046.0000
Family
Athyriaceae Alston; Taxon 5: 25 (1956)
6-7 genera, ca. 760 spp. Circumscription following Rothfels et al. (2012). According to Fraser-Jenkins et al. (2018) it should be included in Woodsiaceae s. lat.
Species listed in database: 758; hybrids: 102
= Athyriinae Milde; Bot. Zeitung (Berlin) 28: 332 (1870)
= Athyrioideae B.K.Nayar; Taxon 19: 235 (1970)
= Physematieae J.Sm.; (1875)
= Deparioideae M.L.Wang & Y.T.Hsieh; Acta Phytotax. Sin. 42: 526 (2004)
= Diplazioideae Shmakov; Turczaninowia 4(1-2): 59 (2001), nom. superfl.
= Diplazioideae Ching & Z.R.Wang; Acta Phytotax. Sin. 35: 35 (1997)
= Diplazieae Pfeiff.; Nomencl. Bot. [Pfeiffer] 1(1): 628 (1873)
= Diplaziinae Payer; Bot. Crypt.: 198 (1850)
046.0008
Genus
Athyrium Roth; Tent. Fl. Germ. 3: 58 (1790)
Distribution: Worldwide, mostly northern hemisphere, many in China (over 100), several in trop. Africa and Madagascar, few in the New World.
Species listed in database: 179; hybrids: 60
= Hypochlamys Fée; Mém. Foug. 5 (Gen. Filic.): 200 (1852)
= Pseudocystopteris Ching; Acta Phytotax. Sin. 9: 76 (1964)
= Solenopteris Zenker ms. ex Kunze; Linnaea 24: 267 (1851)
= Neoathyrium Ching & Z.R.Wang; Acta Phytotax. Sin. 20(1): 76 (1982)
Search for: Athyrium drepanopterum
Nothing found!
species :
Athyrium aberrans
Liang Zhang & Li Bing Zhang; Phytotaxa 533(3): 167 (2022)
Distribution: SE-Tibet (Medog)
36 TIB
species :
Athyrium acutipinnulum
Kodama ex Nakai; Rep. Veg. Ooryongto 13 (1919)
Distribution: South Korea
38 KOR
Remarks: provisionally accepted name, status unconfirmed
species :
Athyrium adpressum
Ching & W.M.Chu; Acta Bot. Boreal.-Occid. Sin. 6(1): 13 (1986)
Distribution: China (S-Yunnan)
36 CHI
species :
Athyrium ammifolium
(Mett.) C.Chr.; Index Filic. 139 (1905)
Distribution: Nigeria; Cameroon; Bioko Isl. [Fernando Poo]; Equatorial Guinea (Rio Muni)
22 NGA 23 BIO CMN EQG
Synonyms: hidden
= Asplenium ammifolium Mett.; Fil. Afr. [Kuhn] 96 (1868)
species :
Athyrium amoenum
C.Chr.; Gard. Bull. Straits Settlem. 7: 267, t. 56 (1934)
Distribution: Borneo (Sabah)
42 BOR
Remarks: Flora of China treats this as synonym of A. rhachidosorum, but the areas are widely separated, so this is quite unlikely
species :
Athyrium angustum
(Willd.) C.Presl; Reliq. Haenk. 1: 39 (1825)
Name: Subarctic ladyfern (EN)
Distribution: Greenland; Canada (Labrador, Manitoba, New Brunswick, Newfoundland, Nova Scotia, Ontario, Prince Edward Isl., Québec, Saskatchewan); St. Pierre et Miquelon; USA (Colorado, Connecticut, District of Columbia, Delaware, Iowa, Illinois, Indiana, Kentucky, Massachusetts, Maryland, Maine, Michigan, Minnesota, Missouri, North Carolina, North Dakota, New Hampshire, New Jersey, New York, Ohio, Pennsylvania, Rhode Island, South Dakota, Tennessee, Virginia, Vermont, Wisconsin, West Virginia)
70 GNL 71 CDN 72 CDN SPM 73 USA 74 USA 75 USA 78 USA
Remarks: often treated within A. filix-femina or as variety of A. asplenioides, but increasingly elevated to full species status
Synonyms: hidden
= Aspidium angustum Willd.; Sp. Pl., ed. 4 [Willdenow] 5(1): 277 (1810)
= Asplenium elatius Link; Fil. Spec. 94 (1841)
= Asplenium filix-femina var. angustum (Willd.) D.C.Eaton; Rep. U.S. Geogr. Surv., Wheeler 6(Botany): 330 (1878)
= Asplenium filix-femina var. exile D.C.Eaton; Rep. U.S. Geogr. Surv., Wheeler 6(Botany): 330 (1878)
= Asplenium michauxii Spreng.; Syst. Veg., ed. 16 [Sprengel] 4(1): 88 (1827)
= Athyrium angustum f. confertum Butters; Rhodora 19: 195 (1917)
= Athyrium angustum f. cristatum (Hopkins) Broun ex Weath.; Amer. Fern J. 26: 133 (1936)
= Athyrium angustum f. elegans (Gilbert) Butters; Rhodora 19: 196 (1917)
= Athyrium angustum f. laciniatum Butters; Rhodora 19: 195 (1917)
= Athyrium angustum var. boreale Jenn.; Amer. Fern J. 8: 82 (1918)
= Athyrium angustum var. cristatum Hopkins; Amer. Fern J. 9: 88 (1919)
= Athyrium angustum var. elatius (Link) Butters; Rhodora 19: 191 (1917)
= Athyrium angustum var. glanduliferum Jenn.; Amer. Fern J. 8: 82 (1918)
= Athyrium angustum var. laurentianum Butters; Rhodora 19: 194 (1917)
= Athyrium angustum var. rubellum (Gilbert) Butters; Rhodora 19: 193 (1917)
= Athyrium asplenioides var. angustum (Willd.) T.Moore; Index Fil. [T. Moore] 179 (1860)
= Athyrium filix-femina f. confertum (Butters) Fernald; Rhodora 48(no. 574): 389 (1946)
= Athyrium filix-femina f. cristatum (Hopkins) Clute; Our Ferns: 224, 384 (1938)
= Athyrium filix-femina f. elatius (Link) Clute; Ferns in Their Haunts 224 (1938)
= Athyrium filix-femina f. elegans (Gilbert) Clute; Ferns in Their Haunts 224 (1938)
= Athyrium filix-femina f. laciniatum (Butters) Fernald; Rhodora 48(no. 574): 389 (1946)
= Athyrium filix-femina f. laurentianum (Butters) Fernald; Rhodora 48(no. 574): 389 (1946)
= Athyrium filix-femina f. plano-rhaeticum Gilbert; List N. Amer. Pterid.: 15, 34 (1901)
= Athyrium filix-femina subsp. angustum (Willd.) Hultén; Kongl. Svenska Vetensk. Acad. Handl. 8(5): 178 (1962), isonym
= Athyrium filix-femina subsp. angustum (Willd.) R.T.Clausen; Cornell Univ. Agric. Exp. Sta. Mem. 291: 7 (1949)
= Athyrium filix-femina subvar. elatius (Link) Farw.; Amer. Midl. Naturalist 10: 203 (1927)
= Athyrium filix-femina subvar. elegans (Gilbert) Farw.; Amer. Midl. Naturalist 10: 203 (1927)
= Athyrium filix-femina var. angustum (Willd.) G.Lawson; Edinburgh New Philos. J., ser. 2, 19: 114 (1864)
= Athyrium filix-femina var. elegans Gilbert; List N. Amer. Pterid.: 33-34 (1901)
= Athyrium filix-femina var. michauxii (Spreng.) Farw.; Rep. (Annual) Michigan Acad. Sci. 18: 79 (1916)
= Athyrium filix-femina var. rubellum Gilbert; List N. Amer. Pterid.: 35 (1901)
= Athyrium michauxii (Spreng.) Fée; Mém. Foug. 5 (Gen. Filic.): 186 (1852)
= Athyrium michauxii f. furcata Hahne; Fern Bull. 12: 116 (1904)
species :
Athyrium anisopterum
Christ; Bull. Herb. Boissier, sér. 1, 6: 962 (1898)
Distribution: China (Guangdong, Guangxi, Guizhou, Hunan, Jiangxi, Sichuan, Yunnan); Tibet; Taiwan; Nepal; Bhutan; India (Arunachal Pradesh, Himachal Pradesh, ?Karnataka, Kerala, Meghalaya, Sikkim, Tamil Nadu, Tripura, Uttarakhand, West Bengal); Sri Lanka; Myanmar (Chin, etc.); N-Thailand; Vietnam; peninsular Malaysia (Cameron Highlands, Pahang); Sumatra; Borneo (Mt. Kinabalu, Sabah); Philippines (Luzon); Java
36 CHI TIB 38 TAI 40 BHU IND NEP SRL 41 MYA THA VIE 42 BOR JAW MLY PHI SUM
Remarks: sect. Polystichoides, anisopterum clade. Not in Nepal
Synonyms: hidden
= Aspidium fauriei var. elatius Christ; Bull. Herb. Boissier, sér. 1, 6(3): 193 (1898)
= Athyrium kumaonicum Holttum ex Punetha; Indian Fern J. 2(1-2): 29 (1985) [fide Fraser-Jenkins et al. 2018]
= Athyrium membranifolium Ching; (1976), nom. nud.
= Athyrium woodsioides Christ; Bull. Acad. Int. Géogr., sér. 3, 16: 124 (1906), nom. illeg. superfl.
= Athyrium ×heterosporum Y.T.Hsieh & Z.R.Wang; Acta Phytotax. Sin. 27(2): 156 (1989) (fide Fraser-Jenkins 1997, 2018)
= Dryopteris thysanocarpa Hayata; Icon. Pl. Formosan. 4: 160 (1914), pro syn.
= Lastrea membranacea Arn. ex Baker; Syn. Fil. [W. J. Hooker & Baker], ed. 2: 498 (1874) (nom. inval.)
= Lastrea sparsa var. membranacea Arn. ex Bedd.; Suppl. Ferns S. Ind. 17 (1876) (nom. inval. in syn.)
= Nephrodium sparsum var. membranaceum Arn. ex C.B.Clarke; (1880) (nom. inval.)
species :
Athyrium annae
Kornas; Kew Bull. 33(1): 99 (1978)
Distribution: Zambia
26 ZAM
Remarks: status doubtful, probably a synonym of A. lewalleanum or not belonging to Athyrium at all (Verdcourt in Fl. Trop. East Africa)
species :
Athyrium appendiculiferum
Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 16: 3 (1914)
Distribution: peninsular Malaysia (Cameron Highlands, Pahang); Sumatra; Java
42 JAW MLY SUM
species :
Athyrium araiostegioides
Ching; Acta Bot. Boreal.-Occid. Sin. 6(2): 103 (1986)
Distribution: China (Yunnan, SW-Sichuan)
36 CHI
Remarks: sect. Strigoathyrium
species :
Athyrium archboldii
Copel.; Univ. Calif. Publ. Bot. 18: 222 (1942)
Distribution: New Guinea (Irian Jaya)
42 NWG
species :
Athyrium arcuatum
Liebm.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 5, 1: 278 (seors. 126) (1849)
Distribution: Mexico (Chiapas, Colima, Ciudad de Mexico, Guanajuato, Guerrero, Hidalgo, Jalisco, Mexico State, Michoacan, Morelos, Nayarit, Oaxaca, Puebla, Queretaro, Veracruz); Guatemala; Honduras
79 MEX 80 GUA HON
Remarks: included by Fl. Mesoamericana in A. filix-femina, kept separate by subsequent authors
Synonyms: hidden
= Asplenium arcuatum (Liebm.) Hook.; Sp. Fil. [W. J. Hooker] 3: 219 (1860)
= Asplenium martensii Kunze; Amer. J. Sci. Arts, ser. 2, 6: 86 (1848), nom. nud.
= Asplenium martensii Kunze ex Mett.; Abh. Senckenberg. Naturf. Ges. 3(1) [Aspl.]: 244 (1859)
= Athyrium barnebyanum Mickel & Beitel; Mem. New York Bot. Gard. 46 [Pterid. Fl. Oaxaca]: 77 (1988)
= Athyrium galeottii Fée; Mém. Foug. 5 (Gen. Filic.): 187 (1852)
= Athyrium martensii (Kunze ex Mett.) T.Moore; Index Fil. [T. Moore] 186 (1860)
= Athyrium pumilio Christ; Bull. Herb. Boissier, sér. 2, 8: 147 (1903)
species :
Athyrium arisanense
(Hayata) Tagawa; Acta Phytotax. Geobot. 2(3): 195 (1933)
Distribution: ?Japan; Taiwan
38 TAI
Remarks: sect. Strigoathyrium, arisanense agg.
Synonyms: hidden
= Diplazium arisanense Hayata; Icon. Pl. Formosan. 4: 212, f. 144 (1914)
species :
Athyrium asplenioides
(Michx.) Desv.; Mem. Soc. Linn. Paris 6(2): 264 (1827)
Name: Asplenium ladyfern
Distribution: USA (Alabama, Arkansas, Connecticut, District of Columbia, Delaware, Florida, Georgia, Illinois, Indiana, Kansas, Kentucky, Louisiana, Massachusetts, Maryland, Missouri, Mississippi, North Carolina, New Jersey, New York, Ohio, Oklahoma, Pennsylvania, Rhode Island, South Carolina, Tennessee, Texas, Virginia, West Virginia)
74 USA 75 USA 77 USA 78 USA
Remarks: separate from A. filix-femina fide Fraser-Jenkins (2008)
Synonyms: hidden
= Aspidium asplenioides (Michx.) Sw.; Syn. Fil. [Swartz] 60 (1806)
= Asplenium athyrium Spreng.; Anleit. Kenntn. Gew. 3: 106 (1804)
= Athyrium asplenioides f. ellipticum Wherry; Amer. Fern J. 38: 158 (1949)
= Athyrium asplenioides f. subtripinnatum Butters; Rhodora 19: 190 (1917)
= Athyrium asplenioides var. subtripinnatum (Butters) Wherry; Bartonia 21: 14 (1942)
= Athyrium filix-femina f. ellipticum (Wherry) C.F.Reed; Castanea 16: 71 (1951)
= Athyrium filix-femina f. rectangulare Gilbert; List N. Amer. Pterid.: 15, 35 (1901)
= Athyrium filix-femina f. subtripinnatum (Butters) Fernald; Rhodora 48(no. 574): 389 (1946)
= Athyrium filix-femina subsp. asplenioides (Michx.) Hultén; Kongl. Svenska Vetensk. Acad. Handl. 8(5): 178 (1962)
= Athyrium filix-femina var. asplenioides (Michx.) Farw.; Pap. Michigan Acad. Sci. 2: 13 (1923)
= Nephrodium asplenioides Michx.; Fl. Bor.-Amer. (Michaux) 2: 268 (1803)
species :
Athyrium atkinsonii
Bedd.; Ferns Brit. India Suppl. 11, t. 369 (1876)
Distribution: China (Chongqing, Fujian, Gansu, Guizhou, Henan, Hubei, Hunan, Jiangxi, S-Shaanxi, Shanxi, Sichuan, Yunnan); Tibet; Japan; Taiwan; Pakistan (Swat); Nepal; Bhutan; India (Jammu & Kashmir, Arunachal Pradesh, Himachal Pradesh, Sikkim, Uttarakhand, West Bengal); Myanmar (Chin, etc.); Vietnam
36 CHI TIB 38 JAP TAI 40 BHU IND NEP PAK 41 MYA VIE
Synonyms: hidden
= Aspidium lastreoides Baker; J. Bot. 26(7): 227 (1888)
= Aspidium senanense Franch. & Sav.; Enum. Pl. Jap. 2: 241, 636 (1877)
= Asplenium andersonii C.B.Clarke; J. Linn. Soc., Bot. 16: 154 (1876)
= Asplenium atkinsonii (Bedd.) C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 487 (1880)
= Asplenium atkinsonii var. andersonii (C.B.Clarke) C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 487 (1880)
= Asplenium fimbriatum var. sphaeropteroides C.B.Clarke; (1880)
= Asplenium lastreoides Baker; J. Bot. 26: 227 (1888)
= Asplenium microsorum Makino; Bot. Mag. (Tokyo) 6(60): 46 (1892)
= Athyrium andersonii (C.B.Clarke) Panigrahi & Basu; Proc. Ind. Sci. Congr., Lucknow, Bot., Abstr. 72: 14 (1985) (fide Fraser-Jenkins 1997)
= Athyrium atkinsonii var. andersonii (C.B.Clarke) C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 494 (1880)
= Athyrium filix-femina var. deltoideum Christ; Mém. Soc. Bot. France 1 [Hist. Acrostich.]: 46 (1905), nom. illeg.
