290.0000
Family
Cleomaceae Bercht. & J.Presl; Prir. Rostlin 2(64): 253 (1825)
28 genera, ca. 270 spp. Phylogeny: Saunders et al. 2024
Species listed in database: 251; hybrids: 2
= Cleomeae DC.; Prodr. [A. P. de Candolle] 1: 237 (1824)
= Cleomineae J.Presl; Nowoceská Bibl. [Wsobecný Rostl.] 7: 91 (1846)
= Cleomoideae Burnett; Outlines Bot.: 867 (1835)
= Dipterygieae Kuntze; T. E. von Post & C. E. O. Kuntze, Lex. Gen. Phan.: 646 (1903)
= Dipterygiinae Boiss.; Fl. Orient. 1: 417 (1867)
= Dipterygioideae Pax; Nat. Pflanzenfam. [Engler & Prantl] III, 2: 220, 225 (1891)
= Oxystylidaceae Hutch.; Evol. Phylog. Fl. Pl.: 516 (1969)
290.0002
Genus
Cleome L.; Sp. Pl.: 671 (1753)
Distribution: Tropics and subtropics, few north into Mediterranean and C-Asia. More species will need separation into split genera
Species listed in database: 31; hybrids: 1
= Buhsea Bunge; Del. Sem. Hort. Dorpat. 41 (1859)
= Diorimasperma Raf.; Sylva Tellur. 111 (1838)
= Micambe Adans.; Fam. 2: 407 (1763)
= Siliquaria Forssk.; Fl. Aegypt. Arab. 78 (1775)
= Sinapistrum Mill.; Gard. Dict. Abridg., ed. 4 (1754)
= Stylista Raf.; Sylva Tellur. 112 (1838)
species :
Cleome albescens
Franch.; Revoil. Comali, 11 (1882)
Distribution: NE-Somalia
24 SOM
species :
Cleome amblyocarpa
Barratte & Murb.; Lunds Univ. Arsskr. N. S. 2, 1 4: 25 (1905)
Distribution: Morocco; Western Sahara; Algeria; Tunisia; Libya; Egypt (Eastern Desert, Great Southwestern Desert, NE-Egypt, SE-Egypt); Mauritania; Mali; Niger; Chad; Sudan; Eritrea; Iraq (S-Iraq, W-Iraq: Desert); Iran (S-Iran); Israel (Rift Valley, N-Negev Desert); Sinai peninsula (C-Sinai, N-Sinai, S-Sinai); Jordania (S-Jordania); Saudi Arabia (Hejaz); Qatar; United Arab Emirates; Oman; Yemen (Aden Desert, Inner Hadhramaut, N-Inner Yemen, SW-Yemen, W-Yemen)
20 ALG EGY LBY MOR TUN WSA 22 MLI MTN NGR 24 CHA ERI SUD 34 IRN IRQ ISR JOR SIN 35 OMA QAT SAU UAE YEM
Synonyms: hidden
= Cleome africana Botsch.; Novosti Sist. Vyss. Rast. 1: 130 (1964) (nom. superfl.)
= Cleome amblyocarpa var. glandulosa (Forssk.) Botsch.; Novit. Syst. pl. Vasc., Acad. URSS, 1968: 327 (1968)
= Cleome arabica subsp. amblyocarpa (Barratte & Murb.) Ozenda; Fl. Sahara: 247 (1958) (comb. inval.)
= Cleome arabica var. lutea Parsa; Kew Bull. 2: 17 (1947)
= Cleome daryoushiana Parsa; Fl. Iran [Parsa] 1(1): 909 (1951)
= Cleome hexandra Poir. ex Steud.; Nomencl. Bot. 1: 206 (1821), pro syn.
