral Isl. (Rapa Iti)
42 NWG 43 BIS SOL 50 AUS LHI 60 ASA FIJ NWC SAM VAN 61 AUI
subspecies :
subsp. carolinensis
Brownsey; Blumea 32(2): 266 (1987)
Distribution: Palau; Fed. States of Micronesia (Pohnpei)
62 MIC PLU
species :
Hypolepis feeana
Schwartsb.; Amer. Fern J. 108(4): 157 (2018)
Distribution: Mexico (Veracruz); Costa Rica; Panama; Jamaica; Hispaniola (Haiti, Dominican Republic); Colombia; Venezuela (Aragua, Falcon, Lara, Yaracuy)
79 MEX 80 COS PAN 81 HIS JAM 82 CLM VEN
Remarks: previously treated as ´H. stuebelii´
species :
Hypolepis fimbriata
Maxon ex Proctor; Amer. Fern J. 72(4): 113 (1982)
Distribution: Jamaica
81 JAM
species :
Hypolepis flexuosa
Sodiro; Crypt. Vasc. Quit.: 634 (1893)
Distribution: Honduras; Costa Rica; Colombia (Cundinamarca, Cauca, Norte de Santander, Nariño); ?Venezuela (?Aragua); Ecuador; Peru; Bolivia (Cochabamba, La Paz, Santa Cruz)
80 COS HON 82 CLM 83 BOL ECU PER
Remarks: contains most records for ´Hypolepis bogotensis´ outside of Colombia (Schwartsb. & J. Prado 2015)
Synonyms: hidden
= Hypolepis inermis (Hook.) Christ; Bull. Herb. Boissier, sér. 2, 4: 1095 (1904)
= Hypolepis repens var. inermis Hook.; Sp. Fil. [W. J. Hooker] 2: 64 (1852)
species :
Hypolepis forzzae
Schwartsb.; Amer. Fern J. 108(4): 164 (2018)
Distribution: Brazil (Amazonas)
84 BRA
species :
Hypolepis galapagensis
Schwartsb. & J.Prado; Amer. Fern J. 105(4): 298 (2015)
Distribution: Galapagos
83 GAL
species :
Hypolepis glabrescens
Ching; Fl. Reipubl. Popularis Sin. [Fl. China Ed. Committee] 2: 371 (1959)
Distribution: China (W-Yunnan)
36 CHI
species :
Hypolepis glandulosopilosa
H.G.Zhou & H.Li; Guihaia 11(1): 41 (1991)
Distribution: China (Guangxi)
36 CHI
Remarks: status doubtful, not treated in Flora of China due to the lack of material.
species :
Hypolepis grandis
Lellinger; Proc. Biol. Soc. Wash. 98(2): 385 (1985)
Distribution: Costa Rica; Panama
80 COS PAN
species :
Hypolepis guianensis
Klotzsch; Linnaea 20: 339 (1847)
Distribution: ?SE-Venezuela (Mt. Roraima); Guyana (Mt. Roraima); Brazil (Roraima)
82 GUY 84 BRA
Remarks: known only from type collection
species :
Hypolepis hawaiiensis
Brownsey; Blumea 32(2): 240 (1987), nom. nov.
Distribution: Hawaii (Kauai, Oahu, Molokai, Lanai, Maui, Hawaii Isl.)
63 HAW
Synonyms: hidden
= Phegopteris punctata var. glabra Hillebr.; Fl. Hawaiian Isl. [Hillebrand] 562 (1888)
variant :
var. mauiensis
(Hillebr.) D.D.Palmer; Amer. Fern J. 92(2): 102 (2002)
Distribution: Hawaii (Maui)
63 HAW
Synonyms: hidden
= Phegopteris punctata var. mauiensis Hillebr.; Fl. Hawaiian Isl. [Hillebrand] 563 (1888)
species :
Hypolepis hostilis
(Kunze) C.Presl; Tent. Pterid. 162 (1836)
Distribution: Nicaragua; Costa Rica; Panama; Colombia (Amazonas, Antioquia, Caquetá, Cauca, Meta, Nariño, Norte de Santander, Quindío, Santander, Vaupés); Venezuela (Amazonas, Apure, Barinas, Falcon, Lara, Miranda, Portuguesa, Tachira, Yaracuy); Guyana; Surinam; French Guiana; Ecuador; Peru; Bolivia (Cochabamba); Brazil (Amazonas)
80 COS NIC PAN 82 CLM FRG GUY SUR VEN 83 BOL ECU PER 84 BRA
Synonyms: hidden
= Cheilanthes hostilis Kunze; Linnaea 9: 86 (1834)
= Hypolepis hostilis var. major Hook.; Sp. Fil. [W. J. Hooker] 2: 69 (1852)
= Hypolepis parviloba Fée; Crypt. Vasc. Brésil 1: 53, t. 20, f. 1 (1869)
= Hypolepis repens var. hostilis (Kunze) Baker; Fl. Bras. [Martius] 1(2): 383 (1870)
species :
Hypolepis jamaicensis
Maxon ex Proctor; Amer. Fern J. 72(4): 113 (1982)
Distribution: Jamaica
81 JAM
species :
Hypolepis krameri
Schwartsb., Boudrie & Cremers; Fern Gaz. 19(1): 3 (fig.) (2012)