= Athyrium fimbriatum var. sphaeropteroides (C.B.Clarke) Bedd.; Trans. Linn. Soc. London, Bot., ser. 2, 1: 495 (1883)
= Athyrium lastreoides (Baker) Diels; Nat. Pflanzenfam. [Engler & Prantl] I, 4: 224 (1899)
= Athyrium microsorum Makino; Bot. Mag. (Tokyo) 18: 25 (1899)
= Athyrium monticola Rosenst.; Repert. Spec. Nov. Regni Veg. 13: 123 (1913)
= Athyrium senanense (Franch. & Sav.) Koidz. & Tagawa; Tagawa, Acta Phytotax. Geobot. 3: 34 (1934)
= Cystopteris grandis C.Chr.; H. Lév., Cat. Pl. Yun-Nan 100 (1916)
= Davallia athyriifolia Baker; Ann. Bot. (Oxford) 6: 200 (1891)
= Dryopteris gracilifrons C.Chr.; Index Filic. 268 (1906) (nom. nov. for Polypodium leptophyllum Baker)
= Dryopteris senanensis (Franch. & Sav) C.Chr.; Index Filic. 291 (1905)
= Polypodium leptophyllum Baker; Ann. Bot. (Oxford) 5: 458 (1891) (non L. (1753) nec Moritz ex Mett. (1864))
= Pseudocystopteris andersonii (C.B.Clarke) Ching; Acta Phytotax. Sin. 9: 79 (1964) (fide Fraser-Jenkins 1997)
= Pseudocystopteris atkinsonii (Bedd.) Ching; Acta Phytotax. Sin. 9(1): 78 (1964)
species :
Athyrium atratum
Bedd.; Ferns Brit. India Suppl. 33 (1892)
Distribution: India (Arunachal Pradesh, Manipur); Vietnam; Sumatra; Java
40 IND 41 VIE 42 JAW SUM
Remarks: sect. Strigoathyrium, non-setose group
Conservation Status: IUCN EN for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Asplenium gedeanum Racib.; Pteridoph. Buitenzorg 223 (1898)
= Athyrium gedeanum (Racib.) Christ; J. Bot. (Morot) 19: 68 (1905) [fide Fraser-Jenkins et al. 2018]
species :
Athyrium attenuatum
(C.B.Clarke) Tagawa; Acta Phytotax. Geobot. 16: 177 (1956)
Distribution: E-Afghanistan; China (SE-Gansu, Guizhou, W-Sichuan, N-Yunnan); Tibet; Pakistan (Azad Kashmir, Kurram, Swat, Hazara); Nepal; N-India (Jammu & Kashmir, ?Arunachal Pradesh, Himachal Pradesh, Sikkim, Uttarakhand, West Bengal); Myanmar
34 AFG 36 CHI TIB 40 IND NEP PAK 41 MYA
Remarks: A. dentigerum included here by Fraser-Jenkins et al. (2015 etc.) but kept separate by Flora of China
Synonyms: hidden
= Asplenium filix-femina f. dentigerum (Wall. ex C.B.Clarke) C.Hope; J. Bombay Nat. Hist. Soc. 14: 125 (1902)
= Asplenium filix-femina var. attenuatum C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 492, t. 59(1) (1880)
= Asplenium filix-femina var. dentigerum C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 491 (1880) (fide Fraser-Jenkins 1997)
= Athyrium angustifrons Ching; (1976) (nom. nud.) (fide Fraser-Jenkins 2008)
= Athyrium attenuatum f. dentigerum (Wall. ex C.B.Clarke) Khullar & Fraser-Jenk.; Aspects Pl. Sci. 6: 102 (1983)
= Athyrium attenuatum f. intermedium Mehra & Khullar ex Fraser-Jenk. & Khullar; Aspects Pl. Sci. 6: 99 (1983)
= Athyrium dentigerum (C.B.Clarke) Mehra & Bir; Amer. Fern J. 50: 279 (1960)
= Athyrium ensiferum Ching & H.S.Kung; Acta Bot. Boreal.-Occid. Sin. 6(2): 102 (1986) (fide Fraser-Jenkins 2008)
= Athyrium excelsius Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 20 (1986) (non Nakai 1914) (fide Fraser-Jenkins 2008)
= Athyrium filix-femina var. attenuatum (C.B.Clarke) Bedd.; Ferns Brit. India 169 (1883)
= Athyrium filix-femina var. dentigerum (C.B.Clarke) Bedd.; Ferns Brit. India 169 (1883)
= Athyrium leiboense Ching & Y.T.Hsieh; Acta Bot. Boreal.-Occid. Sin. 6(3): 158 (1986)
= Athyrium pseudofilix-femina Ching; Acta Bot. Austro Sin. 1: 17 (1983) (fide Fraser-Jenkins 1997)
= Athyrium tatsienluense Ching; (1976) (nom. nud.) (fide Fraser-Jenkins 2008)
= Polypodium dentigerum Wall.; Numer. List. [Wallich] no. 334 (1829), nomen (fide Fraser-Jenkins 1997)
species :
Athyrium auriculatum
Seriz.; J. Jap. Bot. 46: 280 (1971)
Distribution: Taiwan
38 TAI
Remarks: sect. Strigoathyrium, non-setose group. Often included in A. clivicola but genetically separate
species :
Athyrium austroorientale
Ching; Fl. Xizangica 1: 141 (1983)
Distribution: China (Sichuan, Yunnan); Tibet
36 CHI TIB
Synonyms: hidden
= Athyrium aculeatum Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 15 (1986)
= Athyrium dajinense Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 17 (1986)
= Athyrium takingense Ching; (1976), nom. nud.
species :
Athyrium baoxingense
Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 14 (1986)
Distribution: China (SW-Sichuan)
36 CHI
Synonyms: hidden
= Athyrium paoshingense Ching; (1976), nom. nud.
species :
Athyrium bipinnatum
K.Hori; PhytoKeys 148: 97 (2020)
Distribution: Japan (Shikoko, Honshu, Kyushu)
38 JAP
species :
Athyrium biserrulatum
Christ; Bull. Acad. Int. Géogr. Bot. 17: 135-136 (1907)
Distribution: China (Sichuan, W-Guizhou, Yunnan); Tibet; Pakistan (Swat, Pakistani Punjab, Hazara); Nepal; Bhutan; India (Arunachal Pradesh, Himachal Pradesh, Madhya Pradesh, Nagaland, Rajasthan, Sikkim, Uttarakhand, West Bengal, Jammu & Kashmir); Myanmar
36 CHI TIB 40 BHU IND NEP PAK 41 MYA
Synonyms: hidden
= Asplenium filix-femina var. polysporum C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 493 (1880)
= Athyrium filix-femina var. polysporum (C.B.Clarke) Bedd.; Ferns Brit. India 170 (1883)
= Athyrium polysporum (C.B.Clarke) Ching ex Mehra & Bir; Amer. Fern J. 50: 289 (1960)
= Athyrium schimperi subsp. biserrulatum (Christ) Fraser-Jenk.; New Species Syndr. Indian Pteridol. 60 (1997)
= Athyrium wumonshanicum Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 20 (1986) (fide Fraser-Jenkins (2008) but kept separate by Flora of China)
species :
Athyrium bomicola
Ching; Fl. Xizangica 1: 136 (1983)
Distribution: Tibet
36 TIB
species :
Athyrium bourgaei
E.Fourn.; Mexic. Pl. 1: 102 (1872)
Distribution: Mexico (Chihuahua, Durango, Nayarit, Jalisco, Michoacan, Mexico State, Oaxaca, Veracruz, Hidalgo, Chiapas, Baja California Sur, México Distrito Federal, Guanajuato, Morelos, Nuevo Leon, Puebla, Sonora); Guatemala; El Salvador
79 MEX 80 ELS GUA
Remarks: included by Fl. Mesoamericana in A. filix-femina, kept separate by later Mexican floras
Synonyms: hidden
= Athyrium paucifrons C.Chr.; Repert. Spec. Nov. Regni Veg. 9: 371 (1911)
species :
Athyrium brevipinnulum
Copel.; Philipp. J. Sci. 8: 291 (1908)
Distribution: Philippines
42 PHI
Remarks: sect. Strigoathyrium, valid species and not a synonym of A. nitidulum fide Wardani et al. 2024
species :
Athyrium brevistipes
Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 12 (1986)
Distribution: China (Chongqing, Guizhou, Hubei, Hunan, S-Shaanxi, Sichuan)
36 CHI
species :
Athyrium caudatum
Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 21 (1986)
Distribution: China (NW-Yunnan)
36 CHI
species :
Athyrium caudiforme
Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 21 (1986)
Distribution: China (Hunan, W-Sichuan)
36 CHI
species :
Athyrium chingianum
X.C.Zhang ex Z.R.Wang & X.C.Zhang; Bull. Bot. Res., Harbin 11(3): 12 (1991)
Distribution: China (Yunnan)
36 CHI
species :
Athyrium christensenii
Tardieu; Aspl. Tonkin 80, t. 12 (1932)
Distribution: China (SE-Yunnan, SW-Guangxi); Vietnam
36 CHI 41 VIE
Remarks: sect. Strigoathyrium, arisanense agg.
Conservation Status: IUCN R
Synonyms: hidden
= Lunathyrium christensenii (Tardieu) Ching; Acta Phytotax. Sin. 9: 72 (1964)
= Athyrium puberulum Ching; (1976), nom. nud.
species :
Athyrium chungtienense
Ching; Bull. Fan Mem. Inst. Biol. Bot., ser. 2, 1: 277 (1949)
Distribution: China (NW-Yunnan)
36 CHI
Synonyms: hidden
= Athyrium habaense Ching; Acta Bot. Boreal.-Occid. Sin. 6(3): 154 (1986)
= Athyrium confertum Ching; (1976), nom. nud.
species :
Athyrium clarkei
Bedd.; Ferns Brit. India Suppl. II, t. 360 (1876)
Distribution: China (Guizhou, S-Yunnan); Nepal; India (Manipur, Nagaland, Sikkim, West Bengal, Darjeeling); Myanmar
36 CHI 40 IND NEP 41 MYA
Remarks: sect. Strigoathyrium, strigillosum agg.
Synonyms: hidden
= Asplenium clarkei (Bedd.) Atk. ex C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 489 (1880)
= Athyrium nigripes var. clarkei (Bedd.) Bedd.; Ferns Brit. India 166 (1883)
species :
Athyrium clemensiae
Copel.; Philipp. J. Sci., C, 12: 58 (1917)
Distribution: Borneo (Sabah: Mt. Kinabalu, Sabah)
42 BOR
Remarks: sect. Strigoathyrium, scandicinum agg.
species :
Athyrium clivicola
Tagawa; Acta Phytotax. Geobot. 3: 32 (1934)
Distribution: China (Anhui, Chongqing, Fujian, Guangxi, Guizhou, Hubei, Hunan, Jiangxi, Sichuan, Zhejiang); South Korea (incl. Cheju Isl.); Japan; ?Taiwan
36 CHI 38 JAP KOR
Remarks: sect. Strigoathyrium, non-setose group
Synonyms: hidden
= Athyrium clivicola f. incisum (H.Itô) Nakaike; J. Nippon Fernist Club 3(Suppl. 2): 23 (2004)
= Athyrium wardii f. incisum H.Itô; J. Jap. Bot. 11: 428 (1935)
= Athyrium wardii var. clivicola (Tagawa) Sa.Kurata; (1954)
= Athyrium wardii var. majus Ohwi; (1957) (p. p., non Makino 1899)
variant :
var. rotundum
Z.R.Wang; Fl. Reipubl. Popularis Sin. [Fl. China Ed. Committee] 3(2): 508 (1999)
Distribution: China (Chongqing)
36 CHI
Synonyms: hidden
= Athyrium rotundum Ching; in sched.
species :
Athyrium contingens
Ching & S.K.Wu; Fl. Xizangica 1: 134 (1983)
Distribution: China (W-Sichuan); Tibet; Nepal; Bhutan; India (Sikkim)
36 CHI TIB 40 BHU IND NEP
Remarks: valid species fide Fraser-Jenkins (2008) and not a synonym of A. attenuatum. Flora of China keeps the synonyms separate
Synonyms: hidden
= Athyrium adscendens Ching; Acta Bot. Boreal.-Occid. Sin. 6(2): 100 (1986) (fide Fraser-Jenkins 2008, 2018)
= Athyrium dentatum Ching; Acta Bot. Boreal.-Occid. Sin. 6(3): 150 (1986), non (Sw.) Gray (1821) (fide Fraser-Jenkins 2018)
= Athyrium lancipinnulum Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 18 (1986) (fide Fraser-Jenkins 2018)
= Athyrium nudifrons Ching; Acta Bot. Boreal.-Occid. Sin. 6(3): 158 (1986) (fide Fraser-Jenkins 2008, 2018)
= Athyrium oblongum Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 17 (1986)
= Athyrium squamipes Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 17 (1986) (fide Fraser-Jenkins 2018)
= Athyrium supranigrescens Ching; Acta Bot. Boreal.-Occid. Sin. 6(3): 150 (1986) (fide Fraser-Jenkins 2018)
= Athyrium tarulakaense Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 12 (1986) (fide Fraser-Jenkins 2008)
= Athyrium tsaii Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 19 (1986) (fide Fraser-Jenkins 2018)
species :
Athyrium costulalisorum
Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 15 (1986)
Distribution: China (W-Sichuan)
36 CHI
species :
Athyrium crassicaule
J.P.Roux; Kew Bull. 63(3): 433 (2008 publ. 2009)
Distribution: South Africa (KwaZulu-Natal, Free State); Lesotho
27 LES SAF
species :
Athyrium crassipes
Ching; Acta Bot. Boreal.-Occid. Sin. 6(2): 103 (1986)
Distribution: China (W-Sichuan)
36 CHI
species :
Athyrium criticum
Ching; Fl. Tsinling. 2: 216 (1974)
Distribution: China (Shaanxi, W-Sichuan, NW-Yunnan)
36 CHI
species :
Athyrium cryptogrammoides
Hayata; Icon. Pl. Formosan. 6: 156 (1916)
Distribution: China (Zhejiang, Hubei, Hunan, Guangxi, Guizhou); Japan; Taiwan
36 CHI 38 JAP TAI
Synonyms: hidden
= Athyrium dissectifolium Ching; Acta Bot. Boreal.-Occid. Sin. 6(2): 104 (1986)
= Athyrium ×hohuanshanense Yoshik.; J. Phytogeogr. Taxon. 33(2): 80 (1985)
= Athyrium ×pseudocryptogrammoides Yoshik.; J. Phytogeogr. Taxon. 29(1): 27 (1981)
species :
Athyrium cyclosorum
(Rupr.) Gilbert; Fernwort Pap. 27 (1900) (Dec. 20)
Name: Subarctic ladyfern
Distribution: Russian Far East (Kamchatka, Kuril Isl., Magadan, Sakhalin); Alaska; Canada (Alberta, British Columbia, Manitoba, Northern Territories, Ontario, Québec, Saskatchewan, Yukon); USA (Arizona, California, Colorado, Idaho, Michigan, Minnesota, Montana, North Dakota, Nebraska, New Mexico, Nevada, Oregon, South Dakota, Utah, Washington State, Wyoming)
31 RFE 70 ASK CDN 71 CDN 72 CDN 73 USA 74 USA 75 USA 76 USA 77 USA
Remarks: often treated within A. filix-femina or as variety of A. asplenioides, but increasingly elevated to full species status
Synonyms: hidden
= Athyrium cyclosorum (Rupr.) Maxon; Bull. Torrey Bot. Club 27: 637 (1900), isonym (Dec. 29)
= Asplenium cyclosorum (Rupr.) Fernald; Rhodora 9: 158 (1907), isonym
= Asplenium filix-femina var. cyclosorum (Rupr.) D.C.Eaton; Rep. U.S. Geogr. Surv., Wheeler 6(Botany): 331 (1878)
= Athyrium alpestre var. cyclosorum (Rupr.) T.Moore
= Athyrium cyclosorum f. hillii Gilbert; List N. Amer. Pterid.: 14, 32 (1901)
= Athyrium cyclosorum var. strictum Gilbert; List N. Amer. Pterid.: 14, 32 (1901)
= Athyrium filix-femina f. hillii (Gilbert) Butters; Rhodora 19: 199 (1917)
= Athyrium filix-femina f. strictum (Gilbert) Butters; Rhodora 19: 200 (1917)
= Athyrium filix-femina f. strictum (Gilbert) J.T.Howell; Marin Fl., ed. 2, Suppl. 363 (1969), isonym
= Athyrium filix-femina subsp. cyclosorum (Rupr.) C.Chr.; Fl. Aleutian Isl. 50 (1937)
= Athyrium filix-femina var. cyclosorum Rupr.; Distr. Crypt. Vasc. Ross. 41 (1845)
species :
Athyrium davidii
(Franch.) Christ; Mém. Soc. Bot. France 1 [Hist. Acrostich.]: 50 (1903)
Distribution: China (W-Yunnan, W-Sichuan); Tibet; Nepal; Bhutan; India (Uttarakhand, Sikkim, Arunachal Pradesh); Myanmar (Kachin, etc.)
36 CHI TIB 40 BHU IND NEP 41 MYA
Conservation Status: IUCN R
Synonyms: hidden
= Asplenium duthiei Bedd.; J. Bot. 27: 72 (1889)
= Athyrium duthiei (Bedd.) Bedd.; Ferns Brit. India Suppl. 34 (1892) (fide Fraser-Jenkins (2008)
= Phegopteris davidii (Franch.) Bedd.; Suppl. Ferns Brit. Ind. 82 (1892)
= Polypodium davidii Franch.; Nouv. Arch. Mus. Hist. Nat., sér. 2, 10: 119 (1887)
= Pseudocystopteris davidii (Franch.) Z.R.Wang; Vasc. Pl. Hengduan Mount. 1: 77 (1993) (fide Fraser-Jenkins 1997)
species :
Athyrium daxianglingense
Ching & H.S.Kung; Acta Bot. Boreal.-Occid. Sin. 6(1): 16 (1986)
Distribution: China (W-Sichuan, Yunnan)
36 CHI
species :
Athyrium decorum
Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 13 (1986)
Distribution: China (NW-Yunnan); Tibet; Nepal; Bhutan; India (Arunachal Pradesh, Sikkim, West Bengal)
36 CHI TIB 40 BHU IND NEP
Remarks: sect. Strigoathyrium; nitidulum group
species :
Athyrium delavayi
Christ; Mém. Soc. Bot. France 1 [Hist. Acrostich.]: 47 (1905)
Distribution: China (Hubei, Guangxi, Sichuan, Chongqing, Guizhou, Yunnan); Taiwan; India (Meghalaya, Mizoram); Myanmar
36 CHI 38 TAI 40 IND 41 MYA
Remarks: sect. Strigoathyrium, non-setose group
Conservation Status: IUCN CR for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Athyrium costale var. polystichoides T.Moore; Index Fil. [T. Moore] 180 (1860), nom. nud.
