046.0000 Family Athyriaceae Alston; Taxon 5: 25 (1956)

6-7 genera, ca. 760 spp. Circumscription following Rothfels et al. (2012). According to Fraser-Jenkins et al. (2018) it should be included in Woodsiaceae s. lat.

Species listed in database: 758; hybrids: 102

046.0008 Genus Athyrium Roth; Tent. Fl. Germ. 3: 58 (1790)

Distribution: Worldwide, mostly northern hemisphere, many in China (over 100), several in trop. Africa and Madagascar, few in the New World.

Species listed in database: 179; hybrids: 60

species :

Athyrium aberrans Liang Zhang & Li Bing Zhang; Phytotaxa 533(3): 167 (2022)

Distribution: SE-Tibet (Medog)

36 TIB

species :

Athyrium acutipinnulum Kodama ex Nakai; Rep. Veg. Ooryongto 13 (1919)

Distribution: South Korea

38 KOR

Remarks: provisionally accepted name, status unconfirmed

species :

Athyrium adpressum Ching & W.M.Chu; Acta Bot. Boreal.-Occid. Sin. 6(1): 13 (1986)

Distribution: China (S-Yunnan)

36 CHI

species :

Athyrium ammifolium (Mett.) C.Chr.; Index Filic. 139 (1905)

Distribution: Nigeria; Cameroon; Bioko Isl. [Fernando Poo]; Equatorial Guinea (Rio Muni)

22 NGA 23 BIO CMN EQG

Synonyms: hidden

= Asplenium ammifolium Mett.; Fil. Afr. [Kuhn] 96 (1868)

species :

Athyrium amoenum C.Chr.; Gard. Bull. Straits Settlem. 7: 267, t. 56 (1934)

Distribution: Borneo (Sabah)

42 BOR

Remarks: Flora of China treats this as synonym of A. rhachidosorum, but the areas are widely separated, so this is quite unlikely

species :

Athyrium angustum (Willd.) C.Presl; Reliq. Haenk. 1: 39 (1825)

Name: Subarctic ladyfern (EN)

Distribution: Greenland; Canada (Labrador, Manitoba, New Brunswick, Newfoundland, Nova Scotia, Ontario, Prince Edward Isl., Québec, Saskatchewan); St. Pierre et Miquelon; USA (Colorado, Connecticut, District of Columbia, Delaware, Iowa, Illinois, Indiana, Kentucky, Massachusetts, Maryland, Maine, Michigan, Minnesota, Missouri, North Carolina, North Dakota, New Hampshire, New Jersey, New York, Ohio, Pennsylvania, Rhode Island, South Dakota, Tennessee, Virginia, Vermont, Wisconsin, West Virginia)

70 GNL 71 CDN 72 CDN SPM 73 USA 74 USA 75 USA 78 USA

Remarks: often treated within A. filix-femina or as variety of A. asplenioides, but increasingly elevated to full species status

Synonyms: hidden

= Aspidium angustum Willd.; Sp. Pl., ed. 4 [Willdenow] 5(1): 277 (1810)

= Asplenium elatius Link; Fil. Spec. 94 (1841)

= Asplenium filix-femina var. angustum (Willd.) D.C.Eaton; Rep. U.S. Geogr. Surv., Wheeler 6(Botany): 330 (1878)

= Asplenium filix-femina var. exile D.C.Eaton; Rep. U.S. Geogr. Surv., Wheeler 6(Botany): 330 (1878)

= Asplenium michauxii Spreng.; Syst. Veg., ed. 16 [Sprengel] 4(1): 88 (1827)

= Athyrium angustum f. confertum Butters; Rhodora 19: 195 (1917)

= Athyrium angustum f. cristatum (Hopkins) Broun ex Weath.; Amer. Fern J. 26: 133 (1936)

= Athyrium angustum f. elegans (Gilbert) Butters; Rhodora 19: 196 (1917)

= Athyrium angustum f. laciniatum Butters; Rhodora 19: 195 (1917)

= Athyrium angustum var. boreale Jenn.; Amer. Fern J. 8: 82 (1918)

= Athyrium angustum var. cristatum Hopkins; Amer. Fern J. 9: 88 (1919)

= Athyrium angustum var. elatius (Link) Butters; Rhodora 19: 191 (1917)

= Athyrium angustum var. glanduliferum Jenn.; Amer. Fern J. 8: 82 (1918)

= Athyrium angustum var. laurentianum Butters; Rhodora 19: 194 (1917)

= Athyrium angustum var. rubellum (Gilbert) Butters; Rhodora 19: 193 (1917)

= Athyrium asplenioides var. angustum (Willd.) T.Moore; Index Fil. [T. Moore] 179 (1860)

= Athyrium filix-femina f. confertum (Butters) Fernald; Rhodora 48(no. 574): 389 (1946)

= Athyrium filix-femina f. cristatum (Hopkins) Clute; Our Ferns: 224, 384 (1938)

= Athyrium filix-femina f. elatius (Link) Clute; Ferns in Their Haunts 224 (1938)

= Athyrium filix-femina f. elegans (Gilbert) Clute; Ferns in Their Haunts 224 (1938)

= Athyrium filix-femina f. laciniatum (Butters) Fernald; Rhodora 48(no. 574): 389 (1946)

= Athyrium filix-femina f. laurentianum (Butters) Fernald; Rhodora 48(no. 574): 389 (1946)

= Athyrium filix-femina f. plano-rhaeticum Gilbert; List N. Amer. Pterid.: 15, 34 (1901)

= Athyrium filix-femina subsp. angustum (Willd.) Hultén; Kongl. Svenska Vetensk. Acad. Handl. 8(5): 178 (1962), isonym

= Athyrium filix-femina subsp. angustum (Willd.) R.T.Clausen; Cornell Univ. Agric. Exp. Sta. Mem. 291: 7 (1949)

= Athyrium filix-femina subvar. elatius (Link) Farw.; Amer. Midl. Naturalist 10: 203 (1927)

= Athyrium filix-femina subvar. elegans (Gilbert) Farw.; Amer. Midl. Naturalist 10: 203 (1927)

= Athyrium filix-femina var. angustum (Willd.) G.Lawson; Edinburgh New Philos. J., ser. 2, 19: 114 (1864)

= Athyrium filix-femina var. elegans Gilbert; List N. Amer. Pterid.: 33-34 (1901)

= Athyrium filix-femina var. michauxii (Spreng.) Farw.; Rep. (Annual) Michigan Acad. Sci. 18: 79 (1916)

= Athyrium filix-femina var. rubellum Gilbert; List N. Amer. Pterid.: 35 (1901)

= Athyrium michauxii (Spreng.) Fée; Mém. Foug. 5 (Gen. Filic.): 186 (1852)

= Athyrium michauxii f. furcata Hahne; Fern Bull. 12: 116 (1904)

species :

Athyrium anisopterum Christ; Bull. Herb. Boissier, sér. 1, 6: 962 (1898)

Distribution: China (Guangdong, Guangxi, Guizhou, Hunan, Jiangxi, Sichuan, Yunnan); Tibet; Taiwan; Nepal; Bhutan; India (Arunachal Pradesh, Himachal Pradesh, ?Karnataka, Kerala, Meghalaya, Sikkim, Tamil Nadu, Tripura, Uttarakhand, West Bengal); Sri Lanka; Myanmar (Chin, etc.); N-Thailand; Vietnam; peninsular Malaysia (Cameron Highlands, Pahang); Sumatra; Borneo (Mt. Kinabalu, Sabah); Philippines (Luzon); Java

36 CHI TIB 38 TAI 40 BHU IND NEP SRL 41 MYA THA VIE 42 BOR JAW MLY PHI SUM

Remarks: sect. Polystichoides, anisopterum clade. Not in Nepal

Synonyms: hidden

= Aspidium fauriei var. elatius Christ; Bull. Herb. Boissier, sér. 1, 6(3): 193 (1898)

= Athyrium kumaonicum Holttum ex Punetha; Indian Fern J. 2(1-2): 29 (1985) [fide Fraser-Jenkins et al. 2018]

= Athyrium membranifolium Ching; (1976), nom. nud.

= Athyrium woodsioides Christ; Bull. Acad. Int. Géogr., sér. 3, 16: 124 (1906), nom. illeg. superfl.

= Athyrium ×heterosporum Y.T.Hsieh & Z.R.Wang; Acta Phytotax. Sin. 27(2): 156 (1989) (fide Fraser-Jenkins 1997, 2018)

= Dryopteris thysanocarpa Hayata; Icon. Pl. Formosan. 4: 160 (1914), pro syn.

= Lastrea membranacea Arn. ex Baker; Syn. Fil. [W. J. Hooker & Baker], ed. 2: 498 (1874) (nom. inval.)

= Lastrea sparsa var. membranacea Arn. ex Bedd.; Suppl. Ferns S. Ind. 17 (1876) (nom. inval. in syn.)

= Nephrodium sparsum var. membranaceum Arn. ex C.B.Clarke; (1880) (nom. inval.)

species :

Athyrium annae Kornas; Kew Bull. 33(1): 99 (1978)

Distribution: Zambia

26 ZAM

Remarks: status doubtful, probably a synonym of A. lewalleanum or not belonging to Athyrium at all (Verdcourt in Fl. Trop. East Africa)

species :

Athyrium appendiculiferum Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 16: 3 (1914)

Distribution: peninsular Malaysia (Cameron Highlands, Pahang); Sumatra; Java

42 JAW MLY SUM

species :

Athyrium araiostegioides Ching; Acta Bot. Boreal.-Occid. Sin. 6(2): 103 (1986)

Distribution: China (Yunnan, SW-Sichuan)

36 CHI

Remarks: sect. Strigoathyrium

species :

Athyrium archboldii Copel.; Univ. Calif. Publ. Bot. 18: 222 (1942)

Distribution: New Guinea (Irian Jaya)

42 NWG

species :

Athyrium arcuatum Liebm.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 5, 1: 278 (seors. 126) (1849)

Distribution: Mexico (Chiapas, Colima, Ciudad de Mexico, Guanajuato, Guerrero, Hidalgo, Jalisco, Mexico State, Michoacan, Morelos, Nayarit, Oaxaca, Puebla, Queretaro, Veracruz); Guatemala; Honduras

79 MEX 80 GUA HON

Remarks: included by Fl. Mesoamericana in A. filix-femina, kept separate by subsequent authors

Synonyms: hidden

= Asplenium arcuatum (Liebm.) Hook.; Sp. Fil. [W. J. Hooker] 3: 219 (1860)

= Asplenium martensii Kunze; Amer. J. Sci. Arts, ser. 2, 6: 86 (1848), nom. nud.

= Asplenium martensii Kunze ex Mett.; Abh. Senckenberg. Naturf. Ges. 3(1) [Aspl.]: 244 (1859)

= Athyrium barnebyanum Mickel & Beitel; Mem. New York Bot. Gard. 46 [Pterid. Fl. Oaxaca]: 77 (1988)

= Athyrium galeottii Fée; Mém. Foug. 5 (Gen. Filic.): 187 (1852)

= Athyrium martensii (Kunze ex Mett.) T.Moore; Index Fil. [T. Moore] 186 (1860)

= Athyrium pumilio Christ; Bull. Herb. Boissier, sér. 2, 8: 147 (1903)

species :

Athyrium arisanense (Hayata) Tagawa; Acta Phytotax. Geobot. 2(3): 195 (1933)

Distribution: ?Japan; Taiwan

38 TAI

Remarks: sect. Strigoathyrium, arisanense agg.

Synonyms: hidden

= Diplazium arisanense Hayata; Icon. Pl. Formosan. 4: 212, f. 144 (1914)

species :

Athyrium asplenioides (Michx.) Desv.; Mem. Soc. Linn. Paris 6(2): 264 (1827)

Name: Asplenium ladyfern

Distribution: USA (Alabama, Arkansas, Connecticut, District of Columbia, Delaware, Florida, Georgia, Illinois, Indiana, Kansas, Kentucky, Louisiana, Massachusetts, Maryland, Missouri, Mississippi, North Carolina, New Jersey, New York, Ohio, Oklahoma, Pennsylvania, Rhode Island, South Carolina, Tennessee, Texas, Virginia, West Virginia)

74 USA 75 USA 77 USA 78 USA

Remarks: separate from A. filix-femina fide Fraser-Jenkins (2008)

Synonyms: hidden

= Aspidium asplenioides (Michx.) Sw.; Syn. Fil. [Swartz] 60 (1806)

= Asplenium athyrium Spreng.; Anleit. Kenntn. Gew. 3: 106 (1804)

= Athyrium asplenioides f. ellipticum Wherry; Amer. Fern J. 38: 158 (1949)

= Athyrium asplenioides f. subtripinnatum Butters; Rhodora 19: 190 (1917)

= Athyrium asplenioides var. subtripinnatum (Butters) Wherry; Bartonia 21: 14 (1942)

= Athyrium filix-femina f. ellipticum (Wherry) C.F.Reed; Castanea 16: 71 (1951)

= Athyrium filix-femina f. rectangulare Gilbert; List N. Amer. Pterid.: 15, 35 (1901)

= Athyrium filix-femina f. subtripinnatum (Butters) Fernald; Rhodora 48(no. 574): 389 (1946)

= Athyrium filix-femina subsp. asplenioides (Michx.) Hultén; Kongl. Svenska Vetensk. Acad. Handl. 8(5): 178 (1962)

= Athyrium filix-femina var. asplenioides (Michx.) Farw.; Pap. Michigan Acad. Sci. 2: 13 (1923)

= Nephrodium asplenioides Michx.; Fl. Bor.-Amer. (Michaux) 2: 268 (1803)

species :

Athyrium atkinsonii Bedd.; Ferns Brit. India Suppl. 11, t. 369 (1876)

Distribution: China (Chongqing, Fujian, Gansu, Guizhou, Henan, Hubei, Hunan, Jiangxi, S-Shaanxi, Shanxi, Sichuan, Yunnan); Tibet; Japan; Taiwan; Pakistan (Swat); Nepal; Bhutan; India (Jammu & Kashmir, Arunachal Pradesh, Himachal Pradesh, Sikkim, Uttarakhand, West Bengal); Myanmar (Chin, etc.); Vietnam

36 CHI TIB 38 JAP TAI 40 BHU IND NEP PAK 41 MYA VIE

Synonyms: hidden

= Aspidium lastreoides Baker; J. Bot. 26(7): 227 (1888)

= Aspidium senanense Franch. & Sav.; Enum. Pl. Jap. 2: 241, 636 (1877)

= Asplenium andersonii C.B.Clarke; J. Linn. Soc., Bot. 16: 154 (1876)

= Asplenium atkinsonii (Bedd.) C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 487 (1880)

= Asplenium atkinsonii var. andersonii (C.B.Clarke) C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 487 (1880)

= Asplenium fimbriatum var. sphaeropteroides C.B.Clarke; (1880)

= Asplenium lastreoides Baker; J. Bot. 26: 227 (1888)

= Asplenium microsorum Makino; Bot. Mag. (Tokyo) 6(60): 46 (1892)

= Athyrium andersonii (C.B.Clarke) Panigrahi & Basu; Proc. Ind. Sci. Congr., Lucknow, Bot., Abstr. 72: 14 (1985) (fide Fraser-Jenkins 1997)

= Athyrium atkinsonii var. andersonii (C.B.Clarke) C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 494 (1880)

= Athyrium filix-femina var. deltoideum Christ; Mém. Soc. Bot. France 1 [Hist. Acrostich.]: 46 (1905), nom. illeg.

= Athyrium fimbriatum var. sphaeropteroides (C.B.Clarke) Bedd.; Trans. Linn. Soc. London, Bot., ser. 2, 1: 495 (1883)

= Athyrium lastreoides (Baker) Diels; Nat. Pflanzenfam. [Engler & Prantl] I, 4: 224 (1899)

= Athyrium microsorum Makino; Bot. Mag. (Tokyo) 18: 25 (1899)

= Athyrium monticola Rosenst.; Repert. Spec. Nov. Regni Veg. 13: 123 (1913)

= Athyrium senanense (Franch. & Sav.) Koidz. & Tagawa; Tagawa, Acta Phytotax. Geobot. 3: 34 (1934)

= Cystopteris grandis C.Chr.; H. Lév., Cat. Pl. Yun-Nan 100 (1916)

= Davallia athyriifolia Baker; Ann. Bot. (Oxford) 6: 200 (1891)

= Dryopteris gracilifrons C.Chr.; Index Filic. 268 (1906) (nom. nov. for Polypodium leptophyllum Baker)

= Dryopteris senanensis (Franch. & Sav) C.Chr.; Index Filic. 291 (1905)

= Polypodium leptophyllum Baker; Ann. Bot. (Oxford) 5: 458 (1891) (non L. (1753) nec Moritz ex Mett. (1864))

= Pseudocystopteris andersonii (C.B.Clarke) Ching; Acta Phytotax. Sin. 9: 79 (1964) (fide Fraser-Jenkins 1997)

= Pseudocystopteris atkinsonii (Bedd.) Ching; Acta Phytotax. Sin. 9(1): 78 (1964)

species :

Athyrium atratum Bedd.; Ferns Brit. India Suppl. 33 (1892)

Distribution: India (Arunachal Pradesh, Manipur); Vietnam; Sumatra; Java

40 IND 41 VIE 42 JAW SUM

Remarks: sect. Strigoathyrium, non-setose group

Conservation Status: IUCN EN for India (Fraser-Jenkins 2012)

Synonyms: hidden

= Asplenium gedeanum Racib.; Pteridoph. Buitenzorg 223 (1898)

= Athyrium gedeanum (Racib.) Christ; J. Bot. (Morot) 19: 68 (1905) [fide Fraser-Jenkins et al. 2018]

species :

Athyrium attenuatum (C.B.Clarke) Tagawa; Acta Phytotax. Geobot. 16: 177 (1956)

Distribution: E-Afghanistan; China (SE-Gansu, Guizhou, W-Sichuan, N-Yunnan); Tibet; Pakistan (Azad Kashmir, Kurram, Swat, Hazara); Nepal; N-India (Jammu & Kashmir, ?Arunachal Pradesh, Himachal Pradesh, Sikkim, Uttarakhand, West Bengal); Myanmar

34 AFG 36 CHI TIB 40 IND NEP PAK 41 MYA

Remarks: A. dentigerum included here by Fraser-Jenkins et al. (2015 etc.) but kept separate by Flora of China

Synonyms: hidden

= Asplenium filix-femina f. dentigerum (Wall. ex C.B.Clarke) C.Hope; J. Bombay Nat. Hist. Soc. 14: 125 (1902)

= Asplenium filix-femina var. attenuatum C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 492, t. 59(1) (1880)

= Asplenium filix-femina var. dentigerum C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 491 (1880) (fide Fraser-Jenkins 1997)

= Athyrium angustifrons Ching; (1976) (nom. nud.) (fide Fraser-Jenkins 2008)

= Athyrium attenuatum f. dentigerum (Wall. ex C.B.Clarke) Khullar & Fraser-Jenk.; Aspects Pl. Sci. 6: 102 (1983)

= Athyrium attenuatum f. intermedium Mehra & Khullar ex Fraser-Jenk. & Khullar; Aspects Pl. Sci. 6: 99 (1983)

= Athyrium dentigerum (C.B.Clarke) Mehra & Bir; Amer. Fern J. 50: 279 (1960)

= Athyrium ensiferum Ching & H.S.Kung; Acta Bot. Boreal.-Occid. Sin. 6(2): 102 (1986) (fide Fraser-Jenkins 2008)

= Athyrium excelsius Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 20 (1986) (non Nakai 1914) (fide Fraser-Jenkins 2008)

= Athyrium filix-femina var. attenuatum (C.B.Clarke) Bedd.; Ferns Brit. India 169 (1883)

= Athyrium filix-femina var. dentigerum (C.B.Clarke) Bedd.; Ferns Brit. India 169 (1883)

= Athyrium leiboense Ching & Y.T.Hsieh; Acta Bot. Boreal.-Occid. Sin. 6(3): 158 (1986)

= Athyrium pseudofilix-femina Ching; Acta Bot. Austro Sin. 1: 17 (1983) (fide Fraser-Jenkins 1997)

= Athyrium tatsienluense Ching; (1976) (nom. nud.) (fide Fraser-Jenkins 2008)

= Polypodium dentigerum Wall.; Numer. List. [Wallich] no. 334 (1829), nomen (fide Fraser-Jenkins 1997)

species :

Athyrium auriculatum Seriz.; J. Jap. Bot. 46: 280 (1971)

Distribution: Taiwan

38 TAI

Remarks: sect. Strigoathyrium, non-setose group. Often included in A. clivicola but genetically separate

species :