Distribution: ?SE-Venezuela (Mt. Roraima); Guyana (Mt. Roraima); Brazil (Roraima)
82 GUY 84 BRA
Remarks: known only from type collection
species :
Hypolepis lellingeri
A.Rojas; Revista Biol. Trop. 49(2): 439 (2001)
Distribution: Isla del Coco
80 COC
species :
Hypolepis malesiana
Brownsey; Blumea 32(2): 248 (1987)
Distribution: ?Sumatra; Philippines; New Guinea (Papua New Guinea)
42 NWG PHI
species :
Hypolepis melanochlaena
A.R.Sm.; Proc. Calif. Acad. Sci. ser. 4, 40: 221 (1975)
Distribution: Mexico (Chiapas, ?Veracruz); Venezuela (Barinas, Lara, Yaracuy)
79 MEX 82 VEN
species :
Hypolepis microchlaena
Mickel & Beitel; Mem. New York Bot. Gard. 46 [Pterid. Fl. Oaxaca]: 220 (1988)
Distribution: Mexico (Oaxaca)
79 MEX
Remarks: records for Colombia refer probably to H. rigescens fide Schwartsb. & J. Prado (2015)
species :
Hypolepis millefolium
Hook.; Sp. Fil. [W. J. Hooker] 2: 68, t. 95 B (1852)
Name: Dissected ground fern, Thousand-leaved fern
Distribution: New Zealand (North Isl., South Isl., Stewart Isl.); Chatham Isl.; Antipodes Isl.; Auckland Isl.; Campbell Isl.
51 ATP AUC CTM NZL 91 CMP
species :
Hypolepis minima
M.Kessler & A.R.Sm.; Brittonia 59(2): 194 (2007)
Distribution: Bolivia (La Paz)
83 BOL
Remarks: known only from type collection
species :
Hypolepis miodelii
Schwartsb.; Amer. Fern J. 108(4): 165 (2018)
Distribution: Brazil (Rio de Janeiro)
84 BRA
species :
Hypolepis mitis
Kunze ex Kuhn; Linnaea 36: 105 (1869)
Distribution: Brazil (Espírito Santo, Minas Gerais, Rio de Janeiro, Rio Grande do Sul, São Paulo)
84 BRA
species :
Hypolepis moraniana
A.Rojas; Revista Biol. Trop. 49(2): 442 (2001)
Distribution: Costa Rica
80 COS
species :
Hypolepis muelleri
Wakef.; Vict. Naturalist 60: 42, f. 2 (1943)
Name: Harsh Ground fern
Distribution: Australia (Australian Capital Territory, E-Queensland, E-New South Wales, E-Victoria, NE-Tasmania)
50 AUS
species :
Hypolepis muenchii
(Christ) Mickel; Mem. New York Bot. Gard. 88 [Pterid. Mexico]: 330 (2004)
Distribution: Mexico (Chiapas, Guerrero, Oaxaca); El Salvador
79 MEX 80 ELS
Synonyms: hidden
= Dennstaedtia muenchii Christ; Bull. Herb. Boissier, sér. 2, 5: 732 (1905)
= Hypolepis eurychlaena Mickel & Beitel; Mem. New York Bot. Gard. 46 [Pterid. Fl. Oaxaca]: 220 (1988)
species :
Hypolepis nuda
Mett.; Kuhn, Linnaea 36: 105 (1869)
Distribution: Costa Rica; Panama; Jamaica; Hispaniola (Haiti, Dominican Republic); Colombia; Venezuela (Apure, Aragua, Merida, Yaracuy)
80 COS PAN 81 HIS JAM 82 CLM VEN
Remarks: nuda agg.
species :
Hypolepis obtusata
(C.Presl) Kuhn; Festschr. 50jähr. Jubiläum Königstädt. Realschule zu Berlin [Chaetopterides] 347 (1882)
Distribution: Colombia (Arauca, Boyacá, Caldas, Nariño, Risaralda, Santander, Valle); Ecuador; Peru; Bolivia (La Paz)
82 CLM 83 BOL ECU PER
Remarks: at very high elevations > 3800 m
Synonyms: hidden
= Cheilanthes obtusata C.Presl; Reliq. Haenk. 1: 64, t. 11, f. 1 (1825)
= Hypolepis purdieana Hook.; Sp. Fil. [W. J. Hooker] 2: 69, t. 91 (1852)
= Plecosorus peruvianus Fée; Mém. Foug. 5 (Gen. Filic.): 151 t. 13 A, f. 2 (1850-52)
= Polypodium fulvescens Hook. & Grev.; Bot. Misc. 239 (1831)
species :
Hypolepis pallida
(Blume) Hook.; Sp. Fil. [W. J. Hooker] 2: 64 (1852)
Distribution: Bonin Isl. (Hahajima); Taiwan; ?Vietnam; peninsular Malaysia (Perak, Selangor); Sumatra; Borneo (Mt. Kinabalu, Sabah); ?Philippines
38 OGA TAI 42 BOR MLY SUM
Remarks: not in India. The record for Bonin Isl. refers possibly to H. punctata
Synonyms: hidden
= Cheilanthes pallida Blume; Enum. Pl. Javae 2: 139 (1828)
= Hypolepis beddomei N.C.Nair & S.R.Ghosh; J. Jap. Bot. 50: 93 (1975) (nom. nov. superfl.)