= Athyrium latibasis Ching; Acta Bot. Boreal.-Occid. Sin. 6(2): 104 (1986)
= Athyrium mairei Rosenst.; Repert. Spec. Nov. Regni Veg. 13: 125 (1914)
= Athyrium neodelavayi Ching & H.S.Kung; Acta Bot. Boreal.-Occid. Sin. 6(1): 22 (1986)
= Athyrium pseudodelavayi Ching; (1976), nom. nud.
species :
Athyrium deltoidofrons
Makino; Bot. Mag. (Tokyo) 28: 178 (1914)
Distribution: China (Zhejiang, Jiangxi, Fujian, Hunan, Sichuan, Guizhou); South Korea; Japan
36 CHI 38 JAP KOR
Synonyms: hidden
= Athyrium acutissimum Kodama; Matsum., Icon. Pl. Koisikav. 3: 127, t. 209 (1917)
= Athyrium deltoidofrons f. ohmurae Sa.Kurata; J. Geobot. 10(1): 10 (1961)
= Athyrium deltoidofrons f. reflexipinnulum (H.Itô) Nakaike; J. Nippon Fernist Club 3(Suppl. 2): 24 (2004)
= Athyrium deltoidofrons var. acutissimum (Kodama) Sa.Kurata; (1955)
= Athyrium deltoidofrons var. gracillimum (Ching) Z.R.Wang; Fl. Reipubl. Popularis Sin. [Fl. China Ed. Committee] 3(2): 186 (1999)
= Athyrium deltoidofrons var. ohmurae (Sa.Kurata) Seriz.; Shidekobushi 3(1): 51 (2015)
= Athyrium filix-femina var. deltoideum Makino; Bot. Mag. (Tokyo) 18: 30 (1899)
= Athyrium gracillimum Ching; Acta Bot. Boreal.-Occid. Sin. 6(3): 149 (1986)
= Athyrium jiulungshanense Ching; Bull. Bot. Res., Harbin 2(2): 72 (1982)
= Athyrium rotundilobum Ching; Fl. Fujianica 1: 597 (1982)
= Athyrium vidalii f. reflexipinnulum H.Itô; J. Jap. Bot. 11: 427 (1935)
= Athyrium ×multifidum var. deltoideum (Makino) Nakai; Bot. Mag. (Tokyo) 44: 507 (1930)
species :
Athyrium dentilobum
Ching & S.K.Wu; Fl. Xizangica 1: 139 (1983)
Distribution: Tibet
36 TIB
Remarks: probably a synonym of A. distans fide Fraser-Jenkins et al. 2018
species :
Athyrium devolii
Ching; Sunyatsenia 3: 1, t. 1 (1935)
Distribution: China (Chongqing, N-Fujian, Guangxi, Guizhou, Jiangxi, Sichuan, Yunnan, Zhejiang); Tibet; E-Nepal; Bhutan; India (Manipur, Meghalaya, Nagaland, Sikkim, West Bengal); Myanmar; Vietnam
36 CHI TIB 40 BHU IND NEP 41 MYA VIE
Remarks: A. delicatulum (which is anyway an invalid name) is included here by Fraser-Jenkins et al. (2018). Also, reports of A. solenopteris from the Himalaya belong here fide Fraser-Jenkins (2008)
Synonyms: hidden
= Athyrium aridum Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 10 (1986)
= Athyrium austrooccidentale Ching; Acta Bot. Boreal.-Occid. Sin. 6(3): 152 (1986)
= Athyrium baishanzuense Ching & Y.T.Hsieh; Acta Bot. Boreal.-Occid. Sin. 6(3): 157 (1986) (fide Fraser-Jenkins et al. 2018)
= Athyrium bambusicola Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 18 (1986)
= Athyrium delicatulum Ching & S.K.Wu; Fl. Xizangica 1: 139 (1983), non Ôishi (1941)
= Athyrium filix-femina var. fissidens Christ; (1905) (non Döll 1855)
= Athyrium fujianense Ching; Fl. Fujianica 1: 597 (1982)
= Athyrium giganteum De Vol; Lingnan Sci. J. 21: 79, t. 5 (1945) (fide Fraser-Jenkins 2018)
= Athyrium nanchuanense Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 3(4): 17 (1983)
= Athyrium nitidum Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 19 (1986)
= Athyrium praticola Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 18 (1986)
= Athyrium remotisorum Ching & H.S.Kung; Acta Bot. Boreal.-Occid. Sin. 6(3): 153 (1986)
= Athyrium remotum Ching; Acta Bot. Boreal.-Occid. Sin. 6(3): 149 (1986)
= Athyrium yaanense Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 19 (1986)
species :
Athyrium dissitifolium
(Baker) C.Chr.; Contr. U. S. Natl. Herb. 26: 296, t. 18 (1931)
Distribution: China (Guangxi, Guizhou, Hunan, Sichuan, Yunnan); NE-India (Arunachal Pradesh, Manipur, Mizoram); Myanmar (Kachin, Chin, etc.); N-Thailand; Vietnam
36 CHI 40 IND 41 MYA THA VIE
Conservation Status: IUCN CR for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Athyrium dissitifolium var. funebre (Christ) Ching & Z.R.Wang; Fl. Reipubl. Popularis Sin. [Fl. China Ed. Committee] 3(2): 124 (1999)
= Athyrium drepanopteron var. funebre Christ; Notul. Syst. (Paris) 1: 46 (1909)
= Athyrium drepanopteron var. brevicaudatum Christ; Notul. Syst. (Paris) 1: 46 (1909)
= Athyrium fasciculatum Hand.-Mazz.; Symb. Sin. 6: 31, t. 2, f. 5 (1929)
= Dryopteris apicidens (Baker) C.Chr.; Index Filic. 252 (1905)
= Dryopteris aristulata Rosenst.; Repert. Spec. Nov. Regni Veg. 13: 132 (1914)
= Dryopteris dissitifolia (Baker) C.Chr.; Index Filic. 262 (1905)
= Dryopteris incrassata (Christ) C.Chr.; Index Filic. 272 (1905)
= Nephrodium dissitifolium (Baker) Baker; Bull. Misc. Inform. Kew 1899: 120 (1899)
= Phegopteris incrassata Christ; Bull. Herb. Boissier, sér. 1, 6: 963 (1898)
= Polypodium apicidens Baker; Bull. Misc. Inform. Kew 1894: 54 (1894)
= Polypodium dissitifolium Baker; Bull. Misc. Inform. Kew 1895: 54 (1895)
species :
Athyrium distans
(D.Don) T.Moore; Index Fil. [T. Moore] 125, 181 (1860)
Distribution: China (Chongqing, Guizhou, Sichuan); Japan; Nepal; Bhutan; N-India (Jammu & Kashmir, Arunachal Pradesh, Assam State, Himachal Pradesh, Manipur, Meghalaya, Mizoram, Nagaland, Sikkim, Uttarakhand, West Bengal); Myanmar (Kachin, etc.)
36 CHI 38 JAP 40 BHU IND NEP 41 MYA
Remarks: sect. Strigoathyrium, synonymy following Fraser-Jenkins (2008, 2018). Flora of China keeps A. imbricatum and A. nyalamense separate
Synonyms: hidden
= Asplenium distans D.Don; Prodr. Fl. Nepal. 9 (1826)
= Athyrium frangulum f. viride Sa.Kurata; Hokuriko J. Bot. 4: 17 (1955)
= Athyrium frangulum Tagawa; J. Jap. Bot. 12: 749 (1936)
= Athyrium imbricatum Christ; Bull. Acad. Int. Géogr., sér. 3, 16: 1906: 123 (1906)
= Athyrium imbricatum f. viride (Sa.Kurata) Nakaike; Checkl. Lycophytes Ferns Japan: 17 (2013)
= Athyrium iseanum var. fragile Tagawa; Acta Phytotax. Geobot. 2: 15 (1933)
= Athyrium nigripes var. dissectum C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 491 (1880)
species :
Athyrium nyalamense
Y.T.Hsieh & C.R.Wang; Fl. Xizangica 1: 138 (1983)
Distribution: SE-Tibet
36 TIB
Remarks: sect. Strigoathyrium, sometimes included in A. distans, but more close to A. araiostegioides
Synonyms: hidden
= Athyrium nyalamense var. puberulum Z.R.Wang; Vasc. Pl. Hengduan Mount. 1: 87 (1993) (without latin descr. or type), et Acta Phytotax. Sin. 32(1): 84 (1994)
species :
Athyrium dombeyi
Desv.; Mem. Soc. Linn. Paris 6(2): 266 (1827) (dombei auct.)
Distribution: Colombia (Antioquia, Boyacá, Caldas, Cundinamarca, Norte de Santander, Tolima); Venezuela (Aragua, Merida, Tachira, Trujillo); Ecuador; Peru; Bolivia (Cochabamba, La Paz, Tarija, Santa Cruz); Brazil (Espírito Santo, Minas Gerais, Rio de Janeiro, Rio Grande do Sul, Santa Catarina, São Paulo); Argentina (Buenos Aires, Cordoba, Salta, Tucuman)
82 CLM VEN 83 BOL ECU PER 84 BRA 85 ARG
Synonyms: hidden
= Asplenium dombeyi (Desv.) Mett.; Ann. Sci. Nat., Bot., sér. 5, 2: 238 (1864)
= Athyrium filix-femina var. dombeyi (Desv.) Hieron.; Hedwigia 47: 210 (1908)
species :
Athyrium drepanopteron
(Kunze) A.Braun ex Milde; Fil. Eur. 49 (1867)
Distribution: China (Yunnan); Tibet; Nepal; Bhutan; India (Arunachal Pradesh, Manipur, Meghalaya, Nagaland, Sikkim, Tripura, Uttarakhand, West Bengal); Myanmar (Kachin, Chin, etc.); Thailand; Vietnam; Sumatra; Philippines
36 CHI TIB 40 BHU IND NEP 41 MYA THA VIE 42 PHI SUM
Remarks: synonymy unclear due to confusion with A. eburneum (mupinense). Both species have been separated by Fraser-Jenkins (2008) and Liu (2009) but are united by Flora of China. They are kept separate here - see also under A. eburneum
Synonyms: hidden
= Aspidium drepanopteron (Kunze) A.Braun ex Mett.; Fil. Hort. Bot. Lips. 93, t. 19, f. 1-4 (1856)
= Aspidium drepanopteron var. decompositum Christ; (1909)
= Asplenium drepanopteron (Kunze) A.Braun; Index Sem. (B, Berolinensis) 17 (1856)
= Asplenium oxyphyllum f. kulhaitense Atk. ex C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 494 (1880)
= Asplenium oxyphyllum Wall. ex Hook.; Sp. Fil. [W. J. Hooker] 3: 221 (1860), nom. superfl.
= Athyrium dissitifolium var. kulhaitense (Atk. ex C.B.Clarke) Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 22 (1986)
= Athyrium exindusiatum Ching; Acta Bot. Boreal.-Occid. Sin. 6(2): 102 (1986) (fide Fraser-Jenkins (2008), kept separate by Flora of China)
= Athyrium oxyphyllum (Hook.) T.Moore; Index Fil. [T. Moore] 49 (1857) ex Bedd. 1866
= Polypodium drepanopteron Kunze; Linnaea 23: 278, 318 (1850)
= Polypodium kulhaitense Atk. ex C.B.Clarke; (1880) nom. inval. in syn.
= Polypodium oxyphyllum Wall.; Numer. List. [Wallich] no. 324 (1829), nomen
species :
Athyrium dubium
Ching; Bull. Fan Mem. Inst. Biol. Bot., ser. 2, 1: 281 (1949)
Distribution: China (Yunnan, Guizhou, Sichuan); Tibet; Nepal; Bhutan; India (Arunachal Pradesh, Himachal Pradesh, Sikkim, Uttarakhand, West Bengal, Jammu & Kashmir)
36 CHI TIB 40 BHU IND NEP
Synonyms: hidden
= Asplenium filix-femina subvar. rubricaule Edgew. ex C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 492, t. 59, f. 2 (1880)
= Athyrium acutum Ching; Acta Bot. Boreal.-Occid. Sin. 6(2): 101 (1986)
= Athyrium aequilaterale Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 20 (1986)
= Athyrium juxtapositum Ching; (1976), nom. nud.
= Athyrium longipinna Ching; Acta Bot. Boreal.-Occid. Sin. 6: 15 (1986)
= Athyrium machangense Ching; Acta Bot. Boreal.-Occid. Sin. 6(2): 100 (1986)
= Athyrium mackinnoniorum var. yigongense Ching & S.K.Wu; Fl. Xizangica 1: 142 (1983)
= Athyrium rubricaule (Edgew. ex C.B.Clarke) Bir; Nova Hedwigia 4: 169, t. 54 (1962)
= Athyrium serratodentatum Ching; Acta Bot. Boreal.-Occid. Sin. 6(2): 99 (1986)
= Athyrium sublineare Ching; Acta Bot. Boreal.-Occid. Sin. 6(2): 99 (1986)
= Athyrium uniseriatum Ching; Acta Bot. Boreal.-Occid. Sin. 6(2): 99 (1986)
species :
Athyrium eburneum
(J.Sm. ex Mett.) J.Sm.; Hist. Fil. 328 (1875)
Distribution: China (W-Sichuan, Guizhou, Yunnan); Tibet; Japan; Taiwan; Nepal; Bhutan; India (Arunachal Pradesh, Meghalaya, Manipur, Nagaland, Sikkim, West Bengal); Myanmar; Vietnam; Philippines
36 CHI TIB 38 JAP TAI 40 BHU IND NEP 41 MYA VIE 42 PHI
Remarks: listed as A. drepanopterum in Fl. China etc. fide Fraser-Jenkins 2008 but kept separate by Liu (2009) and Knapp (2013).
Synonyms: hidden
= Aspidium eburneum Wall.; Numer. List. [Wallich] no. 389 (1829), nomen
= Asplenium eburneum J.Sm. ex Mett.; Cat. Cult. Ferns: 47 (1857), et Abh. Senckenberg. Naturf. Ges. 3(1) [Aspl.]: 238 (1859)
= Athyrium micans Tagawa; Acta Phytotax. Geobot. 4: 143 (1935)
= Athyrium mupinense Christ; Bot. Gaz. 51: 355 (1911) (moupinense auct.)
= Athyrium niponicum var. elatius Christ; (1905)
= Athyrium takeoi (Hayata) Tagawa; Acta Phytotax. Geobot. 2(3): 18 (1933)
= Athyrium veitchii Christ; Bull. Acad. Int. Géogr., sér. 3, 16: 123 (1906)
= Dryopteris takeoi Hayata; Icon. Pl. Formosan. 6: 289, f. 114 (1915)
= Lastrea eburnea (Wall.) J.Sm.; Compan. Bot. Mag. 34 (1846), nom. inval.
species :
Athyrium elmeri
Copel.; Philipp. J. Sci. 3: 292 (1908)
Distribution: Philippines
42 PHI
Synonyms: hidden
= Diplazium elmeri (Copel.) C.Chr.; Index Filic., Suppl. Tert.: 73 (1934)
= Phegopteris elmeri Alderw.; Malayan Ferns Fern Allies, Suppl. 314 (1917)
species :
Athyrium elongatum
Ching; Acta Bot. Boreal.-Occid. Sin. 6(2): 101 (1986)
Distribution: China (N-Guangxi, Jiangxi)
36 CHI
Synonyms: hidden
= Athyrium dolichophyllum P.S.Wang; (1989), nom. nud.
species :
Athyrium emeicola
Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 13 (1986)
Distribution: China (Sichuan)
36 CHI
species :
Athyrium epirachis
(Christ) Ching; C. Chr., Index Filic., Suppl. Tert.: 41 (1934)
Distribution: China (Chongqing, Fujian, S-Guangdong, Guangxi, Guizhou, SW-Hubei, Hunan, Sichuan, Yunnan); South Korea (Cheju Isl.); Japan (Honshu); Taiwan
36 CHI 38 JAP KOR TAI
Remarks: sect. Strigoathyrium, non-setose group. In Taiwan very rare. A. eremicola (Japan) is sometimes kept separate but genetically very close
Synonyms: hidden
= Athyrium eremicola Oka & Sa.Kurata ex Sa.Kurata; J. Geobot. 8: 83 (1960)
= Athyrium lilacinum Ching; Bull. Fan Mem. Inst. Biol. Bot., ser. 2, 1: 283 (1949)
= Athyrium muticum Christ; Bull. Acad. Int. Géogr., sér. 3, 17: 147 (1907)
= Athyrium rubristipes Ching; (1976), nom. nud.
