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249.0000 Family Cannabaceae Martinov; Tekhno-Bot. Slovar 99 (1820), nom. cons.
10 genera, ca. 105 spp. Phylogeny: Yang et al. (2013)
Species listed in database: 120
= Cannabeae Dumort.; Fl. Belg. [Dumortier] 24. Dumort., Fl. Belg. 24 (1827)
= Cannaboideae Beilschm.; Flora 16(Beibl. 7): 60, 105 (1833)
= Celtidaceae Endl.; Enchir. Bot. [Endlicher] 171 (1841), Family
= Celtideae Dumort.; Fl. Belg. [Dumortier] 25 (1827)
= Celtidinae Dumort.; Fl. Belg. [Dumortier] 25 (1827)
= Celtidoideae A.Rich. ex Darl.; Agric. Bot. 151 (1847)
= Humulaceae Bercht. & J.Presl; Prír. Rostlin 258 (1820)
= Humuleae Dumort.; Anal. Fam. Pl. 17 (1829)
= Humulineae J.Presl; Nowoceská Bibl. [Wseobecný Rostl.]: 7: 1359 [´1258´], 1363 (1846)
= Lupulaceae Schultz Sch.; Nat. Syst. Pflanzenr. 370 (1832)
= Lupuleae Rchb.; Fl. Germ. Excurs. 1(2): 179 (1831)
= Treminae A.V.Lazarev; Ukrayins´k. Bot. Zhurn. 50(1): 36 (1993), without latin descr.
249.1005 Genus Humulus L.; Sp. Pl. [Linnaeus] 2: 1028 (1753)
Distribution: Temperate Northern Hemisphere south to SW-USA and Indochina
Species listed in database: 6
= Lupulus Mill.; Gard. Dict. Abr., ed. 4 (1754)
= Humulopsis Grudz.; Bot. Zhurn. (Moscow & Leningrad) 73(4): 592 (1988)
= Waldensia Lavy; État Général, Vég. Orig. 14 (1830)
species :
Humulus americanus Nutt.; J. Acad. Nat. Sci. Philadelphia n. s. 1: 181 (1847), et Proc. Acad. Nat. Sci. Philadelphia 4: 23 (Mart.-Apr. 1848)
Distribution: Canada (Alberta, Manitoba, New Brunswick, Newfoundland, Nova Scotia, Ontario, Prince Edward Isl., Québec, Saskatchewan); USA (Colorado, Connecticut, Delaware, Georgia, Iowa, Idaho, Illinois, Indiana, Kansas, Kentucky, Massachusetts, Maryland, Maine, Michigan, Minnesota, Montana, North Carolina, North Dakota, Nebraska, New Hampshire, New Jersey, New York, Ohio, Pennsylvania, Rhode Island, South Carolina, South Dakota, Tennessee, Utah, Virginia, Vermont, Wisconsin, West Virginia, Wyoming)
71 CDN 72 CDN 73 USA 74 USA 75 USA 76 USA 78 USA
Synonyms: hidden
= Humulus lupuloides (E.Small) Tembrock; J. Bot. Res. Inst. Texas 10(1): 24 (2016)
= Humulus lupulus subsp. americanus (Nutt.) Á.Löve & D.Löve; Taxon 31(1): 121 (1982)
= Humulus lupulus var. lupuloides E.Small; Syst. Bot. 3(1): 63 (1979)
species :
Humulus cordifolius Miq.; Ann. Mus. Bot. Lugduno-Batavi 2: 133 (1865)
Distribution: Russian Far East (Kuril Isl., Sakhalin); China; Japan (Hokkaido, Honshu)
31 RFE 36 CHI 38 JAP
Synonyms: hidden
= Humulus lupulus var. cordifolius (Miq.) Maxim. ex Franch. & Sav.; Enum. Pl. Jap. 2: 489 (1878)
species :
Humulus lupulus L.; Sp. Pl. [Linnaeus] 2: 1028 (1753)
Name: Gewöhnlicher Hopfen (DE), Common Hop (EN)