Athyrium austroorientale Ching; Fl. Xizangica 1: 141 (1983)

Distribution: China (Sichuan, Yunnan); Tibet

36 CHI TIB

Synonyms: hidden

= Athyrium aculeatum Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 15 (1986)

= Athyrium dajinense Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 17 (1986)

= Athyrium takingense Ching; (1976), nom. nud.

species :

Athyrium baoxingense Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 14 (1986)

Distribution: China (SW-Sichuan)

36 CHI

Synonyms: hidden

= Athyrium paoshingense Ching; (1976), nom. nud.

species :

Athyrium bipinnatum K.Hori; PhytoKeys 148: 97 (2020)

Distribution: Japan (Shikoko, Honshu, Kyushu)

38 JAP

species :

Athyrium biserrulatum Christ; Bull. Acad. Int. Géogr. Bot. 17: 135-136 (1907)

Distribution: China (Sichuan, W-Guizhou, Yunnan); Tibet; Pakistan (Swat, Pakistani Punjab, Hazara); Nepal; Bhutan; India (Arunachal Pradesh, Himachal Pradesh, Madhya Pradesh, Nagaland, Rajasthan, Sikkim, Uttarakhand, West Bengal, Jammu & Kashmir); Myanmar

36 CHI TIB 40 BHU IND NEP PAK 41 MYA

Synonyms: hidden

= Asplenium filix-femina var. polysporum C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 493 (1880)

= Athyrium filix-femina var. polysporum (C.B.Clarke) Bedd.; Ferns Brit. India 170 (1883)

= Athyrium polysporum (C.B.Clarke) Ching ex Mehra & Bir; Amer. Fern J. 50: 289 (1960)

= Athyrium schimperi subsp. biserrulatum (Christ) Fraser-Jenk.; New Species Syndr. Indian Pteridol. 60 (1997)

= Athyrium wumonshanicum Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 20 (1986) (fide Fraser-Jenkins (2008) but kept separate by Flora of China)

species :

Athyrium bomicola Ching; Fl. Xizangica 1: 136 (1983)

Distribution: Tibet

36 TIB

species :

Athyrium bourgaei E.Fourn.; Mexic. Pl. 1: 102 (1872)

Distribution: Mexico (Chihuahua, Durango, Nayarit, Jalisco, Michoacan, Mexico State, Oaxaca, Veracruz, Hidalgo, Chiapas, Baja California Sur, México Distrito Federal, Guanajuato, Morelos, Nuevo Leon, Puebla, Sonora); Guatemala; El Salvador

79 MEX 80 ELS GUA

Remarks: included by Fl. Mesoamericana in A. filix-femina, kept separate by later Mexican floras

Synonyms: hidden

= Athyrium paucifrons C.Chr.; Repert. Spec. Nov. Regni Veg. 9: 371 (1911)

species :

Athyrium brevipinnulum Copel.; Philipp. J. Sci. 8: 291 (1908)

Distribution: Philippines

42 PHI

Remarks: sect. Strigoathyrium, valid species and not a synonym of A. nitidulum fide Wardani et al. 2024

species :

Athyrium brevistipes Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 12 (1986)

Distribution: China (Chongqing, Guizhou, Hubei, Hunan, S-Shaanxi, Sichuan)

36 CHI

species :

Athyrium caudatum Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 21 (1986)

Distribution: China (NW-Yunnan)

36 CHI

species :

Athyrium caudiforme Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 21 (1986)

Distribution: China (Hunan, W-Sichuan)

36 CHI

species :

Athyrium chingianum X.C.Zhang ex Z.R.Wang & X.C.Zhang; Bull. Bot. Res., Harbin 11(3): 12 (1991)

Distribution: China (Yunnan)

36 CHI

species :

Athyrium christensenii Tardieu; Aspl. Tonkin 80, t. 12 (1932)

Distribution: China (SE-Yunnan, SW-Guangxi); Vietnam

36 CHI 41 VIE

Remarks: sect. Strigoathyrium, arisanense agg.

Conservation Status: IUCN R

Synonyms: hidden

= Lunathyrium christensenii (Tardieu) Ching; Acta Phytotax. Sin. 9: 72 (1964)

= Athyrium puberulum Ching; (1976), nom. nud.

species :

Athyrium chungtienense Ching; Bull. Fan Mem. Inst. Biol. Bot., ser. 2, 1: 277 (1949)

Distribution: China (NW-Yunnan)

36 CHI

Synonyms: hidden

= Athyrium habaense Ching; Acta Bot. Boreal.-Occid. Sin. 6(3): 154 (1986)

= Athyrium confertum Ching; (1976), nom. nud.

species :

Athyrium clarkei Bedd.; Ferns Brit. India Suppl. II, t. 360 (1876)

Distribution: China (Guizhou, S-Yunnan); Nepal; India (Manipur, Nagaland, Sikkim, West Bengal, Darjeeling); Myanmar

36 CHI 40 IND NEP 41 MYA

Remarks: sect. Strigoathyrium, strigillosum agg.

Synonyms: hidden

= Asplenium clarkei (Bedd.) Atk. ex C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 489 (1880)

= Athyrium nigripes var. clarkei (Bedd.) Bedd.; Ferns Brit. India 166 (1883)

species :

Athyrium clemensiae Copel.; Philipp. J. Sci., C, 12: 58 (1917)

Distribution: Borneo (Sabah: Mt. Kinabalu, Sabah)

42 BOR

Remarks: sect. Strigoathyrium, scandicinum agg.

species :

Athyrium clivicola Tagawa; Acta Phytotax. Geobot. 3: 32 (1934)

Distribution: China (Anhui, Chongqing, Fujian, Guangxi, Guizhou, Hubei, Hunan, Jiangxi, Sichuan, Zhejiang); South Korea (incl. Cheju Isl.); Japan; ?Taiwan

36 CHI 38 JAP KOR

Remarks: sect. Strigoathyrium, non-setose group

Synonyms: hidden

= Athyrium clivicola f. incisum (H.Itô) Nakaike; J. Nippon Fernist Club 3(Suppl. 2): 23 (2004)

= Athyrium wardii f. incisum H.Itô; J. Jap. Bot. 11: 428 (1935)

= Athyrium wardii var. clivicola (Tagawa) Sa.Kurata; (1954)

= Athyrium wardii var. majus Ohwi; (1957) (p. p., non Makino 1899)

variant :

var. rotundum Z.R.Wang; Fl. Reipubl. Popularis Sin. [Fl. China Ed. Committee] 3(2): 508 (1999)

Distribution: China (Chongqing)

36 CHI

Synonyms: hidden

= Athyrium rotundum Ching; in sched.

species :

Athyrium contingens Ching & S.K.Wu; Fl. Xizangica 1: 134 (1983)

Distribution: China (W-Sichuan); Tibet; Nepal; Bhutan; India (Sikkim)

36 CHI TIB 40 BHU IND NEP

Remarks: valid species fide Fraser-Jenkins (2008) and not a synonym of A. attenuatum. Flora of China keeps the synonyms separate

Synonyms: hidden

= Athyrium adscendens Ching; Acta Bot. Boreal.-Occid. Sin. 6(2): 100 (1986) (fide Fraser-Jenkins 2008, 2018)

= Athyrium dentatum Ching; Acta Bot. Boreal.-Occid. Sin. 6(3): 150 (1986), non (Sw.) Gray (1821) (fide Fraser-Jenkins 2018)

= Athyrium lancipinnulum Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 18 (1986) (fide Fraser-Jenkins 2018)

= Athyrium nudifrons Ching; Acta Bot. Boreal.-Occid. Sin. 6(3): 158 (1986) (fide Fraser-Jenkins 2008, 2018)

= Athyrium oblongum Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 17 (1986)

= Athyrium squamipes Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 17 (1986) (fide Fraser-Jenkins 2018)

= Athyrium supranigrescens Ching; Acta Bot. Boreal.-Occid. Sin. 6(3): 150 (1986) (fide Fraser-Jenkins 2018)

= Athyrium tarulakaense Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 12 (1986) (fide Fraser-Jenkins 2008)

= Athyrium tsaii Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 19 (1986) (fide Fraser-Jenkins 2018)

species :

Athyrium costulalisorum Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 15 (1986)

Distribution: China (W-Sichuan)

36 CHI

species :

Athyrium crassicaule J.P.Roux; Kew Bull. 63(3): 433 (2008 publ. 2009)

Distribution: South Africa (KwaZulu-Natal, Free State); Lesotho

27 LES SAF

species :

Athyrium crassipes Ching; Acta Bot. Boreal.-Occid. Sin. 6(2): 103 (1986)

Distribution: China (W-Sichuan)

36 CHI

species :

Athyrium criticum Ching; Fl. Tsinling. 2: 216 (1974)

Distribution: China (Shaanxi, W-Sichuan, NW-Yunnan)

36 CHI

species :

Athyrium cryptogrammoides Hayata; Icon. Pl. Formosan. 6: 156 (1916)

Distribution: China (Zhejiang, Hubei, Hunan, Guangxi, Guizhou); Japan; Taiwan

36 CHI 38 JAP TAI

Synonyms: hidden

= Athyrium dissectifolium Ching; Acta Bot. Boreal.-Occid. Sin. 6(2): 104 (1986)

= Athyrium ×hohuanshanense Yoshik.; J. Phytogeogr. Taxon. 33(2): 80 (1985)

= Athyrium ×pseudocryptogrammoides Yoshik.; J. Phytogeogr. Taxon. 29(1): 27 (1981)

species :

Athyrium cyclosorum (Rupr.) Gilbert; Fernwort Pap. 27 (1900) (Dec. 20)

Name: Subarctic ladyfern

Distribution: Russian Far East (Kamchatka, Kuril Isl., Magadan, Sakhalin); Alaska; Canada (Alberta, British Columbia, Manitoba, Northern Territories, Ontario, Québec, Saskatchewan, Yukon); USA (Arizona, California, Colorado, Idaho, Michigan, Minnesota, Montana, North Dakota, Nebraska, New Mexico, Nevada, Oregon, South Dakota, Utah, Washington State, Wyoming)

31 RFE 70 ASK CDN 71 CDN 72 CDN 73 USA 74 USA 75 USA 76 USA 77 USA

Remarks: often treated within A. filix-femina or as variety of A. asplenioides, but increasingly elevated to full species status

Synonyms: hidden

= Athyrium cyclosorum (Rupr.) Maxon; Bull. Torrey Bot. Club 27: 637 (1900), isonym (Dec. 29)

= Asplenium cyclosorum (Rupr.) Fernald; Rhodora 9: 158 (1907), isonym

= Asplenium filix-femina var. cyclosorum (Rupr.) D.C.Eaton; Rep. U.S. Geogr. Surv., Wheeler 6(Botany): 331 (1878)

= Athyrium alpestre var. cyclosorum (Rupr.) T.Moore

= Athyrium cyclosorum f. hillii Gilbert; List N. Amer. Pterid.: 14, 32 (1901)

= Athyrium cyclosorum var. strictum Gilbert; List N. Amer. Pterid.: 14, 32 (1901)

= Athyrium filix-femina f. hillii (Gilbert) Butters; Rhodora 19: 199 (1917)

= Athyrium filix-femina f. strictum (Gilbert) Butters; Rhodora 19: 200 (1917)

= Athyrium filix-femina f. strictum (Gilbert) J.T.Howell; Marin Fl., ed. 2, Suppl. 363 (1969), isonym

= Athyrium filix-femina subsp. cyclosorum (Rupr.) C.Chr.; Fl. Aleutian Isl. 50 (1937)

= Athyrium filix-femina var. cyclosorum Rupr.; Distr. Crypt. Vasc. Ross. 41 (1845)

species :

Athyrium davidii (Franch.) Christ; Mém. Soc. Bot. France 1 [Hist. Acrostich.]: 50 (1903)

Distribution: China (W-Yunnan, W-Sichuan); Tibet; Nepal; Bhutan; India (Uttarakhand, Sikkim, Arunachal Pradesh); Myanmar (Kachin, etc.)

36 CHI TIB 40 BHU IND NEP 41 MYA

Conservation Status: IUCN R

Synonyms: hidden

= Asplenium duthiei Bedd.; J. Bot. 27: 72 (1889)

= Athyrium duthiei (Bedd.) Bedd.; Ferns Brit. India Suppl. 34 (1892) (fide Fraser-Jenkins (2008)

= Phegopteris davidii (Franch.) Bedd.; Suppl. Ferns Brit. Ind. 82 (1892)

= Polypodium davidii Franch.; Nouv. Arch. Mus. Hist. Nat., sér. 2, 10: 119 (1887)

= Pseudocystopteris davidii (Franch.) Z.R.Wang; Vasc. Pl. Hengduan Mount. 1: 77 (1993) (fide Fraser-Jenkins 1997)

species :

Athyrium daxianglingense Ching & H.S.Kung; Acta Bot. Boreal.-Occid. Sin. 6(1): 16 (1986)

Distribution: China (W-Sichuan, Yunnan)

36 CHI

species :

Athyrium decorum Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 13 (1986)

Distribution: China (NW-Yunnan); Tibet; Nepal; Bhutan; India (Arunachal Pradesh, Sikkim, West Bengal)

36 CHI TIB 40 BHU IND NEP

Remarks: sect. Strigoathyrium; nitidulum group

species :

Athyrium delavayi Christ; Mém. Soc. Bot. France 1 [Hist. Acrostich.]: 47 (1905)

Distribution: China (Hubei, Guangxi, Sichuan, Chongqing, Guizhou, Yunnan); Taiwan; India (Meghalaya, Mizoram); Myanmar

36 CHI 38 TAI 40 IND 41 MYA

Remarks: sect. Strigoathyrium, non-setose group

Conservation Status: IUCN CR for India (Fraser-Jenkins 2012)

Synonyms: hidden

= Athyrium costale var. polystichoides T.Moore; Index Fil. [T. Moore] 180 (1860), nom. nud.

= Athyrium latibasis Ching; Acta Bot. Boreal.-Occid. Sin. 6(2): 104 (1986)

= Athyrium mairei Rosenst.; Repert. Spec. Nov. Regni Veg. 13: 125 (1914)

= Athyrium neodelavayi Ching & H.S.Kung; Acta Bot. Boreal.-Occid. Sin. 6(1): 22 (1986)

= Athyrium pseudodelavayi Ching; (1976), nom. nud.

species :

Athyrium deltoidofrons Makino; Bot. Mag. (Tokyo) 28: 178 (1914)

Distribution: China (Zhejiang, Jiangxi, Fujian, Hunan, Sichuan, Guizhou); South Korea; Japan

36 CHI 38 JAP KOR

Synonyms: hidden

= Athyrium acutissimum Kodama; Matsum., Icon. Pl. Koisikav. 3: 127, t. 209 (1917)

= Athyrium deltoidofrons f. ohmurae Sa.Kurata; J. Geobot. 10(1): 10 (1961)

= Athyrium deltoidofrons f. reflexipinnulum (H.Itô) Nakaike; J. Nippon Fernist Club 3(Suppl. 2): 24 (2004)

= Athyrium deltoidofrons var. acutissimum (Kodama) Sa.Kurata; (1955)

= Athyrium deltoidofrons var. gracillimum (Ching) Z.R.Wang; Fl. Reipubl. Popularis Sin. [Fl. China Ed. Committee] 3(2): 186 (1999)

= Athyrium deltoidofrons var. ohmurae (Sa.Kurata) Seriz.; Shidekobushi 3(1): 51 (2015)

= Athyrium filix-femina var. deltoideum Makino; Bot. Mag. (Tokyo) 18: 30 (1899)

= Athyrium gracillimum Ching; Acta Bot. Boreal.-Occid. Sin. 6(3): 149 (1986)

= Athyrium jiulungshanense Ching; Bull. Bot. Res., Harbin 2(2): 72 (1982)

= Athyrium rotundilobum Ching; Fl. Fujianica 1: 597 (1982)

= Athyrium vidalii f. reflexipinnulum H.Itô; J. Jap. Bot. 11: 427 (1935)

= Athyrium ×multifidum var. deltoideum (Makino) Nakai; Bot. Mag. (Tokyo) 44: 507 (1930)

species :

Athyrium dentilobum Ching & S.K.Wu; Fl. Xizangica 1: 139 (1983)

Distribution: Tibet

36 TIB

Remarks: probably a synonym of A. distans fide Fraser-Jenkins et al. 2018

species :

Athyrium devolii Ching; Sunyatsenia 3: 1, t. 1 (1935)

Distribution: China (Chongqing, N-Fujian, Guangxi, Guizhou, Jiangxi, Sichuan, Yunnan, Zhejiang); Tibet; E-Nepal; Bhutan; India (Manipur, Meghalaya, Nagaland, Sikkim, West Bengal); Myanmar; Vietnam

36 CHI TIB 40 BHU IND NEP 41 MYA VIE

Remarks: A. delicatulum (which is anyway an invalid name) is included here by Fraser-Jenkins et al. (2018). Also, reports of A. solenopteris from the Himalaya belong here fide Fraser-Jenkins (2008)

Synonyms: hidden

= Athyrium aridum Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 10 (1986)

= Athyrium austrooccidentale Ching; Acta Bot. Boreal.-Occid. Sin. 6(3): 152 (1986)

= Athyrium baishanzuense Ching & Y.T.Hsieh; Acta Bot. Boreal.-Occid. Sin. 6(3): 157 (1986) (fide Fraser-Jenkins et al. 2018)

= Athyrium bambusicola Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 18 (1986)

= Athyrium delicatulum Ching & S.K.Wu; Fl. Xizangica 1: 139 (1983), non Ôishi (1941)

= Athyrium filix-femina var. fissidens Christ; (1905) (non Döll 1855)

= Athyrium fujianense Ching; Fl. Fujianica 1: 597 (1982)

= Athyrium giganteum De Vol; Lingnan Sci. J. 21: 79, t. 5 (1945) (fide Fraser-Jenkins 2018)

= Athyrium nanchuanense Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 3(4): 17 (1983)

= Athyrium nitidum Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 19 (1986)

= Athyrium praticola Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 18 (1986)

= Athyrium remotisorum Ching & H.S.Kung; Acta Bot. Boreal.-Occid. Sin. 6(3): 153 (1986)

= Athyrium remotum Ching; Acta Bot. Boreal.-Occid. Sin. 6(3): 149 (1986)

= Athyrium yaanense Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 19 (1986)

species :

Athyrium dissitifolium (Baker) C.Chr.; Contr. U. S. Natl. Herb. 26: 296, t. 18 (1931)

Distribution: China (Guangxi, Guizhou, Hunan, Sichuan, Yunnan); NE-India (Arunachal Pradesh, Manipur, Mizoram); Myanmar (Kachin, Chin, etc.); N-Thailand; Vietnam

36 CHI 40 IND 41 MYA THA VIE

Conservation Status: IUCN CR for India (Fraser-Jenkins 2012)

Synonyms: hidden

= Athyrium dissitifolium var. funebre (Christ) Ching & Z.R.Wang; Fl. Reipubl. Popularis Sin. [Fl. China Ed. Committee] 3(2): 124 (1999)

= Athyrium drepanopteron var. funebre Christ; Notul. Syst. (Paris) 1: 46 (1909)

= Athyrium drepanopteron var. brevicaudatum Christ; Notul. Syst. (Paris) 1: 46 (1909)

= Athyrium fasciculatum Hand.-Mazz.; Symb. Sin. 6: 31, t. 2, f. 5 (1929)

= Dryopteris apicidens (Baker) C.Chr.; Index Filic. 252 (1905)

= Dryopteris aristulata Rosenst.; Repert. Spec. Nov. Regni Veg. 13: 132 (1914)

= Dryopteris dissitifolia (Baker) C.Chr.; Index Filic. 262 (1905)

= Dryopteris incrassata (Christ) C.Chr.; Index Filic. 272 (1905)

= Nephrodium dissitifolium (Baker) Baker; Bull. Misc. Inform. Kew 1899: 120 (1899)

= Phegopteris incrassata Christ; Bull. Herb. Boissier, sér. 1, 6: 963 (1898)

= Polypodium apicidens Baker; Bull. Misc. Inform. Kew 1894: 54 (1894)

= Polypodium dissitifolium Baker; Bull. Misc. Inform. Kew 1895: 54 (1895)

species :

Athyrium distans (D.Don) T.Moore; Index Fil. [T. Moore] 125, 181 (1860)

Distribution: China (Chongqing, Guizhou, Sichuan); Japan; Nepal; Bhutan; N-India (Jammu & Kashmir, Arunachal Pradesh, Assam State, Himachal Pradesh, Manipur, Meghalaya, Mizoram, Nagaland, Sikkim, Uttarakhand, West Bengal); Myanmar (Kachin, etc.)