= Athyrium subcoriaceum Ching; Acta Bot. Boreal.-Occid. Sin. 6(3): 151 (1986)
= Athyrium wardii var. elongatum Christ; (1905)
= Diplazium epirachis Christ; Mém. Soc. Bot. France 1 [Hist. Acrostich.]: 51 (1905)
species :
Athyrium erythropodum
Hayata; Icon. Pl. Formosan. 4: 233 t 163 (1914)
Distribution: Taiwan; Vietnam; ?Philippines
38 TAI 41 VIE
Remarks: sect. Strigoathyrium, non-setose group, erythropodum clade. Not in Japan
Conservation Status: IUCN R
species :
Athyrium falcatum
Bedd.; Ferns S. India t. 151 (1865)
Distribution: W-Nepal; India (Chhattisgarh, Gujarata, Himachal Pradesh, Jharkand, Karnataka, Kerala, Madhya Pradesh, Maharashtra, Meghalaya, Mizoram, Odisha, Rajasthan, Tamil Nadu, Tripura, Uttarakhand); Myanmar
40 IND NEP 41 MYA
Synonyms: hidden
= Asplenium drepanophyllum Baker; Syn. Fil. [W. J. Hooker & Baker], ed. 2: 226 (1874) (non Kunze (1834))
= Athyrium drepanophyllum (Baker) Bedd.; Ferns Brit. India Suppl. 32 (1892)
= Athyrium grammitoides Bedd.; Ferns Brit. India Correct. 11 (1870)
= Athyrium ×keralense Manickam & Irudayaraj; Pterid. Fl. West. Ghats, S. India: 238 (1992) (as ´keralensis´, nom. inval., sine type specimen, fide Fraser-Jenkins 1997)
species :
Athyrium fallaciosum
Milde; Fil. Eur. 6: 54 (1867)
Distribution: Russian Far East (Kuril Isl., Sakhalin); China (Gansu, Hebei, Heilongjiang, Henan, Hubei, Jilin, Liaoning, Nei Mongol, Ningxia, Shaanxi, Shanxi, N-Sichuan); North Korea; Japan (Hokkaido, Honshu); ?Philippines (?Luzon)
31 RFE 36 CHI 38 JAP NKO
Remarks: A. fauriei is kept separate by Flora of China but included here by Fraser-Jenkins (2008)
Synonyms: hidden
= Aspidium fauriei (Christ) Christ; Bull. Herb. Boissier, sér. 1, 6: 193 (1898)
= Asplenium mongolicum Franch.; Nouv. Arch. Mus. Hist. Nat., sér. 2, 7: 161 (1883)
= Asplenium nikkoense Makino; Bot. Mag. (Tokyo) 17: 43 (1903)
= Athyrium fauriei (Christ) Makino; Bot. Mag. (Tokyo) 17: 160 (1903)
= Athyrium mongolicum (Franch.) Diels; Nat. Pflanzenfam. [Engler & Prantl] I, 4: 224 (1899)
= Athyrium mongolicum var. purdomii C.Chr.; Bot. Gaz. 56(4): 333 (1913)
= Athyrium rupestre Kodama; Matsum., Icon. Pl. Koisikav. 4: 63, t. 244 (1919) (fide Fraser-Jenkins 2008)
= Athyrium yokoscense var. fauriei (Christ) Tagawa; Acta Phytotax. Geobot. 2: 20 (1933)
= Dryopteris mongolica (Franch.) C.Chr.; Index Filic. 278 (1905)
= Nephrodium fauriei Christ; Bull. Herb. Boissier, sér. 1, 4: 671 (1896)
= Nephrodium mongolicum (Franch.) Baker; Ann. Bot. (Oxford) 6: 317 (1891)
species :
Athyrium fangii
Ching; Bull. Fan Mem. Inst. Biol. Bot., ser. 2, 1: 282 (1949)
Distribution: China (Sichuan, Yunnan); ?Tibet; Nepal; Bhutan; N-India (Arunachal Pradesh, Sikkim, Uttarakhand, West Bengal)
36 CHI 40 BHU IND NEP
Synonyms: hidden
= Athyrium paranigripes X.C.Zhang; Bull. Bot. Res., Harbin 11(3): 9 (1991) (nom. nov. for A. sikkimense Ching) (fide Fraser-Jenkins et al. 2015)
= Athyrium sikkimense Ching; Acta Bot. Austro Sin. 1: 19 (1983) (non (Bir) Á. Löve & D. Löve) (fide Fraser-Jenkins et al. 2015)
species :
Athyrium filix-femina
(L.) Roth; Tent. Fl. Germ. 3: 65 (1799)
subspecies :
subsp. filix-femina
Name: Wald-Frauenfarn (DE), Lady Fern (EN); Helecho hembra (ES)
Distribution: Iceland; Faeroe Isl.; England (England, Wales, Scotland, Outer Hebrides, Orkney Isl., Shetland Isl., Isle of Man); Ireland (Ireland, Northern Ireland); Denmark; Norway; Sweden; Finland; Netherlands; Belgium; Luxembourg; Germany (Brandenburg, Berlin, Baden-Württemberg, Bayern, Hessen, Hamburg, Mecklenburg-Vorpommern, Niedersachsen, Nordrhein-Westfalen, Rheinland-Pfalz, Schleswig-Holstein, Saarland, Sachsen, Sachsen-Anhalt, Thüringen); Switzerland; Liechtenstein; Austria; Poland; Czech Republic; Slovakia; Hungary; Portugal; Spain; Andorra; France; Channel Isl. (Alderney, Guernsey, Jersey); Corsica; Sardinia; Italy; Sicily; Malta; Slovenia; Croatia; Bosnia & Hercegovina; Montenegro; Serbia; Kosovo; North Macedonia; Albania; Romania; Bulgaria; European Turkey; Greece (widespread, incl. Kiklades, Samothraki); Crete; Estonia; Latvia; Lithuania; Belarus; C-European Russia; E-European Russia; N-European Russia; W-European Russia; Moldova; Ukraine; Crimea; Morocco; Algeria; Tunisia; Azores (Santa Maria Isl., Sao Miguel Isl., Terceira, Graciosa, Sao Jorge, Pico, Faial, Flores Isl., Corvo Isl.); Madeira (Madeira Isl.); Canary Isl. (Gran Canaria, Tenerife, La Gomera, Hierro, La Palma Isl.); Siberia (Altai, Buryatia, Chita, Irkutsk, Krasnoyarsk, Tuva, W-Siberia, Yakutia); Russian Far East (Kamchatka, Khabarovsk, Kuril Isl., Magadan, Primorye, Sakhalin); Kazakhstan; Turkmenistan; ?Uzbekistan; Northern Caucasus; Georgia [Caucasus]; Armenia; Azerbaijan (incl. Talysh); East Aegaean Isl. (Lesvos, Irakia, Samos); Turkey (Anatolia); Iran; Lebanon; Mongolia; ?Japan; New Zealand [I]; Mexico (Baja California Norte, Baja California Sur, Chiapas, Chihuahua, Colima, Ciudad de Mexico, Durango, Guerrero, Hidalgo, Jalisco, Mexico State, Michoacan, Morelos, Nayarit, Nuevo Leon, Oaxaca, Puebla, Queretaro, Sonora, Tamaulipas, Veracruz, Zacatecas); Guatemala; ?Belize; ?El Salvador; Costa Rica; Panama
10 DEN FIN FOR GRB ICE IRE NOR SWE 11 AUT BGM CZE GER HUN LIE LUX NET POL SLO SWI 12 ADO CIS COR FRA POR SAR SPA 13 ALB BIH BUL CRO GRC ITA KOS KRI MLT MNT NMA ROM SIC SLV SRB TUE 14 BLR EST KRY LAT LIT MLD RUS UKR 20 ALG MOR TUN 21 AZO CNY MDR 30 SIB 31 RFE 32 KAZ TKM 33 ARM AZE GEO NCS 34 EAI IRN LEB TUR 37 MON 51 nzl 79 MEX 80 COS GUA PAN
Remarks: records from South America are referrable to A. dombeyi
Synonyms: hidden
= Allantodia axillaris (Aiton) Kaulf.; Enum. Filic. [Kaulfuss] 178 (1824)
= Aspidium axillare (Aiton) Sw.; J. Bot. (Schrader) 1800(2): 42 (1801)
= Aspidium cordatum Steud.; Nomencl. Bot. [Steudel] 2: 61 (1824)
= Aspidium filix-femina (L.) Sw.; J. Bot. (Schrader) 1800(2): 41 (1801)
= Aspidium filix-femina var. dentatum (Döll) Döll; Fl. Baden [Döll] 1(1): 23 (1855)
= Aspidium filix-femina var. incisum Hartm.; Handb. Skand. Fl. 369 (1820)
= Aspidium filix-femina var. multidentatum (Döll) Döll; Gefässkrypt. Baden 24 (1855)
= Aspidium filix-femina var. multifidum Döll; Fl. Baden [Döll] 1(1): 25 25 (1855)
= Aspidium intermedium Link; Enum. Hort. Berol. Alt. 2: 459 (1822), nom. illeg.
= Aspidium irriguum Sm.; Engl. Bot. 81, t. 2199 (1810)
= Aspidium pontederae Willd.; Sp. Pl., ed. 4 [Willdenow] 5(1): 273 (1810)
= Aspidium rhaeticum (L.) Sw.; J. Bot. (Schrader) 1800(2): 41 (1801), as ´rheticum´
= Asplenium axillare (Aiton) J.Sm.; Compan. Bot. Mag. 30 (1846)
= Asplenium cyathioides Bernh.; J. Bot. (Schrader) 1799(1): 313 (1799), nom. illeg.
= Asplenium filix-femina (L.) Bernh.; Neues J. Bot. 12: 26, 48, t. 2, f. 7 (1806)
= Asplenium filix-femina f. dentatum (Döll) Döll; Fl. Baden [Döll] 1(1): 23 (1855)
= Asplenium filix-femina f. fissidens Döll; Fl. Baden [Döll] 1(1): 24 (1855)
= Asplenium filix-femina f. multidentatum (Döll) Döll; Fl. Baden [Döll] 1(1): 24 (1855)
= Asplenium filix-femina var. bidentatum Döll; Rhein. Fl. 12 (1843)
= Asplenium filix-femina var. molle (Schreb.) Hook.f.; 462 (1870)
= Asplenium filix-femina var. purpureum Regel; Index Sem. (LE, Petropolitanus) 1866(Suppl.): 25 (1867)
= Asplenium filix-femina var. rhaeticum (L.) Hook.f.; 462 (1870)
= Asplenium monstrosum Kunze; Linnaea 23: 235, 305 (1850)
= Asplenium rhaeticum (L.) Brügger; Naturg. Beitr. Chur: 47 (1874)
= Athyrium acrostichoideum Bory ex Mérat; Nouv. Fl. Env. Paris, ed. 4, 1: 372 (1836)
= Athyrium alpestre Clairv.; Man. Herbor. Suisse 301 (1811) non (Hoppe)
= Athyrium axillare (Aiton) C.Presl; Tent. Pterid. 98 (1836)
= Athyrium azoricum Fée; Mém. Foug. 5 (Gen. Filic.): 186 (1852)
= Athyrium cassum Chiov.; Fl. Alpi Lepont. 2: 46 (1929)
= Athyrium convexum (Bab.) Newman; Phytologist 4(App.): XIII (1851)
= Athyrium corsicum Fée; Mém. Foug. 5 (Gen. Filic.): 186 (1852)
= Athyrium depauperatum Schumach.; Enum. Pl. [Schumacher] 2: 17 (1803)
= Athyrium filix-femina proles acrostichoideum (Bory ex Mérat) Rouy; Fl. France [Rouy & Foucaud] 14: 428 (1913)
= Athyrium filix-femina f. rhaeticum (L.) Murr; 5 (1923)
= Athyrium filix-femina var. bidentatum Briq.; Prodr. Fl. Corse 1: 12 (1910)
= Athyrium filix-femina var. convexum Bab.; Man. Brit. Bot. [Babington] 388 (1843)
= Athyrium filix-femina var. curtum-cristatum Ivery ex R.Hogg; Florist & Pomol. 27: 135 (1874)
= Athyrium filix-femina var. dentatum (Döll) Milde; Fil. Eur. 50 (1867)
= Athyrium filix-femina var. dubpulayae Lowe; Gard. Chron., ser. 2, 4: 81 (1875)
= Athyrium filix-femina var. erectum Syme & N.E.Br.; Engl. Bot., 3rd ed., vol. XII: 109 (1886)
= Athyrium filix-femina var. fissidens (Doll) Milde; Fil. Eur. 50 (1867)
= Athyrium filix-femina var. frizelliae W.Bull ex J.Dix; Proc. Roy. Hort. Soc. London 1: 652 (1861)
= Athyrium filix-femina var. incisum (Fée) Hieron.; Bot. Jahrb. Syst. 22(3): 363 (1896), nom. illeg., non Hartm. 1820
= Athyrium filix-femina var. incisum Hartm.; Handb. Skand. Fl. 369 (1820)
= Athyrium filix-femina var. incisum Newman; Hist. Brit. Ferns, ed. II: 243 (1844), nom. illeg.
= Athyrium filix-femina var. iveryanum Ivery ex J.Dix; Proc. Roy. Hort. Soc. London 2: 675 (1862)
= Athyrium filix-femina var. kingscotei Lowe; Gard. Chron., ser. 2, 4: 81 (1875)
= Athyrium filix-femina var. latipes T.Moore; Handb. Brit. Ferns 2: 30 (1860)
= Athyrium filix-femina var. lineare-reflexum Lowe; Gard. Chron., ser. 2, 4: 81 (1875)
= Athyrium filix-femina var. minus Payot; Fl. Mont-Blanc 2: 10 (1882)
= Athyrium filix-femina var. molle (Schreb.) Briq.; Prodr. Fl. Corse 1: 12 (1910)
= Athyrium filix-femina var. molle (Schreb.) Schur; 168 (1851)
= Athyrium filix-femina var. mucronatum Ivery ex J.Dix; Proc. Roy. Hort. Soc. London 2: 675 (1862)
= Athyrium filix-femina var. multiceps T.Moore ex J.Dix; Proc. Roy. Hort. Soc. London 1: 70 (1859)
= Athyrium filix-femina var. multidentatum (Döll) Milde; Fil. Eur. 50 (1867)
= Athyrium filix-femina var. ovatum (Roth) Schur; 168 (1851)
= Athyrium filix-femina var. parsonsiae J.Dix; Proc. Roy. Hort. Soc. London 2: 459 (1862)
= Athyrium filix-femina var. pectinatum Millsp.; Living Fl. W. Virginia 5A(1): 193 (1913), nom. illeg.
= Athyrium filix-femina var. plumosum T.Moore ex J.Dix; Proc. Roy. Hort. Soc. London 1: 239 (1860)
= Athyrium filix-femina var. pulchellum-plumosum Anon.; Gard. Chron., ser. 2, 2: 49 (1874)
= Athyrium filix-femina var. qulterianum Anon.; Gard. Chron., ser. 2, 2: 49 (1874)
= Athyrium filix-femina var. rhaeticum (L.) Schur; 168 (1851)
= Athyrium filix-femina var. tripinnatum Rupr.; Dist. Crypt. Vasc. Ross. 41 (1845)
= Athyrium filix-femina var. watsonii Syme & N.E.Br.; Engl. Bot., 3rd ed., vol. XII: 110 (1886)
= Athyrium fimbriatum Dulac; Fl. Hautes-Pyrenées 33 (1867) non (Wall.) T. Moore
= Athyrium incisum (Hoffm.) Newman; Phytologist 4: app. XIII (1851)
= Athyrium incisum Fée; Mém. Foug. 5 (Gen. Filic.): 187 (1852), nom. illeg., non (Hoffm.) Newman 1851
= Athyrium irriguum (Sm.) Gray; Nat. Arr. Brit. Pl. 2: 10 (1821)
= Athyrium laetum (Salisb.) Gray; Nat. Arr. Brit. Pl. 2: 10 (1821), nom. superfl.
= Athyrium laxum Schumach.; Enum. Pl. [Schumacher] 2: 16 (1803)
= Athyrium molle (Schreb.) Roth; Tent. Fl. Germ. 3: 61 (1799)
= Athyrium molliusculum Baumg.; Enum. Stirp. Transsilv. 4: 31 (1846)
= Athyrium officinale Bubani; Fl. Pyren. (Bubani) 4: 430 (1901)
= Athyrium ovatum Roth; Tent. Fl. Germ. 3(1.1): 64 (1799)
= Athyrium paramicola L.D.Gómez; Phytologia 52(3): 154 (1982)
= Athyrium rhaeticum (L.) Dalla Torre; Anleit. Beob. Alpenpfl. 2: 348 (1882), nom. illeg.
= Athyrium rhaeticum (L.) Roth; Tent. Fl. Germ. 3: 67 (1799)
= Athyrium rhaeticum var. minus Doerr. ex Roth; 68 (1799)
= Athyrium scorpioideum Chauv.; Mém. Soc. Linn. Calvados 2: 502 (1825)
= Athyrium trifidum (Hoffm.) Roth; Tent. Fl. Germ. 3: 63 (1799)
= Cyathea filix-femina (L.) Bertol.; Amoen. Ital. 429 (1819)
= Cystopteris filix-femina (L.) Coss. & Germ.; Fl. Env. Paris: 676 (1845)
= Dryopteris rigida var. rhiphaea Pau & Font Quer; Iter Marocc. 1927: no. 3 (1928)
= Filix bipinnata Gilib.; Exerc. Phyt. 2: 557 (1792), opus utique oppr.
= Lastrea filix-femina (L.) Colomb; Compt. Rend. Hebd. Séances Acad. Sci. 107: 1013 (1888)
= Lastrea filix-femina var. bollandiae W.Bull ex J.Dix; Proc. Roy. Hort. Soc. London 2: 458 (1862)
= Nephrodium filix-femina (L.) Michx.; Fl. Bor.-Amer. (Michaux) 2: 268 (1803)
= Phegopteris plumosa J.Sm.; Ferns Brit. Foreign 289 (1866)
= Phegopteris rhaetica (L.) Pérard; Bull. Soc. Bot. France 16: 258 (1869), nom. illeg.
= Phegopteris rhaetica (L.) Watt; Canad. Naturalist & Quart. J. Sci., ser. 2, 13: 159 (1867)
= Polypodium axillare Aiton; Hort. Kew. [W. Aiton] 3: 466 (1789)
= Polypodium bifidum Hoffm.; Bot. Mag. (Römer & Usteri) 9: 10 (1790)
= Polypodium crenatum Schrank; Schrank & Moll, Naturhist. Briefe Oestreich 2: 295 (1785), nom. illeg.
= Polypodium dentatum Schrank; Schrank & Moll, Naturhist. Briefe Oestreich 2: 295 (1785), nom. illeg.