= Pedicellaria glandulosa (Forssk.) P.Lima; Broteria, Ser. Bot. 19: 117 (1921)
= Siliquaria glandulosa Forssk.; Fl. Aegypt. Arab. 78 (1775)
species :
Cleome arabica
L.; Cent. Pl. 1: 20 (1775)
Distribution: Egypt (Eastern Desert, NE-Egypt, SE-Egypt); Iran (S-Iran); Israel (coastal W-Israel, Rift Valley, N-Negev Desert, Judean Desert); Sinai peninsula (C-Sinai, S-Sinai); Jordania (S-Jordania); Saudi Arabia (C-Saudi Arabia, NE-Saudi Arabia, N-Saudi Arabia, Nafud Desert, Hejaz, Nejd Desert)
20 EGY 34 IRN ISR JOR SIN 35 SAU
Synonyms: hidden
= Cleome aschersoniana Pfund; Flora 57: 413 (1874)
= Cleome siliquaria R.Br.; Salt, Voy. Abyss., App.: lxv (1814)
= Cleome trinervia Fresen.; Mus. Senckenb. 1: 177, t. 11 (1834)
= Siliquaria arabica (L.) Raf.; Sylva Tellur. 111 (1838)
= Sinapistrum lanatum Moench; Methodus: 250 (1794)
species :
Cleome ariana
Hedge & Lamond; Rech.f., Fl. Iran., Lief. 68, 17 (1970)
Distribution: Tajikistan; Afghanistan (Baghlan, Bamyan, Ghazni, Kabul, Wardak, Orozgan / Daykundi, Paktia / Khost, Parwan, Takhar); Pakistan (Baluchistan, Waziristan, Kurram, Chitral); India (Jammu & Kashmir)
32 TZK 34 AFG 40 IND PAK
species :
Cleome coluteoides
Boiss.; Diagn. ser. 1, 1: 3 (1843)
Distribution: Turkmenistan; Iraq (W-Iraq: Desert); Iran (EC-Iran, E-Iran, NE-Iran: Mts., N-Iran, Iranian Aserbaijan, S-Iran, W-Iran); Afghanistan (Farah, Herat)
32 TKM 34 AFG IRN IRQ
Synonyms: hidden
= Buhsea coluteoides (Boiss.) Bunge; Index Seminum (TU, Dorpatensis) 1859: 41 (1895)
= Buhsea raddeana (Trautv.) A.Boriss.; Fl. Turkmen. 3: 164 (1948)
= Buhsea raddeana (Trautv.) Kamelin; Bot. Zhurn. S.S.S.R. 55: 1462 (1970)
= Buhsea trinervia (DC.) Stapf; Denkschr. Kaiserl. Akad. Wiss., Wien. Math.-Naturwiss. Kl. 51: 306 (1886)
= Buhsea trinervia var. ramosa Bornm.; Bull. Herb. Boissier, sér. 2, 5: 57 (1904)
= Buhsia raddeana (Trautv.) Kamelin; Bot. J., URSS 55: 1462 (1970)
= Cadaba trinervia DC.; Prodr. [A. P. de Candolle] 1: 244 (1824)
= Capparis coluteoides Boiss.; Diagn. Pl. Orient. 13: 83 (1854)
= Cleome raddeana Trautv.; Act. Hort. Petrop. 1: 19 (1871)
= Stroemia trinervia (DC.) D.Dietr.; Syn. Plant. 1: 802 (1839)
species :
Cleome foliolosa
DC.; Prodr. [A. P. de Candolle] 1: 240 (1824)
Distribution: Iran (EC-Iran, SW-Iran)
34 IRN
Synonyms: hidden
= Cleome melilotoides Turcz.; Bull. Soc. Imp. Naturalistes Moscou 27(II): 320 (1854 publ. 1855)
species :
Cleome frutescens
Aubl.; Pl. Gui. 2: 677 (1775)
Distribution: French Guiana
82 FRG
species :
Cleome glaucescens
DC.; Prodr. [A. P. de Candolle] 1: 239 (1824)
Distribution: Iraq (NE-Iraq, NW-Iraq, SE-Iraq: Mesopotamia, W-Iraq: Desert); Iran (W-Iran); Syria (Jazira, W-Syrian Mountains); Saudi Arabia (C-Saudi Arabia, N-Saudi Arabia, Hejaz); Bahrain; Oman (Mascat & Oman)