= Hypolepis punctata Bedd.; Suppl. Ferns Brit. Ind. 19 (1892) (non Mett. 1868)
species :
Hypolepis parallelogramma
(Kunze) C.Presl; Tent. Pterid. 162 (1836)
Distribution: Colombia (Antioquia, Boyacá, Chocó, Santander); Venezuela (Aragua, Barinas, Falcon, Lara, Merida, Portuguesa, Trujillo, Yaracuy); Ecuador; Peru; Bolivia (Cochabamba, La Paz)
82 CLM VEN 83 BOL ECU PER
Synonyms: hidden
= Cheilanthes parallelogramma Kunze; Linnaea 9: 85 (1834)
= Cheilanthes radula Kunze ex Hook. & Baker; Syn. Fil. [W. J. Hooker & Baker] 130 (1867) nom. nud.
= Cheilanthes scabra H.Karst.; Bot. Zeitung (Berlin) 855 (1854) ex Syn. Fil. [W. J. Hooker & Baker] 130 (1867) nom. nud.
species :
Hypolepis pedropaloensis
Schwartsb. & J.Prado; Amer. Fern J. 106(1): 5 (2016)
Distribution: Colombia (Cundinamarca, Antioquia)
82 CLM
species :
Hypolepis periculosa
Schwartsb.; Phytotaxa 332(3): 257 (2017)
Distribution: Bolivia (La Paz)
83 BOL
Remarks: known only from type
species :
Hypolepis poeppigii
(Kunze) R.Rodr.; Gayana, Bot. 46(3-4): 202 (1989)
Name: Wilel-lawen, pesebre [Chile]
Distribution: Bolivia (Chuquisaca, Santa Cruz, Tarija); Argentina (Jujuy, Salta, Tucuman, Catamarca); Chile (Coquimbo, Valparaíso, Metropolitana de Santiago, O´Higgins, Maule, Ñuble, Biobío, Araucanía, Los Rios, Los Lagos, Aisén, Magallanes); Juan Fdz. Isl.
83 BOL 85 ARG CLE JNF
Remarks: not to be confused with H. rugosula ssp. poeppigiana with almost the same distribution
Synonyms: hidden
= Cystopteris fragilis var. pubescens Phil.; Anales Univ. Chile 43: 582 (1873)
= Dryopteris sturmii (Phil.) C.Chr.; Index Filic. 295 (1905)
= Hypolepis chilensis Fée; Mém. Foug. 8 (Icon. Esp. Nouv.): 76 (1857), p. p., nom. superfl.
= Hypolepis poeppigii (Kunze) Mett. ex Maxon; Lilloa 6: 289 (1941), comb. inval.
= Hypolepis rugosula var. poeppigii (Kunze) C.Chr. & Skottsb.; Nat. Hist. Juan Fernandez (Bot.) 2: 31 (1920), nom. illeg.
= Phegopteris poeppigii (Kunze) Fée ex Gay; Fl. Chil. [Gay] 6: 506 (1853)
= Phegopteris poeppigii var. hirsuta Phil.; Anales Univ. Chile 43: 583 (1873)
= Phegopteris sturmii Phil.; Anales Univ. Chile 94: 358 (1896)
= Polypodium poeppigii Kunze; Linnaea 9: 50 (1834)
species :
Hypolepis polypodioides
(Blume) Hook.; Sp. Fil. [W. J. Hooker] 2: 63 (1852)
Distribution: ?China; ?Tibet; Taiwan; Nepal; Bhutan; India (Jammu & Kashmir, Arunachal Pradesh, Assam State, Himachal Pradesh, Manipur, Meghalaya, Nagaland, Sikkim, Tamil Nadu, Uttarakhand, Uttar Pradesh, West Bengal); Bangladesh; Myanmar; Laos; Cambodia; Vietnam; peninsular Malaysia; Sulawesi; Philippines
38 TAI 40 BAN BHU IND NEP 41 CBD LAO MYA VIE 42 MLY PHI SUL
Synonyms: hidden
= Cheilanthes polypodioides Blume; Enum. Pl. Javae 2: 139 (1828)
= Hypolepis coerulescens A.Biswas; J. Econ. Taxon. Bot. 7(1): 121 (1985) (fide Fraser-Jenkins 2008)
= Hypolepis gamblei A.Biswas; J. Econ. Taxon. Bot. 7(1): 124 (1985) (fide Fraser-Jenkins 2008)
= Hypolepis indica A.Biswas; J. Econ. Taxon. Bot. 7(1): 112 (1985) (fide Fraser-Jenkins 2008)
= Hypolepis sikkimensis A.Biswas; J. Econ. Taxon. Bot. 7(1): 122 (1985) (fide Fraser-Jenkins 2008)
= Hypolepis viridula A.Biswas; J. Econ. Taxon. Bot. 7(1): 123 (1985) (fide Fraser-Jenkins 2008)
species :
Hypolepis pteroides
Mett.; Fil. Lechl. 1: 17, t. 3, f. 7-13 (1856)
Distribution: Colombia (Tolima); Ecuador; Peru
82 CLM 83 ECU PER
species :
Hypolepis punctata
(Thunb. ex Murray) Mett.; Fil. Afr. [Kuhn] 120 (1868)
Distribution: China (Anhui, Fujian, Guangdong, Guizhou, Jiangsu, Jiangxi, Sichuan, Yunnan, Zhejiang); South Korea (Cheju Isl.); Ryukyu Isl.; Taiwan; ?Philippines (?Luzon)
36 CHI 38 KOR NNS TAI
Remarks: limited distribution fide Fraser-Jenkins (2008). See also under H. resistens and H. polypodioides.