= Polypodium femina L.; Mant. Pl. 2: 506 (1771)
= Polypodium filix-femina L.; Sp. Pl. 2: 1090 (1753)
= Polypodium filix-femina var. molle (Schreb.) Lej.; Rev. Fl. Spa: 212 (1825)
= Polypodium filix-femina var. nanum Gren.; Bull. Soc. Bot. France 14: 63 (1867)
= Polypodium filix-femina var. purpurascens Bory ex Lej.; Rev. Fl. Spa: 212 (1825)
= Polypodium incisum Hoffm.; Bot. Mag. (Römer & Usteri) 9: 10 (1790), nom. illeg.
= Polypodium laetum Salisb.; Prodr. Stirp. Chap. Allerton 403 (1796), nom. superfl.
= Polypodium leseblii Mérat; Nouv. Fl. Env. Paris, ed. 2: 276 (1821)
= Polypodium molle Schreb.; Spic. Fl. Lips. 70 (1771)
= Polypodium oblongodentatum Hoffm.; Bot. Mag. (Römer & Usteri) 9: 10 (1790)
= Polypodium ovatocrenatum Hoffm.; Bot. Mag. (Römer & Usteri) 9: 10 (1790)
= Polypodium rhaeticum L.; Sp. Pl. 2: 1091 (1753)
= Polypodium trifidum Hoffm.; Bot. Mag. (Römer & Usteri) 9: 10 (1790)
= Tectaria filix-femina (L.) Cav.; Anales Ci. Nat. 4: 100 (1801)
subspecies :
subsp. californicum
(Butters) Hultén; Kongl. Svenska Vetensk. Acad. Handl. 8(5): 178 (1962)
Distribution: W-USA (Arizona, California, Colorado, Idaho, Nevada, New Mexico, Oregon, South Dakota, Utah, Wyoming)
73 USA 74 USA 76 USA 77 USA
Remarks: status unconfirmed, might belong to one of the species formerly included in A. filix-femina s. l. from North America
Synonyms: hidden
= Athyrium filix-femina var. californicum Butters; Rhodora 19: 201 (1917)
subspecies :
subsp. sitchense
(Rupr.) Tzvelev; Sosud. Rast. Sovet. Dal´nego Vostoka 5: 73 (1991)
Distribution: Alaska (Sitka Isl.)
70 ASK
Synonyms: hidden
= Athyrium sitchense (Rupr.) Tzvelev; Novosti Sist. Vyssh. Rast. 26: 8 (1989)
= Athyrium filix-femina var. sitchense Rupr.; Distr. Crypt. Vasc. Ross. 41 (1845)
species :
Athyrium fimbriatum
Wall. ex T.Moore; Index Fil. [T. Moore] 185 (1860)
Distribution: China (Yunnan, Sichuan, Hunan); Tibet; Nepal; Bhutan; India (Himachal Pradesh, Uttarakhand, Sikkim, Darjeeling, Arunachal Pradesh, Meghalaya, Nagaland, West Bengal); Myanmar (Kachin, etc.)
36 CHI TIB 40 BHU IND NEP 41 MYA
Remarks: sect. Polystichoides
Synonyms: hidden
= Araiostegia yaklaensis (Bedd.) B.K.Nayar & S.Kaur; Companion Ferns Brit. India 17 (1974)
= Aspidium fimbriatum Wall.; Numer. List. [Wallich] no. 339 (1829), nomen, non Willd. 1810
= Asplenium fimbriatum var. foliolosum (Wall.) C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 495 (1880)
= Asplenium fimbriatum Wall. ex Hook.; Sp. Fil. [W. J. Hooker] 3: 234 (1860), nom. illeg.
= Asplenium foliolosum (C.B.Clarke) C.Hope; J. Bombay Nat. Hist. Soc. 14: 255 (1902), nom. illeg.
= Athyrium bucahwangense Ching; Fl. Xizangica 1: 132 (1983) (fide Fraser-Jenkins et al. 2015)
= Athyrium decompositum Ching; Sporae Pterid. Sin. 215, t. 44, f. 5-6 (1976), nom. nud.
= Athyrium fimbriatum (Wall. ex Hook.) Bedd.; Ferns Brit. India t. 295 (1866), nom. superfl.
= Athyrium fimbriatum var. foliolosa Wall.; Numer. List. [Wallich] no. 359 (1829), nomen
= Athyrium fimbriatum var. foliolosum (C.B.Clarke) Bedd.; Ferns Brit. India 172 (1880)
= Athyrium fimbriatum var. squamatum Bedd.; Suppl. Ferns Brit. India 37 (1892)
= Athyrium foliolosum (C.B.Clarke) Bedd.; Suppl. Ferns Brit. Ind. 37 (1892), nom. illeg.
= Athyrium yaklaense (Bedd.) Panigrahi & Basu; (1984) (fide Fraser-Jenkins 2008)
= Davallia yaklaensis (Bedd.) C.Chr.; Index Filic. 215 (1905) (fide Fraser-Jenkins 2008)
= Leucostegia yaklaensis Bedd.; Suppl. Ferns Brit. Ind. 13 (1892) (fide Fraser-Jenkins 2008)
= Polystichum fimbriatum Wall. ex C.Presl; Abh. Königl. Böhm. Ges. Wiss., ser. 5, 6 [Epim. Bot.]: 58 (1851)
species :
Athyrium flabellulatum
(C.B.Clarke) Tardieu; Aspl. Tonkin 81, t. 12 (1932)
Distribution: Tibet; Nepal; Bhutan; India (Himachal Pradesh, Uttarakhand, Sikkim, Darjeeling, Arunachal Pradesh, West Bengal); Myanmar; Vietnam
36 TIB 40 BHU IND NEP 41 MYA VIE
Synonyms: hidden
= Asplenium filix-femina var. flabellulatum C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: t. 60 (1880)
= Athyrium filix-femina var. deltoidea Christ; (1905)
= Athyrium filix-femina var. flabellulatum (C.B.Clarke) Bedd.; Ferns Brit. India 170 (1883)
= Athyrium tenuifolium Y.T.Hsieh & C.R.Wang; Fl. Xizangica 1: 141 (1983), non Copel. 1929
species :
Athyrium foliolosum
(Wall.) T.Moore ex R.Sim; Cat. Brit. Ferns 6: 22 (1859)
Distribution: China (Yunnan); Tibet; Nepal; Bhutan; India (Arunachal Pradesh, Himachal Pradesh, ?Karnataka, Kerala, Manipur, Meghalaya, Mizoram, Nagaland, Rajasthan, Sikkim, Tamil Nadu, Uttarakhand, West Bengal); N-Myanmar (Kachin, etc.); Vietnam
36 CHI TIB 40 BHU IND NEP 41 MYA VIE
Remarks: sect. Polystichoides. Not in Taiwan
Synonyms: hidden
= Aspidium foliolosum Wall.; Numer. List. [Wallich] no. 359 (1829), nomen
= Aspidium squarrosum Wall.; Numer. List. [Wallich] no. 356 (1829), nomen, non Don 1825
= Asplenium decipiens Mett.; Abh. Senckenberg. Naturf. Ges. 3(1) [Aspl.]: 195, no. 237 (1859) (non Zipp. ex Kunze 1849, nec Kuhn 1879)
= Asplenium foliolosum (T.Moore ex R.Sim) E.J.Lowe; Nat. Hist. Ferns: 107 (1861)
= Asplenium foliolosum Wall.; Numer. List. [Wallich] no. 2205 (1829), nomen
= Athyrium austroyunnanense Ching; Acta Bot. Austro Sin. 1: 19 (1983), et Fl. Xizangica 1: 132 (1983) (fide Fraser-Jenkins 1997)
= Athyrium submacrocarpum Ching & S.K.Wu; Fl. Xizangica 1: 131 (1983)
= Polypodium foliolosum Wall.; Numer. List. [Wallich] no. 359 (1824), nomen
species :
Athyrium guangnanense
Ching; Acta Bot. Boreal.-Occid. Sin. 6(2): 103 (1986), nom. nov.
Distribution: China (SE-Yunnan, Guizhou)
36 CHI
Synonyms: hidden
= Athyrium acrocarpum W.M.Chu; Acta Bot. Yunnan., Suppl. 5: 41 (1992) (non (Rosenst.) Copel. 1951)
= Athyrium guangnanense var. acrocarpum W.M.Chu; Fl. Yunnanica 20: 395 (2006)
= Athyrium yunnanicum Ching; Bull. Fan Mem. Inst. Biol. Bot., ser. 2, 1: 278 (1949) (non Christ 1907)
species :
Athyrium gymnogrammoides
(Klotzsch ex Mett.) Bedd.; Ferns S. India 52 (1864)
Distribution: S-India (Kerala, Tamil Nadu); Sri Lanka
40 IND SRL
Synonyms: hidden
= Allantodia gymnogrammoides (Klotzsch ex Mett.) Ching; Acta Phytotax. Sin. 9: 54 (1964)
= Allantodia solenopteris var. pusilla Kunze; Linnaea 24: 267 (1851)
= Asplenium gymnogrammoides Klotzsch ex Mett.; Abh. Senckenberg. Naturf. Ges. 3(1) [Aspl.]: 193, no. 234, t. 16, f. 13-14 (1859)
= Athyrium solenopteris var. pusillum (Kunze) T.Moore; Index Fil. [T. Moore] 187 (1860)
species :
Athyrium hahajimense
Nakai; Bot. Mag. (Tokyo) 43: 4 (1929)
Distribution: Bonin Isl. (Hahajima)
38 OGA
species :
Athyrium hainanense
Ching; Acta Bot. Boreal.-Occid. Sin. 6(2): 101 (1986)
Distribution: China (Hainan)
36 CHI
Remarks: sect. Strigoathyrium, arisanense agg.
species :
Athyrium haleakalae
K.R.Wood & W.L.Wagner; PhytoKeys 76: 115-124 (2017)
Distribution: Hawaii (Maui, Kauai)
63 HAW
species :
Athyrium himalaicum
Ching ex Mehra & Bir; Res. Bull. Panjab Univ. Sci. 15: 137, f. 26-29 (1964)
Distribution: China (SW-Sichuan, N-Guizhou, NW-Yunnan); Tibet; Nepal; Bhutan; N-India (Sikkim, Uttarakhand, Arunachal Pradesh); Myanmar (Kachin)
36 CHI TIB 40 BHU IND NEP 41 MYA
Synonyms: hidden
= Athyrium birii Ching ex Mehra & Bir; Amer. Fern J. 50: 276-295 (1960) (nom. nud., non (Ching ex Bir) Seriz., fide Fraser-Jenkins 1997)
= Athyrium suprapubescens Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 10 (1986) (fide Fraser-Jenkins 2018)
species :
Athyrium hirtirachis
Ching & Y.P.Hsu; Fl. Tsinling. 2: 215 (1974)
Distribution: China (Sichuan, SE-Gansu)
36 CHI
Synonyms: hidden
= Athyrium gonggaense Z.R.Wang & Li Bing Zhang; Li Bing Zhang, Z. R. Wang & X. S. Guo, Acta Bot. Yunnan. 17(3): 301- 303, t. 1, f. 5-8 (1995)
species :
Athyrium hohenackerianum
(Kunze) T.Moore; Index Fil. [T. Moore] 49 (1857)
Distribution: India (Andhra Pradesh, Chhattisgarh, Goa, Gujarat, Karnataka, Kerala, Madhya Pradesh, Maharashtra, Odisha, Rajasthan, Tamil Nadu); Sri Lanka
40 IND SRL
Synonyms: hidden
= Allantodia hohenackeriana (Kunze) Kunze; Farnkräuter 2: 63, t. 126 (1850)
= Asplenium hohenackerianum Kunze; Bot. Zeitung (Berlin) 771 (1849)
species :
Athyrium infrapuberulum
Ching; Acta Bot. Boreal.-Occid. Sin. 6(2): 100 (1986)
Distribution: China (NW-Guangxi)
36 CHI
Remarks: sect. Strigoathyrium, arisanense agg.
species :
Athyrium interjectum
Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 15 (1986)
Distribution: China (NW-Yunnan)
36 CHI
Synonyms: hidden
= Athyrium heqingense Ching; (1986), nom. nud.
= Athyrium acutiserratum Ching; Acta Bot. Boreal.-Occid. Sin. 6(3): 149 (1986)
species :
Athyrium intermixtum
Ching & P.S.Chui; Acta Bot. Boreal.-Occid. Sin. 6(1): 21 (1986)
Distribution: China (Anhui, Zhejiang)
36 CHI
species :
Athyrium iseanum
Rosenst.; Repert. Spec. Nov. Regni Veg. 13: 124 (1913)
Distribution: China (Anhui, Chongqing, Fujian, Guangdong, Guangxi, Guizhou, Hubei, Hunan, Jiangsu, Jiangxi, Sichuan, Yunnan, Zhejiang); Tibet; South Korea (Cheju Isl.); Japan
36 CHI TIB 38 JAP KOR
Synonyms: hidden
= Athyrium filix-femina var. filipes Christ; (1905)
= Athyrium wuyishanense Ching; Acta Bot. Boreal.-Occid. Sin. 6(2): 104 (1986)
variant :
var. angustisectum
Tagawa; Acta Phytotax. Geobot. 2: 15 (1933)
Distribution: Taiwan
38 TAI
Synonyms: hidden
= Athyrium iseanum f. angustisectum (Tagawa) Tagawa; Acta Phytotax. Geobot. 6: 42 (1957)
variant :
var. chuanqianense
Z.R.Wang; Fl. Reipubl. Popularis Sin. [Fl. China Ed. Committee] 3(2): 509 (1999)
Distribution: China (Guangxi, Sichuan, Guizhou, Yunnan); Tibet
36 CHI TIB
species :
Athyrium jinshajiangense
Ching & K.H.Shing; Acta Bot. Boreal.-Occid. Sin. 6(1): 14 (1986)
Distribution: China (C-Yunnan)
36 CHI
species :
Athyrium kandelii
Fraser-Jenk.; Annot. Checkl. Ind. Pterid. 2: 36 (2018), nom. nov.
Distribution: S-China; Nepal; Bhutan; India (Himachal Pradesh, Sikkim, Uttarakhand, Jammu & Kashmir)
36 CHI 40 BHU IND NEP
Synonyms: hidden
= Athyrium mackinnoniorum var. glabratum Y.T.Hsieh & C.R.Wang; Fl. Xizangica 1: 143 (1983)
species :
Athyrium kenzo-satakei
Sa.Kurata; J. Geobot. 7: 75 (1958)
Distribution: China (Guangdong, Guangxi, Sichuan, Guizhou); Japan (incl. Yakushima Isl.)
36 CHI 38 JAP NNS
Remarks: sect. Strigoathyrium, arisanense agg. Very close to A. arisanense and possibly the same.
Conservation Status: CR in Japan
Synonyms: hidden
= Athyrium arisanense var. kenzo-satakei (Sa.Kurata) Seriz.; J. Jap. Bot. 45: 117 (1970)
variant :
var. jieguishanense
(Ching) Z.R.Wang; Fl. Reipubl. Popularis Sin. [Fl. China Ed. Committee] 3(2): 225 (1999)
Distribution: China (Guangxi)
36 CHI
Synonyms: hidden
= Athyrium jieguishanense Ching; Acta Bot. Boreal.-Occid. Sin. 6(3): 157 (1986)
species :
Athyrium khasimontanum
Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 419 (2008)
Distribution: India (Meghalaya)
40 IND
Conservation Status: IUCN CR for India (Fraser-Jenkins 2012)
species :
Athyrium kirishimaense
Tagawa; Acta Phytotax. Geobot. 2(1): 22 (1932)
Distribution: Japan (Izu Isl.)
38 JAP
Synonyms: hidden
= Athyrium yokoscense var. kirishimaense (Tagawa) Li & J.Z.Wang; Clav. Pl. Chin. Bor.-Or. 36 (1995)
species :
Athyrium kuratae
Seriz.; J. Jap. Bot. 45: 117 (1970)
Distribution: China (Yunnan); S-Japan
36 CHI 38 JAP
Remarks: not in Taiwan. possibly a synonym of A. otophorum
species :
Athyrium kutaiense
C.Chr.; Dansk Bot. Ark. 9(3): 55 (1937)
Distribution: Borneo
42 BOR
Remarks: provisionally accepted name
species :
Athyrium latinervatum
M.Kessler & A.R.Sm.; Brittonia 59(2): 186 (2007)
Distribution: Bolivia (La Paz)
83 BOL
Remarks: known only from type
species :
Athyrium leiopodum
(Hayata) Tagawa; Acta Phytotax. Geobot. 2(3): 195 (1933)
Distribution: Taiwan
38 TAI
Remarks: not included in Fl. of China but confirmed for Taiwan by Knapp (2011) and Liu et al. (2009). Included in A. roseum by Fraser-Jenkins (2008)
Synonyms: hidden
= Diplazium leiopodum Hayata; Icon. Pl. Formosan. 4: 217, f. 148 (1914)
species :
Athyrium lewalleanum
Pic.Serm.; Webbia 27: 440 (1972 publ. 1973)
Distribution: Burundi; Tanzania
23 BUR 25 TAN
species :
Athyrium lineare
Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 11 (1986)
Distribution: China (NW-Yunnan)
36 CHI
species :
Athyrium longius
Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 178 (1940)
Distribution: China (Guizhou, Hunan)
36 CHI
Synonyms: hidden
= Athyrium hengshanense Ching; (1976) (nom. nud.
species :
Athyrium ludingense
Z.R.Wang & Li Bing Zhang; Acta Bot. Yunnan. 17(3): 303, t. 1, f. 1-4 (1995)
Distribution: China (Sichuan)
36 CHI
species :
Athyrium mackinnoniorum
(C.Hope) C.Chr.; Index Filic. 143 (1905) (´mackinnoni´, emend. Fraser-Jenkins 1997)
Distribution: E-Afghanistan; Pakistan (Azad Kashmir); N-India (Himachal Pradesh, Uttarakhand, Jammu & Kashmir)
34 AFG 40 IND PAK
Remarks: all of the Chinese and Indochina records belong to other species fide Fraser-Jenkins 2018, but identity is not yet clear
Synonyms: hidden
= Asplenium mackinnoniorum C.Hope; J. Bot. 34: 124 (1896), as ´mackinnonii´
= Athyrium caudipinna Ching; Acta Bot. Austro Sin. 1: 20 (1983) (fide Fraser-Jenkins 1997)
= Athyrium neomackinnonii Ching; Sporae Pterid. Sin. 211, pl. 43, f. 9-10 (1976), nom. nud.