Distribution: England (England, Wales, Isle of Man [I], Scotland [I], Outer Hebrides [I], Orkney Isl. [I], Shetland Isl. [I], Isles of Scilly); Ireland [I] (Ireland [I], Northern Ireland [I]); Denmark; Norway; Sweden; Finland; Netherlands; Belgium; Luxembourg; Germany (Brandenburg, Berlin, Baden-Württemberg, Bayern, Hessen, Hamburg, Mecklenburg-Vorpommern, Niedersachsen, Nordrhein-Westfalen, Rheinland-Pfalz, Schleswig-Holstein, Saarland, Sachsen, Sachsen-Anhalt, Thüringen); Switzerland; Liechtenstein; Austria; Poland; Czech Republic; Slovakia; Hungary; Portugal; Spain; Andorra; Baleares; France; Channel Isl. (Alderney, Guernsey, Jersey, Sark); Corsica; Sardinia; Italy; +Sicily; San Marino; Slovenia; Croatia; Bosnia & Hercegovina; Montenegro; Serbia; Kosovo; Serbia; Kosovo; North Macedonia; Albania; Romania; Bulgaria; European Turkey; Greece (scattered mainland, Kerkyra); Estonia; Latvia; Lithuania; Belarus; European Russia (widespread); Moldova; Ukraine; Crimea; Morocco; Siberia (Altai, Buryatia, Chita, Irkutsk, Krasnoyarsk, W-Siberia); Russian Far East (Amur [I], Primorye [I], Sakhalin [I]); Kazakhstan; Uzbekistan; Kyrgyzstan; Northern Caucasus; Georgia [Caucasus]; Armenia; Azerbaijan; Turkey (N-Anatolia, NE-Anatolia, NW-Anatolia: Bithynia); Iran (N-Iran); Afghanistan [c]; Syria; South Korea [I]; NW-India [I] (Himachal Pradesh [I], Jammu & Kashmir [I], Uttarakhand [I]); Myanmar [I]; Vietnam [I]; Australia [I] (New South Wales [I]); New Zealand [I]; Chatham Isl. [I]; Canada [I] (Manitoba [I], New Brunswick [I], Newfoundland [I], Nova Scotia [I], Ontario [I], Prince Edward Isl. [I], Québec [I], Saskatchewan [I]); USA [I] (California [I], Connecticut [I], Delaware [I], Illinois [I], Indiana [I], Kentucky [I], Massachusetts [I], Maryland [I], Maine [I], Michigan [I], Missouri [I], New Hampshire [I], New Jersey [I], New York [I], Ohio [I], Pennsylvania [I], Virginia [I], Vermont [I], Wisconsin [I], West Virginia [I]); Argentina [I]
10 DEN FIN GRB ire NOR SWE 11 AUT BGM CZE GER HUN LIE LUX NET POL SLO SWI 12 ADO BAL CIS COR FRA POR SAR SPA 13 ALB BIH BUL CRO GRC ITA KOS MNT NMA ROM SIC SLV SMA SRB TUE 14 BLR EST KRY LAT LIT MLD RUS UKR 20 MOR 30 SIB 31 RFE 32 KAZ KGZ UZB 33 ARM AZE GEO NCS 34 afg IRN SYR TUR 38 kor 40 ind 41 mya vie 50 aus 51 ctm nzl 71 cdn 72 cdn 74 usa 75 usa 76 usa 78 usa 85 arg
Synonyms: hidden
= Cannabis lupulus (L.) Scop.; Fl. Carniol., ed. 2, 2: 263 (1772)
= Humulus lupulus var. acrocarpus Alef.; Landw. Fl. 389 (1866)
= Humulus lupulus var. brachystachyus Zapal.; Rozpr. Wydz. Mat.-Przyr. Akad. Umiejetn., Dzial B, Nauki Biol., Ser. 3, 7B(47B): 276 (1907 publ. 1908)
= Humulus lupulus var. effusus Alef.; Landw. Fl. 389 (1866)
= Humulus lupulus var. fengxianensis J.Q.Fu; Acta Bot. Boreal.-Occid. Sin. 12(6): 70 (1992), contrary to Art. 37.3 ICBN (2000)
= Humulus lupulus var. silvestris Alef.; Landw. Fl. 389 (1866)
= Humulus lupulus var. spaltensis Alef.; Landw. Fl. 389 (1866)
= Humulus volubilis Salisb.; Prodr. Stirp. Chap. Allerton 176 (1796), nom. superfl.
= Humulus vulgaris Gilib.; Hist. Pl. Europe 2: 343 (1798)
= Humulus yunnanensis Hu; Bull. Fan Mem. Inst. Biol., Bot. 7: 211 (1936)
= Lupulus amarus Gilib.; Exerc. Phyt. 2: 451 (1792), opus utique oppr.
= Lupulus communis Gaertn.; Fruct. Sem. Pl. 1: 358, t. 75 (1788)
= Lupulus communis Moench; Methodus [Moench] 332 (1794)
= Lupulus humulus Mill.; Gard. Dict., ed. 8: (1768)
= Lupulus scandens Lam.; Fl. Franç. [Lamarck] 2: 217 (1779), nom. superfl.