Synonyms: hidden
= Dryopteris punctata (Thunb. ex Murray) C.Chr.; Index Filic. 287 (1905) (excl. syn. Hypolepis punctata Bedd.), non Kuntze (1891)
= Hypolepis yunnanensis Ching; Fl. Reipubl. Popularis Sin. [Fl. China Ed. Committee] 2: 370 (1959)
= Nephrodium punctatum (Thunb. ex Murray) Diels; Nat. Pflanzenfam. [Engler & Prantl] I, 4: (1899)
= Phegopteris punctata (Thunb. ex Murray) Mett.; Ann. Mus. Bot. Lugduno-Batavi 1: 222 (1863)
= Polypodium punctatum Thunb.; Fl. Jap. [Thunberg] 337 (1784), et Syst. Veg., ed. 14 (J. A. Murray): 939 (1784) (non Sw. (1801))
species :
Hypolepis repens
(L.) C.Presl; Tent. Pterid. 162 (1836)
Distribution: Mexico (Chiapas, Guerrero, Hidalgo, Oaxaca, Puebla, Veracruz); Guatemala; Belize; El Salvador; Honduras; Nicaragua; Costa Rica; Panama; Hispaniola (Haiti, Dominican Republic); Puerto Rico; St. Kitts; Guadeloupe; Dominica; Martinique; St. Lucia; Tobago; Trinidad; Colombia (Antioquia, Caquetá, Chocó, Nariño, Norte de Santander, Valle); Venezuela (Bolívar, Falcon, Miranda, Tachira); Guyana; Surinam; French Guiana; Ecuador; Bolivia (Cochabamba, La Paz); Brazil (Alagoas, Bahia, Ceará, Mato Grosso do Sul, Pará, Pernambuco, Rio de Janeiro, São Paulo); Argentina (Catamarca, Cordoba, Jujuy, Misiones, Salta, Tucuman)
79 MEX 80 BLZ COS ELS GUA HON NIC PAN 81 DOM GDL HIS MAR PUE STK STL TRT 82 CLM FRG GUY SUR VEN 83 BOL ECU 84 BRA 85 ARG
Synonyms: hidden
= Cheilanthes aculeata (Spreng.) Kaulf.; Flora 1828(1): 367 (1828)
= Cheilanthes repens (L.) Kaulf.; Enum. Filic. [Kaulfuss] 215 (1824)
= Dicksonia aculeata Spreng.; Neue Entdeck. Pflanzenk. 3: 7 (1822)
= Hypolepis aculeata (Spreng.) J.Sm.; J. Bot. (Hooker) 4: 157 (1841)
= Hypolepis delicatula Fée; Mém. Foug. 11 (Hist. Foug. Antil.): 28 (1866)
= Hypolepis dicksonioides Fée; Mém. Foug. 11 (Hist. Foug. Antil.): t. 21, f. 1 (1866), nom. nov. superfl. et illeg.