36 CHI 38 JAP 40 BHU IND NEP 41 MYA

Remarks: sect. Strigoathyrium, synonymy following Fraser-Jenkins (2008, 2018). Flora of China keeps A. imbricatum and A. nyalamense separate

Synonyms: hidden

= Asplenium distans D.Don; Prodr. Fl. Nepal. 9 (1826)

= Athyrium frangulum f. viride Sa.Kurata; Hokuriko J. Bot. 4: 17 (1955)

= Athyrium frangulum Tagawa; J. Jap. Bot. 12: 749 (1936)

= Athyrium imbricatum Christ; Bull. Acad. Int. Géogr., sér. 3, 16: 1906: 123 (1906)

= Athyrium imbricatum f. viride (Sa.Kurata) Nakaike; Checkl. Lycophytes Ferns Japan: 17 (2013)

= Athyrium iseanum var. fragile Tagawa; Acta Phytotax. Geobot. 2: 15 (1933)

= Athyrium nigripes var. dissectum C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 491 (1880)

species :

Athyrium nyalamense Y.T.Hsieh & C.R.Wang; Fl. Xizangica 1: 138 (1983)

Distribution: SE-Tibet

36 TIB

Remarks: sect. Strigoathyrium, sometimes included in A. distans, but more close to A. araiostegioides

Synonyms: hidden

= Athyrium nyalamense var. puberulum Z.R.Wang; Vasc. Pl. Hengduan Mount. 1: 87 (1993) (without latin descr. or type), et Acta Phytotax. Sin. 32(1): 84 (1994)

species :

Athyrium dombeyi Desv.; Mem. Soc. Linn. Paris 6(2): 266 (1827) (dombei auct.)

Distribution: Colombia (Antioquia, Boyacá, Caldas, Cundinamarca, Norte de Santander, Tolima); Venezuela (Aragua, Merida, Tachira, Trujillo); Ecuador; Peru; Bolivia (Cochabamba, La Paz, Tarija, Santa Cruz); Brazil (Espírito Santo, Minas Gerais, Rio de Janeiro, Rio Grande do Sul, Santa Catarina, São Paulo); Argentina (Buenos Aires, Cordoba, Salta, Tucuman)

82 CLM VEN 83 BOL ECU PER 84 BRA 85 ARG

Synonyms: hidden

= Asplenium dombeyi (Desv.) Mett.; Ann. Sci. Nat., Bot., sér. 5, 2: 238 (1864)

= Athyrium filix-femina var. dombeyi (Desv.) Hieron.; Hedwigia 47: 210 (1908)

species :

Athyrium drepanopteron (Kunze) A.Braun ex Milde; Fil. Eur. 49 (1867)

Distribution: China (Yunnan); Tibet; Nepal; Bhutan; India (Arunachal Pradesh, Manipur, Meghalaya, Nagaland, Sikkim, Tripura, Uttarakhand, West Bengal); Myanmar (Kachin, Chin, etc.); Thailand; Vietnam; Sumatra; Philippines

36 CHI TIB 40 BHU IND NEP 41 MYA THA VIE 42 PHI SUM

Remarks: synonymy unclear due to confusion with A. eburneum (mupinense). Both species have been separated by Fraser-Jenkins (2008) and Liu (2009) but are united by Flora of China. They are kept separate here - see also under A. eburneum

Synonyms: hidden

= Aspidium drepanopteron (Kunze) A.Braun ex Mett.; Fil. Hort. Bot. Lips. 93, t. 19, f. 1-4 (1856)

= Aspidium drepanopteron var. decompositum Christ; (1909)

= Asplenium drepanopteron (Kunze) A.Braun; Index Sem. (B, Berolinensis) 17 (1856)

= Asplenium oxyphyllum f. kulhaitense Atk. ex C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 494 (1880)

= Asplenium oxyphyllum Wall. ex Hook.; Sp. Fil. [W. J. Hooker] 3: 221 (1860), nom. superfl.

= Athyrium dissitifolium var. kulhaitense (Atk. ex C.B.Clarke) Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 22 (1986)

= Athyrium exindusiatum Ching; Acta Bot. Boreal.-Occid. Sin. 6(2): 102 (1986) (fide Fraser-Jenkins (2008), kept separate by Flora of China)

= Athyrium oxyphyllum (Hook.) T.Moore; Index Fil. [T. Moore] 49 (1857) ex Bedd. 1866

= Polypodium drepanopteron Kunze; Linnaea 23: 278, 318 (1850)

= Polypodium kulhaitense Atk. ex C.B.Clarke; (1880) nom. inval. in syn.

= Polypodium oxyphyllum Wall.; Numer. List. [Wallich] no. 324 (1829), nomen

species :

Athyrium dubium Ching; Bull. Fan Mem. Inst. Biol. Bot., ser. 2, 1: 281 (1949)

Distribution: China (Yunnan, Guizhou, Sichuan); Tibet; Nepal; Bhutan; India (Arunachal Pradesh, Himachal Pradesh, Sikkim, Uttarakhand, West Bengal, Jammu & Kashmir)

36 CHI TIB 40 BHU IND NEP

Synonyms: hidden

= Asplenium filix-femina subvar. rubricaule Edgew. ex C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 492, t. 59, f. 2 (1880)

= Athyrium acutum Ching; Acta Bot. Boreal.-Occid. Sin. 6(2): 101 (1986)

= Athyrium aequilaterale Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 20 (1986)

= Athyrium juxtapositum Ching; (1976), nom. nud.

= Athyrium longipinna Ching; Acta Bot. Boreal.-Occid. Sin. 6: 15 (1986)

= Athyrium machangense Ching; Acta Bot. Boreal.-Occid. Sin. 6(2): 100 (1986)

= Athyrium mackinnoniorum var. yigongense Ching & S.K.Wu; Fl. Xizangica 1: 142 (1983)

= Athyrium rubricaule (Edgew. ex C.B.Clarke) Bir; Nova Hedwigia 4: 169, t. 54 (1962)

= Athyrium serratodentatum Ching; Acta Bot. Boreal.-Occid. Sin. 6(2): 99 (1986)

= Athyrium sublineare Ching; Acta Bot. Boreal.-Occid. Sin. 6(2): 99 (1986)

= Athyrium uniseriatum Ching; Acta Bot. Boreal.-Occid. Sin. 6(2): 99 (1986)

species :

Athyrium eburneum (J.Sm. ex Mett.) J.Sm.; Hist. Fil. 328 (1875)

Distribution: China (W-Sichuan, Guizhou, Yunnan); Tibet; Japan; Taiwan; Nepal; Bhutan; India (Arunachal Pradesh, Meghalaya, Manipur, Nagaland, Sikkim, West Bengal); Myanmar; Vietnam; Philippines

36 CHI TIB 38 JAP TAI 40 BHU IND NEP 41 MYA VIE 42 PHI

Remarks: listed as A. drepanopterum in Fl. China etc. fide Fraser-Jenkins 2008 but kept separate by Liu (2009) and Knapp (2013).

Synonyms: hidden

= Aspidium eburneum Wall.; Numer. List. [Wallich] no. 389 (1829), nomen

= Asplenium eburneum J.Sm. ex Mett.; Cat. Cult. Ferns: 47 (1857), et Abh. Senckenberg. Naturf. Ges. 3(1) [Aspl.]: 238 (1859)

= Athyrium micans Tagawa; Acta Phytotax. Geobot. 4: 143 (1935)

= Athyrium mupinense Christ; Bot. Gaz. 51: 355 (1911) (moupinense auct.)

= Athyrium niponicum var. elatius Christ; (1905)

= Athyrium takeoi (Hayata) Tagawa; Acta Phytotax. Geobot. 2(3): 18 (1933)

= Athyrium veitchii Christ; Bull. Acad. Int. Géogr., sér. 3, 16: 123 (1906)

= Dryopteris takeoi Hayata; Icon. Pl. Formosan. 6: 289, f. 114 (1915)

= Lastrea eburnea (Wall.) J.Sm.; Compan. Bot. Mag. 34 (1846), nom. inval.

species :

Athyrium elmeri Copel.; Philipp. J. Sci. 3: 292 (1908)

Distribution: Philippines

42 PHI

Synonyms: hidden

= Diplazium elmeri (Copel.) C.Chr.; Index Filic., Suppl. Tert.: 73 (1934)

= Phegopteris elmeri Alderw.; Malayan Ferns Fern Allies, Suppl. 314 (1917)

species :

Athyrium elongatum Ching; Acta Bot. Boreal.-Occid. Sin. 6(2): 101 (1986)

Distribution: China (N-Guangxi, Jiangxi)

36 CHI

Synonyms: hidden

= Athyrium dolichophyllum P.S.Wang; (1989), nom. nud.

species :

Athyrium emeicola Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 13 (1986)

Distribution: China (Sichuan)

36 CHI

species :

Athyrium epirachis (Christ) Ching; C. Chr., Index Filic., Suppl. Tert.: 41 (1934)

Distribution: China (Chongqing, Fujian, S-Guangdong, Guangxi, Guizhou, SW-Hubei, Hunan, Sichuan, Yunnan); South Korea (Cheju Isl.); Japan (Honshu); Taiwan

36 CHI 38 JAP KOR TAI

Remarks: sect. Strigoathyrium, non-setose group. In Taiwan very rare. A. eremicola (Japan) is sometimes kept separate but genetically very close

Synonyms: hidden

= Athyrium eremicola Oka & Sa.Kurata ex Sa.Kurata; J. Geobot. 8: 83 (1960)

= Athyrium lilacinum Ching; Bull. Fan Mem. Inst. Biol. Bot., ser. 2, 1: 283 (1949)

= Athyrium muticum Christ; Bull. Acad. Int. Géogr., sér. 3, 17: 147 (1907)

= Athyrium rubristipes Ching; (1976), nom. nud.

= Athyrium subcoriaceum Ching; Acta Bot. Boreal.-Occid. Sin. 6(3): 151 (1986)

= Athyrium wardii var. elongatum Christ; (1905)

= Diplazium epirachis Christ; Mém. Soc. Bot. France 1 [Hist. Acrostich.]: 51 (1905)

species :

Athyrium erythropodum Hayata; Icon. Pl. Formosan. 4: 233 t 163 (1914)

Distribution: Taiwan; Vietnam; ?Philippines

38 TAI 41 VIE

Remarks: sect. Strigoathyrium, non-setose group, erythropodum clade. Not in Japan

Conservation Status: IUCN R

species :

Athyrium falcatum Bedd.; Ferns S. India t. 151 (1865)

Distribution: W-Nepal; India (Chhattisgarh, Gujarata, Himachal Pradesh, Jharkand, Karnataka, Kerala, Madhya Pradesh, Maharashtra, Meghalaya, Mizoram, Odisha, Rajasthan, Tamil Nadu, Tripura, Uttarakhand); Myanmar

40 IND NEP 41 MYA

Synonyms: hidden

= Asplenium drepanophyllum Baker; Syn. Fil. [W. J. Hooker & Baker], ed. 2: 226 (1874) (non Kunze (1834))

= Athyrium drepanophyllum (Baker) Bedd.; Ferns Brit. India Suppl. 32 (1892)

= Athyrium grammitoides Bedd.; Ferns Brit. India Correct. 11 (1870)

= Athyrium ×keralense Manickam & Irudayaraj; Pterid. Fl. West. Ghats, S. India: 238 (1992) (as ´keralensis´, nom. inval., sine type specimen, fide Fraser-Jenkins 1997)

species :

Athyrium fallaciosum Milde; Fil. Eur. 6: 54 (1867)

Distribution: Russian Far East (Kuril Isl., Sakhalin); China (Gansu, Hebei, Heilongjiang, Henan, Hubei, Jilin, Liaoning, Nei Mongol, Ningxia, Shaanxi, Shanxi, N-Sichuan); North Korea; Japan (Hokkaido, Honshu); ?Philippines (?Luzon)

31 RFE 36 CHI 38 JAP NKO

Remarks: A. fauriei is kept separate by Flora of China but included here by Fraser-Jenkins (2008)

Synonyms: hidden

= Aspidium fauriei (Christ) Christ; Bull. Herb. Boissier, sér. 1, 6: 193 (1898)

= Asplenium mongolicum Franch.; Nouv. Arch. Mus. Hist. Nat., sér. 2, 7: 161 (1883)

= Asplenium nikkoense Makino; Bot. Mag. (Tokyo) 17: 43 (1903)

= Athyrium fauriei (Christ) Makino; Bot. Mag. (Tokyo) 17: 160 (1903)

= Athyrium mongolicum (Franch.) Diels; Nat. Pflanzenfam. [Engler & Prantl] I, 4: 224 (1899)

= Athyrium mongolicum var. purdomii C.Chr.; Bot. Gaz. 56(4): 333 (1913)

= Athyrium rupestre Kodama; Matsum., Icon. Pl. Koisikav. 4: 63, t. 244 (1919) (fide Fraser-Jenkins 2008)

= Athyrium yokoscense var. fauriei (Christ) Tagawa; Acta Phytotax. Geobot. 2: 20 (1933)

= Dryopteris mongolica (Franch.) C.Chr.; Index Filic. 278 (1905)

= Nephrodium fauriei Christ; Bull. Herb. Boissier, sér. 1, 4: 671 (1896)

= Nephrodium mongolicum (Franch.) Baker; Ann. Bot. (Oxford) 6: 317 (1891)

species :

Athyrium fangii Ching; Bull. Fan Mem. Inst. Biol. Bot., ser. 2, 1: 282 (1949)

Distribution: China (Sichuan, Yunnan); ?Tibet; Nepal; Bhutan; N-India (Arunachal Pradesh, Sikkim, Uttarakhand, West Bengal)

36 CHI 40 BHU IND NEP

Synonyms: hidden

= Athyrium paranigripes X.C.Zhang; Bull. Bot. Res., Harbin 11(3): 9 (1991) (nom. nov. for A. sikkimense Ching) (fide Fraser-Jenkins et al. 2015)

= Athyrium sikkimense Ching; Acta Bot. Austro Sin. 1: 19 (1983) (non (Bir) Á. Löve & D. Löve) (fide Fraser-Jenkins et al. 2015)

species :

Athyrium filix-femina (L.) Roth; Tent. Fl. Germ. 3: 65 (1799)

subspecies :

subsp. filix-femina

Name: Wald-Frauenfarn (DE), Lady Fern (EN); Helecho hembra (ES)

Distribution: Iceland; Faeroe Isl.; England (England, Wales, Scotland, Outer Hebrides, Orkney Isl., Shetland Isl., Isle of Man); Ireland (Ireland, Northern Ireland); Denmark; Norway; Sweden; Finland; Netherlands; Belgium; Luxembourg; Germany (Brandenburg, Berlin, Baden-Württemberg, Bayern, Hessen, Hamburg, Mecklenburg-Vorpommern, Niedersachsen, Nordrhein-Westfalen, Rheinland-Pfalz, Schleswig-Holstein, Saarland, Sachsen, Sachsen-Anhalt, Thüringen); Switzerland; Liechtenstein; Austria; Poland; Czech Republic; Slovakia; Hungary; Portugal; Spain; Andorra; France; Channel Isl. (Alderney, Guernsey, Jersey); Corsica; Sardinia; Italy; Sicily; Malta; Slovenia; Croatia; Bosnia & Hercegovina; Montenegro; Serbia; Kosovo; North Macedonia; Albania; Romania; Bulgaria; European Turkey; Greece (widespread, incl. Kiklades, Samothraki); Crete; Estonia; Latvia; Lithuania; Belarus; C-European Russia; E-European Russia; N-European Russia; W-European Russia; Moldova; Ukraine; Crimea; Morocco; Algeria; Tunisia; Azores (Santa Maria Isl., Sao Miguel Isl., Terceira, Graciosa, Sao Jorge, Pico, Faial, Flores Isl., Corvo Isl.); Madeira (Madeira Isl.); Canary Isl. (Gran Canaria, Tenerife, La Gomera, Hierro, La Palma Isl.); Siberia (Altai, Buryatia, Chita, Irkutsk, Krasnoyarsk, Tuva, W-Siberia, Yakutia); Russian Far East (Kamchatka, Khabarovsk, Kuril Isl., Magadan, Primorye, Sakhalin); Kazakhstan; Turkmenistan; ?Uzbekistan; Northern Caucasus; Georgia [Caucasus]; Armenia; Azerbaijan (incl. Talysh); East Aegaean Isl. (Lesvos, Irakia, Samos); Turkey (Anatolia); Iran; Lebanon; Mongolia; ?Japan; New Zealand [I]; Mexico (Baja California Norte, Baja California Sur, Chiapas, Chihuahua, Colima, Ciudad de Mexico, Durango, Guerrero, Hidalgo, Jalisco, Mexico State, Michoacan, Morelos, Nayarit, Nuevo Leon, Oaxaca, Puebla, Queretaro, Sonora, Tamaulipas, Veracruz, Zacatecas); Guatemala; ?Belize; ?El Salvador; Costa Rica; Panama

10 DEN FIN FOR GRB ICE IRE NOR SWE 11 AUT BGM CZE GER HUN LIE LUX NET POL SLO SWI 12 ADO CIS COR FRA POR SAR SPA 13 ALB BIH BUL CRO GRC ITA KOS KRI MLT MNT NMA ROM SIC SLV SRB TUE 14 BLR EST KRY LAT LIT MLD RUS UKR 20 ALG MOR TUN 21 AZO CNY MDR 30 SIB 31 RFE 32 KAZ TKM 33 ARM AZE GEO NCS 34 EAI IRN LEB TUR 37 MON 51 nzl 79 MEX 80 COS GUA PAN

Remarks: records from South America are referrable to A. dombeyi

Synonyms: hidden

= Allantodia axillaris (Aiton) Kaulf.; Enum. Filic. [Kaulfuss] 178 (1824)

= Aspidium axillare (Aiton) Sw.; J. Bot. (Schrader) 1800(2): 42 (1801)

= Aspidium cordatum Steud.; Nomencl. Bot. [Steudel] 2: 61 (1824)

= Aspidium filix-femina (L.) Sw.; J. Bot. (Schrader) 1800(2): 41 (1801)

= Aspidium filix-femina var. dentatum (Döll) Döll; Fl. Baden [Döll] 1(1): 23 (1855)

= Aspidium filix-femina var. incisum Hartm.; Handb. Skand. Fl. 369 (1820)

= Aspidium filix-femina var. multidentatum (Döll) Döll; Gefässkrypt. Baden 24 (1855)

= Aspidium filix-femina var. multifidum Döll; Fl. Baden [Döll] 1(1): 25 25 (1855)

= Aspidium intermedium Link; Enum. Hort. Berol. Alt. 2: 459 (1822), nom. illeg.

= Aspidium irriguum Sm.; Engl. Bot. 81, t. 2199 (1810)

= Aspidium pontederae Willd.; Sp. Pl., ed. 4 [Willdenow] 5(1): 273 (1810)

= Aspidium rhaeticum (L.) Sw.; J. Bot. (Schrader) 1800(2): 41 (1801), as ´rheticum´

= Asplenium axillare (Aiton) J.Sm.; Compan. Bot. Mag. 30 (1846)

= Asplenium cyathioides Bernh.; J. Bot. (Schrader) 1799(1): 313 (1799), nom. illeg.