34 IRN IRQ SYR 35 BAR OMA SAU
Synonyms: hidden
= Cleome kotschyana Boiss.; Fl. Orient. 1: 413 (1867)
= Cleome glauca DC.; Prodr. [A. P. de Candolle] 1: 239 (1824)
species :
Cleome gobica
Grubov; Opred. Sosud. Rast. Mongolii (Key Vasc. Pl. Mongolia): 118 (1982)
Distribution: Mongolia
37 MON
species :
Cleome gordjaginii
Popov; Korovin, Kult. & Popov, Descr. Pl. Nov. Turkmenistan. (Bodenk. u. Pflanzengeogr. Beob. u. Unters. 1: Flussgeb. d. Amu Darya u. Syr Darya, II.), 51 (1916)
Distribution: Turkmenistan; Uzbekistan
32 TKM UZB
species :
Cleome hadramautica
Thulin; Nordic J. Bot. 22(2): 216 (2002 publ. 2003)
Distribution: S-Yemen
35 YEM
species :
Cleome heratensis
Bunge & Bien. ex Boiss.; Fl. Orient. 1: 412 (1867)
Distribution: Iran (E-Iran, NE-Iran: Mts., S-Iran); Afghanistan (Herat)
34 AFG IRN
Synonyms: hidden
= Cleome gaubae Rech.f.; Repert. Spec. Nov. Regni Veg. 48: 38 (1940)
species :
Cleome iberica
DC.; Prodr. [A. P. de Candolle] 1: 240 (1824)
Name: Kaukasus-Spinnenblume (DE); Caucasian Beeplant (EN)
Distribution: France [I]; European Russia; Crimea; Northern Caucasus; Georgia [Caucasus]; Armenia; Azerbaijan; East Aegaean Isl. (Rhodos); Turkey (E-Anatolia, N-Anatolia, NE-Anatolia, SE-Anatolia); Iraq (NE-Iraq, NW-Iraq); Iran (EC-Iran, N-Iran, Iranian Aserbaijan, S-Iran, W-Iran); Cyprus (C-Mountains); Lebanon (Antilebanon); USA [I] (Kentucky [I])
12 fra 14 KRY RUS 33 ARM AZE GEO NCS 34 CYP EAI IRN IRQ LEB TUR 78 usa
Remarks: ornithopodioides agg.; the plants from Rhodos and SW-Anatolia might represent an undescribed, local taxon fide Strid (2024)
Synonyms: hidden
= Cleome canescens Steven ex DC.; Prodr. [A. P. de Candolle] 1: 241 (1824)
= Cleome circassica Tzvelev; Notul. Syst. Herb. Inst. Bot. Acad. Sci. URSS 22: 132 (1963)
= Cleome daghestanica (Rupr.) Tzvelev; Bot. Mater. Gerb. Bot. Inst. Komarova Akad. Nauk S.S.S.R. 22: 128 (1963)
= Cleome donetzica Tzvelev; Bot, Mater. Gerb. Bot. Inst. Komarova Akad. Nauk SSSR 22: 131 (1963)
= Cleome ornithopodioides f. daghestanica Rupr.; Mem. Acad. Sci. Petersb., ser. 7, 15 2: 137 (1869)
= Cleome ornithopodioides subsp. canescens (DC.) Tzvelev; Fedor., Fl. Evr. Casti SSSR 4: 30 (1979)
= Cleome ornithopodioides subsp. donetzica (Tzvelev) Tzvelev; Fedor., Fl. Evr. Casti SSSR 4: 30 (1979)
= Cleome ornithopodioides var. sessilis Boiss.; Fl. Orient. 1: 411 (1867)
species :
Cleome karachiensis
S.Riaz, Abid & Qaiser; Acta Bot. Croat. 78(1): 103 (2019)
Distribution: Pakistan
40 PAK
species :
Cleome karjaginii
Tzvelev; Conspectus Fl. Kavk. 3(2): 370 (2012)
Distribution: Armenia
33 ARM
species :
Cleome khorassanica
Bunge & Bien. ex Boiss.; Fl. Orient. 1: 412 (1867)
Distribution: Turkmenistan; Iran (NE-Iran: Mts.)