= Hypolepis mexicana Liebm.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 5, 1: 260 (seors. 108) (1849)
= Hypolepis repens var. selloana Baker; Fl. Bras. [Martius] 1(2): 384 (1870)
= Lonchitis repens L.; Sp. Pl. 2: 1078 (1753)
species :
Hypolepis resistens
(Kunze) Hook.; Sp. Fil. [W. J. Hooker] 2: 64 (1852)
Name: Downy Ground fern
Distribution: China (Hainan); India (Karnataka, Kerala, Tamil Nadu); Sri Lanka; Myanmar; Vietnam; peninsular Malaysia (Gunung Hijau, Perak, Cameron Highlands, Pahang); Borneo (Mt. Kinabalu, Sabah); Philippines; New Guinea (Papua New Guinea); Australia (Australian Capital Territory, Queensland, New South Wales, Victoria, Tasmania); New Caledonia
36 CHI 40 IND SRL 41 MYA VIE 42 BOR MLY NWG PHI 50 AUS 60 NWC
Remarks: taxonomy fide Fraser-Jenkins 2008, 2017
Synonyms: hidden
= Cheilanthes dicksonioides var. phyllochaena Kunze; Linnaea 24: (1851)
= Cheilanthes resistens Kunze; Linnaea 24: 275 (1851)
= Hypolepis glandulifera Brownsey & Chinnock; J. Adelaide Bot. Gard. 10(1): 16 (1987) (fide Fraser-Jenkins 2008)
= Hypolepis longa A.Biswas; J. Econ. Taxon. Bot. 7(1): 112 (1985) (fide Fraser-Jenkins 2008)
species :
Hypolepis rigescens
(Kunze) T.Moore; Index Fil. [T. Moore] 252 (1861)
Distribution: Costa Rica; Panama; Colombia (Bolívar, Antioquia, Chocó, Cundinamarca, Boyacá, Arauca, Meta); Venezuela (Tachira, Falcon, Mt. Roraima); ?Guyana (?Mt. Roraima); Surinam; Ecuador (Morona-Santiago); Peru (Huanuco, Junin); Bolivia (Cochabamba, La Paz, Santa Cruz); Brazil (Bahia, Espírito Santo, Minas Gerais, Pernambuco, Paraná, Rio de Janeiro, São Paulo)
80 COS PAN 82 CLM SUR VEN 83 BOL ECU PER 84 BRA
Synonyms: hidden
= Cheilanthes rigescens Kunze; Flora 1889(1). Beibl. 51 (1889)
= Hypolepis rubiginosopilosula Lellinger; Amer. Fern J. 93(3): 147 (2003)
= Polypodium punctatum f. rigescens (Kunze ex Mart.) Baker; Fl. Bras. [Martius] 1(2): 503, tab. 65 (1870)
= Polypodium punctatum var. rigescens (Kunze) Baker; Fl. Bras. [Martius] 1(2): 503, t. 65 (1870)
variant :
var. buchtienii
(Rosenst.) Schwartsb.; Phytotaxa 332(3): 259 (2017), et Phytotaxa 394(2): 174 (2019)
Distribution: Bolivia (Cochabamba, La Paz)
83 BOL
Synonyms: hidden
= Hypolepis buchtienii Rosenst.; Repert. Spec. Nov. Regni Veg. 25: 58 (1928)
variant :
var. paranaensis
Schwartsb.; Amer. Fern J. 108(4): 166 (2018)
Distribution: Brazil (Paraná)
84 BRA
species :
Hypolepis rugosula
(Labill.) J.Sm.; Compan. Bot. Mag. 8 (1846)
subspecies :
subsp. rugosula
Name: Ruddy Ground fern
Distribution: Australia (SE-Western Australia, SE-South Australia, SE-Queensland, E-New South Wales, Victoria, Tasmania, Australian Capital Territory)
50 AUS
Synonyms: hidden
= Dryopteris punctata subsp. rugosula (Labill.) C.Chr.; index Filic. 287 (1905)
= Dryopteris punctata var. rugosula (Labill.) Domin; Biblioth. Bot. 20: 41 (1914)
= Nephrodium rugulosum (Labill.) Copel.; Polypodiaceae Philipp. 26 (1905)
= Nephrodium viscidum Watson; Proc. Amer. Acad. Arts 26: 163 (1891)
= Phegopteris punctata var. rugosula (Labill.) Hillebr.; Fl. Hawaiian Isl. [Hillebrand] 563 (1888)
= Phegopteris rugosula (Labill.) Mett.; Ann. Mus. Bot. Lugduno-Batavi 1: 222 (1863)
= Phegopteris rugulosa (Labill.) Fée; Mém. Foug. 5 (Gen. Filic.): 243 (1852)
= Polypodium punctatum var. rugosulum (Labill.) Hook. & Baker; Syn. Fil. [W. J. Hooker & Baker] 312 (1868)
= Polypodium rugosulum Labill.; Nov. Holl. Pl. 2: 92, t. 241 (1806)
subspecies :
subsp. africana
(C.Chr.) Schwartsb. & J.Prado; Webbia 66(2): 161 (2011)
Distribution: South Africa; Madagascar; Mauritius; La Réunion
27 SAF 29 MAU MDG REU
Synonyms: hidden
= Hypolepis rugosula var. africana C.Chr.; Dansk Bot. Ark. 7: 121 (1932)
= Phegopteris boryana Mett. ex Ettingsh.; Farnkr. Jetztw. [Ettingshausen]: 162 t. 98 f. 2, t. 102 f. 2, t. 103 f. 4 (1864)
subspecies :
subsp. archboldii
(Copel.) Schwartsb.; Acta Bot. Brasil. 28(2): 216 (2014)
Distribution: New Guinea (Irian Jaya, Papua New Guinea)
42 NWG
Synonyms: hidden
= Hypolepis archboldii Copel.; Univ. Calif. Publ. Bot. 18: 218 (1942)
subspecies :
subsp. colorata
(Christ) Schwartsb. & J.Prado; Acta Bot. Brasil. 28(2): 213 (2014)
Distribution: Costa Rica
80 COS
Synonyms: hidden
= Hypolepis repens var. colorata Christ; Pittier, Prim. Fl. Costaric. 3: 37 (1901)
subspecies :
subsp. formosa
Schwartsb. & M.H.Oliveira; Folia Geobot. 51(4): 377 (2017)
Distribution: Equatorial Guinea
23 EQG
subspecies :
subsp. lactea
(Brownsey & Chinnock) Schwartsb.; Acta Bot. Brasil. 28(2): 216 (2014)
Distribution: New Zealand (North Isl., South Isl.); Chatham Isl.