= Asplenium filix-femina (L.) Bernh.; Neues J. Bot. 12: 26, 48, t. 2, f. 7 (1806)

= Asplenium filix-femina f. dentatum (Döll) Döll; Fl. Baden [Döll] 1(1): 23 (1855)

= Asplenium filix-femina f. fissidens Döll; Fl. Baden [Döll] 1(1): 24 (1855)

= Asplenium filix-femina f. multidentatum (Döll) Döll; Fl. Baden [Döll] 1(1): 24 (1855)

= Asplenium filix-femina var. bidentatum Döll; Rhein. Fl. 12 (1843)

= Asplenium filix-femina var. molle (Schreb.) Hook.f.; 462 (1870)

= Asplenium filix-femina var. purpureum Regel; Index Sem. (LE, Petropolitanus) 1866(Suppl.): 25 (1867)

= Asplenium filix-femina var. rhaeticum (L.) Hook.f.; 462 (1870)

= Asplenium monstrosum Kunze; Linnaea 23: 235, 305 (1850)

= Asplenium rhaeticum (L.) Brügger; Naturg. Beitr. Chur: 47 (1874)

= Athyrium acrostichoideum Bory ex Mérat; Nouv. Fl. Env. Paris, ed. 4, 1: 372 (1836)

= Athyrium alpestre Clairv.; Man. Herbor. Suisse 301 (1811) non (Hoppe)

= Athyrium axillare (Aiton) C.Presl; Tent. Pterid. 98 (1836)

= Athyrium azoricum Fée; Mém. Foug. 5 (Gen. Filic.): 186 (1852)

= Athyrium cassum Chiov.; Fl. Alpi Lepont. 2: 46 (1929)

= Athyrium convexum (Bab.) Newman; Phytologist 4(App.): XIII (1851)

= Athyrium corsicum Fée; Mém. Foug. 5 (Gen. Filic.): 186 (1852)

= Athyrium depauperatum Schumach.; Enum. Pl. [Schumacher] 2: 17 (1803)

= Athyrium filix-femina proles acrostichoideum (Bory ex Mérat) Rouy; Fl. France [Rouy & Foucaud] 14: 428 (1913)

= Athyrium filix-femina f. rhaeticum (L.) Murr; 5 (1923)

= Athyrium filix-femina var. bidentatum Briq.; Prodr. Fl. Corse 1: 12 (1910)

= Athyrium filix-femina var. convexum Bab.; Man. Brit. Bot. [Babington] 388 (1843)

= Athyrium filix-femina var. curtum-cristatum Ivery ex R.Hogg; Florist & Pomol. 27: 135 (1874)

= Athyrium filix-femina var. dentatum (Döll) Milde; Fil. Eur. 50 (1867)

= Athyrium filix-femina var. dubpulayae Lowe; Gard. Chron., ser. 2, 4: 81 (1875)

= Athyrium filix-femina var. erectum Syme & N.E.Br.; Engl. Bot., 3rd ed., vol. XII: 109 (1886)

= Athyrium filix-femina var. fissidens (Doll) Milde; Fil. Eur. 50 (1867)

= Athyrium filix-femina var. frizelliae W.Bull ex J.Dix; Proc. Roy. Hort. Soc. London 1: 652 (1861)

= Athyrium filix-femina var. incisum (Fée) Hieron.; Bot. Jahrb. Syst. 22(3): 363 (1896), nom. illeg., non Hartm. 1820

= Athyrium filix-femina var. incisum Hartm.; Handb. Skand. Fl. 369 (1820)

= Athyrium filix-femina var. incisum Newman; Hist. Brit. Ferns, ed. II: 243 (1844), nom. illeg.

= Athyrium filix-femina var. iveryanum Ivery ex J.Dix; Proc. Roy. Hort. Soc. London 2: 675 (1862)

= Athyrium filix-femina var. kingscotei Lowe; Gard. Chron., ser. 2, 4: 81 (1875)

= Athyrium filix-femina var. latipes T.Moore; Handb. Brit. Ferns 2: 30 (1860)

= Athyrium filix-femina var. lineare-reflexum Lowe; Gard. Chron., ser. 2, 4: 81 (1875)

= Athyrium filix-femina var. minus Payot; Fl. Mont-Blanc 2: 10 (1882)

= Athyrium filix-femina var. molle (Schreb.) Briq.; Prodr. Fl. Corse 1: 12 (1910)

= Athyrium filix-femina var. molle (Schreb.) Schur; 168 (1851)

= Athyrium filix-femina var. mucronatum Ivery ex J.Dix; Proc. Roy. Hort. Soc. London 2: 675 (1862)

= Athyrium filix-femina var. multiceps T.Moore ex J.Dix; Proc. Roy. Hort. Soc. London 1: 70 (1859)

= Athyrium filix-femina var. multidentatum (Döll) Milde; Fil. Eur. 50 (1867)

= Athyrium filix-femina var. ovatum (Roth) Schur; 168 (1851)

= Athyrium filix-femina var. parsonsiae J.Dix; Proc. Roy. Hort. Soc. London 2: 459 (1862)

= Athyrium filix-femina var. pectinatum Millsp.; Living Fl. W. Virginia 5A(1): 193 (1913), nom. illeg.

= Athyrium filix-femina var. plumosum T.Moore ex J.Dix; Proc. Roy. Hort. Soc. London 1: 239 (1860)

= Athyrium filix-femina var. pulchellum-plumosum Anon.; Gard. Chron., ser. 2, 2: 49 (1874)

= Athyrium filix-femina var. qulterianum Anon.; Gard. Chron., ser. 2, 2: 49 (1874)

= Athyrium filix-femina var. rhaeticum (L.) Schur; 168 (1851)

= Athyrium filix-femina var. tripinnatum Rupr.; Dist. Crypt. Vasc. Ross. 41 (1845)

= Athyrium filix-femina var. watsonii Syme & N.E.Br.; Engl. Bot., 3rd ed., vol. XII: 110 (1886)

= Athyrium fimbriatum Dulac; Fl. Hautes-Pyrenées 33 (1867) non (Wall.) T. Moore

= Athyrium incisum (Hoffm.) Newman; Phytologist 4: app. XIII (1851)

= Athyrium incisum Fée; Mém. Foug. 5 (Gen. Filic.): 187 (1852), nom. illeg., non (Hoffm.) Newman 1851

= Athyrium irriguum (Sm.) Gray; Nat. Arr. Brit. Pl. 2: 10 (1821)

= Athyrium laetum (Salisb.) Gray; Nat. Arr. Brit. Pl. 2: 10 (1821), nom. superfl.

= Athyrium laxum Schumach.; Enum. Pl. [Schumacher] 2: 16 (1803)

= Athyrium molle (Schreb.) Roth; Tent. Fl. Germ. 3: 61 (1799)

= Athyrium molliusculum Baumg.; Enum. Stirp. Transsilv. 4: 31 (1846)

= Athyrium officinale Bubani; Fl. Pyren. (Bubani) 4: 430 (1901)

= Athyrium ovatum Roth; Tent. Fl. Germ. 3(1.1): 64 (1799)

= Athyrium paramicola L.D.Gómez; Phytologia 52(3): 154 (1982)

= Athyrium rhaeticum (L.) Dalla Torre; Anleit. Beob. Alpenpfl. 2: 348 (1882), nom. illeg.

= Athyrium rhaeticum (L.) Roth; Tent. Fl. Germ. 3: 67 (1799)

= Athyrium rhaeticum var. minus Doerr. ex Roth; 68 (1799)

= Athyrium scorpioideum Chauv.; Mém. Soc. Linn. Calvados 2: 502 (1825)

= Athyrium trifidum (Hoffm.) Roth; Tent. Fl. Germ. 3: 63 (1799)

= Cyathea filix-femina (L.) Bertol.; Amoen. Ital. 429 (1819)

= Cystopteris filix-femina (L.) Coss. & Germ.; Fl. Env. Paris: 676 (1845)

= Dryopteris rigida var. rhiphaea Pau & Font Quer; Iter Marocc. 1927: no. 3 (1928)

= Filix bipinnata Gilib.; Exerc. Phyt. 2: 557 (1792), opus utique oppr.

= Lastrea filix-femina (L.) Colomb; Compt. Rend. Hebd. Séances Acad. Sci. 107: 1013 (1888)

= Lastrea filix-femina var. bollandiae W.Bull ex J.Dix; Proc. Roy. Hort. Soc. London 2: 458 (1862)

= Nephrodium filix-femina (L.) Michx.; Fl. Bor.-Amer. (Michaux) 2: 268 (1803)

= Phegopteris plumosa J.Sm.; Ferns Brit. Foreign 289 (1866)

= Phegopteris rhaetica (L.) Pérard; Bull. Soc. Bot. France 16: 258 (1869), nom. illeg.

= Phegopteris rhaetica (L.) Watt; Canad. Naturalist & Quart. J. Sci., ser. 2, 13: 159 (1867)

= Polypodium axillare Aiton; Hort. Kew. [W. Aiton] 3: 466 (1789)

= Polypodium bifidum Hoffm.; Bot. Mag. (Römer & Usteri) 9: 10 (1790)

= Polypodium crenatum Schrank; Schrank & Moll, Naturhist. Briefe Oestreich 2: 295 (1785), nom. illeg.

= Polypodium dentatum Schrank; Schrank & Moll, Naturhist. Briefe Oestreich 2: 295 (1785), nom. illeg.

= Polypodium femina L.; Mant. Pl. 2: 506 (1771)

= Polypodium filix-femina L.; Sp. Pl. 2: 1090 (1753)

= Polypodium filix-femina var. molle (Schreb.) Lej.; Rev. Fl. Spa: 212 (1825)

= Polypodium filix-femina var. nanum Gren.; Bull. Soc. Bot. France 14: 63 (1867)

= Polypodium filix-femina var. purpurascens Bory ex Lej.; Rev. Fl. Spa: 212 (1825)

= Polypodium incisum Hoffm.; Bot. Mag. (Römer & Usteri) 9: 10 (1790), nom. illeg.

= Polypodium laetum Salisb.; Prodr. Stirp. Chap. Allerton 403 (1796), nom. superfl.

= Polypodium leseblii Mérat; Nouv. Fl. Env. Paris, ed. 2: 276 (1821)

= Polypodium molle Schreb.; Spic. Fl. Lips. 70 (1771)

= Polypodium oblongodentatum Hoffm.; Bot. Mag. (Römer & Usteri) 9: 10 (1790)

= Polypodium ovatocrenatum Hoffm.; Bot. Mag. (Römer & Usteri) 9: 10 (1790)

= Polypodium rhaeticum L.; Sp. Pl. 2: 1091 (1753)

= Polypodium trifidum Hoffm.; Bot. Mag. (Römer & Usteri) 9: 10 (1790)

= Tectaria filix-femina (L.) Cav.; Anales Ci. Nat. 4: 100 (1801)

subspecies :

subsp. californicum (Butters) Hultén; Kongl. Svenska Vetensk. Acad. Handl. 8(5): 178 (1962)

Distribution: W-USA (Arizona, California, Colorado, Idaho, Nevada, New Mexico, Oregon, South Dakota, Utah, Wyoming)

73 USA 74 USA 76 USA 77 USA

Remarks: status unconfirmed, might belong to one of the species formerly included in A. filix-femina s. l. from North America

Synonyms: hidden

= Athyrium filix-femina var. californicum Butters; Rhodora 19: 201 (1917)

subspecies :

subsp. sitchense (Rupr.) Tzvelev; Sosud. Rast. Sovet. Dal´nego Vostoka 5: 73 (1991)

Distribution: Alaska (Sitka Isl.)

70 ASK

Synonyms: hidden

= Athyrium sitchense (Rupr.) Tzvelev; Novosti Sist. Vyssh. Rast. 26: 8 (1989)

= Athyrium filix-femina var. sitchense Rupr.; Distr. Crypt. Vasc. Ross. 41 (1845)

species :

Athyrium fimbriatum Wall. ex T.Moore; Index Fil. [T. Moore] 185 (1860)

Distribution: China (Yunnan, Sichuan, Hunan); Tibet; Nepal; Bhutan; India (Himachal Pradesh, Uttarakhand, Sikkim, Darjeeling, Arunachal Pradesh, Meghalaya, Nagaland, West Bengal); Myanmar (Kachin, etc.)

36 CHI TIB 40 BHU IND NEP 41 MYA

Remarks: sect. Polystichoides

Synonyms: hidden

= Araiostegia yaklaensis (Bedd.) B.K.Nayar & S.Kaur; Companion Ferns Brit. India 17 (1974)

= Aspidium fimbriatum Wall.; Numer. List. [Wallich] no. 339 (1829), nomen, non Willd. 1810

= Asplenium fimbriatum var. foliolosum (Wall.) C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 495 (1880)

= Asplenium fimbriatum Wall. ex Hook.; Sp. Fil. [W. J. Hooker] 3: 234 (1860), nom. illeg.

= Asplenium foliolosum (C.B.Clarke) C.Hope; J. Bombay Nat. Hist. Soc. 14: 255 (1902), nom. illeg.

= Athyrium bucahwangense Ching; Fl. Xizangica 1: 132 (1983) (fide Fraser-Jenkins et al. 2015)

= Athyrium decompositum Ching; Sporae Pterid. Sin. 215, t. 44, f. 5-6 (1976), nom. nud.

= Athyrium fimbriatum (Wall. ex Hook.) Bedd.; Ferns Brit. India t. 295 (1866), nom. superfl.

= Athyrium fimbriatum var. foliolosa Wall.; Numer. List. [Wallich] no. 359 (1829), nomen

= Athyrium fimbriatum var. foliolosum (C.B.Clarke) Bedd.; Ferns Brit. India 172 (1880)

= Athyrium fimbriatum var. squamatum Bedd.; Suppl. Ferns Brit. India 37 (1892)

= Athyrium foliolosum (C.B.Clarke) Bedd.; Suppl. Ferns Brit. Ind. 37 (1892), nom. illeg.

= Athyrium yaklaense (Bedd.) Panigrahi & Basu; (1984) (fide Fraser-Jenkins 2008)

= Davallia yaklaensis (Bedd.) C.Chr.; Index Filic. 215 (1905) (fide Fraser-Jenkins 2008)

= Leucostegia yaklaensis Bedd.; Suppl. Ferns Brit. Ind. 13 (1892) (fide Fraser-Jenkins 2008)

= Polystichum fimbriatum Wall. ex C.Presl; Abh. Königl. Böhm. Ges. Wiss., ser. 5, 6 [Epim. Bot.]: 58 (1851)

species :

Athyrium flabellulatum (C.B.Clarke) Tardieu; Aspl. Tonkin 81, t. 12 (1932)

Distribution: Tibet; Nepal; Bhutan; India (Himachal Pradesh, Uttarakhand, Sikkim, Darjeeling, Arunachal Pradesh, West Bengal); Myanmar; Vietnam

36 TIB 40 BHU IND NEP 41 MYA VIE

Synonyms: hidden

= Asplenium filix-femina var. flabellulatum C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: t. 60 (1880)

= Athyrium filix-femina var. deltoidea Christ; (1905)

= Athyrium filix-femina var. flabellulatum (C.B.Clarke) Bedd.; Ferns Brit. India 170 (1883)

= Athyrium tenuifolium Y.T.Hsieh & C.R.Wang; Fl. Xizangica 1: 141 (1983), non Copel. 1929

species :

Athyrium foliolosum (Wall.) T.Moore ex R.Sim; Cat. Brit. Ferns 6: 22 (1859)

Distribution: China (Yunnan); Tibet; Nepal; Bhutan; India (Arunachal Pradesh, Himachal Pradesh, ?Karnataka, Kerala, Manipur, Meghalaya, Mizoram, Nagaland, Rajasthan, Sikkim, Tamil Nadu, Uttarakhand, West Bengal); N-Myanmar (Kachin, etc.); Vietnam

36 CHI TIB 40 BHU IND NEP 41 MYA VIE

Remarks: sect. Polystichoides. Not in Taiwan

Synonyms: hidden

= Aspidium foliolosum Wall.; Numer. List. [Wallich] no. 359 (1829), nomen

= Aspidium squarrosum Wall.; Numer. List. [Wallich] no. 356 (1829), nomen, non Don 1825

= Asplenium decipiens Mett.; Abh. Senckenberg. Naturf. Ges. 3(1) [Aspl.]: 195, no. 237 (1859) (non Zipp. ex Kunze 1849, nec Kuhn 1879)

= Asplenium foliolosum (T.Moore ex R.Sim) E.J.Lowe; Nat. Hist. Ferns: 107 (1861)

= Asplenium foliolosum Wall.; Numer. List. [Wallich] no. 2205 (1829), nomen

= Athyrium austroyunnanense Ching; Acta Bot. Austro Sin. 1: 19 (1983), et Fl. Xizangica 1: 132 (1983) (fide Fraser-Jenkins 1997)

= Athyrium submacrocarpum Ching & S.K.Wu; Fl. Xizangica 1: 131 (1983)

= Polypodium foliolosum Wall.; Numer. List. [Wallich] no. 359 (1824), nomen

species :

Athyrium guangnanense Ching; Acta Bot. Boreal.-Occid. Sin. 6(2): 103 (1986), nom. nov.

Distribution: China (SE-Yunnan, Guizhou)

36 CHI

Synonyms: hidden

= Athyrium acrocarpum W.M.Chu; Acta Bot. Yunnan., Suppl. 5: 41 (1992) (non (Rosenst.) Copel. 1951)

= Athyrium guangnanense var. acrocarpum W.M.Chu; Fl. Yunnanica 20: 395 (2006)

= Athyrium yunnanicum Ching; Bull. Fan Mem. Inst. Biol. Bot., ser. 2, 1: 278 (1949) (non Christ 1907)

species :

Athyrium gymnogrammoides (Klotzsch ex Mett.) Bedd.; Ferns S. India 52 (1864)

Distribution: S-India (Kerala, Tamil Nadu); Sri Lanka

40 IND SRL

Synonyms: hidden

= Allantodia gymnogrammoides (Klotzsch ex Mett.) Ching; Acta Phytotax. Sin. 9: 54 (1964)

= Allantodia solenopteris var. pusilla Kunze; Linnaea 24: 267 (1851)

= Asplenium gymnogrammoides Klotzsch ex Mett.; Abh. Senckenberg. Naturf. Ges. 3(1) [Aspl.]: 193, no. 234, t. 16, f. 13-14 (1859)

= Athyrium solenopteris var. pusillum (Kunze) T.Moore; Index Fil. [T. Moore] 187 (1860)

species :

Athyrium hahajimense Nakai; Bot. Mag. (Tokyo) 43: 4 (1929)

Distribution: Bonin Isl. (Hahajima)

38 OGA

species :

Athyrium hainanense Ching; Acta Bot. Boreal.-Occid. Sin. 6(2): 101 (1986)

Distribution: China (Hainan)

36 CHI

Remarks: sect. Strigoathyrium, arisanense agg.

species :

Athyrium haleakalae K.R.Wood & W.L.Wagner; PhytoKeys 76: 115-124 (2017)

Distribution: Hawaii (Maui, Kauai)

63 HAW

species :

Athyrium himalaicum Ching ex Mehra & Bir; Res. Bull. Panjab Univ. Sci. 15: 137, f. 26-29 (1964)

Distribution: China (SW-Sichuan, N-Guizhou, NW-Yunnan); Tibet; Nepal; Bhutan; N-India (Sikkim, Uttarakhand, Arunachal Pradesh); Myanmar (Kachin)

36 CHI TIB 40 BHU IND NEP 41 MYA

Synonyms: hidden

= Athyrium birii Ching ex Mehra & Bir; Amer. Fern J. 50: 276-295 (1960) (nom. nud., non (Ching ex Bir) Seriz., fide Fraser-Jenkins 1997)

= Athyrium suprapubescens Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 10 (1986) (fide Fraser-Jenkins 2018)

species :

Athyrium hirtirachis Ching & Y.P.Hsu; Fl. Tsinling. 2: 215 (1974)

Distribution: China (Sichuan, SE-Gansu)

36 CHI

Synonyms: hidden

= Athyrium gonggaense Z.R.Wang & Li Bing Zhang; Li Bing Zhang, Z. R. Wang & X. S. Guo, Acta Bot. Yunnan. 17(3): 301- 303, t. 1, f. 5-8 (1995)

species :

Athyrium hohenackerianum (Kunze) T.Moore; Index Fil. [T. Moore] 49 (1857)

Distribution: India (Andhra Pradesh, Chhattisgarh, Goa, Gujarat, Karnataka, Kerala, Madhya Pradesh, Maharashtra, Odisha, Rajasthan, Tamil Nadu); Sri Lanka

40 IND SRL

Synonyms: hidden

= Allantodia hohenackeriana (Kunze) Kunze; Farnkräuter 2: 63, t. 126 (1850)

= Asplenium hohenackerianum Kunze; Bot. Zeitung (Berlin) 771 (1849)

species :

Athyrium infrapuberulum Ching; Acta Bot. Boreal.-Occid. Sin. 6(2): 100 (1986)

Distribution: China (NW-Guangxi)

36 CHI

Remarks: sect. Strigoathyrium, arisanense agg.

species :

Athyrium interjectum Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 15 (1986)

Distribution: China (NW-Yunnan)

36 CHI

Synonyms: hidden

= Athyrium heqingense Ching; (1986), nom. nud.