32 TKM 34 IRN
Remarks: belongs together with C. turkmena in an early Cleomaceae clade and needs to be spun off fide Patchell et al. 2014
species :
Cleome moricandii
Briq.; Annuaire Conserv. & Jard. Bot. Geneve 17: 376 (1914)
Distribution: Peru
83 PER
species :
Cleome omanensis
(D.F.Chamb. & Lamond) Thulin; Nordic J. Bot. 22(2): 217 (2002 publ. 2003)
Distribution: Oman; S-Yemen
35 OMA YEM
Synonyms: hidden
= Cleome albescens subsp. omanensis D.F.Chamb. & Lamond; Edinburgh J. Bot. 51(1): 52 (1994)
species :
Cleome ornithopodioides
L.; Sp. Pl.: 672 (1753)
Name: Bird spiderflower
Distribution: North Macedonia; Albania; Bulgaria; Greece (NE-Greece, rare NW-Greece); East Aegaean Isl. (Lesvos); Turkey (Inner Anatolia, NW-Anatolia: Bithynia, S-Anatolia, SE-Anatolia, SSW-Anatolia, SW-Anatolia); Cyprus (C-Mountains, N-Cyprus); USA [I] (Maryland [I], Pennsylvania [I])
13 ALB BUL GRC NMA 34 CYP EAI TUR 75 usa 78 usa
Remarks: ornithopodioides agg.
Synonyms: hidden
= Cleome aurea Celak.; Oesterr. Bot. Z. 34: 113 (1884), nom. illeg.
= Cleome cypria Celak.; Österr. Bot. Z. 34: 114 (1884)
= Cleome dilleniana DC.; Prodr. [A. P. de Candolle] 1: 240 (1824)
= Cleome macedonica Heldr. ex Nadji; Emp. Ottom. Geogr. Bot.: 11 (1892)
= Cleome ornithopodioides subsp. cypria (Celak.) Holmboe; Bergens Mus. Skr. ser. 2, 1(2): 93 (1914)
= Micambe ornithopodioides (L.) Medik.; Phil. Bot. 1: 43 (1789)
species :
Cleome oxypetala
Boiss.; Diagn. ser. 1, 4: 20 (1844)
Distribution: Iran (E-Iran, S-Iran); Afghanistan (Farah, Helmand, Herat, Kandahar); Saudi Arabia (C-Saudi Arabia, NE-Saudi Arabia); Oman (Mascat & Oman); Pakistan (Baluchistan)
34 AFG IRN 35 OMA SAU 40 PAK
Synonyms: hidden
= Cleome hotsonii Blatt. & Hallb.; J. Indian Bot. 1: 58 (1919)
species :
Cleome pakistanica
(Jafri) Khatoon & A.Perveen; Pakistan J. Bot. 35(2): 145 (2003)
Distribution: Pakistan (Baluchistan)
40 PAK
Synonyms: hidden
= Cleome heratensis subsp. pakistanica Jafri; Fl. W. Pakistan 34: 25 (1973)
species :
Cleome postrata
D.Subram.; Pl. Arch. 17(2): 1785, t. 4 (2017)
Distribution: India (Tamil Nadu)
40 IND
species :
Cleome potosina
B.L.Rob.; Proc. Amer. Acad. Arts 27: 165 (1893)
Distribution: Mexico (Queretaro, San Luis Potosi)
79 MEX
species :
Cleome pulchella
(Lindl.) Schult. & Schult.f.; Roem. & Schult., Syst. Veg. ed. 15[bis]. 7: 1617 (1830)
Distribution: Brazil (Maranhao)
84 BRA
Remarks: provisionally accepted name
Synonyms: hidden
= Gynandropsis pulchella Lindl.; Trans. Hort. Soc. London 7: 65 (1830)
species :
Cleome ramosissima
Parl. ex Webb; Fragm. Fl. Aethiop.-Aegypt.: 22 (1854)