= Athyrium varians Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(3): 9 (1984)
species :
Athyrium majus
(Makino) Makino; Bot. Mag. (Tokyo) 28: 178 (1914)
Distribution: Japan
38 JAP
Remarks: possibly a synonym of A. wardii
Synonyms: hidden
= Athyrium wardii var. majus Makino; Bot. Mag. (Tokyo) 18: 28 (1899)
species :
Athyrium maoshanense
Ching & P.S.Chiu; Acta Bot. Boreal.-Occid. Sin. 6(3): 157 (1986)
Distribution: China (S-Zhejiang)
36 CHI
species :
Athyrium masamunei
Seriz.; Acta Phytotax. Geobot. 32(5-6): 176 (1981)
Distribution: S-Japan (Yakushima Isl.)
38 JAP NNS
Conservation Status: IUCN R. CR in Japan
species :
Athyrium mearnsianum
(Copel.) Alderw.; Malayan Ferns Fern Allies, Suppl. 279 (1917)
Distribution: Philippines
42 PHI
Remarks: sect. Strigoathyrium, valid species and not a synonym of A. nitidulum fide Wardani et al. 2024
Synonyms: hidden
= Athyrium nigripes var. mearnsianum Copel.; Philipp. J. Sci., C, 3: 211 (1908)
species :
Athyrium medium
(Carmich.) T.Moore; Index Fil. [T. Moore] 186 (1860)
Distribution: Tristan da Cunha (incl. Inaccessible Isl.)
91 TDC
Synonyms: hidden
= Aspidium medium Carmich.; Trans. Linn. Soc. London 12: 611 (1818)
= Asplenium medium (Carmich.) Hook.; Sp. Fil. [W. J. Hooker] 3: 227 (1860)
species :
Athyrium mehrae
Bir; Nova Hedwigia 4: 165, t. 52 (1962)
Distribution: China (Gansu, Hubei, Sichuan, Yunnan); Tibet; Nepal; Bhutan; N-India (Sikkim, Uttarakhand, Arunachal Pradesh); N-Myanmar
36 CHI TIB 40 BHU IND NEP 41 MYA
Remarks: bona sp. and not a synonym of A. nephrodioides fide Fraser-Jenkins et al. (2015)
Synonyms: hidden
= Athyrium stenopodum Ching & S.K.Wu; Fl. Xizangica 1: 137 (1983) (fide Fraser-Jenkins 2008)
= Athyrium tibeticum Ching; Fl. Xizangica 1: 137 (1983) (fide Fraser-Jenkins 2008)
species :
Athyrium melanolepis
(Franch. & Sav.) Christ; Bull. Herb. Boissier, sér. 1, 4: 668 (1896)
Distribution: China (Heilongjiang, Jilin); Japan
36 CHI 38 JAP
Synonyms: hidden
= Asplenium melanolepis Franch. & Sav.; Enum. Pl. Jap. 2: 226 (1877)
= Athyrium filix-femina subsp. melanolepis (Franch. & Sav.) Á.Löve & D.Löve; Taxon 26(2-3): 326 (1977)
= Athyrium filix-femina var. melanolepis (Franch. & Sav.) Makino; (1899)
= Athyrium filix-femina var. nigropaleaceum Makino; Bot. Mag. (Tokyo) 13: 29 (1899)
species :
Athyrium mengtzeense
Hieron.; Hedwigia 59: 319 (1918), nom. nov.
Distribution: China (Yunnan, W-Guizhou, SW-Sichuan); Nepal; India (West Bengal, Sikkim, Arunachal Pradesh, Uttarakhand)
36 CHI 40 IND NEP
Remarks: kept separate from A. roseum by Flora of China and also by Fraser-Jenkins et al. (2015)
Synonyms: hidden
= Asplenium sinense Baker; Bull. Misc. Inform. Kew 1906: 9 (1906)
= Athyrium confertipinnum Ching; Acta Bot. Boreal.-Occid. Sin. 6(3): 155 (1986)
= Athyrium liangwangshanicum Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 21 (1986)
= Athyrium sinense (Baker) C.Chr.; Index Filic., Suppl. 2 (1906-1912): 15 (1913) (non Rupr. (1845))
= Athyrium tenuicaule (Hayata) Tagawa; Acta Phytotax. Geobot. 2(3): 195 (1933) (included in A. roseum by Fraser-Jenkins (2008))
= Athyrium thelypteroides var. henryi Christ; Bull. Herb. Boissier, sér. 1, 6: 961 (1898)
= Diplazium tenuicaule Hayata; Icon. Pl. Formosan. 4: 220, f. 150 (1914) (included in A. roseum by Fraser-Jenkins (2008))
species :
Athyrium microphyllum
(Sm.) Alston; Philipp. J. Sci. 50: 178 (descr. orig.) (1933)
Distribution: Hawaii (Kauai, Oahu, Molokai, Lanai, Maui, Hawaii Isl.)
63 HAW
Synonyms: hidden
= Asplenium baldwinii Hillebr.; Fl. Hawaiian Isl. [Hillebrand] 618 (1888)
= Asplenium multisectum Brack.; U.S. Expl. Exped., Filic. 16: 174 (1854)
= Asplenium poiretianum Gaudich.; Voy. Uranie [Freycinet], Bot. 321, t. 13 (1827)
= Asplenium vexans Underw. ex A.Heller; Minnesota Bot. Stud. 1: 776 (1897)
= Athyrium baldwinii (Hillebr.) C.Chr.; Index Filic. 140 (1905)
= Athyrium poiretianum (Gaudich.) C.Presl; Tent. Pterid. 98 (1836)
= Darea microphylla Sm.; Cycl. [A. Rees] (London ed.) 11, no. 9 (1808)
species :
Athyrium micropterum
Fraser-Jenk.; New Species Syndr. Indian Pteridol. 58 (1997)
Distribution: China (Himalayas); Tibet; Nepal; Bhutan; India (Uttarakhand, Himachal Pradesh, Sikkim, Darjeeling, Arunachal Pradesh, Nagaland, West Bengal)
36 CHI TIB 40 BHU IND NEP
Synonyms: hidden
= Asplenium macrocarpon var. atkinsonii C.B.Clarke ex Baker; Syn. Fil. [W. J. Hooker & Baker], ed. 2: 489 (1874)
= Athyrium macrocarpon var. atkinsonii (C.B.Clarke ex Baker) Tardieu; (1940)
species :
Athyrium minimum
Ching; Acta Bot. Boreal.-Occid. Sin. 6(3): 151 (1986)
Distribution: Japan; Taiwan
38 JAP TAI
Conservation Status: EN in Taiwan
species :
Athyrium minutum
Copel.; Univ. Calif. Publ. Bot. 18: 221 (1942)
Distribution: New Guinea (Irian Jaya, Papua New Guinea)
42 NWG
species :
Athyrium mohillense
(Fée) Tardieu; Bull. Mus. Natl. Hist. Nat., sér. 2, 29: 290 (1957)
Distribution: Mauritius; La Réunion
29 MAU REU
Synonyms: hidden
= Asplenium mohillense (Fée) Kuhn; Filic. Afr. 107 (1868)
= Diplazium mohillense Fée; Mém. Foug. 8 (Icon. Esp. Nouv.): 85 (1857)
species :
Athyrium monomachii
(Kom.) Kom.; Izv. Kievsk. Bot. Sada 13: 145 (1931)
Distribution: Siberia (Altai, Buryatia, Chita, Irkutsk, Krasnoyarsk, Yakutia); Russian Far East (Amur, Kamchatka, Khabarovsk, Magadan, Primorye); China (Hebei, Heilongjiang, Jilin, Liaoning, Nei Mongol, N-Shandong, Shanxi); Mongolia; North Korea; South Korea; Japan
30 SIB 31 RFE 36 CHI 37 MON 38 JAP KOR NKO
Remarks: included in A. sinense sensu lato by Fraser-Jenkins (2008) but kept separate by Flora of China
Synonyms: hidden
= Aspidium filix-femina var. multidentatum Döll; Gefässkrypt. Baden 24 (1855)
= Athyrium acutidentatum Ching; Acta Bot. Boreal.-Occid. Sin. 6(3): 150 (1986)
= Athyrium brevifrons Nakai ex Kitag.; Rep. First Sci. Exped. Manchoukuo 4(2): 75 (1935) [fide Fraser-Jenkins (2008)]
= Athyrium brevifrons var. angustifrons (Kodama) T.Mori; Enum. Pl. Corea: 4 (1922)
= Athyrium changbaishanense Ching & J.J.Chien; Bull. Bot. Res., Harbin 4(1): 128 (1984) ex Vorosch. (nom. inval.)
= Athyrium dailingense Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 18 (1986)
= Athyrium extensum Ching; (1976), nom. nud.
= Athyrium filix-femina subsp. jacutense (Petrov) Sipliv.; Novosti Sist. Vyssh. Rast. 11: 328 (1974)
= Athyrium filix-femina subsp. longipes (H.Hara) Tzvelev; Sosud. Rast. Sovet. Dal´nego Vostoka 5: 73 (1991)
= Athyrium filix-femina var. jacutense Petrov; Fl. Iakut. 1: 18 (1930)
= Athyrium filix-femina var. longipes H.Hara; Bot. Mag. (Tokyo) 48: 691 (1934) (non Athyrium longipes Christ)
= Athyrium filix-femina var. monomachii Kom.; Izv. Imp. Bot. Sada Petra Velikago 16: 149 (1916)
= Athyrium hebeiense Ching; Acta Bot. Boreal.-Occid. Sin. 6(3): 158 (1986)
= Athyrium melanolepis var. angustifrons Kodama; (1917)
= Athyrium multidentatum (Döll) Ching; Wang et al., Fl. Pl. Herb. Chin. Bor.-Or. 1: 40. (1958)
= Athyrium sinense f. multidentatum (Döll) Kitag.; Neo-Lineam. Fl. Manshur.: 32 (1979)
= Athyrium sinense subsp. longipes (H.Hara) Vorosch.; Florist. Issl. Razn. Raionakh SSSR [A. K. Svortsov]: 144 (1985)
= Athyrium submelanolepis Tzvelev; Novosti Sist. Vyssh. Rast. 26: 8 (1989), nom. nov.
species :
Athyrium morobense
Copel.; Philipp. J. Sci. 78: 464, t. 40 (1951)
Distribution: New Guinea (Papua New Guinea)
42 NWG
species :
Athyrium multipinnum
Y.T.Hsieh & Z.B.Wang; Acta Bot. Boreal.-Occid. Sin. 7(1): 55 (1987)
Distribution: China (Guizhou, Hunan, Jiangxi, Zhejiang)
36 CHI
species :
Athyrium myer-dreesii
Copel.; Univ. Calif. Publ. Bot. 18: 222 (1942)
Distribution: Moluccas (Seram); New Guinea (Irian Jaya)
42 MOL NWG
species :
Athyrium nakanoi
Makino; Bot. Mag. (Tokyo) 23: 247 (1909)
Distribution: China (Yunnan); Tibet; Japan (Yakushima Isl.); Taiwan; Bhutan; N-India (Arunachal Pradesh, Sikkim); Myanmar (Kachin); Philippines (Mindanao)
36 CHI TIB 38 JAP NNS TAI 40 BHU IND 41 MYA 42 PHI
Remarks: sect. Polystichoides, anisopterum clade. Not in Nepal
Conservation Status: IUCN NT for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Asplenium macrocarpon var. unipinnatum C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 489 (1880)
= Athyrium macrocarpon var. unipinnatum (C.B.Clarke) Bedd.; Ferns Brit. India 165 (1883)
= Athyrium obtusifolium Rosenst.; Hedwigia 66: 335 (1915)
= Athyrium subfalcatum Ching; (1976), nom. nud.
= Athyrium tenuissimum (Hayata) Kodama; Gen. Ind. Fl. Taiwan 4: 202 (1914 publ. 1915)
= Athyrium tenuissimum (Hayata) Merr.; Philipp. J. Sci., C, 13: 126 (1918), isonym
= Nephrolepis tenuissima Hayata; Icon. Pl. Formosan. 4: 202, f. 137 (1914)
species :
Athyrium nanyueense
Ching; Acta Bot. Boreal.-Occid. Sin. 6(3): 152 (1986)
Distribution: China (Hunan)
36 CHI
Remarks: probably a synonym of A. distans fide Fraser-Jenkins et al. 2018
species :
Athyrium neglectum
Seriz.; J. Jap. Bot. 55(8): 250 (1980)
Distribution: Japan
38 JAP
Conservation Status: CR in Japan
species :
Athyrium nephrodioides
(Baker) Christ; Mém. Soc. Bot. France 1 [Hist. Acrostich.]: 47 (1905)
Distribution: China
36 CHI
Remarks: status unclear, possibly a synonym or aberrant specimen of A. mehrae
Synonyms: hidden
= Asplenium nephrodioides Baker; J. Bot. 25: 170 (1887)
species :
Athyrium newtonii
(Baker) Diels; Nat. Pflanzenfam. [Engler & Prantl] I, 4: 224 (1899)
Distribution: Guinea; Liberia; São Tomé; D.R.Congo [Zaire]; Rwanda; Burundi; Sudan; Uganda; Kenya; Tanzania; Malawi; E-Zimbabwe; Mozambique; South Africa (Limpopo, Mpulamanga, KwaZulu-Natal); Lesotho; Eswatini [Swaziland]; ?Madagascar
22 GUI LBR 23 BUR RWA SAO ZAI 24 SUD 25 KEN TAN UGA 26 MLW MOZ ZIM 27 LES SAF SWZ
Remarks: sect. Strigoathyrium, scandicinum agg.
Synonyms: hidden
= Allantodia aspidioides (Willd.) Kunze; Bot. Zeitung (Berlin) 6: 191 (1848)
= Aspidium aspidioides (Willd.) Christ; Farnkr. Erde: 224, 225 (1897)
= Asplenium aspidioides (Willd.) Schltdl.; Adumbr. Pl. 24, t. 13 (1825)
= Asplenium aspidioides (Willd.) Spreng.; Syst. Veg., ed. 16 [Sprengel] 4(1): 90 (1827), isonym
= Asplenium laxum (Pappe & Rawson) Kuhn; Filic. Afr. 105 (1868) (non R. Br. (1810))
= Asplenium newtonii Baker; Ann. Bot. (Oxford) 6: 307 (1891)
= Athyrium aspidioides (Willd.) Christ; Farnkr. Erde 224 (1897)
= Athyrium laxum Pappe & Rawson; Syn. Filic. Afr. Austr. 16 (1858) (non Schumach. (1803))
= Athyrium scandicinum subsp. newtonii (Baker) Verdc.; Fl. Trop. E. Africa, Woods. 10 (2003)
= Athyrium scandicinum var. rhodesianum Schelpe; Bol. Soc. Brot., ser. 2(41): 211 (1967)
= Caenopteris aspidioides (Willd.) Desv.; Mem. Soc. Linn. Paris 6(2): 268 (1827)
= Darea aspidioides Willd.; Enum. Pl. Hort. Berol. 1072 (1809)
= Dryopteris schnellii Tardieu; Notul. Syst. (Paris) 13: 370 (1949)
species :
Athyrium nigripes
(Blume) T.Moore; Index Fil. [T. Moore] 49, 98 (1858)
Distribution: Sulawesi; Philippines (Luzon); Java; Lesser Sunda Isl. (Bali, Flores, Timor); New Guinea (Irian Jaya, Papua New Guinea)
42 JAW LSI NWG PHI SUL
Remarks: sect. Strigoathyrium, nigripes clade
Synonyms: hidden
= Allantodia paludosa Zippel; Kunze, Bot. Zeitung (Berlin) 191 (1848)
= Aspidium costale Blume; Enum. Pl. Javae 2: 170 (1828)
= Aspidium nigripes Blume; Enum. Pl. Javae 2: 162 (1828)
= Asplenium denticulatum J.Sm.; Cat. Cult. Ferns: 47 (1857), nom. illeg.
= Asplenium incisum J.Sm.; J. Bot. (Hooker) 4: 174 (1841), nom. illeg.
= Asplenium nigripes (Blume) Hook.; Sp. Fil. [W. J. Hooker] 3: 222 (1860), nom. illeg.