= Athyrium acutiserratum Ching; Acta Bot. Boreal.-Occid. Sin. 6(3): 149 (1986)

species :

Athyrium intermixtum Ching & P.S.Chui; Acta Bot. Boreal.-Occid. Sin. 6(1): 21 (1986)

Distribution: China (Anhui, Zhejiang)

36 CHI

species :

Athyrium iseanum Rosenst.; Repert. Spec. Nov. Regni Veg. 13: 124 (1913)

Distribution: China (Anhui, Chongqing, Fujian, Guangdong, Guangxi, Guizhou, Hubei, Hunan, Jiangsu, Jiangxi, Sichuan, Yunnan, Zhejiang); Tibet; South Korea (Cheju Isl.); Japan

36 CHI TIB 38 JAP KOR

Synonyms: hidden

= Athyrium filix-femina var. filipes Christ; (1905)

= Athyrium wuyishanense Ching; Acta Bot. Boreal.-Occid. Sin. 6(2): 104 (1986)

variant :

var. angustisectum Tagawa; Acta Phytotax. Geobot. 2: 15 (1933)

Distribution: Taiwan

38 TAI

Synonyms: hidden

= Athyrium iseanum f. angustisectum (Tagawa) Tagawa; Acta Phytotax. Geobot. 6: 42 (1957)

variant :

var. chuanqianense Z.R.Wang; Fl. Reipubl. Popularis Sin. [Fl. China Ed. Committee] 3(2): 509 (1999)

Distribution: China (Guangxi, Sichuan, Guizhou, Yunnan); Tibet

36 CHI TIB

species :

Athyrium jinshajiangense Ching & K.H.Shing; Acta Bot. Boreal.-Occid. Sin. 6(1): 14 (1986)

Distribution: China (C-Yunnan)

36 CHI

species :

Athyrium kandelii Fraser-Jenk.; Annot. Checkl. Ind. Pterid. 2: 36 (2018), nom. nov.

Distribution: S-China; Nepal; Bhutan; India (Himachal Pradesh, Sikkim, Uttarakhand, Jammu & Kashmir)

36 CHI 40 BHU IND NEP

Synonyms: hidden

= Athyrium mackinnoniorum var. glabratum Y.T.Hsieh & C.R.Wang; Fl. Xizangica 1: 143 (1983)

species :

Athyrium kenzo-satakei Sa.Kurata; J. Geobot. 7: 75 (1958)

Distribution: China (Guangdong, Guangxi, Sichuan, Guizhou); Japan (incl. Yakushima Isl.)

36 CHI 38 JAP NNS

Remarks: sect. Strigoathyrium, arisanense agg. Very close to A. arisanense and possibly the same.

Conservation Status: CR in Japan

Synonyms: hidden

= Athyrium arisanense var. kenzo-satakei (Sa.Kurata) Seriz.; J. Jap. Bot. 45: 117 (1970)

variant :

var. jieguishanense (Ching) Z.R.Wang; Fl. Reipubl. Popularis Sin. [Fl. China Ed. Committee] 3(2): 225 (1999)

Distribution: China (Guangxi)

36 CHI

Synonyms: hidden

= Athyrium jieguishanense Ching; Acta Bot. Boreal.-Occid. Sin. 6(3): 157 (1986)

species :

Athyrium khasimontanum Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 419 (2008)

Distribution: India (Meghalaya)

40 IND

Conservation Status: IUCN CR for India (Fraser-Jenkins 2012)

species :

Athyrium kirishimaense Tagawa; Acta Phytotax. Geobot. 2(1): 22 (1932)

Distribution: Japan (Izu Isl.)

38 JAP

Synonyms: hidden

= Athyrium yokoscense var. kirishimaense (Tagawa) Li & J.Z.Wang; Clav. Pl. Chin. Bor.-Or. 36 (1995)

species :

Athyrium kuratae Seriz.; J. Jap. Bot. 45: 117 (1970)

Distribution: China (Yunnan); S-Japan

36 CHI 38 JAP

Remarks: not in Taiwan. possibly a synonym of A. otophorum

species :

Athyrium kutaiense C.Chr.; Dansk Bot. Ark. 9(3): 55 (1937)

Distribution: Borneo

42 BOR

Remarks: provisionally accepted name

species :

Athyrium latinervatum M.Kessler & A.R.Sm.; Brittonia 59(2): 186 (2007)

Distribution: Bolivia (La Paz)

83 BOL

Remarks: known only from type

species :

Athyrium leiopodum (Hayata) Tagawa; Acta Phytotax. Geobot. 2(3): 195 (1933)

Distribution: Taiwan

38 TAI

Remarks: not included in Fl. of China but confirmed for Taiwan by Knapp (2011) and Liu et al. (2009). Included in A. roseum by Fraser-Jenkins (2008)

Synonyms: hidden

= Diplazium leiopodum Hayata; Icon. Pl. Formosan. 4: 217, f. 148 (1914)

species :

Athyrium lewalleanum Pic.Serm.; Webbia 27: 440 (1972 publ. 1973)

Distribution: Burundi; Tanzania

23 BUR 25 TAN

species :

Athyrium lineare Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 11 (1986)

Distribution: China (NW-Yunnan)

36 CHI

species :

Athyrium longius Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 178 (1940)

Distribution: China (Guizhou, Hunan)

36 CHI

Synonyms: hidden

= Athyrium hengshanense Ching; (1976) (nom. nud.

species :

Athyrium ludingense Z.R.Wang & Li Bing Zhang; Acta Bot. Yunnan. 17(3): 303, t. 1, f. 1-4 (1995)

Distribution: China (Sichuan)

36 CHI

species :

Athyrium mackinnoniorum (C.Hope) C.Chr.; Index Filic. 143 (1905) (´mackinnoni´, emend. Fraser-Jenkins 1997)

Distribution: E-Afghanistan; Pakistan (Azad Kashmir); N-India (Himachal Pradesh, Uttarakhand, Jammu & Kashmir)

34 AFG 40 IND PAK

Remarks: all of the Chinese and Indochina records belong to other species fide Fraser-Jenkins 2018, but identity is not yet clear

Synonyms: hidden

= Asplenium mackinnoniorum C.Hope; J. Bot. 34: 124 (1896), as ´mackinnonii´

= Athyrium caudipinna Ching; Acta Bot. Austro Sin. 1: 20 (1983) (fide Fraser-Jenkins 1997)

= Athyrium neomackinnonii Ching; Sporae Pterid. Sin. 211, pl. 43, f. 9-10 (1976), nom. nud.

= Athyrium varians Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(3): 9 (1984)

species :

Athyrium majus (Makino) Makino; Bot. Mag. (Tokyo) 28: 178 (1914)

Distribution: Japan

38 JAP

Remarks: possibly a synonym of A. wardii

Synonyms: hidden

= Athyrium wardii var. majus Makino; Bot. Mag. (Tokyo) 18: 28 (1899)

species :

Athyrium maoshanense Ching & P.S.Chiu; Acta Bot. Boreal.-Occid. Sin. 6(3): 157 (1986)

Distribution: China (S-Zhejiang)

36 CHI

species :

Athyrium masamunei Seriz.; Acta Phytotax. Geobot. 32(5-6): 176 (1981)

Distribution: S-Japan (Yakushima Isl.)

38 JAP NNS

Conservation Status: IUCN R. CR in Japan

species :

Athyrium mearnsianum (Copel.) Alderw.; Malayan Ferns Fern Allies, Suppl. 279 (1917)

Distribution: Philippines

42 PHI

Remarks: sect. Strigoathyrium, valid species and not a synonym of A. nitidulum fide Wardani et al. 2024

Synonyms: hidden

= Athyrium nigripes var. mearnsianum Copel.; Philipp. J. Sci., C, 3: 211 (1908)

species :

Athyrium medium (Carmich.) T.Moore; Index Fil. [T. Moore] 186 (1860)

Distribution: Tristan da Cunha (incl. Inaccessible Isl.)

91 TDC

Synonyms: hidden

= Aspidium medium Carmich.; Trans. Linn. Soc. London 12: 611 (1818)

= Asplenium medium (Carmich.) Hook.; Sp. Fil. [W. J. Hooker] 3: 227 (1860)

species :

Athyrium mehrae Bir; Nova Hedwigia 4: 165, t. 52 (1962)

Distribution: China (Gansu, Hubei, Sichuan, Yunnan); Tibet; Nepal; Bhutan; N-India (Sikkim, Uttarakhand, Arunachal Pradesh); N-Myanmar

36 CHI TIB 40 BHU IND NEP 41 MYA

Remarks: bona sp. and not a synonym of A. nephrodioides fide Fraser-Jenkins et al. (2015)

Synonyms: hidden

= Athyrium stenopodum Ching & S.K.Wu; Fl. Xizangica 1: 137 (1983) (fide Fraser-Jenkins 2008)

= Athyrium tibeticum Ching; Fl. Xizangica 1: 137 (1983) (fide Fraser-Jenkins 2008)

species :

Athyrium melanolepis (Franch. & Sav.) Christ; Bull. Herb. Boissier, sér. 1, 4: 668 (1896)

Distribution: China (Heilongjiang, Jilin); Japan

36 CHI 38 JAP

Synonyms: hidden

= Asplenium melanolepis Franch. & Sav.; Enum. Pl. Jap. 2: 226 (1877)

= Athyrium filix-femina subsp. melanolepis (Franch. & Sav.) Á.Löve & D.Löve; Taxon 26(2-3): 326 (1977)

= Athyrium filix-femina var. melanolepis (Franch. & Sav.) Makino; (1899)

= Athyrium filix-femina var. nigropaleaceum Makino; Bot. Mag. (Tokyo) 13: 29 (1899)

species :

Athyrium mengtzeense Hieron.; Hedwigia 59: 319 (1918), nom. nov.

Distribution: China (Yunnan, W-Guizhou, SW-Sichuan); Nepal; India (West Bengal, Sikkim, Arunachal Pradesh, Uttarakhand)

36 CHI 40 IND NEP

Remarks: kept separate from A. roseum by Flora of China and also by Fraser-Jenkins et al. (2015)

Synonyms: hidden

= Asplenium sinense Baker; Bull. Misc. Inform. Kew 1906: 9 (1906)

= Athyrium confertipinnum Ching; Acta Bot. Boreal.-Occid. Sin. 6(3): 155 (1986)

= Athyrium liangwangshanicum Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 21 (1986)

= Athyrium sinense (Baker) C.Chr.; Index Filic., Suppl. 2 (1906-1912): 15 (1913) (non Rupr. (1845))

= Athyrium tenuicaule (Hayata) Tagawa; Acta Phytotax. Geobot. 2(3): 195 (1933) (included in A. roseum by Fraser-Jenkins (2008))

= Athyrium thelypteroides var. henryi Christ; Bull. Herb. Boissier, sér. 1, 6: 961 (1898)

= Diplazium tenuicaule Hayata; Icon. Pl. Formosan. 4: 220, f. 150 (1914) (included in A. roseum by Fraser-Jenkins (2008))

species :

Athyrium microphyllum (Sm.) Alston; Philipp. J. Sci. 50: 178 (descr. orig.) (1933)

Distribution: Hawaii (Kauai, Oahu, Molokai, Lanai, Maui, Hawaii Isl.)

63 HAW

Synonyms: hidden

= Asplenium baldwinii Hillebr.; Fl. Hawaiian Isl. [Hillebrand] 618 (1888)

= Asplenium multisectum Brack.; U.S. Expl. Exped., Filic. 16: 174 (1854)

= Asplenium poiretianum Gaudich.; Voy. Uranie [Freycinet], Bot. 321, t. 13 (1827)

= Asplenium vexans Underw. ex A.Heller; Minnesota Bot. Stud. 1: 776 (1897)

= Athyrium baldwinii (Hillebr.) C.Chr.; Index Filic. 140 (1905)

= Athyrium poiretianum (Gaudich.) C.Presl; Tent. Pterid. 98 (1836)

= Darea microphylla Sm.; Cycl. [A. Rees] (London ed.) 11, no. 9 (1808)

species :

Athyrium micropterum Fraser-Jenk.; New Species Syndr. Indian Pteridol. 58 (1997)

Distribution: China (Himalayas); Tibet; Nepal; Bhutan; India (Uttarakhand, Himachal Pradesh, Sikkim, Darjeeling, Arunachal Pradesh, Nagaland, West Bengal)

36 CHI TIB 40 BHU IND NEP

Synonyms: hidden

= Asplenium macrocarpon var. atkinsonii C.B.Clarke ex Baker; Syn. Fil. [W. J. Hooker & Baker], ed. 2: 489 (1874)

= Athyrium macrocarpon var. atkinsonii (C.B.Clarke ex Baker) Tardieu; (1940)

species :

Athyrium minimum Ching; Acta Bot. Boreal.-Occid. Sin. 6(3): 151 (1986)

Distribution: Japan; Taiwan

38 JAP TAI

Conservation Status: EN in Taiwan

species :

Athyrium minutum Copel.; Univ. Calif. Publ. Bot. 18: 221 (1942)

Distribution: New Guinea (Irian Jaya, Papua New Guinea)

42 NWG

species :

Athyrium mohillense (Fée) Tardieu; Bull. Mus. Natl. Hist. Nat., sér. 2, 29: 290 (1957)

Distribution: Mauritius; La Réunion

29 MAU REU

Synonyms: hidden

= Asplenium mohillense (Fée) Kuhn; Filic. Afr. 107 (1868)

= Diplazium mohillense Fée; Mém. Foug. 8 (Icon. Esp. Nouv.): 85 (1857)

species :

Athyrium monomachii (Kom.) Kom.; Izv. Kievsk. Bot. Sada 13: 145 (1931)

Distribution: Siberia (Altai, Buryatia, Chita, Irkutsk, Krasnoyarsk, Yakutia); Russian Far East (Amur, Kamchatka, Khabarovsk, Magadan, Primorye); China (Hebei, Heilongjiang, Jilin, Liaoning, Nei Mongol, N-Shandong, Shanxi); Mongolia; North Korea; South Korea; Japan

30 SIB 31 RFE 36 CHI 37 MON 38 JAP KOR NKO

Remarks: included in A. sinense sensu lato by Fraser-Jenkins (2008) but kept separate by Flora of China

Synonyms: hidden

= Aspidium filix-femina var. multidentatum Döll; Gefässkrypt. Baden 24 (1855)

= Athyrium acutidentatum Ching; Acta Bot. Boreal.-Occid. Sin. 6(3): 150 (1986)

= Athyrium brevifrons Nakai ex Kitag.; Rep. First Sci. Exped. Manchoukuo 4(2): 75 (1935) [fide Fraser-Jenkins (2008)]

= Athyrium brevifrons var. angustifrons (Kodama) T.Mori; Enum. Pl. Corea: 4 (1922)

= Athyrium changbaishanense Ching & J.J.Chien; Bull. Bot. Res., Harbin 4(1): 128 (1984) ex Vorosch. (nom. inval.)

= Athyrium dailingense Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 18 (1986)

= Athyrium extensum Ching; (1976), nom. nud.

= Athyrium filix-femina subsp. jacutense (Petrov) Sipliv.; Novosti Sist. Vyssh. Rast. 11: 328 (1974)

= Athyrium filix-femina subsp. longipes (H.Hara) Tzvelev; Sosud. Rast. Sovet. Dal´nego Vostoka 5: 73 (1991)

= Athyrium filix-femina var. jacutense Petrov; Fl. Iakut. 1: 18 (1930)

= Athyrium filix-femina var. longipes H.Hara; Bot. Mag. (Tokyo) 48: 691 (1934) (non Athyrium longipes Christ)

= Athyrium filix-femina var. monomachii Kom.; Izv. Imp. Bot. Sada Petra Velikago 16: 149 (1916)

= Athyrium hebeiense Ching; Acta Bot. Boreal.-Occid. Sin. 6(3): 158 (1986)

= Athyrium melanolepis var. angustifrons Kodama; (1917)

= Athyrium multidentatum (Döll) Ching; Wang et al., Fl. Pl. Herb. Chin. Bor.-Or. 1: 40. (1958)

= Athyrium sinense f. multidentatum (Döll) Kitag.; Neo-Lineam. Fl. Manshur.: 32 (1979)

= Athyrium sinense subsp. longipes (H.Hara) Vorosch.; Florist. Issl. Razn. Raionakh SSSR [A. K. Svortsov]: 144 (1985)

= Athyrium submelanolepis Tzvelev; Novosti Sist. Vyssh. Rast. 26: 8 (1989), nom. nov.

species :

Athyrium morobense Copel.; Philipp. J. Sci. 78: 464, t. 40 (1951)

Distribution: New Guinea (Papua New Guinea)

42 NWG

species :

Athyrium multipinnum Y.T.Hsieh & Z.B.Wang; Acta Bot. Boreal.-Occid. Sin. 7(1): 55 (1987)

Distribution: China (Guizhou, Hunan, Jiangxi, Zhejiang)

36 CHI

species :

Athyrium myer-dreesii Copel.; Univ. Calif. Publ. Bot. 18: 222 (1942)

Distribution: Moluccas (Seram); New Guinea (Irian Jaya)

42 MOL NWG

species :

Athyrium nakanoi Makino; Bot. Mag. (Tokyo) 23: 247 (1909)

Distribution: China (Yunnan); Tibet; Japan (Yakushima Isl.); Taiwan; Bhutan; N-India (Arunachal Pradesh, Sikkim); Myanmar (Kachin); Philippines (Mindanao)

36 CHI TIB 38 JAP NNS TAI 40 BHU IND 41 MYA 42 PHI

Remarks: sect. Polystichoides, anisopterum clade. Not in Nepal

Conservation Status: IUCN NT for India (Fraser-Jenkins 2012)

Synonyms: hidden

= Asplenium macrocarpon var. unipinnatum C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 489 (1880)

= Athyrium macrocarpon var. unipinnatum (C.B.Clarke) Bedd.; Ferns Brit. India 165 (1883)

= Athyrium obtusifolium Rosenst.; Hedwigia 66: 335 (1915)

= Athyrium subfalcatum Ching; (1976), nom. nud.

= Athyrium tenuissimum (Hayata) Kodama; Gen. Ind. Fl. Taiwan 4: 202 (1914 publ. 1915)

= Athyrium tenuissimum (Hayata) Merr.; Philipp. J. Sci., C, 13: 126 (1918), isonym

= Nephrolepis tenuissima Hayata; Icon. Pl. Formosan. 4: 202, f. 137 (1914)

species :

Athyrium nanyueense Ching; Acta Bot. Boreal.-Occid. Sin. 6(3): 152 (1986)

Distribution: China (Hunan)

36 CHI

Remarks: probably a synonym of A. distans fide Fraser-Jenkins et al. 2018

species :

Athyrium neglectum Seriz.; J. Jap. Bot. 55(8): 250 (1980)

Distribution: Japan

38 JAP

Conservation Status: CR in Japan

species :

Athyrium nephrodioides (Baker) Christ; Mém. Soc. Bot. France 1 [Hist. Acrostich.]: 47 (1905)

Distribution: China

36 CHI

Remarks: status unclear, possibly a synonym or aberrant specimen of A. mehrae

Synonyms: hidden

= Asplenium nephrodioides Baker; J. Bot. 25: 170 (1887)

species :

Athyrium newtonii (Baker) Diels; Nat. Pflanzenfam. [Engler & Prantl] I, 4: 224 (1899)

Distribution: Guinea; Liberia; São Tomé; D.R.Congo [Zaire]; Rwanda; Burundi; Sudan; Uganda; Kenya; Tanzania; Malawi; E-Zimbabwe; Mozambique; South Africa (Limpopo, Mpulamanga, KwaZulu-Natal); Lesotho; Eswatini [Swaziland]; ?Madagascar

22 GUI LBR 23 BUR RWA SAO ZAI 24 SUD 25 KEN TAN UGA 26 MLW MOZ ZIM 27 LES SAF SWZ

Remarks: sect. Strigoathyrium, scandicinum agg.

Synonyms: hidden

= Allantodia aspidioides (Willd.) Kunze; Bot. Zeitung (Berlin) 6: 191 (1848)

= Aspidium aspidioides (Willd.) Christ; Farnkr. Erde: 224, 225 (1897)

= Asplenium aspidioides (Willd.) Schltdl.; Adumbr. Pl. 24, t. 13 (1825)

= Asplenium aspidioides (Willd.) Spreng.; Syst. Veg., ed. 16 [Sprengel] 4(1): 90 (1827), isonym

= Asplenium laxum (Pappe & Rawson) Kuhn; Filic. Afr. 105 (1868) (non R. Br. (1810))

= Asplenium newtonii Baker; Ann. Bot. (Oxford) 6: 307 (1891)

= Athyrium aspidioides (Willd.) Christ; Farnkr. Erde 224 (1897)

= Athyrium laxum Pappe & Rawson; Syn. Filic. Afr. Austr. 16 (1858) (non Schumach. (1803))

= Athyrium scandicinum subsp. newtonii (Baker) Verdc.; Fl. Trop. E. Africa, Woods. 10 (2003)

= Athyrium scandicinum var. rhodesianum Schelpe; Bol. Soc. Brot., ser. 2(41): 211 (1967)

= Caenopteris aspidioides (Willd.) Desv.; Mem. Soc. Linn. Paris 6(2): 268 (1827)

= Darea aspidioides Willd.; Enum. Pl. Hort. Berol. 1072 (1809)

= Dryopteris schnellii Tardieu; Notul. Syst. (Paris) 13: 370 (1949)

species :

Athyrium nigripes (Blume) T.Moore; Index Fil. [T. Moore] 49, 98 (1858)

Distribution: Sulawesi; Philippines (Luzon); Java; Lesser Sunda Isl. (Bali, Flores, Timor); New Guinea (Irian Jaya, Papua New Guinea)

42 JAW LSI NWG PHI SUL

Remarks: sect. Strigoathyrium, nigripes clade

Synonyms: hidden

= Allantodia paludosa Zippel; Kunze, Bot. Zeitung (Berlin) 191 (1848)

= Aspidium costale Blume; Enum. Pl. Javae 2: 170 (1828)

= Aspidium nigripes Blume; Enum. Pl. Javae 2: 162 (1828)

= Asplenium denticulatum J.Sm.; Cat. Cult. Ferns: 47 (1857), nom. illeg.