51 CTM NZL
Remarks: treated at species level in recent NZ floras
Synonyms: hidden
= Hypolepis lactea Brownsey & Chinnock; New Zealand J. Bot. 22(1): 55 (1984)
subspecies :
subsp. pichi-sermolliana
Schwartsb. & J.Prado; Webbia 66(2): 158 (2011)
Distribution: Bioko Isl. [Fernando Poo]; D.R.Congo [Zaire]; Rwanda; Ethiopia; Uganda; Kenya; Tanzania
23 BIO RWA ZAI 24 ETH 25 KEN TAN UGA
subspecies :
subsp. poeppigiana
(Mett.) Schwartsb. & J.Prado; Acta Bot. Brasil. 28(2): 213 (2014)
Distribution: Argentina (Buenos Aires, Neuquen, Rio Negro); Chile (Coquimbo, Bio Bio, La Araucania, Aisen, Magallanes, Chiloe Isl., Los Rios, Valparaiso); Juan Fdz. Isl.
85 ARG CLE JNF
Remarks: see also H. poeppigii (Kunze) Rodr., which is a different species with almost the same distribution
Synonyms: hidden
= Hypolepis hauman-merckii Hicken; Anales Soc. Ci. Argent. 68. (22.) c. tab. (1907)
= Hypolepis poeppigiana Mett.; Fil. Lechl. 1: 18 (1856)
subspecies :
subsp. pradoana
Schwartsb.; Kew Bull. 67(4): 818 (2012)
Distribution: Brazil (Espírito Santo, Minas Gerais, Paraná, Rio de Janeiro, Rio Grande do Sul, Santa Catarina, São Paulo)
84 BRA
Synonyms: hidden
= Hypolepis juergensii Rosenst.; nom. nud.
subspecies :
subsp. pulcherrima
(J.K.Underw. & Maxon) Schwartsb. & J.Prado; Acta Bot. Brasil. 28(2): 213 (2014)
Distribution: Mexico (Chiapas); Costa Rica; Panama; Jamaica; Hispaniola (Haiti)
79 MEX 80 COS PAN 81 HIS JAM
Synonyms: hidden
= Hypolepis pulcherrima Underw. & Maxon; Proc. Biol. Soc. Wash. 43: 84 (1930)
subspecies :
subsp. rouxii
Schwartsb. & M.H.Oliveira; Folia Geobot. 51(4): 375 (2017)
Distribution: South Africa (W-Cape Prov., E-Cape Prov.)
27 SAF
subspecies :
subsp. rudis
(Kunze) Schwartsb.; Acta Bot. Brasil. 28(2): 216 (2014)
Distribution: Philippines (Luzon)
42 PHI
Synonyms: hidden
= Cheilanthes rudis Kunze; Bot. Zeitung (Berlin) 213 (1848)
= Hypolepis rudis (Kunze) Mett.; Ettingsh., Farnkr. Jetztw. 160 (1865)
subspecies :
subsp. rufobarbata
(Colenso) Schwartsb.; Acta Bot. Brasil. 28(2): 217 (2014)
Distribution: New Zealand (North Isl., South Isl., Stewart Isl.); Chatham Isl.
51 CTM NZL
Remarks: treated at species level in recent NZ floras
Synonyms: hidden
= Hypolepis rufobarbata (Colenso) Wakef.; Vict. Naturalist 72: 159 (1956)
= Polypodium rufobarbatum Colenso; Trans. & Proc. New Zealand Inst. 16: 347 (1844)
= Polypodium viscidum Colenso; Tasmanian J. Nat. Sci. 2: 164 (1845) non Roxb. 1816
subspecies :
subsp. villosoviscida
(Thouars) Schwartsb. & J.Prado; Acta Bot. Brasil. 28(2): 214 (2014)
Distribution: Gough Isl.; Tristan da Cunha (incl. Nightingale Isl., Inaccessible Isl.)
91 TDC
Synonyms: hidden
= Hypolepis villosoviscida (Thouars) Tardieu; Fl. Madagasc. Fam. 5, 1: 6, f. 1(3-5) (1958)
= Hypolepis rugosula var. villosoviscida (Thouars) C.Chr.; Sci. Results Norweg. Antarct. Exped 1927-1928, no. 16: 15 (1937)
= Polypodium villosoviscidum Thouars; Fl. Tristan d´Acugna 33 (1804)
= Cheilanthes viscosa Carmich.; Trans. Linn. Soc. London 12(2): 511 (1818 publ. 1819) nom. superfl.
subspecies :
subsp. viscida
(Roxb.) Schwartsb. & J.Prado; Acta Bot. Brasil. 28(2): 215 (2014)
Distribution: St. Helena; +Ascension Isl.
28 ASC STH
Synonyms: hidden
= Cheilanthes remota Kunze; Analecta Pteridogr. 36 (1837) nom. superfl.