Distribution: Sudan; Ethiopia; Eritrea; Djibouti; N-Somalia; Saudi Arabia; Yemen (W-Yemen, Inner Hadhramaut)
24 DJI ERI ETH SOM SUD 35 SAU YEM
Synonyms: hidden
= Cleome arabica var. stenocarpa Franch.; Sert. Somal.: 11 (1882)
= Cleome schweinfurthii Gilg; Notizbl. Bot. Gart. Berlin-Dahlem 1: 62 (1895)
= Polanisia ramosissima (Parl. ex Webb) T.Durand & Schinz; Consp. Fl. Afric. 1(2): 163 (1898)
species :
Cleome rupicola
Vicary; J. Asiat. Soc. Bengal 16: 1158 (1847), et Ann. Nat. Hist. Ser. 2, 1: 425 (1848)
Distribution: Iraq (NW-Iraq); Iran (S-Iran); Saudi Arabia; United Arab Emirates; Oman; Pakistan (Sind, Baluchistan, N. W. Frontier Prov., Waziristan)
34 IRN IRQ 35 OMA SAU UAE 40 PAK
Synonyms: hidden
= Cleome oxypetala var. micrantha Boiss.; Fl. Orient. 1: 415 (1867)
= Cleome stocksiana Boiss.; Diagn. ser. 2, 1: 47 (1853)
species :
Cleome socotrana
Balf.f.; Proc. Roy. Soc. Edinb. 11: 501 (1882)
Distribution: Soqotra
24 SOC
species :
Cleome steveniana
Schult. & Schult.f.; Roem. & Schult., Syst. Veg. 7: 40 (1829)
Distribution: Georgia [Caucasus]; Armenia; Azerbaijan; Iran (EC-Iran, W-Iran, W-Iran)
33 ARM AZE GEO 34 IRN
Remarks: ornithopodioides agg.
Synonyms: hidden
= Cleome ornithopodioides var. lactea Bornm.; Bull. Herb. Boissier, sér. 2, 5: 57 (1904)
= Cleome ornithopodioides var. stipitata Boiss.; Fl. Orient. 1: 411 (1867)
= Cleome virgata Steven; Prodr. [A. P. de Candolle] 1: 240 (1824) (non Thunb. 1800)
species :
Cleome turkmena
Bobrov; Kom., Fl. URSS 8: 6, 633 (1939)
Distribution: Turkmenistan
32 TKM
Remarks: belongs together with C. khorassanica in an early Cleomaceae clade and needs to be spun off fide Patchell et al. 2014
species :
Cleome violacea
L.; Sp. Pl.: 672 (1753)
Distribution: Portugal; Spain; Morocco
12 POR SPA 20 MOR
Synonyms: hidden
= Cleome canadensis Schult. & Schult.f.; Roem. & Schult., Syst. Veg. ed. 15[bis]. 7: 38 (1829)
= Cleome cardaminoides Hoffmanns.; Verz. Pfl.-Kult., Nachtr. 2: 93 (1826)
= Cleome deflexa Ruiz & Pav. ex DC.; Prodr. [A. P. de Candolle] 1: 240 (1824)
= Cleome secundiflora Schrad. ex Schult. & Schult.f.; Roem. & Schult., Syst. Veg. ed. 15[bis]. 7: 37 (1829)
= Cleome violacea f. unicolor Cout.; Fl. Portugal: 273 (1913)
= Cleome violacea var. ortegae Pau & Merino; Fl. Galicia 3: 516 (1909)
= Diorimasperma violacea (L.) Raf.; Sylva Tellur.: 111 (1838)
species :
Cleome yunnanensis
W.W.Sm.; Notes Roy. Bot. Gard. Edinburgh 12: 199 (1920)
Distribution: China (Yunnan)
36 CHI
Remarks: provisionally accepted name
species :
Cleome ×chrysogyna
Gilg ex Heilborn; Ark. Bot. 23A(10): 13 (1931)
Distribution: Ecuador
83 ECU