= Asplenium incisum J.Sm.; J. Bot. (Hooker) 4: 174 (1841), nom. illeg.

= Asplenium nigripes (Blume) Hook.; Sp. Fil. [W. J. Hooker] 3: 222 (1860), nom. illeg.

= Asplenium paludosum T.Moore; Index Fil. [T. Moore] 162 (1859)

= Athyrium aristulatum Copel.; Philipp. J. Sci. 1, Suppl. IV: 253 (1906)

= Athyrium purpureum Lowe ex Milde; Fil. Eur. 49 (1867)

species :

Athyrium nikkoense Makino; Bot. Mag. (Tokyo) 17: 43 (1903)

Distribution: Japan

38 JAP

species :

Athyrium nitidulum (Kunze) Milde; Bot. Zeitung (Berlin) 370 (1870)

Distribution: Myanmar (Kachin); Philippines; Java

41 MYA 42 JAW PHI

Remarks: sect. Strigoathyrium, non-setose group, erythropodum clade

Synonyms: hidden

= Allantodia nitidula Kunze; Bot. Zeitung (Berlin) 191 (1848)

= Asplenium nitidulum (Kunze) T.Moore; Index Fil. [T. Moore] 43 (1857)

= Athyrium philippinense Christ; Copel., Philipp. J. Sci. 1, Suppl. IV: 254 (1906)

= Athyrium sarasinorum var. philippinense Christ; Bull. Herb. Boissier, sér. 1, 6: 154 (1898)

species :

Athyrium oblitescens Sa.Kurata; J. Geobot. 17: 3 (1969)

Distribution: Japan

38 JAP

Remarks: sect. Strigoathyrium, non-setose group

species :

Athyrium obtusilimbum Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 11 (1986)

Distribution: China (NW-Yunnan)

36 CHI

species :

Athyrium omeiense Ching; Bull. Fan Mem. Inst. Biol. Bot., ser. 2, 1: 282 (1949)

Distribution: China (Chongqing, SE-Gansu, NW-Guizhou, NW-Hubei, Hunan, Shaanxi, Sichuan, N-Yunnan)

36 CHI

Synonyms: hidden

= Athyrium amplissimum Ching, Boufford & K.H.Shing; J. Arnold Arbor. 64(1): 20 (1983) (non (Baker) Holttum 1955)

= Athyrium crassiusculum Ching; (1976), nom. nud.

= Athyrium filix-femina var. flavicoma Christ; (1905)

= Athyrium flavicoma (Christ) Ching; (1986)

= Athyrium magnum Ching; (1976), nom. nud.

= Athyrium parapellucidum Ching; Fl. Tsinling. 2: 218 (1974)

= Athyrium pellucidum Ching; Fl. Tsinling. 2: 217 (1974)

= Athyrium subtripinnatum Ching; (1976), nom. nud.

species :

Athyrium oosorum (Baker) Christ; Bot. Jahrb. Syst. 28: 349 (1896)

Distribution: [Western] Samoa (Savai´i)

60 SAM

Synonyms: hidden

= Asplenium oosorum Baker; J. Bot. 14: 343 (1876)

species :

Athyrium oppositipennum Hayata; J. Coll. Sci. Imp. Univ. Tokyo 30: 441 (1911) (oppositipinnum auct.)

Distribution: Taiwan; Philippines (Luzon)

38 TAI 42 PHI

Remarks: not in Japan

Synonyms: hidden

= Athyrium oreopteris Copel.; Philipp. J. Sci. 40: 302, t. 5 (1929)

= Athyrium taiwanense Tagawa; Acta Phytotax. Geobot. 2: 18 (1932)

= Dryopteris oppositipenna (Hayata) Hayata; J. Coll. Sci. Imp. Univ. Tokyo 30: 450 (1911)

variant :

var. pubescens (Tagawa) Tagawa; Acta Phytotax. Geobot. 4: 142-143 (1935)

Distribution: Japan (Kyushu, Yakushima Isl.); Taiwan

38 JAP NNS TAI

Synonyms: hidden

= Athyrium oppositipennum var. pubescens (Tagawa) Y.C.Liu, W.L.Chiou & H.Y.Liu; Fern Fl. Taiwan Athyrium 36 (2009), isonym (isonym)

= Athyrium reflexipinnum Hayata; Icon. Pl. Formosan. 4: 234, t. 164 (1914)

= Athyrium taiwanense var. pubescens Tagawa; Acta Phytotax. Geobot. 2: 18 (1933)

species :

Athyrium otophorum (Miq.) Koidz.; Fl. Symb. Orient.-Asiat. 40 (1930)

Distribution: China (Anhui, Chongqing, Fujian, Guangdong, Guangxi, Guizhou, Hubei, Hunan, Sichuan, Yunnan, Zhejiang); South Korea (Cheju Isl.); Japan; Taiwan; N-India (Uttarakhand, Pithoragarh, Banlekh, Champawat, Meghalaya)

36 CHI 38 JAP KOR TAI 40 IND

Remarks: sect. Strigoathyrium, non-setose group

Conservation Status: IUCN EN for India (Fraser-Jenkins 2012)

Synonyms: hidden

= Asplenium otophorum Miq.; Ann. Mus. Bot. Lugduno-Batavi 8: 175 (1867)

= Athyrium erythrocaulon Ching; Acta Bot. Boreal.-Occid. Sin. 6(3): 154 (1986)

= Athyrium otophorum f. okanum (Sa.Kurata) Ebihara; Bull. Nat. Mus. Nat. Sci., Ser. B, 43(1): 19 (2017)

= Athyrium otophorum var. okanum Sa.Kurata; J. Geobot. 17: 3 (1969)

= Athyrium rigescens Makino; Bot. Mag. (Tokyo) 18: 27 (1899)

= Athyrium sinovidalii Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(3): 8 (1984)

= Athyrium tsingyingense Ching; (1976), nom. nud.

= Athyrium vidalii var. chinense Koidz.

= Athyrium violascens Diels; Bot. Jahrb. Syst. 29: 196 (1900)

= Athyrium wardii Christ; (1905) (non (Hook.) Makino 1857)

= Diplazium otophorum (Miq.) C.Chr.; Index Filic. 236 (1905)

species :

Athyrium pachyphyllum Ching; Acta Bot. Boreal.-Occid. Sin. 6(2): 102 (1986)

Distribution: China (Hunan, N-Guangxi, S-Guizhou, SE-Yunnan)

36 CHI

Synonyms: hidden

= Athyrium xiangxiense S.F.Wu; Keys Vasc. Pl. Wuling Mts. 563 (1995)

species :

Athyrium palustre Seriz.; J. Jap. Bot. 45: 264 (1970)

Distribution: Japan; Taiwan

38 JAP TAI

Remarks: included by Fraser-Jenkins et al. (2018) in A. devolii

Conservation Status: EN in Japan

species :

Athyrium parasnathense (C.B.Clarke) Ching ex Mehra & Bir; Res. Bull. Panjab Univ. Sci. 15: 141 (1964)

Distribution: India (?Chhattisgarh, Gujarat, Jharkhand, Karnataka, Madhya Pradesh, Maharashtra, Odisha, Rajasthan, Tamil Nadu)

40 IND

Synonyms: hidden

= Asplenium filix-femina var. parasnathensis C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 493, t. 61, C 2 (1880)

= Athyrium filix-femina var. parasnathensis (C.B.Clarke) Bedd.; Ferns Brit. India 170 (1883)

= Athyrium pectinatum var. tenellum Bedd.; Suppl. Ferns Brit. India 36 (1892)

species :

Athyrium pectinatum (Wall. ex C.Hope) C.Presl; Tent. Pterid. 98 (1836)

Distribution: Tibet; Nepal; Bhutan; India (Arunachal Pradesh, Himachal Pradesh, Jharkand, Madhya Pradesh, Manipur, Meghalaya, Mizoram, +Rajasthan, Uttarakhand, West Bengal, Jammu & Kashmir, Sikkim, Darjeeling)

36 TIB 40 BHU IND NEP

Synonyms: hidden

= Asplenium filix-femina var. pectinatum (Wall. ex Mett.) C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 492 (1880)

= Asplenium pectinatum Wall.; Numer. List. [Wallich] no. 231 (1829), nomen

= Asplenium pectinatum Wall. ex C.Hope; J. Bombay Nat. Hist. Soc. 14: 253 (1902)

= Asplenium pectinatum Wall. ex Mett.; Abh. Senckenberg. Naturf. Ges. 3(1) [Aspl.] 197 (1859)

= Athyrium filix-femina subsp. pectinatum (Wall.) Á.Löve & D.Löve; Taxon 26(2-3): 326 (1977) (fide Fraser-Jenkins 1997)

= Athyrium filix-femina var. pectinatum (Wall. ex Mett.) Bedd.; Ferns Brit. India 169 (1883)

= Athyrium pectinatum (Wall. ex Mett.) Bedd.; Ferns S. India 51, t. 154 (1863), non A. pectinatum (Wall. ex Mett.) T. Moore (1860)

species :

Athyrium pinetorum Tagawa; Acta Phytotax. Geobot. 2(1): 16 (1932)

Distribution: Japan

38 JAP

species :

Athyrium pitcairnense Copel.; Occas. Pap. Bernice P. Bishop Mus. 14: 60, t. 11 (1938)

Distribution: Pitcairn Isl.

61 PIT

species :

Athyrium praetermissum Sledge; Ann. Mag. Nat. Hist. XII, 9: 457, t. 16 (1956)

Distribution: S-India (Kerala, Tamil Nadu); Sri Lanka

40 IND SRL

Synonyms: hidden

= Athyrium praetermissum var. tripinnatum Sledge; Ann. Mag. Nat. Hist. XII, 9: 453 (1956)

species :

Athyrium pubicostatum Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(3): 7 (1984)

Distribution: China (Hubei, Hunan, Guangxi, Sichuan, Chongqing, Guizhou, Yunnan); Taiwan

36 CHI 38 TAI

Remarks: sect. Strigoathyrium, non-setose group

Synonyms: hidden

= Athyrium guizhouense Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 11 (1986)

= Athyrium hirtirachis Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(3): 6 (1984) (non Ching & Y. P. Hsu 1974)

= Athyrium kweichowense Ching; (1976), nom. nud.

= Athyrium pseudoepirachis Ching; Acta Bot. Boreal.-Occid. Sin. 6(3): 156 (1986)

= Athyrium sessile Ching; Acta Bot. Boreal.-Occid. Sin. 6(3): 154 (1986)

= Athyrium simulans Ching; (1976), nom. nud.

= Athyrium subpubicostatum Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(3): 6 (1984)

species :

Athyrium pulcherrimum Copel.; Philipp. J. Sci. 8: 141, t. 3 (1913)

Distribution: Borneo (Mt. Kinabalu, Sabah); Java

42 BOR JAW

Remarks: sect. Strigoathyrium, nigripes clade

species :

Athyrium puncticaule (Blume) T.Moore; Index Fil. [T. Moore] 186 (1860)

Distribution: Japan; Taiwan; India (Arunachal Pradesh, Kerala, Meghalaya, Nagaland, Tamil Nadu); Sri Lanka; Myanmar (Kachin, etc.); Thailand; Vietnam; Sumatra; Sulawesi; Philippines (Luzon); Java; Lesser Sunda Isl. (Lombok); Moluccas (Seram)

38 JAP TAI 40 IND SRL 41 MYA THA VIE 42 JAW LSI MOL PHI SUL SUM

Remarks: sect. Polystichoides, anisopterum clade. Confirmed for Taiwan by Knapp (2011) and Liu et al. (2009)

Synonyms: hidden

= Aspidium lanceum Kunze; Bot. Zeitung (Berlin) 473 (1846) (fide Fraser-Jenkins 2008)

= Aspidium macrocarpon Blume; Enum. Pl. Javae 2: 162 (1828)

= Aspidium puncticaule Blume; Enum. Pl. Javae 2: 156 (1828)

= Asplenium fallax Mett.; Abh. Senckenberg. Naturf. Ges. 3(1) [Aspl.]: 194, no. 236 (1859)

= Asplenium macrocarpon (Blume) Hook.; Sp. Fil. [W. J. Hooker] 3: 222 (1860), isonym

= Asplenium macrocarpon (Blume) J.Sm.; Cat. Cult. Ferns: 47 (1857), as ´macrocarpum´

= Asplenium oxyphyllum J.Sm.; J. Bot. 8: 408 (1841) ex Kunze, Bot. Zeitung 1846: 441 (1846), non Hook. 1860

= Asplenium puncticaule (Blume) Mett.; Ann. Mus. Bot. Lugduno-Batavi 2: 240 (1866)

= Athyrium fallax Milde; Fil. Eur. 49 (1867)

= Athyrium halconense Christ; Philipp. J. Sci. 8: 273 (1908)

= Athyrium hochreutineri Christ; Annuaire Conserv. Jard. Bot. Genève 15-16: 197 (1912)

= Athyrium lanceum (Kunze) T.Moore; Index Fil. [T. Moore] 185 (1860) (fide Fraser-Jenkins 2008)

= Athyrium macrocarpum (Blume) Bedd.; Ferns S. India 51, t. 153 (1865)

= Lastrea macrocarpa (Blume) T.Moore; Index Fil. [T. Moore] 95 (1858)

= Nephrodium lanceum (Kunze) T.Moore; Index Fil. [T. Moore] 95 (1858) (fide Fraser-Jenkins 2008)

= Nephrodium sparsum var. membranaceum Baker; Syn. Fil. [W. J. Hooker & Baker], ed. 2: 498 (1874)

= Tarachia oxyphylla (J.Sm.) C.Presl; Abh. Königl. Böhm. Ges. Wiss., ser. 5, 6 [Epim. Bot.]: 77 (1851)

species :

Athyrium repens (Ching) Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 238 (2008)

Distribution: China (Sichuan); ?Tibet; Nepal; Bhutan; India (Sikkim)

36 CHI 40 BHU IND NEP

Conservation Status: IUCN EN for India (Fraser-Jenkins 2012)

Synonyms: hidden

= Pseudocystopteris repens Ching; Acta Phytotax. Sin. 9: 80 (1964)

species :

Athyrium rhachidosorum (Hand.-Mazz.) Ching; C. Chr., Index Filic., Suppl. Tert.: 43 (1934)

Distribution: China (Yunnan, SW-Sichuan); Tibet; Myanmar

36 CHI TIB 41 MYA

Remarks: sect. Strigoathyrium, nigripes clade

Synonyms: hidden

= Asplenium rhachidosorum Hand.-Mazz.; Symb. Sin. 6: 33, t. 2, f. 2 (1929)

= Athyrium densisorum X.C.Zhang; Bull. Bot. Res., Harbin 11(3): 8 (1991)

= Athyrium wuliangshanense Ching; Acta Bot. Boreal.-Occid. Sin. 6(3): 153 (1986)

species :

Athyrium rondoense Verdc.; Fl. Trop. E. Africa, Woods. 12 (2003)

Distribution: Tanzania

25 TAN

species :

Athyrium roseum Christ; Bull. Herb. Boissier, sér. 1, 6: 961 (1898)

Distribution: China (Yunnan); Nepal; India (Darjeeling); Myanmar (Chin)

36 CHI 40 IND NEP 41 MYA

Remarks: sect. Strigoathyrium, non-setose group

Conservation Status: IUCN CR for India (Fraser-Jenkins 2012)

variant :

var. fugongense Z.R.Wang; Vasc. Pl. Hengduan Mount. 1: 85 (1993), without latin descr. or type, et Fl. Reipubl. Popularis Sin. [Fl. China Ed. Committee] 3(2): 233 (1999)

Distribution: China (Yunnan)

36 CHI

species :

Athyrium rubripes (Kom.) Kom.; Izv. Kievsk. Bot. Sada 13: 145 (1931) [fide Fraser-Jenkins (2008)]

Distribution: Siberia (Altai, Buryatia, Chita, Irkutsk, Krasnoyarsk, W-Siberia, Yakutia); Russian Far East (Amur, Kamchatka, Khabarovsk, Kuril Isl., Magadan, Primorye, Sakhalin); ?Kazakhstan; China (NE-Heilongjiang)

30 SIB 31 RFE 36 CHI

Remarks: included in A. sinense sensu lato by Fraser-Jenkins (2008) but kept separate by Flora of China

Synonyms: hidden

= Athyrium filix-femina var. rubripes Kom.; Izv. Imp. Bot. Sada Petra Velikago 16: 149 (1916)

species :

Athyrium ruilicolum W.M.Chu; Acta Bot. Yunnan., Suppl. 5: 41 (1992), as ´ruilicola´

Distribution: China (Yunnan)

36 CHI

species :

Athyrium rupicola (C.Hope) C.Chr.; Index Filic. 145 (1905)

Distribution: China (Yunnan, Sichuan); Tibet; Pakistan (Hazara, Baltistan); Nepal; Bhutan; N-India (Himachal Pradesh, Sikkim, Uttarakhand, West Bengal, Jammu & Kashmir); Myanmar (Kachin, etc.)

36 CHI TIB 40 BHU IND NEP PAK 41 MYA

Synonyms: hidden

= Asplenium filix-femina subvar. elongatum C.B.Clarke ex Blanf.; J. Asiat. Soc. Bengal, Pt. 2, Nat. Hist. 57: 306 (1888)

= Asplenium filix-femina var. retusum C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 492 (1880) (p. p.)

= Asplenium rupestre C.Hope; (1899), nom. nud.

= Asplenium rupicola Edgew. ex C.Hope; J. Bombay Nat. Hist. Soc. 12: 531, t. 5 (1899)

= Athyrium falcipinna Ching; Sporae Pterid. Sin. 214, pl. 43, f. 21-22 (1976) (nom. nud. inval.)

= Athyrium filix-femina var. retusum (C.B.Clarke) Bedd.; Ferns Brit. India 170 (1883) (p. p.)

= Athyrium glandulosum Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 19 (1986)

species :

Athyrium sarasinorum Christ; Verh. Naturf. Ges. Basel 11: 434 (1896)

Distribution: Sulawesi

42 SUL

Synonyms: hidden

= Asplenium sarasinorum (Christ) Copel.; Polypodiaceae Philipp. 87 (1905)

species :

Athyrium satowii H.Itô; J. Jap. Bot. 27: 337, f. 1-2 (1952)

Distribution: Japan

38 JAP

species :

Athyrium scandicinum (Willd.) C.Presl; Tent. Pterid. 98 (1836)

Distribution: Madagascar; La Réunion

29 MDG REU

Remarks: sect. Strigoathyrium, scandicinum agg.