= Waldensia lupulina Lavy; État Général, Vég. Orig. 14 (1830)
species :
Humulus neomexicanus (A.Nelson & Cockerell) Rydb.; Fl. Rocky Mts. 208 (1917)
Distribution: Canada (British Columbia, Manitoba, Saskatchewan); USA (Arizona, California, Colorado, District of Columbia, Idaho, Kansas, Montana, North Dakota, Nebraska, New Mexico, Nevada, Oregon, South Dakota, Texas, Utah, Washington State, Wisconsin, Wyoming)
71 CDN 73 USA 74 USA 76 USA 77 USA 78 USA
Synonyms: hidden
= Humulus lupulus var. neomexicanus A.Nelson & Cockerell; Proc. Biol. Soc. Washington 16: 45 (1903)
species :
Humulus pubescens (E.Small) Tembrock; J. Bot. Res. Inst. Texas 10(1): 20 (2016)
Distribution: USA (Alabama, Arkansas, Georgia, Iowa, Illinois, Indiana, Kansas, Kentucky, Maryland, Minnesota, Missouri, North Carolina, Nebraska, New York, Ohio, Oklahoma, Pennsylvania, Virginia, Wisconsin, West Virginia)
74 USA 75 USA 78 USA
Synonyms: hidden
= Humulus lupulus var. pubescens E.Small; Syst. Bot. 3(1): 63 (1979)
species :
Humulus scandens (Lour.) Merr.; Trans. Amer. Philos. Soc., n. s. 24(2): 138 (1935)
Name: Japanischer Hopfen (DE), Japanese hop (EN)
Distribution: Denmark [I]; Sweden [I]; Germany [I]; Austria [I]; Poland [I]; Hungary [I]; France [I]; Italy [I]; Slovenia [I]; Croatia [I]; former Yugoslavia [I]; Serbia; Kosovo [I]; Romania [I]; Ukraine [I]; Russian Far East (Khabarovsk, Primorye); Uzbekistan [I]; North Korea; South Korea (incl. Cheju Isl., Ulleung Isl.); Japan (Hokkaido, Honshu, Shikoku, Kyushu); Ryukyu Isl.; Taiwan; Vietnam [I]; Canada [I] (Ontario [I], Québec [I]); USA [I] (Alabama [I], Arkansas [I], Connecticut [I], District of Columbia [I], Delaware [I], Georgia [I], Iowa [I], Illinois [I], Indiana [I], Kansas [I], Kentucky [I], Massachusetts [I], Maryland [I], Maine [I], Michigan [I], Minnesota [I], Missouri [I], North Carolina [I], North Dakota [I], Nebraska [I], New Hampshire [I], New Jersey [I], New York [I], Ohio [I], Pennsylvania [I], Rhode Island [I], South Carolina [I], South Dakota [I], Tennessee [I], Virginia [I], Vermont [I], Wisconsin [I], West Virginia [I]); Argentina [I]
10 den swe 11 aut ger hun pol 12 fra 13 cro ita kos rom slv SRB 14 ukr 31 RFE 32 uzb 38 JAP KOR NKO NNS TAI 41 vie 72 cdn 74 usa 75 usa 78 usa 85 arg
Synonyms: hidden
= Antidesma scandens Lour.; Fl. Cochinch. 617 (1790), nom. rej. prop.
= Humulopsis scandens (Lour.) Grudz.; Bot. Zhurn. (Moscow & Leningrad) 73(4): 592 (1988)
= Humulus aculeatus Nutt.; J. Acad. Nat. Sci. Philadelphia, Ser. 2, 1: 182 (1848)
= Humulus japonicus Siebold & Zucc.; Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. 4(3): 213 (1846)
= Humulus japonicus var. minor Nakai; Saishu-to Kuan-to Shokubutsu Hokoku-sho [Fl. Quelpaert Isl.]: 38 (1914)
= Humulus japonicus var. variegatus F.Roem.; Amer. Florist (Chicago, 1885-1931) 8: 489 (1892)
= Humulus lupulus Thunb.; Fl. Jap. [Thunberg] 113 (1784) non L.
= Humulus scandens var. variegatus (Siebold & Zucc.) Moldenke; Phytologia 1: 169 (1935)
= Lupulus scandens (Lour.) St.-Lag.; Étude Fl. [A.Cariot], éd. 8, 2: 733 (1889)
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A complete, synonymic checklist of the Higher Plants of the World

Version 25.04 (April 17th, 2025)

Covering 356,375 accepted species,
8,782 hybrids, 48,090 accepted infraspecific taxa and 1,002,897 synonyms,
in total 1,416,144* names

[*about 25,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus" are deliberately omitted]

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