= Asplenium paludosum T.Moore; Index Fil. [T. Moore] 162 (1859)
= Athyrium aristulatum Copel.; Philipp. J. Sci. 1, Suppl. IV: 253 (1906)
= Athyrium purpureum Lowe ex Milde; Fil. Eur. 49 (1867)
species :
Athyrium nikkoense
Makino; Bot. Mag. (Tokyo) 17: 43 (1903)
Distribution: Japan
38 JAP
species :
Athyrium nitidulum
(Kunze) Milde; Bot. Zeitung (Berlin) 370 (1870)
Distribution: Myanmar (Kachin); Philippines; Java
41 MYA 42 JAW PHI
Remarks: sect. Strigoathyrium, non-setose group, erythropodum clade
Synonyms: hidden
= Allantodia nitidula Kunze; Bot. Zeitung (Berlin) 191 (1848)
= Asplenium nitidulum (Kunze) T.Moore; Index Fil. [T. Moore] 43 (1857)
= Athyrium philippinense Christ; Copel., Philipp. J. Sci. 1, Suppl. IV: 254 (1906)
= Athyrium sarasinorum var. philippinense Christ; Bull. Herb. Boissier, sér. 1, 6: 154 (1898)
species :
Athyrium oblitescens
Sa.Kurata; J. Geobot. 17: 3 (1969)
Distribution: Japan
38 JAP
Remarks: sect. Strigoathyrium, non-setose group
species :
Athyrium obtusilimbum
Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 11 (1986)
Distribution: China (NW-Yunnan)
36 CHI
species :
Athyrium omeiense
Ching; Bull. Fan Mem. Inst. Biol. Bot., ser. 2, 1: 282 (1949)
Distribution: China (Chongqing, SE-Gansu, NW-Guizhou, NW-Hubei, Hunan, Shaanxi, Sichuan, N-Yunnan)
36 CHI
Synonyms: hidden
= Athyrium amplissimum Ching, Boufford & K.H.Shing; J. Arnold Arbor. 64(1): 20 (1983) (non (Baker) Holttum 1955)
= Athyrium crassiusculum Ching; (1976), nom. nud.
= Athyrium filix-femina var. flavicoma Christ; (1905)
= Athyrium flavicoma (Christ) Ching; (1986)
= Athyrium magnum Ching; (1976), nom. nud.
= Athyrium parapellucidum Ching; Fl. Tsinling. 2: 218 (1974)
= Athyrium pellucidum Ching; Fl. Tsinling. 2: 217 (1974)
= Athyrium subtripinnatum Ching; (1976), nom. nud.
species :
Athyrium oosorum
(Baker) Christ; Bot. Jahrb. Syst. 28: 349 (1896)
Distribution: [Western] Samoa (Savai´i)
60 SAM
Synonyms: hidden
= Asplenium oosorum Baker; J. Bot. 14: 343 (1876)
species :
Athyrium oppositipennum
Hayata; J. Coll. Sci. Imp. Univ. Tokyo 30: 441 (1911) (oppositipinnum auct.)
Distribution: Taiwan; Philippines (Luzon)
38 TAI 42 PHI
Remarks: not in Japan
Synonyms: hidden
= Athyrium oreopteris Copel.; Philipp. J. Sci. 40: 302, t. 5 (1929)
= Athyrium taiwanense Tagawa; Acta Phytotax. Geobot. 2: 18 (1932)
= Dryopteris oppositipenna (Hayata) Hayata; J. Coll. Sci. Imp. Univ. Tokyo 30: 450 (1911)
variant :
var. pubescens
(Tagawa) Tagawa; Acta Phytotax. Geobot. 4: 142-143 (1935)
Distribution: Japan (Kyushu, Yakushima Isl.); Taiwan
38 JAP NNS TAI
Synonyms: hidden
= Athyrium oppositipennum var. pubescens (Tagawa) Y.C.Liu, W.L.Chiou & H.Y.Liu; Fern Fl. Taiwan Athyrium 36 (2009), isonym (isonym)
= Athyrium reflexipinnum Hayata; Icon. Pl. Formosan. 4: 234, t. 164 (1914)
= Athyrium taiwanense var. pubescens Tagawa; Acta Phytotax. Geobot. 2: 18 (1933)
species :
Athyrium otophorum
(Miq.) Koidz.; Fl. Symb. Orient.-Asiat. 40 (1930)
Distribution: China (Anhui, Chongqing, Fujian, Guangdong, Guangxi, Guizhou, Hubei, Hunan, Sichuan, Yunnan, Zhejiang); South Korea (Cheju Isl.); Japan; Taiwan; N-India (Uttarakhand, Pithoragarh, Banlekh, Champawat, Meghalaya)
36 CHI 38 JAP KOR TAI 40 IND
Remarks: sect. Strigoathyrium, non-setose group
Conservation Status: IUCN EN for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Asplenium otophorum Miq.; Ann. Mus. Bot. Lugduno-Batavi 8: 175 (1867)
= Athyrium erythrocaulon Ching; Acta Bot. Boreal.-Occid. Sin. 6(3): 154 (1986)
= Athyrium otophorum f. okanum (Sa.Kurata) Ebihara; Bull. Nat. Mus. Nat. Sci., Ser. B, 43(1): 19 (2017)
= Athyrium otophorum var. okanum Sa.Kurata; J. Geobot. 17: 3 (1969)
= Athyrium rigescens Makino; Bot. Mag. (Tokyo) 18: 27 (1899)
= Athyrium sinovidalii Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(3): 8 (1984)
= Athyrium tsingyingense Ching; (1976), nom. nud.
= Athyrium vidalii var. chinense Koidz.
= Athyrium violascens Diels; Bot. Jahrb. Syst. 29: 196 (1900)
= Athyrium wardii Christ; (1905) (non (Hook.) Makino 1857)
= Diplazium otophorum (Miq.) C.Chr.; Index Filic. 236 (1905)
species :
Athyrium pachyphyllum
Ching; Acta Bot. Boreal.-Occid. Sin. 6(2): 102 (1986)
Distribution: China (Hunan, N-Guangxi, S-Guizhou, SE-Yunnan)
36 CHI
Synonyms: hidden
= Athyrium xiangxiense S.F.Wu; Keys Vasc. Pl. Wuling Mts. 563 (1995)
species :
Athyrium palustre
Seriz.; J. Jap. Bot. 45: 264 (1970)
Distribution: Japan; Taiwan
38 JAP TAI
Remarks: included by Fraser-Jenkins et al. (2018) in A. devolii
Conservation Status: EN in Japan
species :
Athyrium parasnathense
(C.B.Clarke) Ching ex Mehra & Bir; Res. Bull. Panjab Univ. Sci. 15: 141 (1964)
Distribution: India (?Chhattisgarh, Gujarat, Jharkhand, Karnataka, Madhya Pradesh, Maharashtra, Odisha, Rajasthan, Tamil Nadu)
40 IND
Synonyms: hidden
= Asplenium filix-femina var. parasnathensis C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 493, t. 61, C 2 (1880)
= Athyrium filix-femina var. parasnathensis (C.B.Clarke) Bedd.; Ferns Brit. India 170 (1883)
= Athyrium pectinatum var. tenellum Bedd.; Suppl. Ferns Brit. India 36 (1892)
species :
Athyrium pectinatum
(Wall. ex C.Hope) C.Presl; Tent. Pterid. 98 (1836)
Distribution: Tibet; Nepal; Bhutan; India (Arunachal Pradesh, Himachal Pradesh, Jharkand, Madhya Pradesh, Manipur, Meghalaya, Mizoram, +Rajasthan, Uttarakhand, West Bengal, Jammu & Kashmir, Sikkim, Darjeeling)
36 TIB 40 BHU IND NEP
Synonyms: hidden
= Asplenium filix-femina var. pectinatum (Wall. ex Mett.) C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 492 (1880)
= Asplenium pectinatum Wall.; Numer. List. [Wallich] no. 231 (1829), nomen
= Asplenium pectinatum Wall. ex C.Hope; J. Bombay Nat. Hist. Soc. 14: 253 (1902)
= Asplenium pectinatum Wall. ex Mett.; Abh. Senckenberg. Naturf. Ges. 3(1) [Aspl.] 197 (1859)
= Athyrium filix-femina subsp. pectinatum (Wall.) Á.Löve & D.Löve; Taxon 26(2-3): 326 (1977) (fide Fraser-Jenkins 1997)
= Athyrium filix-femina var. pectinatum (Wall. ex Mett.) Bedd.; Ferns Brit. India 169 (1883)
= Athyrium pectinatum (Wall. ex Mett.) Bedd.; Ferns S. India 51, t. 154 (1863), non A. pectinatum (Wall. ex Mett.) T. Moore (1860)
species :
Athyrium pinetorum
Tagawa; Acta Phytotax. Geobot. 2(1): 16 (1932)
Distribution: Japan
38 JAP
species :
Athyrium pitcairnense
Copel.; Occas. Pap. Bernice P. Bishop Mus. 14: 60, t. 11 (1938)
Distribution: Pitcairn Isl.
61 PIT
species :
Athyrium praetermissum
Sledge; Ann. Mag. Nat. Hist. XII, 9: 457, t. 16 (1956)
Distribution: S-India (Kerala, Tamil Nadu); Sri Lanka
40 IND SRL
Synonyms: hidden
= Athyrium praetermissum var. tripinnatum Sledge; Ann. Mag. Nat. Hist. XII, 9: 453 (1956)
species :
Athyrium pubicostatum
Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(3): 7 (1984)
Distribution: China (Hubei, Hunan, Guangxi, Sichuan, Chongqing, Guizhou, Yunnan); Taiwan
36 CHI 38 TAI
Remarks: sect. Strigoathyrium, non-setose group
Synonyms: hidden
= Athyrium guizhouense Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 11 (1986)
= Athyrium hirtirachis Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(3): 6 (1984) (non Ching & Y. P. Hsu 1974)
= Athyrium kweichowense Ching; (1976), nom. nud.
= Athyrium pseudoepirachis Ching; Acta Bot. Boreal.-Occid. Sin. 6(3): 156 (1986)
= Athyrium sessile Ching; Acta Bot. Boreal.-Occid. Sin. 6(3): 154 (1986)
= Athyrium simulans Ching; (1976), nom. nud.
= Athyrium subpubicostatum Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(3): 6 (1984)
species :
Athyrium pulcherrimum
Copel.; Philipp. J. Sci. 8: 141, t. 3 (1913)
Distribution: Borneo (Mt. Kinabalu, Sabah); Java
42 BOR JAW
Remarks: sect. Strigoathyrium, nigripes clade
species :
Athyrium puncticaule
(Blume) T.Moore; Index Fil. [T. Moore] 186 (1860)
Distribution: Japan; Taiwan; India (Arunachal Pradesh, Kerala, Meghalaya, Nagaland, Tamil Nadu); Sri Lanka; Myanmar (Kachin, etc.); Thailand; Vietnam; Sumatra; Sulawesi; Philippines (Luzon); Java; Lesser Sunda Isl. (Lombok); Moluccas (Seram)
38 JAP TAI 40 IND SRL 41 MYA THA VIE 42 JAW LSI MOL PHI SUL SUM
Remarks: sect. Polystichoides, anisopterum clade. Confirmed for Taiwan by Knapp (2011) and Liu et al. (2009)
Synonyms: hidden
= Aspidium lanceum Kunze; Bot. Zeitung (Berlin) 473 (1846) (fide Fraser-Jenkins 2008)
= Aspidium macrocarpon Blume; Enum. Pl. Javae 2: 162 (1828)
= Aspidium puncticaule Blume; Enum. Pl. Javae 2: 156 (1828)
= Asplenium fallax Mett.; Abh. Senckenberg. Naturf. Ges. 3(1) [Aspl.]: 194, no. 236 (1859)
= Asplenium macrocarpon (Blume) Hook.; Sp. Fil. [W. J. Hooker] 3: 222 (1860), isonym
= Asplenium macrocarpon (Blume) J.Sm.; Cat. Cult. Ferns: 47 (1857), as ´macrocarpum´
= Asplenium oxyphyllum J.Sm.; J. Bot. 8: 408 (1841) ex Kunze, Bot. Zeitung 1846: 441 (1846), non Hook. 1860
= Asplenium puncticaule (Blume) Mett.; Ann. Mus. Bot. Lugduno-Batavi 2: 240 (1866)
= Athyrium fallax Milde; Fil. Eur. 49 (1867)
= Athyrium halconense Christ; Philipp. J. Sci. 8: 273 (1908)
= Athyrium hochreutineri Christ; Annuaire Conserv. Jard. Bot. Genève 15-16: 197 (1912)
= Athyrium lanceum (Kunze) T.Moore; Index Fil. [T. Moore] 185 (1860) (fide Fraser-Jenkins 2008)
= Athyrium macrocarpum (Blume) Bedd.; Ferns S. India 51, t. 153 (1865)
= Lastrea macrocarpa (Blume) T.Moore; Index Fil. [T. Moore] 95 (1858)
= Nephrodium lanceum (Kunze) T.Moore; Index Fil. [T. Moore] 95 (1858) (fide Fraser-Jenkins 2008)
= Nephrodium sparsum var. membranaceum Baker; Syn. Fil. [W. J. Hooker & Baker], ed. 2: 498 (1874)
= Tarachia oxyphylla (J.Sm.) C.Presl; Abh. Königl. Böhm. Ges. Wiss., ser. 5, 6 [Epim. Bot.]: 77 (1851)
species :
Athyrium repens
(Ching) Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 238 (2008)
Distribution: China (Sichuan); ?Tibet; Nepal; Bhutan; India (Sikkim)
36 CHI 40 BHU IND NEP
Conservation Status: IUCN EN for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Pseudocystopteris repens Ching; Acta Phytotax. Sin. 9: 80 (1964)
species :
Athyrium rhachidosorum
(Hand.-Mazz.) Ching; C. Chr., Index Filic., Suppl. Tert.: 43 (1934)
Distribution: China (Yunnan, SW-Sichuan); Tibet; Myanmar
36 CHI TIB 41 MYA
Remarks: sect. Strigoathyrium, nigripes clade
Synonyms: hidden
= Asplenium rhachidosorum Hand.-Mazz.; Symb. Sin. 6: 33, t. 2, f. 2 (1929)
= Athyrium densisorum X.C.Zhang; Bull. Bot. Res., Harbin 11(3): 8 (1991)
= Athyrium wuliangshanense Ching; Acta Bot. Boreal.-Occid. Sin. 6(3): 153 (1986)
species :
Athyrium rondoense
Verdc.; Fl. Trop. E. Africa, Woods. 12 (2003)
Distribution: Tanzania
25 TAN
species :
Athyrium roseum
Christ; Bull. Herb. Boissier, sér. 1, 6: 961 (1898)
Distribution: China (Yunnan); Nepal; India (Darjeeling); Myanmar (Chin)
36 CHI 40 IND NEP 41 MYA
Remarks: sect. Strigoathyrium, non-setose group
Conservation Status: IUCN CR for India (Fraser-Jenkins 2012)
variant :
var. fugongense
Z.R.Wang; Vasc. Pl. Hengduan Mount. 1: 85 (1993), without latin descr. or type, et Fl. Reipubl. Popularis Sin. [Fl. China Ed. Committee] 3(2): 233 (1999)
Distribution: China (Yunnan)
36 CHI
species :
Athyrium rubripes
(Kom.) Kom.; Izv. Kievsk. Bot. Sada 13: 145 (1931) [fide Fraser-Jenkins (2008)]
Distribution: Siberia (Altai, Buryatia, Chita, Irkutsk, Krasnoyarsk, W-Siberia, Yakutia); Russian Far East (Amur, Kamchatka, Khabarovsk, Kuril Isl., Magadan, Primorye, Sakhalin); ?Kazakhstan; China (NE-Heilongjiang)
30 SIB 31 RFE 36 CHI
Remarks: included in A. sinense sensu lato by Fraser-Jenkins (2008) but kept separate by Flora of China
Synonyms: hidden
= Athyrium filix-femina var. rubripes Kom.; Izv. Imp. Bot. Sada Petra Velikago 16: 149 (1916)
species :
Athyrium ruilicolum
W.M.Chu; Acta Bot. Yunnan., Suppl. 5: 41 (1992), as ´ruilicola´
Distribution: China (Yunnan)
36 CHI
species :
Athyrium rupicola
(C.Hope) C.Chr.; Index Filic. 145 (1905)
Distribution: China (Yunnan, Sichuan); Tibet; Pakistan (Hazara, Baltistan); Nepal; Bhutan; N-India (Himachal Pradesh, Sikkim, Uttarakhand, West Bengal, Jammu & Kashmir); Myanmar (Kachin, etc.)
36 CHI TIB 40 BHU IND NEP PAK 41 MYA
Synonyms: hidden
= Asplenium filix-femina subvar. elongatum C.B.Clarke ex Blanf.; J. Asiat. Soc. Bengal, Pt. 2, Nat. Hist. 57: 306 (1888)
= Asplenium filix-femina var. retusum C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 492 (1880) (p. p.)
= Asplenium rupestre C.Hope; (1899), nom. nud.
= Asplenium rupicola Edgew. ex C.Hope; J. Bombay Nat. Hist. Soc. 12: 531, t. 5 (1899)
= Athyrium falcipinna Ching; Sporae Pterid. Sin. 214, pl. 43, f. 21-22 (1976) (nom. nud. inval.)
= Athyrium filix-femina var. retusum (C.B.Clarke) Bedd.; Ferns Brit. India 170 (1883) (p. p.)
= Athyrium glandulosum Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 19 (1986)
species :
Athyrium sarasinorum
Christ; Verh. Naturf. Ges. Basel 11: 434 (1896)
Distribution: Sulawesi
42 SUL
Synonyms: hidden
= Asplenium sarasinorum (Christ) Copel.; Polypodiaceae Philipp. 87 (1905)
species :
Athyrium satowii
H.Itô; J. Jap. Bot. 27: 337, f. 1-2 (1952)
Distribution: Japan
38 JAP
species :
Athyrium scandicinum
(Willd.) C.Presl; Tent. Pterid. 98 (1836)
Distribution: Madagascar; La Réunion
29 MDG REU
Remarks: sect. Strigoathyrium, scandicinum agg.