Synonyms: hidden

= Allantodia scandicina (Willd.) Kaulf.; Enum. Filic. [Kaulfuss] 179 (1824)

= Aspidium inquinans (Fée) Kuhn; Filic. Afr. 135 (1868)

= Aspidium scandicinum Willd.; Sp. Pl., ed. 4 [Willdenow] 5(1): 286 (1810)

= Asplenium scandicinum (Willd.) A.Heller; Minnesota Bot. Stud. 1: 775 (1897) (non R. Br. (1824))

= Athyrium scandicinum f. bipinnatum Rakotondr.; Bull. Mus. Nat. Hist. Nat., B, Adansonia, ser. 4, 10(4): 375 (1988 publ. 1989)

= Chamaefilix scandicina (Willd.) Farw.; Amer. Midl. Naturalist 12(8): 275 (1931)

= Cystopteris scandicina (Willd.) Desv.; Mem. Soc. Linn. Paris 6(2): 264 (1827)

= Nephrodium inquinans Fée; Mém. Soc. Sci. Nat. Strasbourg 6(1): 43, t. 44, fig. 1 (1866)

= Nephrodium scandicinum (Willd.) Bory; Voy. Indes Or. [Bélanger] 2: 63 (1833)

= Polypodium multifissum Goldm.; Nov. Actorum Acad. Caes. Leop.-Carol. Nat. Cur. 19 (Suppl. 1): 453 (1843)

species :

Athyrium schimperi Moug. ex Fée; Mém. Foug. 5 (Gen. Filic.): 187 (1852)

Distribution: Ghana; Nigeria; Cameroon; D.R.Congo [Zaire]; Rwanda; Burundi; Sudan; Ethiopia; Uganda; Kenya; Tanzania; Angola; Zambia; Malawi; Zimbabwe; South Africa (Mpulamanga, KwaZulu-Natal, E-Cape Prov.); Lesotho; Madagascar

22 GHA NGA 23 BUR CMN RWA ZAI 24 ETH SUD 25 KEN TAN UGA 26 ANG MLW ZAM ZIM 27 LES SAF 29 MDG

Synonyms: hidden

= Aspidium abyssinicum Döll; Gefässkrypt. Baden 23 (1855)

= Aspidium repens Döll; Jahresber. Mannheimer Vereins Naturk. 28: 39 (1862)

= Asplenium schimperi (Moug. ex Fée) A.Braun; Beitr. Fl. Aethiop. 1: 224 (1867), nom. illeg.

= Athyrium filix-femina var. schimperi (Moug. ex Fée) C.B.Clarke & Baker; (1888)

= Athyrium solenopteris var. madagascarica Bonap.; Notes Ptéridol. 5: 59 (1917)

species :

Athyrium schizochlamys (Ching) K.Iwats., H.Ohba & S.B.Malla; Bull. Univ. Mus. Univ. Tokyo 31 (Himalayan Plants 1): 313 (1988)

Distribution: China (Sichuan, Yunnan); Tibet; Nepal; Bhutan

36 CHI TIB 40 BHU NEP

Synonyms: hidden

= Athyrium atuntzeense (Ching) Z.R.Wang & Z.R.He; Fl. China 2-3: 498 (2013) (fide Fraser-Jenkins 2018)

= Pseudocystopteris atuntzeensis Ching; Acta Phytotax. Sin. 9: 83 (1964) (fide Fraser-Jenkins 2018)

= Pseudocystopteris lanpingensis Ching; Acta Phytotax. Sin. 9: 82 (1964)

= Pseudocystopteris laterepens Ching; Acta Phytotax. Sin. 9: 82 (1964)

= Pseudocystopteris schizochlamys Ching; Acta Phytotax. Sin. 9: 79 (1964) (fide Fraser-Jenkins 1997)

= Pseudocystopteris sinica Ching; Acta Phytotax. Sin. 9: 81 (1964)

species :

Athyrium sericellum Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 17 (1986)

Distribution: China (Chongqing)

36 CHI

species :

Athyrium sessilipinnum X.C.Zhang & R.Wei; Brittonia 68(4): 444 (2016)

Distribution: China (Guangxi)

36 CHI

Remarks: sect. Strigoathyrium, arisanense agg.

species :

Athyrium setiferum C.Chr.; Index Filic. 146 (1905), nom. nov. for Asplenium tenellum (T. Moore ex R. Sim) C. Hope

Distribution: China (Yunnan, Guangdong); Tibet; Japan; Nepal; Bhutan; India (Arunachal Pradesh, Himachal Pradesh, Kerala, Madhya Pradesh, Manipur, Meghalaya, Sikkim, Tamil Nadu, Uttarakhand, West Bengal); Sri Lanka; Myanmar (Kachin, etc.); N-Thailand; peninsular Malaysia; Sumatra; Borneo; ?Philippines (?Luzon); Moluccas (Seram); New Guinea (Irian Jaya, Papua New Guinea)

36 CHI TIB 38 JAP 40 BHU IND NEP SRL 41 MYA THA 42 BOR MLY MOL NWG SUM

Remarks: included in A. strigillosum by Flora of China but kept separate by Fraser-Jenkins (2008, 2015, 2018). A. tozanense is included here by Fraser-Jenkins et al. (2018)

Synonyms: hidden

= Asplenium gracile D.Don; Prodr. Fl. Nepal. 8 (1825) non E. Fourn. 1872

= Asplenium tenellum (Wall. ex T.Moore ex R.Sim) C.Hope; J. Bombay Nat. Hist. Soc. 12: 259, t. 4 (1899) (non Roxb. (1816) nec Fée (1852))

= Asplenium tenuifrons var. tenellum T.Moore; Index Fil. [T. Moore] 188 (1860), isonym, nom. nov.

= Asplenium tenuifrons var. tenellum Wall. ex T.Moore ex R.Sim; Priced Cat. Stove Greenh. Hardy Exot. Ferns 6: 22 (1859)

= Athyrium medogense X.C.Zhang; Bull. Bot. Res., Harbin 11(3): 11 (1991) (fide Fraser-Jenkins et al. 2018)

= Athyrium supraspinescens C.Chr.; Contr. U. S. Natl. Herb. 297, t. 19 (1931) (fide Fraser-Jenkins et al. 2018)

species :

Athyrium setuligerum Sa.Kurata; J. Geobot. 15: 6 (1966)

Distribution: Japan

38 JAP

species :

Athyrium silvicola Tagawa; Acta Phytotax. Geobot. 2(3): 17 (1933)

Distribution: Japan; Taiwan

38 JAP TAI

Remarks: sect. Strigoathyrium, nitidulum-group

species :

Athyrium sinense Rupr.; Distr. Crypt. Vasc. Ross. 41 (1845)

Distribution: Russian Far East (Primorye); China (SE-Gansu, Hebei, W-Henan, Nei Mongol, Ningxia, Shaanxi, Shandong, Shanxi); Mongolia

31 RFE 36 CHI 37 MON

Synonyms: hidden

= Athyrium filix-femina subsp. sinense (Rupr.) Tzvelev; Novosti Sist. Vyssh. Rast. 32: 182 (2000)

= Athyrium huhsienense Ching & Y.P.Hsu; Fl. Tsinling. 2: 218 (1974)

= Athyrium laochingshanense Ching; (1976), nom. nud.

= Athyrium shaanxiense Ching & Y.T.Hsieh; Acta Bot. Boreal.-Occid. Sin. 6(3): 150 (1986) (fide Fraser-Jenkins (2008)

species :

Athyrium sohayakiense Nakaike; New Fl. Japan, Pterid. 839 (1992), nom. nov.

Distribution: Japan

38 JAP

Synonyms: hidden

= Athyrium neglectum subsp. australe Seriz.; J. Phytogeogr. Taxon. 33(2): 76 (1985)

species :

Athyrium solenopteris (Kunze) T.Moore; Index Fil. [T. Moore] 43 (1857), 187 (1860)

Distribution: S-India (Karnataka, Kerala, Tamil Nadu); Sri Lanka; Java; New Guinea (Irian Jaya, Papua New Guinea)

40 IND SRL 42 JAW NWG

Remarks: sect. Strigoathyrium, non-setose group, erythropodum clade

Synonyms: hidden

= Allantodia solenopteris Kunze; Linnaea 24: 266 (1851)

= Asplenium nigripes var. solenopteris (Kunze) C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 491 (1880)

= Asplenium solenopteris (Kunze) Mett.; Abh. Senckenberg. Naturf. Ges. 3(1) [Aspl.]: 240 (1859), as ´selenopteris´

= Athyrium ceylonense (Klotzsch) T.Moore; Index Fil. [T. Moore] 181 (1860), nom. nud. (fide Fraser-Jenkins 2008)

= Athyrium nigripes Bedd.; Ferns S. India t. 157 (1863), non (Blume) T. Moore

= Athyrium pectinatum Bedd.; Ferns S. India t. 156 (1863) (p. p., non Asplenium pectinatum Wall. ex Mett.)

= Athyrium selenopteris (Kunze) Bedd.; Ferns Brit. India 166 (1883), orth. var.

species :

Athyrium spinescens Sa.Kurata; J. Geobot. 15: 7 (1966)

Distribution: Japan

38 JAP

species :

Athyrium spinulosum (Maxim.) Milde; Bot. Zeitung (Berlin) 376 (1866)

Distribution: Russian Far East (Amur, Khabarovsk, Primorye, Sakhalin); China (Chongqing, Gansu, Heilongjiang, Henan, Hubei, Jilin, Liaoning, Nei Mongol, Qinghai, Shaanxi, Sichuan, Yunnan); Tibet; North Korea; South Korea; Japan; Nepal; Bhutan; India (Sikkim)

31 RFE 36 CHI TIB 38 JAP KOR NKO 40 BHU IND NEP

Remarks: A. subtriangulare is included here by Fraser-Jenkins (2008, 2015, 2018) but kept separate by Flora of China

Synonyms: hidden

= Asplenium spinulosum (Maxim.) Baker; Syn. Fil. [W. J. Hooker & Baker] 225 (1867)

= Asplenium spinulosum (Maxim.) Miq.; Ann. Mus. Bot. Lugduno-Batavi 3: 174 (1867), isonym

= Asplenium spinulosum C.B.Clarke; (1880), not (Blume) Mett. (1859), nor Miq. (1867)

= Asplenium spinulosum var. subtriangulare (Hook.) C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 487 (1880)

= Asplenium subtriangulare Hook.; Syn. Fil. [W. J. Hooker & Baker] 225 (1867)

= Athyrium hookerianum T.Moore; Index Fil. [T. Moore] 185 (1860)

= Athyrium longipes Christ; Mém. Soc. Bot. France 1 [Hist. Acrostich.]: 48 (1905)

= Athyrium sikkimense (Bir) Á.Löve & D.Löve; Taxon 26(2-3): 326 (1977)

= Athyrium spinulosum var. subtriangulare (Hook.) C.Chr.; Index Filic. 3: 146 (1905)

= Athyrium subtriangulare (Hook.) Bedd.; Ferns Brit. India t. 242 (1868)

= Athyrium subtriangulare var. sikkimense Bir; Nova Hedwigia 4: 168, t. 53, f. 9, 10 (1962)

= Cystopteris spinulosa Maxim.; Mém. Acad. Imp. Sci. St. Pétersb. 9: 340 (1859)

= Pseudocystopteris decipiens Ching & S.K.Wu; Fl. Xizangica 1: 125 (1983)

= Pseudocystopteris longipes (Christ) Ching; Acta Phytotax. Sin. 9: 78 (1964)

= Pseudocystopteris purpurascens Ching & S.K.Wu; Fl. Xizangica 1: 128 (1983)

= Pseudocystopteris reflexipinnula Ching & S.K.Wu; Fl. Xizangica 1: 127 (1983)

= Pseudocystopteris remota Ching; Acta Phytotax. Sin. 9: 84 (1964)

= Pseudocystopteris sikkimensis (Bir) Ching; Acta Phytotax. Sin. 9: 79 (1964)

= Pseudocystopteris sparsa Ching & S.K.Wu; Fl. Xizangica 1: 124 (1983)

= Pseudocystopteris spinulosa (Maxim.) Ching; Acta Phytotax. Sin. 9: 78 (1964)

= Pseudocystopteris spinulosa var. taipaishanensis Ching; Fl. Tsinling. 2: 212 (1974)

= Pseudocystopteris subtriangularis (Hook. ex Baker) Ching; Acta Phytotax. Sin. 9: 78 (1964)

= Pseudocystopteris tibetica Ching; Acta Phytotax. Sin. 9: 80 (1964)

species :

Athyrium strigillosum (T.Moore ex Lowe) T.Moore ex Salomon; Nomencl. Gefäßkrypt. 112 (1883)

Distribution: China (Jiangxi, Hunan, Guangdong, Guangxi, Sichuan, Guizhou, Yunnan); Tibet; S-Japan (Yakushima Isl.); Taiwan; Nepal; Bhutan; N-India (Jammu & Kashmir, Himachal Pradesh, Meghalaya, Sikkim, Uttarakhand, West Bengal); Myanmar

36 CHI TIB 38 JAP NNS TAI 40 BHU IND NEP 41 MYA

Remarks: sect. Strigoathyrium, strigillosum agg.

Synonyms: hidden

= Allantodia tenella Wall.; (1828) (p. p., nom. nud.)

= Asplenium setulosum J.Sm.; Cat. Ferns Kew 5 (1856)

= Asplenium strigillosum T.Moore ex E.J.Lowe; Ferns 5, t. 36 (1858)

= Asplenium tenuifrons (T.Moore ex R.Sim) Blanf.; J. Asiat. Soc. Bengal, Pt. 2, Nat. Hist. 57: 306 (1888)

= Athyrium denticulatum J.Sm.; Hist. Fil. 328 (1875)

= Athyrium nigripes var. tenuifrons (T.Moore ex R.Sim) Bedd.; Suppl. Ferns Brit. India 33 (1892)

= Athyrium petiolosum Christ; Bull. Acad. Int. Géogr. Bot. 134 (1907), non Milde (1870)

= Athyrium proliferum T.Moore; Gard. Chron. 1866: 288 (1866), nom. illeg.

= Athyrium punethae Panigrahi; Refresher´s Course Utkal Univ. (Bubaneshwar, Orissa) Bot. (1998), nom. nov. superfl., as ´punethii´

= Athyrium radiciferum Ching; (1976), nom. nud.

= Athyrium tenuifrons T.Moore ex R.Sim; Priced Cat. Stove Greenh. Hardy Exot. Ferns 6: 17 (1859)

= Athyrium tenuifrons Wall. ex T.Moore; Index Fil. [T. Moore] 43, 188 (1857)

species :

Athyrium subrigescens (Hayata) H.Itô; Bot. Mag. (Tokyo) 52: 647 (1938)

Distribution: Japan; Taiwan

38 JAP TAI

Remarks: status disputed, often included as infraspecific taxon in A. delavayi

Synonyms: hidden

= Athyrium delavayi var. subrigescens (Hayata) Yea C.Liu, W.L.Chiou & H.Y.Liu; Fern Fl. Taiwan Athyrium 18 (2009)

= Athyrium elegans Tagawa; Acta Phytotax. Geobot. 3: 33 (1934)

= Athyrium roseum f. pubigerum (Sa.Kurata) Nakaike; Checkl. Lycophytes Ferns Japan: 19 (2013)

= Athyrium subrigescens f. pubigerum Sa.Kurata; Sci. Rep. Yokosuka City Mus. 6: 21 (1961)

= Diplazium subrigescens Hayata; Icon. Pl. Formosan. 4: 219, f. 149 (1914)

species :

Athyrium suprapuberulum Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 14 (1986)

Distribution: China (NW-Yunnan, Sichuan)

36 CHI

Synonyms: hidden

= Athyrium muliense Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 16 (1986)

species :

Athyrium tashiroi Tagawa; Acta Phytotax. Geobot. 2(1): 21 (1932)

Distribution: Japan

38 JAP

species :

Athyrium tenuipaleatum Copel.; Occas. Pap. Bernice P. Bishop Mus. 14: 60, t. 12 (1938)

Distribution: Austral Isl. (Rapa Iti)

61 AUI

species :

Athyrium tozanense (Hayata) Hayata; J. Coll. Sci. Imp. Univ. Tokyo 30: 451 (1911)

Distribution: China (Yunnan); Taiwan

36 CHI 38 TAI

Remarks: sect. Strigoathyrium, nitidulum group; separate from A. nigripes fide molecular results (Wardani et al. 2024)

Synonyms: hidden

= Asplenium tozanense Hayata; J. Coll. Sci. Imp. Univ. Tokyo 30: 440 (1911)

= Athyrium yui Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 6 (1940) (tentative synonym)

species :

Athyrium triangulare Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 20: 8 (1915)

Distribution: Sri Lanka; Java

40 SRL 42 JAW

Synonyms: hidden

= Asplenium praetermissum var. erythrorhachis (Bedd.) Sledge; Ann. Mag. Nat. Hist. XII, 9: 459 (1956) (fide Fraser-Jenkins et al. 2018)

= Athyrium gymnogrammoides var. erythrorhachis Bedd.; Suppl. Ferns S. Ind. 12 (1876) (fide Fraser-Jenkins et al. 2018)

species :

Athyrium tripinnatum Tagawa; Acta Phytotax. Geobot. 6: 163 (1937)

Distribution: Taiwan

38 TAI

Remarks: bona sp. according to Knapp (Ferns of Taiwan) and Liu et al. (2009)

species :

Athyrium uniforme Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 11 (1986)

Distribution: China (NW-Yunnan)

36 CHI

Synonyms: hidden

= Athyrium bijiangense Y.T.Hsieh & W.M.Chu; Acta Bot. Boreal.-Occid. Sin. 7(1): 52 (1987)

= Athyrium gongshanense Ching; Acta Bot. Boreal.-Occid. Sin. 6(3): 153 (1986)

species :

Athyrium venulosum Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 13 (1986)

Distribution: China (NW-Hubei)

36 CHI

species :

Athyrium vermae Fraser-Jenk., Khullar & Pangtey; Taxon. Revis. Indian Subcontinental Pteridophytes 219 (2008)

Distribution: Bhutan; India (Uttarakhand)

40 BHU IND

species :

Athyrium vidalii (Franch. & Sav.) Nakai; Bot. Mag. (Tokyo) 39: 110 (1925)

Distribution: Russian Far East (Kuril Isl.); China (Anhui, Chongqing, Fujian, Gansu, Guangxi, Guizhou, S-Henan, Hubei, Hunan, W-Jiangxi, Shaanxi, Sichuan, Yunnan, Zhejiang); South Korea (incl. Cheju Isl.); Japan; Taiwan; Vietnam

31 RFE 36 CHI 38 JAP KOR TAI 41 VIE

Remarks: not in India

Synonyms: hidden

= Asplenium vidalii Franch. & Sav.; Enum. Pl. Jap. 2: 229, 624 (1877)

= Athyrium commixtum Koidz.; Bot. Mag. (Tokyo) 39: 13 (1925)

= Athyrium glabrescens Ching; Acta Bot. Boreal.-Occid. Sin. 6(3): 156 (1987)

= Athyrium glabrum Ching; Sporae Pterid. Sin. 205, t. 41, f. 31-32 (1976), nom. nud.

= Athyrium kanghsienense Ching & Y.P.Hsu; Fl. Tsinling. 2: 215 (1974)

= Athyrium mabianense Ching & Y.T.Hsieh; Acta Bot. Boreal.-Occid. Sin. 6(3): 157 (1986)

= Athyrium neowardii Ching; Acta Bot. Boreal.-Occid. Sin. 6(3): 155 (1986)

= Athyrium regulare Koidz.; Bot. Mag. (Tokyo) 88: 111 (1924)

= Athyrium taipaishanense Ching; Fl. Tsinling. 2: 217 (1974)

= Athyrium vidalii f. yamadae (Miyabe & Kudô) Sa.Kurata; Enum. Jap. Pterid.: 281, 338 (1961)

= Athyrium vidalii var. confusum Miyabe & Kudô; Fl. Hokkaido 1: 31 (1930)

= Athyrium vidalii var. yamadae (Miyabe & Kudô) Miyabe & Tatew.; Trans. Sapporo Nat. Hist. Soc. 17: 77 (1957)

= Athyrium wugongshanense Ching & Y.T.Hsieh; Acta Bot. Boreal.-Occid. Sin. 6(3): 155 (1986)

= Athyrium yamadae Miyabe & Kudô; Trans. Sapporo Nat. Hist. Soc. 8: 61 (1921)

variant :

var. amabile (Ching) Z.R.Wang; Fl. Reipubl. Popularis Sin. [Fl. China Ed. Committee] 3(2): 199 (1999)

Distribution: China (Zhejiang)

36 CHI

Synonyms: hidden

= Athyrium amabile Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 16 (1986)

species :

Athyrium viridescentipes Sa.Kurata; J. Geobot. 13: 75 (1965)

Distribution: Japan

38 JAP

Remarks: sect. Strigoathyrium, non-setose group

Conservation Status: IUCN R

species :

Athyrium viviparum Christ; Bull. Acad. Int. Géogr. Bot. 13 (1910)

Distribution: China (Jiangxi, Hunan, Guangdong, Guangxi, Sichuan, Chongqing, Guizhou, Yunnan)

36 CHI

Remarks: sect. Strigoathyrium, strigillosum agg.

Synonyms: hidden

= Athyrium spinosissimum Ching; Acta Bot. Boreal.-Occid. Sin. 6(3): 155 (1986)

= Athyrium tianzishanense S.F.Wu & L.F.Zhang; Acta Phytotax. Sin. 28(2): 166 (1990)

= Athyrium yindeense Ching; Acta Bot. Boreal.-Occid. Sin. 6(3): 152 (1986)

species :

Athyrium wallichianum Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 497 (1938), nom. nov.