= Gymnogramma helenensis (Kuhn) Fée; Mém. Foug. 8 (Icon. Esp. Nouv.): 79 (1857)
= Hypolepis goetzei Hieron. ex Reimers; Mildbr., Notizbl. Bot. Gart. Berlin-Dahlem 12: 189 (1934)
= Hypolepis helenensis (Kuhn) Fée; Mém. Foug. 5 (Gen. Filic.): 147 (1852)
= Hypolepis rugosula subsp. brownseyana Schwartsb. & J.Prado; Acta Bot. Brasil. 28(2): 215, fig. 2(map) (2014)
= Phegopteris helenensis Kuhn; Filic. Afr. 122 (1868)
= Polypodium viscidum Roxb.; Beats., St. Helena: 319 (1816)
species :
Hypolepis scandens
M.Kessler & A.R.Sm.; Brittonia 59(2): 193 (2007)
Distribution: Bolivia (Cochabamba, La Paz)
83 BOL
species :
Hypolepis sparsisora
(Schrad.) Kuhn; Filic. Afr. 120 (1868)
Distribution: Guinea; Sierra Leone; Liberia; Cameroon; Bioko Isl. [Fernando Poo]; São Tomé; D.R.Congo [Zaire]; Rwanda; Burundi; Ethiopia; Uganda; Kenya; Tanzania; Malawi; E-Zimbabwe; Mozambique; South Africa (Limpopo, Mpulamanga, KwaZulu-Natal, W-Cape Prov., E-Cape Prov.); Eswatini [Swaziland]; Madagascar; Comores (Anjouan); La Réunion
22 GUI LBR SIE 23 BIO BUR CMN RWA SAO ZAI 24 ETH 25 KEN TAN UGA 26 MLW MOZ ZIM 27 SAF SWZ 29 COM MDG REU
Synonyms: hidden
= Adiantum altissimum Bojer; Hortus Maurit. 405 (1837)
= Cheilanthes anthriscifolia Schltdl.; Adumbr. Pl. 5: 52, t. 32 (1832) (non Willd. (1810))
= Cheilanthes aspera Kaulf.; Linnaea 6: 186 (1831)
= Cheilanthes commutata Kunze; Linnaea 10: 542 (1836)
= Cheilanthes sparsisora Schrad.; Gött. Gel. Anz. 918 (1818)
= Hypolepis anthriscifolia (Schltdl.) C.Presl; Tent. Pterid. 162 (1836)
= Hypolepis aspera (Kaulf.) C.Presl; Tent. Pterid. 162 (1836)
= Hypolepis commutata (Kunze) Fée; Mém. Foug. 5 (Gen. Filic.): 146 (1852)
= Hypolepis eckloniana C.Presl; Tent. Pterid. 162 (1863)
= Phegopteris sparsisora (Schrad.) Keyserl.; Polyp. Herb. Bunge 51 (1873)
species :
Hypolepis stolonifera
Fée; Crypt. Vasc. Brésil 2: 35, t. 91, f. 2 (1873)
Distribution: Brazil (Minas Gerais, Paraná, Rio de Janeiro, Rio Grande do Sul, Santa Catarina, São Paulo); Argentina (Misiones)
84 BRA 85 ARG
variant :
var. delasotae
Arana, Yañez & Schwartsb.; Phytotaxa 188(2): 93 (2014)
Distribution: Brazil (Paraná); Argentina (Misiones)
84 BRA 85 ARG
variant :
var. nebularis
Schwartsb.; Kew Bull. 67(4): 822 (2012)
Distribution: Brazil (Minas Gerais, Paraná, Rio de Janeiro, Rio Grande do Sul, Santa Catarina, São Paulo)
84 BRA
species :
Hypolepis stuebelii
Hieron.; Hedwigia 48: 230 (1909), as ´stübelii´
Distribution: Colombia (Antioquia, Boyacá, Cundinamarca, Magdalena, Putumayo, Santander, Valle); Ecuador; Peru
82 CLM 83 ECU PER
species :
Hypolepis sunduei
Schwartsb.; Amer. Fern J. 108(4): 163 (2018)
Distribution: Peru (Amazonas)
83 PER
species :
Hypolepis tenera
Ching; Fl. Reipubl. Popularis Sin. [Fl. China Ed. Committee] 2: 370 (1959)
Distribution: China (Yunnan)
36 CHI
species :
Hypolepis tenerrima
Maxon; J. Wash. Acad. Sci. 14: 196 (1924)
Distribution: Hispaniola (Haiti, Dominican Republic); Puerto Rico; ?Venezuela
81 HIS PUE
species :
Hypolepis tenuifolia
(G.Forst.) Bernh.; Neues J. Bot. 1(2): 34 (1806)
Name: Soft Ground fern