Distribution: China (Sichuan, NW-Yunnan); Tibet; Pakistan (Swat, Hazara); Nepal; ?Bhutan; India (Jammu & Kashmir, Arunachal Pradesh, Himachal Pradesh, Sikkim, Uttarakhand, West Bengal); Myanmar; Laos

36 CHI TIB 40 IND NEP PAK 41 LAO MYA

Synonyms: hidden

= Aspidium brunonianum Wall.; Numer. List. [Wallich] no. 344 (1829), nomen

= Aspidium brunonianum Wall. ex Mett.; Abh. Senckenberg. Naturf. Ges. 2(2) [Pheg. u. Asp.]: 338 (seors. 54) (1858)

= Dryopteris brunoniana (Wall. ex Mett.) Kuntze; Revis. Gen. Pl. 2: 812 (1891)

= Lastrea brunoniana (Wall. ex Mett.) C.Presl; Tent. Pterid. 76 (1836) (nomen) ex Bedd., Ferns Brit. India t. 37 (1865)

= Nephrodium brunonianum (Wall. ex Mett.) Hook.; Sp. Fil. [W. J. Hooker] 4: 113, t. 251 (1862)

species :

Athyrium wangii Ching; Bull. Fan Mem. Inst. Biol. Bot., ser. 2, 1: 279 (1949)

Distribution: China (S-Yunnan, Hainan)

36 CHI

species :

Athyrium wardii (Hook.) Makino; Bot. Mag. (Tokyo) 18: 15 (1899)

Distribution: Russian Far East (Kuril Isl.); China (Anhui, Chongqing, Fujian, Guangxi, Guizhou, Hubei, Hunan, Jiangxi, Sichuan, Yunnan, Zhejiang); South Korea (Cheju Isl.); Japan

31 RFE 36 CHI 38 JAP KOR

Remarks: sect. Strigoathyrium, non-setose group

Synonyms: hidden

= Asplenium wardii Hook.; Sp. Fil. [W. J. Hooker] 3: 189 (1860)

= Athyrium tsussimense Koidz.; Fl. Symb. Orient.-Asiat. 41 (1930)

= Athyrium wardii f. inadae (Tagawa) Ebihara; Bull. Nat. Mus. Nat. Sci., Ser. B, 43(1): 20 (2017)

= Athyrium wardii var. inadae Tagawa; Acta Phytotax. Geobot. 16: 178 (1956)

variant :

var. densipinnum Z.R.Wang & Li Bing Zhang; Acta Bot. Yunnan. 17(3): 303 (1995)

Distribution: China (Sichuan)

36 CHI

variant :

var. glabratum Y.T.Hsieh & Z.R.Wang; Fl. Reipubl. Popularis Sin. [Fl. China Ed. Committee] 3(2): 508 (1999)

Distribution: China (Zhejiang, Fujian, Hunan)

36 CHI

species :

Athyrium xichouense Y.T.Hsieh & Z.R.Wang; Acta Phytotax. Sin. 27(2): 155 (1989)

Distribution: China (Sichuan, Yunnan); Tibet; Nepal; NE-India (Sikkim, Arunachal Pradesh, Meghalaya, Meghalaya, Mizoram, West Bengal)

36 CHI TIB 40 IND NEP

Remarks: kept separate (again) from A. silvicola by Fraser-Jenkins (2018)

species :

Athyrium yakusimense Tagawa; Acta Phytotax. Geobot. 6: 94 (1937)

Distribution: Japan (Yakushima Isl.)

38 JAP NNS

Conservation Status: CR in Japan

species :

Athyrium yokoscense (Franch. & Sav.) Christ; Bull. Herb. Boissier, sér. 1, 4: 668 (1896)

Distribution: Russian Far East (Kuril Isl., Primorye); China (Anhui, Chongqing, NE-Guizhou, Heilongjiang, Henan, Hunan, Jiangsu, Jiangxi, Jilin, Liaoning, Shandong, Zhejiang); North Korea; South Korea (incl. Cheju Isl.); Japan

31 RFE 36 CHI 38 JAP KOR NKO

Synonyms: hidden

= Aspidium subspinulosum Christ; Bull. Herb. Boissier, sér. 2, 2: 829 (1902)

= Asplenium yokoscense Franch. & Sav.; Enum. Pl. Jap. 2: 225, 622 (1877)

= Athyrium demissum Christ; Repert. Spec. Nov. Regni Veg. 6: 284 (1908)

= Athyrium flaccidum Christ; Repert. Spec. Nov. Regni Veg. 6: 11 (1908)

= Athyrium pachysorum Christ; Notul. Syst. (Paris) 1: 48 (1909)

= Athyrium subspinulosum (Christ) Ching ex Vorosch.

= Dryopteris subspinulosa (Christ) C.Chr.; Index Filic. 296 (1905)

variant :

var. kirisimaense (Tagawa) Li & J.Z.Wang; Clav. Pl. Chin. Bor.-Or., ed. 2 (ed. P. Y. Fu): 36 (1995)

Distribution: ?NE-China; Japan

38 JAP

Synonyms: hidden

= Athyrium kirisimaense Tagawa; Acta Phytotax. Geobot. 2: 22 (1933)

species :

Athyrium yuanyangense Y.T.Hsieh & W.M.Chu; Acta Bot. Boreal.-Occid. Sin. 7(1): 54 (1987)

Distribution: China (S-Yunnan)

36 CHI

species :

Athyrium zayuense Z.R.Wang; Acta Phytotax. Sin. 32(1): 84 (1994)

Distribution: Tibet (Zayu)

36 TIB

Synonyms: hidden

= Athyrium chayuense Z.R.Wang; Vasc. Pl. Hengduan Mount. 1: 84 (1993) (without latin descr. or type)

species :

Athyrium zhenfengense Ching; Acta Bot. Boreal.-Occid. Sin. 6(3): 151 (1986)

Distribution: China (SW-Guizhou)

36 CHI

species :

Athyrium ×akiense Sa.Kurata; J. Geobot. 15: 5 (1966)

Distribution: Japan

38 JAP

Remarks: Athyrium eremicola x otophorum

species :

Athyrium ×amagi-pedis Sa.Kurata; Sci. Rep. Yokosuka City Mus. 6: 23 (1961)

Distribution: Japan

38 JAP

Remarks: Athyrium otophorum x wardii

species :

Athyrium ×anceps Sa.Kurata; J. Geobot. 13: 76 (1965)

Distribution: Japan

38 JAP

Remarks: Athyrium arisanense x kenzo-satakei

species :

Athyrium ×austrojaponense Sa.Kurata; J. Geobot. 16: 99 (1968)

Distribution: Japan

38 JAP

Remarks: Athyrium kuratae x otophorum

species :

Athyrium ×awatae Seriz.; J. Phytogeogr. Taxon. 28(1): 34 (1980)

Distribution: Japan

38 JAP

Remarks: Athyrium oblitescens x otophorum

species :

Athyrium ×bicolor Seriz.; J. Jap. Bot. 55(8): 252 (1980)

Distribution: Japan

38 JAP

Remarks: Athyrium neglectum x vidalii

species :

Athyrium ×boreo-occidentali-indobharaticola-birianum Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 220 (2008)

Distribution: India (Uttarakhand)

40 IND

Remarks: Athyrium pectinatum x schimperi ssp. biserrulatum

species :

Athyrium ×calophyllum Sa.Kurata ex Seriz.; J. Jap. Bot. 55(8): 254 (1980)

Distribution: Japan

38 JAP

Remarks: Athyrium palustre x vidalii

species :

Athyrium ×decurrentiserratum K.Hori; Fern Gaz. 21(6): 262 (2021)

Distribution: Japan (Honshu)

38 JAP

species :

Athyrium ×dixitii Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 225 (2008)

Distribution: India (Uttarakhand)

40 IND

Remarks: Athyrium anisopterum (kumaonicum) x fimbriatum

species :

Athyrium ×drepanopteroides Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 214 (2008)

Distribution: Nepal; India (Meghalaya, West Bengal)

40 IND NEP

Remarks: Athyrium drepanopterum x eburneum (woodsioides)

species :

Athyrium ×flavosorum Seriz.; J. Phytogeogr. Taxon. 33(2): 77 (1985)

Distribution: Japan

38 JAP

Remarks: Athyrium arisanense x delavayi

species :

Athyrium ×fuscopaleaceum Sa.Kurata; Sci. Rep. Yokosuka City Mus. 6: 24 (1961)

Distribution: Japan

38 JAP

Remarks: Athyrium kuratae x clivicola

species :

Athyrium ×gurungae Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 229 (2008)

Distribution: Nepal

40 NEP

Remarks: Athyrium pectinatum x foliolosum

species :

Athyrium ×hisatsuanum Sa.Kurata; J. Geobot. 15: 6 (1966)

Distribution: Japan

38 JAP

Remarks: Athyrium clivicola x iseanum

species :

Athyrium ×ikutae Sa.Kurata; J. Geobot. 16: 33 (1968)

Distribution: Japan

38 JAP

Remarks: Athyrium tozanense x yakusimense

species :

Athyrium ×inabaense Seriz.; J. Phytogeogr. Taxon. 28(1): 34 (1980)

Distribution: Japan

38 JAP

Remarks: Athyrium iseanum x oblitescens

species :

Athyrium ×inouei Sa.Kurata; J. Geobot. 17: 89 (1969)

Distribution: Japan

38 JAP

Remarks: Athyrium iseanum x yokoscense

species :

Athyrium ×kathmanduense Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 211 (2008)

Distribution: China; Nepal; India (Darjeeling, Arunachal Pradesh, West Bengal, Meghalaya)

36 CHI 40 IND NEP

Remarks: Athyrium foliolosum x drepanopterum

Synonyms: hidden

= Athyrium darjeelingensis Fraser-Jenk. ex Subh.Chandra; Ferns India (Enum., Syn. & Dist.) 125 (2000) nom. nud.

species :

Athyrium ×kawabatae Sa.Kurata; Sci. Rep. Yokosuka City Mus. 6: 22 (1961)

Distribution: Japan

38 JAP

Remarks: Athyrium iseanum x subrigescens

species :

Athyrium ×kawabataeoides Sa.Kurata; J. Geobot. 16: 33 (1968)

Distribution: Japan

38 JAP

Remarks: Athyrium subrigescens x tozanense

species :

Athyrium ×kidoanum Sa.Kurata; J. Geobot. 10: 10 (1961)

Distribution: Japan

38 JAP

Remarks: Athyrium iseanum x wardii

species :

Athyrium ×langtangense Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 222 (2008), nom. inval., sine herbarium, et Annot. Checkl. Ind. Pterid. 2: 60 (2018)

Distribution: Nepal

40 NEP

Remarks: Athyrium anisopterum x micropterum

species :

Athyrium ×lobulosoimpolitum Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 212 (2008)

Distribution: Nepal; Bhutan; India (Sikkim, Meghalaya, West Bengal)

40 BHU IND NEP

Remarks: Athyrium eburneum (woodsioides) x foliolosum

species :

Athyrium ×manickamii Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 226 (2008)

Distribution: India (West Bengal)

40 IND

Remarks: Athyrium foliolosum x xichouense

species :

Athyrium ×mantoniae Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 231 (2008)

Distribution: Sri Lanka

40 SRL

Remarks: Athyrium praetermissum x solenopteris

species :

Athyrium ×masayukianum Sa.Kurata; J. Geobot. 13: 76 (1965)

Distribution: Japan

38 JAP

Remarks: Athyrium kuratae x iseanum

species :

Athyrium ×megayakusimense Sa.Kurata; J. Geobot. 14: 5 (1965)

Distribution: Japan

38 JAP

Remarks: Athyrium kuratae x yakusimense

species :

Athyrium ×meghalaicum Fraser-Jenk.; Annot. Checkl. Ind. Pterid. 2: 60 (2018)

Distribution: India (Meghalaya)

40 IND

Remarks: Athyrium foliolosum x puncticaule

species :

Athyrium ×mentiens Sa.Kurata; J. Geobot. 16: 34 (1968)

Distribution: Japan

38 JAP

Remarks: Athyrium vidalii x yokoscense

species :

Athyrium ×minakuchii Sa.Kurata; J. Geobot. 14: 85 (1966)

Distribution: Japan

38 JAP

Remarks: Athyrium otophorum x yokoscense

species :

Athyrium ×multifidum Rosenst.; Repert. Spec. Nov. Regni Veg. 13: 125 (1914)

Distribution: China

36 CHI

Remarks: Athyrium deltoidofrons x vidalii

Synonyms: hidden

= Athyrium deltoidofrons var. latisectum (Rosenst.) Koidz.; Bot. Mag. (Tokyo) 38(449): 109 (1924)

= Athyrium deltoidofrons var. multifidum (Rosenst.) Koidz.; Bot. Mag. (Tokyo) 38(449): 109 (1924)

= Athyrium ×multifidum f. sakuraii (Rosenst.) Sa.Kurata; J. Geobot. 9: 100 (1961)

= Athyrium ×multifidum var. latisectum Rosenst.; Repert. Spec. Nov. Regni Veg. 13: 126 (1914)

= Athyrium ×multifidum var. sakuraii Rosenst.; Repert. Spec. Nov. Regni Veg. 13: 126 (1914)

species :

Athyrium ×neoelegans Sa.Kurata; J. Geobot. 14: 4 (1965)

Distribution: Japan

38 JAP

Remarks: Athyrium subrigescens x yakusimense

species :

Athyrium ×nepalense Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 212 (2008)

Distribution: Nepal; India (West Bengal)

40 IND NEP

Remarks: Athyrium pectinatum x drepanopterum

species :

Athyrium ×paludicola Sa.Kurata ex Seriz.; J. Jap. Bot. 55: 254 (1980)

Distribution: Japan

38 JAP

Remarks: Athyrium deltoidofrons x palustre

species :

Athyrium ×pichisermollianum Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 224 (2008)

Distribution: India (Uttarakhand)

40 IND

Remarks: Athyrium anisopterum (kumaonicum) x foliolosum

Synonyms: hidden

= Athyrium ×pangteyi Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes: 222 (2008)

species :

Athyrium ×pokharense Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 214 (2008)

Distribution: Nepal

40 NEP

Remarks: Athyrium pectinatum x eburneum (woodsioides)

species :

Athyrium ×pseudobipinnatum K.Hori; Fern Gaz. 21(6): 264 (2021)

Distribution: Japan (Honshu, Kyushu, Shikoku)

38 JAP

species :

Athyrium ×pseudoiseanum Sa.Kurata; J. Geobot. 8: 84 (1960)

Distribution: Japan (Kyushu)

38 JAP

Remarks: Athyrium iseanum x otophorum

species :

Athyrium ×pseudopinetorum Seriz.; J. Jap. Bot. 55(8): 253 (1980)

Distribution: Japan

38 JAP

Remarks: Athyrium pinetorum x vidalii

species :

Athyrium ×pseudospinescens Seriz.; J. Phytogeogr. Taxon. 28(1): 35 (1980)

Distribution: Japan

38 JAP

Remarks: Athyrium iseanum x vidalii

species :

Athyrium ×pseudowardii Seriz.; J. Phytogeogr. Taxon. 28(1): 35 (1980)

Distribution: Japan

38 JAP

Remarks: Athyrium vidalii x wardii

species :

Athyrium ×purpurascens (Tagawa) Sa.Kurata; J. Geobot. 14: 4 (1965)

Distribution: Japan

38 JAP

Remarks: Athyrium kuratae (arisanense) x subrigescens

Synonyms: hidden

= Athyrium elegans var. purpurascens Tagawa; Acta Phytotax. Geobot. 3: 33 (1934)

species :

Athyrium ×purpureipes Sa.Kurata; Sci. Rep. Yokosuka City Mus. 6: 23 (1961)

Distribution: Japan

38 JAP

Remarks: Athyrium clivicola x otophorum

species :

Athyrium ×pygmaei-silvae Sa.Kurata; Sci. Rep. Yokosuka City Mus. 6: 20 (1961)

Distribution: Japan

38 JAP

Remarks: Athyrium kuratae x tozanense

species :

Athyrium ×quaesitum Sa.Kurata; J. Geobot. 15: 5 (1966)

Distribution: Japan

38 JAP

Remarks: Athyrium otophorum x vidalii

species :

Athyrium ×sanjappae Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 227 (2008)

Distribution: India (West Bengal)

40 IND

Remarks: Athyrium distans x foliolosum

species :

Athyrium ×satsumense Seriz.; J. Phytogeogr. Taxon. 28(1): 34 (1980)

Distribution: Japan

38 JAP

Remarks: Athyrium kuratae x viridescentipes

species :

Athyrium ×sefuricola Sa.Kurata; J. Geobot. 15: 6 (1966)

Distribution: Japan

38 JAP

Remarks: Athyrium kuratae (arisanense) x wardii

species :

Athyrium ×shiibaense (Seriz.) Nakaike; New Fl. Japan, Pterid. 839 (1992)

Distribution: Japan

38 JAP

Remarks: Athyrium neglectum ssp. australe x vidalii

Synonyms: hidden

= Athyrium ×bicolor nothosubsp. shiibaense Seriz.; J. Phytogeogr. Taxon. 33(2): 77 (1985)

species :

Athyrium ×shikoku-montanum Seriz.; J. Jap. Bot. 55(8): 255 (1980)

Distribution: Japan

38 JAP

Remarks: Athyrium reflexipinnum x vidalii

species :

Athyrium ×simlense Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 231 (2008)

Distribution: India (Himachal Pradesh)

40 IND

Remarks: Athyrium setiferum x strigillosum

species :

Athyrium ×sledgei Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 230 (2008)

Distribution: Sri Lanka

40 SRL

Remarks: Athyrium puncticaule x praetermissum

species :

Athyrium ×subchristensenianum K.Hori; Fern Gaz. 21(6): 265 (2021)

Distribution: Japan (Honshu, Kyushu)

38 JAP

species :

Athyrium ×subcrassipes Seriz.; J. Phytogeogr. Taxon. 33(2): 77 (1985)

Distribution: Japan

38 JAP

Remarks: Athyrium masamunei x tozanense (nigripes)

species :

Athyrium ×tokashikii Sa.Kurata; J. Geobot. 12: 15 (1963)

Distribution: Japan

38 JAP

Remarks: Athyrium clivicola x wardii

species :

Athyrium ×tsurutanum Sa.Kurata; J. Geobot. 13: 77 (1965)

Distribution: Japan

38 JAP

Remarks: Athyrium kenzo-satakei x otophorum

species :

Athyrium ×watanabei Seriz.; J. Jap. Bot. 55(8): 252 (1980)

Distribution: Japan

38 JAP

Remarks: Athyrium deltoidofrons f. ohmurae x neglectum

species :

Athyrium ×yakuinsulare Sa.Kurata; J. Geobot. 13: 77 (1965)

Distribution: Japan

38 JAP

Remarks: Athyrium kenzo-satakei x yakusimense

species :

Athyrium ×yakumonticola Sa.Kurata; J. Geobot. 12: 15 (1963)

Distribution: Japan

38 JAP

Remarks: Athyrium iseanum x tozanense

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A complete, synonymic checklist of the Higher Plants of the World

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A complete, synonymic checklist of the Higher Plants of the World

Version 24.11 (November 5th, 2024)

Covering 355,576 accepted species,
8,715 hybrids, 47,863 accepted infraspecific taxa and 1,000,581 synonyms,
in total 1,412,785* names

[about 25,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus*" are deliberately omitted]

With complete, detailed distribution data (countries and regions) and literature references

Arranged in linear phylogenetic sequence reflecting true relationships

Fully searchable and continously updated

With 38,209 pictures of 15,787 species

Assembled through 40 years independently from other databases

In cooperation with Catalogue of Life

Contact

How to Cite:

Search

+ Family

Filter

Order

Family (28)

Genus (9)

Display options

A complete, synonymic checklist of the Higher Plants of the World

Version 24.11 (November 5th, 2024)

Covering 355,576 accepted species, 8,715 hybrids, 47,863 accepted infraspecific taxa and 1,000,581 synonyms, in total 1,412,785* names

[*about 25,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus" are deliberately omitted]

With complete, detailed distribution data (countries and regions) and literature references

Arranged in linear phylogenetic sequence reflecting true relationships

Fully searchable and continously updated

With 38,209 pictures of 15,787 species

Assembled through 40 years independently from other databases

In cooperation with Catalogue of Life

Covering 355,576 accepted species,
8,715 hybrids, 47,863 accepted infraspecific taxa and 1,000,581 synonyms,
in total 1,412,785* names

[about 25,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus*" are deliberately omitted]