Distribution: China (Hainan); Bonin Isl. (Chichijima); Volcano Isl. (Minami-Iwojima); Taiwan; Vietnam; Borneo (Sabah); Sulawesi; Philippines; Lesser Sunda Isl. (Lombok, Flores, Bali, Sumba, Timor); New Guinea (Papua New Guinea); Australia (NE-Queensland); Norfolk Isl.; New Caledonia; New Caledonia; Vanuatu; Tonga (Tongatapu, ´Eua, Vava´u); Fiji (Viti Levu); [Western] Samoa (Savai´i, ´Upolu); ?American Samoa; Cook Isl. (Rarotonga, Atiu); Society Isl. (Tahiti, Moorea, Raiatea); Gambier Isl. (Mangareva); Austral Isl. (Raivavae, Tubuai, Rurutu, Rapa Iti); Pitcairn Isl.; Fed. States of Micronesia (Truk, Dublon, Tol, Pohnpei)
36 CHI 38 KZN OGA TAI 41 VIE 42 BOR LSI NWG PHI SUL 50 AUS NFK 60 FIJ NWC SAM TON VAN 61 AUI COO GBI PIT SCI 62 MIC
Synonyms: hidden
= Adiantum arborescens (Sw.) Poir.; Encycl. [J. Lamarck et al.] suppl. 1: 144 (1810)
= Cheilanthes arborescens Sw.; Syn. Fil. [Swartz] 129, 336 (1806)
= Cheilanthes dissecta Hook. & Arn.; Bot. Beechey Voy. 75 (1832)
= Hypolepis boryana C.Presl; Tent. Pterid. 162 (1836)
= Hypolepis dissecta (Hook. & Arn.) Brack.; U.S. Expl. Exped., Filic. 16: 89 (1854)
= Hypolepis gigantea Ching; Acta Phytotax. Sin. 8: 165 (1959)
= Hypolepis nausoriensis Brownlie; Nova Hedwigia Beih. 55 (Pterid. Fl. Fiji): 125 (1977)
= Hypolepis neocaledonica Rosenst.; Repert. Spec. Nov. Regni Veg. 10: 159 (1911)
= Lonchitis tenuifolia G.Forst.; Fl. Ins. Austr.: 80 (1786)
= Phegopteris tenuifolia (G.Forst.) Keyserl.; Polyp. Herb. Bunge 51 (1873)
species :
Hypolepis thysanochlaena
Mickel & Beitel; Mem. New York Bot. Gard. 46 [Pterid. Fl. Oaxaca]: 221 (1988)
Distribution: Mexico (Chiapas, Morelos, Oaxaca)
79 MEX
species :
Hypolepis trichobacilliformis
R.C.Moran; Ann. Missouri Bot. Gard. 77: 847 (1990)
Distribution: Costa Rica; Panama
80 COS PAN
Remarks: nuda agg.
species :
Hypolepis trichochlaena
Mickel & Beitel; Mem. New York Bot. Gard. 46 [Pterid. Fl. Oaxaca]: 222 (1988)
Distribution: Mexico (Oaxaca)
79 MEX
species :
Hypolepis trinationalis
Schwartsb.; Fern Gaz. 19(1): 2 (-3, figs.) (2012)
Distribution: SE-Venezuela (Mt. Roraima); Guyana (Mt. Roraima); Brazil (Roraima)
82 GUY VEN 84 BRA
species :
Hypolepis urbani
Brause; Symb. Antill. [Urban] 7: 486 (1913)
Distribution: Hispaniola (Dominican Republic); Puerto Rico
81 HIS PUE
Remarks: nuda agg.
species :
Hypolepis viscosa
(H.Karst.) Mett.; Ann. Sci. Nat., Bot., sér. 5, 2: 238 (1864), nomen, ex H. Karst., Fl. Columb. (H. Karst.) 2: 89, t. 145-46 (1869)
Distribution: Hispaniola (Dominican Republic); Colombia (Antioquia, Boyacá, Cauca, Cundinamarca, Huila, Magdalena, Meta, Nariño, Putumayo, Quindío, Santander, Tolima); Venezuela (Amazonas, Apure, Aragua, Bolívar, Venezuela Distrito Federal, Lara, Merida, Yaracuy); ?Ecuador
81 HIS 82 CLM VEN
Synonyms: hidden
= Cheilanthes viscosa H.Karst.; Bot. Zeitung (Berlin) 855 (1854) (nomen, non Carmich. (1818) nec Link (1833))
species :
Hypolepis woodii
Schwartsb.; Phytotaxa 332(3): 260 (2017)
Distribution: Bolivia (Cochabamba, La Paz)
83 BOL
species :
Hypolepis zimmerae
Schwartsb.; Amer. Fern J. 108(4): 159 (2018)
Distribution: Colombia (Cundinamarca); Ecuador (Carchi); Peru (Amazonas)
82 CLM 83 ECU PER
Remarks: previously treated under H. flexuosa
species :
Hypolepis ×glabra
H.Karst. ex Schwartsb. & J.Prado; Amer. Fern J. 105(4): 299 (2015)
Distribution: Colombia (Cundinamarca)
82 CLM
Remarks: Hypolepis bogotensis x flexuosa
species :
Hypolepis ×paulistana
Schwartsb. & J.Prado; Amer. Fern J. 106(1): 4, Figs. 9F-I, 20B (2016)
Distribution: Brazil (São Paulo)
84 BRA
Remarks: H. stolonifera x rugosula