Synonyms: hidden
= Allantodia scandicina (Willd.) Kaulf.; Enum. Filic. [Kaulfuss] 179 (1824)
= Aspidium inquinans (Fée) Kuhn; Filic. Afr. 135 (1868)
= Aspidium scandicinum Willd.; Sp. Pl., ed. 4 [Willdenow] 5(1): 286 (1810)
= Asplenium scandicinum (Willd.) A.Heller; Minnesota Bot. Stud. 1: 775 (1897) (non R. Br. (1824))
= Athyrium scandicinum f. bipinnatum Rakotondr.; Bull. Mus. Nat. Hist. Nat., B, Adansonia, ser. 4, 10(4): 375 (1988 publ. 1989)
= Chamaefilix scandicina (Willd.) Farw.; Amer. Midl. Naturalist 12(8): 275 (1931)
= Cystopteris scandicina (Willd.) Desv.; Mem. Soc. Linn. Paris 6(2): 264 (1827)
= Nephrodium inquinans Fée; Mém. Soc. Sci. Nat. Strasbourg 6(1): 43, t. 44, fig. 1 (1866)
= Nephrodium scandicinum (Willd.) Bory; Voy. Indes Or. [Bélanger] 2: 63 (1833)
= Polypodium multifissum Goldm.; Nov. Actorum Acad. Caes. Leop.-Carol. Nat. Cur. 19 (Suppl. 1): 453 (1843)
species :
Athyrium schimperi
Moug. ex Fée; Mém. Foug. 5 (Gen. Filic.): 187 (1852)
Distribution: Ghana; Nigeria; Cameroon; D.R.Congo [Zaire]; Rwanda; Burundi; Sudan; Ethiopia; Uganda; Kenya; Tanzania; Angola; Zambia; Malawi; Zimbabwe; South Africa (Mpulamanga, KwaZulu-Natal, E-Cape Prov.); Lesotho; Madagascar
22 GHA NGA 23 BUR CMN RWA ZAI 24 ETH SUD 25 KEN TAN UGA 26 ANG MLW ZAM ZIM 27 LES SAF 29 MDG
Synonyms: hidden
= Aspidium abyssinicum Döll; Gefässkrypt. Baden 23 (1855)
= Aspidium repens Döll; Jahresber. Mannheimer Vereins Naturk. 28: 39 (1862)
= Asplenium schimperi (Moug. ex Fée) A.Braun; Beitr. Fl. Aethiop. 1: 224 (1867), nom. illeg.
= Athyrium filix-femina var. schimperi (Moug. ex Fée) C.B.Clarke & Baker; (1888)
= Athyrium solenopteris var. madagascarica Bonap.; Notes Ptéridol. 5: 59 (1917)
species :
Athyrium schizochlamys
(Ching) K.Iwats., H.Ohba & S.B.Malla; Bull. Univ. Mus. Univ. Tokyo 31 (Himalayan Plants 1): 313 (1988)
Distribution: China (Sichuan, Yunnan); Tibet; Nepal; Bhutan
36 CHI TIB 40 BHU NEP
Synonyms: hidden
= Athyrium atuntzeense (Ching) Z.R.Wang & Z.R.He; Fl. China 2-3: 498 (2013) (fide Fraser-Jenkins 2018)
= Pseudocystopteris atuntzeensis Ching; Acta Phytotax. Sin. 9: 83 (1964) (fide Fraser-Jenkins 2018)
= Pseudocystopteris lanpingensis Ching; Acta Phytotax. Sin. 9: 82 (1964)
= Pseudocystopteris laterepens Ching; Acta Phytotax. Sin. 9: 82 (1964)
= Pseudocystopteris schizochlamys Ching; Acta Phytotax. Sin. 9: 79 (1964) (fide Fraser-Jenkins 1997)
= Pseudocystopteris sinica Ching; Acta Phytotax. Sin. 9: 81 (1964)
species :
Athyrium sericellum
Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 17 (1986)
Distribution: China (Chongqing)
36 CHI
species :
Athyrium sessilipinnum
X.C.Zhang & R.Wei; Brittonia 68(4): 444 (2016)
Distribution: China (Guangxi)
36 CHI
Remarks: sect. Strigoathyrium, arisanense agg.
species :
Athyrium setiferum
C.Chr.; Index Filic. 146 (1905), nom. nov. for Asplenium tenellum (T. Moore ex R. Sim) C. Hope
Distribution: China (Yunnan, Guangdong); Tibet; Japan; Nepal; Bhutan; India (Arunachal Pradesh, Himachal Pradesh, Kerala, Madhya Pradesh, Manipur, Meghalaya, Sikkim, Tamil Nadu, Uttarakhand, West Bengal); Sri Lanka; Myanmar (Kachin, etc.); N-Thailand; peninsular Malaysia; Sumatra; Borneo; ?Philippines (?Luzon); Moluccas (Seram); New Guinea (Irian Jaya, Papua New Guinea)
36 CHI TIB 38 JAP 40 BHU IND NEP SRL 41 MYA THA 42 BOR MLY MOL NWG SUM
Remarks: included in A. strigillosum by Flora of China but kept separate by Fraser-Jenkins (2008, 2015, 2018). A. tozanense is included here by Fraser-Jenkins et al. (2018)
Synonyms: hidden
= Asplenium gracile D.Don; Prodr. Fl. Nepal. 8 (1825) non E. Fourn. 1872
= Asplenium tenellum (Wall. ex T.Moore ex R.Sim) C.Hope; J. Bombay Nat. Hist. Soc. 12: 259, t. 4 (1899) (non Roxb. (1816) nec Fée (1852))
= Asplenium tenuifrons var. tenellum T.Moore; Index Fil. [T. Moore] 188 (1860), isonym, nom. nov.
= Asplenium tenuifrons var. tenellum Wall. ex T.Moore ex R.Sim; Priced Cat. Stove Greenh. Hardy Exot. Ferns 6: 22 (1859)
= Athyrium medogense X.C.Zhang; Bull. Bot. Res., Harbin 11(3): 11 (1991) (fide Fraser-Jenkins et al. 2018)
= Athyrium supraspinescens C.Chr.; Contr. U. S. Natl. Herb. 297, t. 19 (1931) (fide Fraser-Jenkins et al. 2018)
species :
Athyrium setuligerum
Sa.Kurata; J. Geobot. 15: 6 (1966)
Distribution: Japan
38 JAP
species :
Athyrium silvicola
Tagawa; Acta Phytotax. Geobot. 2(3): 17 (1933)
Distribution: Japan; Taiwan
38 JAP TAI
Remarks: sect. Strigoathyrium, nitidulum-group
species :
Athyrium sinense
Rupr.; Distr. Crypt. Vasc. Ross. 41 (1845)
Distribution: Russian Far East (Primorye); China (SE-Gansu, Hebei, W-Henan, Nei Mongol, Ningxia, Shaanxi, Shandong, Shanxi); Mongolia
31 RFE 36 CHI 37 MON
Synonyms: hidden
= Athyrium filix-femina subsp. sinense (Rupr.) Tzvelev; Novosti Sist. Vyssh. Rast. 32: 182 (2000)
= Athyrium huhsienense Ching & Y.P.Hsu; Fl. Tsinling. 2: 218 (1974)
= Athyrium laochingshanense Ching; (1976), nom. nud.
= Athyrium shaanxiense Ching & Y.T.Hsieh; Acta Bot. Boreal.-Occid. Sin. 6(3): 150 (1986) (fide Fraser-Jenkins (2008)
species :
Athyrium sohayakiense
Nakaike; New Fl. Japan, Pterid. 839 (1992), nom. nov.
Distribution: Japan
38 JAP
Synonyms: hidden
= Athyrium neglectum subsp. australe Seriz.; J. Phytogeogr. Taxon. 33(2): 76 (1985)
species :
Athyrium solenopteris
(Kunze) T.Moore; Index Fil. [T. Moore] 43 (1857), 187 (1860)
Distribution: S-India (Karnataka, Kerala, Tamil Nadu); Sri Lanka; Java; New Guinea (Irian Jaya, Papua New Guinea)
40 IND SRL 42 JAW NWG
Remarks: sect. Strigoathyrium, non-setose group, erythropodum clade
Synonyms: hidden
= Allantodia solenopteris Kunze; Linnaea 24: 266 (1851)
= Asplenium nigripes var. solenopteris (Kunze) C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 491 (1880)
= Asplenium solenopteris (Kunze) Mett.; Abh. Senckenberg. Naturf. Ges. 3(1) [Aspl.]: 240 (1859), as ´selenopteris´
= Athyrium ceylonense (Klotzsch) T.Moore; Index Fil. [T. Moore] 181 (1860), nom. nud. (fide Fraser-Jenkins 2008)
= Athyrium nigripes Bedd.; Ferns S. India t. 157 (1863), non (Blume) T. Moore
= Athyrium pectinatum Bedd.; Ferns S. India t. 156 (1863) (p. p., non Asplenium pectinatum Wall. ex Mett.)
= Athyrium selenopteris (Kunze) Bedd.; Ferns Brit. India 166 (1883), orth. var.
species :
Athyrium spinescens
Sa.Kurata; J. Geobot. 15: 7 (1966)
Distribution: Japan
38 JAP
species :
Athyrium spinulosum
(Maxim.) Milde; Bot. Zeitung (Berlin) 376 (1866)
Distribution: Russian Far East (Amur, Khabarovsk, Primorye, Sakhalin); China (Chongqing, Gansu, Heilongjiang, Henan, Hubei, Jilin, Liaoning, Nei Mongol, Qinghai, Shaanxi, Sichuan, Yunnan); Tibet; North Korea; South Korea; Japan; Nepal; Bhutan; India (Sikkim)
31 RFE 36 CHI TIB 38 JAP KOR NKO 40 BHU IND NEP
Remarks: A. subtriangulare is included here by Fraser-Jenkins (2008, 2015, 2018) but kept separate by Flora of China
Synonyms: hidden
= Asplenium spinulosum (Maxim.) Baker; Syn. Fil. [W. J. Hooker & Baker] 225 (1867)
= Asplenium spinulosum (Maxim.) Miq.; Ann. Mus. Bot. Lugduno-Batavi 3: 174 (1867), isonym
= Asplenium spinulosum C.B.Clarke; (1880), not (Blume) Mett. (1859), nor Miq. (1867)
= Asplenium spinulosum var. subtriangulare (Hook.) C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 487 (1880)
= Asplenium subtriangulare Hook.; Syn. Fil. [W. J. Hooker & Baker] 225 (1867)
= Athyrium hookerianum T.Moore; Index Fil. [T. Moore] 185 (1860)
= Athyrium longipes Christ; Mém. Soc. Bot. France 1 [Hist. Acrostich.]: 48 (1905)
= Athyrium sikkimense (Bir) Á.Löve & D.Löve; Taxon 26(2-3): 326 (1977)
= Athyrium spinulosum var. subtriangulare (Hook.) C.Chr.; Index Filic. 3: 146 (1905)
= Athyrium subtriangulare (Hook.) Bedd.; Ferns Brit. India t. 242 (1868)
= Athyrium subtriangulare var. sikkimense Bir; Nova Hedwigia 4: 168, t. 53, f. 9, 10 (1962)
= Cystopteris spinulosa Maxim.; Mém. Acad. Imp. Sci. St. Pétersb. 9: 340 (1859)
= Pseudocystopteris decipiens Ching & S.K.Wu; Fl. Xizangica 1: 125 (1983)
= Pseudocystopteris longipes (Christ) Ching; Acta Phytotax. Sin. 9: 78 (1964)
= Pseudocystopteris purpurascens Ching & S.K.Wu; Fl. Xizangica 1: 128 (1983)
= Pseudocystopteris reflexipinnula Ching & S.K.Wu; Fl. Xizangica 1: 127 (1983)
= Pseudocystopteris remota Ching; Acta Phytotax. Sin. 9: 84 (1964)
= Pseudocystopteris sikkimensis (Bir) Ching; Acta Phytotax. Sin. 9: 79 (1964)
= Pseudocystopteris sparsa Ching & S.K.Wu; Fl. Xizangica 1: 124 (1983)
= Pseudocystopteris spinulosa (Maxim.) Ching; Acta Phytotax. Sin. 9: 78 (1964)
= Pseudocystopteris spinulosa var. taipaishanensis Ching; Fl. Tsinling. 2: 212 (1974)
= Pseudocystopteris subtriangularis (Hook. ex Baker) Ching; Acta Phytotax. Sin. 9: 78 (1964)
= Pseudocystopteris tibetica Ching; Acta Phytotax. Sin. 9: 80 (1964)
species :
Athyrium strigillosum
(T.Moore ex Lowe) T.Moore ex Salomon; Nomencl. Gefäßkrypt. 112 (1883)
Distribution: China (Jiangxi, Hunan, Guangdong, Guangxi, Sichuan, Guizhou, Yunnan); Tibet; S-Japan (Yakushima Isl.); Taiwan; Nepal; Bhutan; N-India (Jammu & Kashmir, Himachal Pradesh, Meghalaya, Sikkim, Uttarakhand, West Bengal); Myanmar
36 CHI TIB 38 JAP NNS TAI 40 BHU IND NEP 41 MYA
Remarks: sect. Strigoathyrium, strigillosum agg.
Synonyms: hidden
= Allantodia tenella Wall.; (1828) (p. p., nom. nud.)
= Asplenium setulosum J.Sm.; Cat. Ferns Kew 5 (1856)
= Asplenium strigillosum T.Moore ex E.J.Lowe; Ferns 5, t. 36 (1858)
= Asplenium tenuifrons (T.Moore ex R.Sim) Blanf.; J. Asiat. Soc. Bengal, Pt. 2, Nat. Hist. 57: 306 (1888)
= Athyrium denticulatum J.Sm.; Hist. Fil. 328 (1875)
= Athyrium nigripes var. tenuifrons (T.Moore ex R.Sim) Bedd.; Suppl. Ferns Brit. India 33 (1892)
= Athyrium petiolosum Christ; Bull. Acad. Int. Géogr. Bot. 134 (1907), non Milde (1870)
= Athyrium proliferum T.Moore; Gard. Chron. 1866: 288 (1866), nom. illeg.
= Athyrium punethae Panigrahi; Refresher´s Course Utkal Univ. (Bubaneshwar, Orissa) Bot. (1998), nom. nov. superfl., as ´punethii´
= Athyrium radiciferum Ching; (1976), nom. nud.
= Athyrium tenuifrons T.Moore ex R.Sim; Priced Cat. Stove Greenh. Hardy Exot. Ferns 6: 17 (1859)
= Athyrium tenuifrons Wall. ex T.Moore; Index Fil. [T. Moore] 43, 188 (1857)
species :
Athyrium subrigescens
(Hayata) H.Itô; Bot. Mag. (Tokyo) 52: 647 (1938)
Distribution: Japan; Taiwan
38 JAP TAI
Remarks: status disputed, often included as infraspecific taxon in A. delavayi
Synonyms: hidden
= Athyrium delavayi var. subrigescens (Hayata) Yea C.Liu, W.L.Chiou & H.Y.Liu; Fern Fl. Taiwan Athyrium 18 (2009)
= Athyrium elegans Tagawa; Acta Phytotax. Geobot. 3: 33 (1934)
= Athyrium roseum f. pubigerum (Sa.Kurata) Nakaike; Checkl. Lycophytes Ferns Japan: 19 (2013)
= Athyrium subrigescens f. pubigerum Sa.Kurata; Sci. Rep. Yokosuka City Mus. 6: 21 (1961)
= Diplazium subrigescens Hayata; Icon. Pl. Formosan. 4: 219, f. 149 (1914)
species :
Athyrium suprapuberulum
Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 14 (1986)
Distribution: China (NW-Yunnan, Sichuan)
36 CHI
Synonyms: hidden
= Athyrium muliense Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 16 (1986)
species :
Athyrium tashiroi
Tagawa; Acta Phytotax. Geobot. 2(1): 21 (1932)
Distribution: Japan
38 JAP
species :
Athyrium tenuipaleatum
Copel.; Occas. Pap. Bernice P. Bishop Mus. 14: 60, t. 12 (1938)
Distribution: Austral Isl. (Rapa Iti)
61 AUI
species :
Athyrium tozanense
(Hayata) Hayata; J. Coll. Sci. Imp. Univ. Tokyo 30: 451 (1911)
Distribution: China (Yunnan); Taiwan
36 CHI 38 TAI
Remarks: sect. Strigoathyrium, nitidulum group; separate from A. nigripes fide molecular results (Wardani et al. 2024)
Synonyms: hidden
= Asplenium tozanense Hayata; J. Coll. Sci. Imp. Univ. Tokyo 30: 440 (1911)
= Athyrium yui Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 6 (1940) (tentative synonym)
species :
Athyrium triangulare
Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 20: 8 (1915)
Distribution: Sri Lanka; Java
40 SRL 42 JAW
Synonyms: hidden
= Asplenium praetermissum var. erythrorhachis (Bedd.) Sledge; Ann. Mag. Nat. Hist. XII, 9: 459 (1956) (fide Fraser-Jenkins et al. 2018)
= Athyrium gymnogrammoides var. erythrorhachis Bedd.; Suppl. Ferns S. Ind. 12 (1876) (fide Fraser-Jenkins et al. 2018)
species :
Athyrium tripinnatum
Tagawa; Acta Phytotax. Geobot. 6: 163 (1937)
Distribution: Taiwan
38 TAI
Remarks: bona sp. according to Knapp (Ferns of Taiwan) and Liu et al. (2009)
species :
Athyrium uniforme
Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 11 (1986)
Distribution: China (NW-Yunnan)
36 CHI
Synonyms: hidden
= Athyrium bijiangense Y.T.Hsieh & W.M.Chu; Acta Bot. Boreal.-Occid. Sin. 7(1): 52 (1987)
= Athyrium gongshanense Ching; Acta Bot. Boreal.-Occid. Sin. 6(3): 153 (1986)
species :
Athyrium venulosum
Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 13 (1986)
Distribution: China (NW-Hubei)
36 CHI
species :
Athyrium vermae
Fraser-Jenk., Khullar & Pangtey; Taxon. Revis. Indian Subcontinental Pteridophytes 219 (2008)
Distribution: Bhutan; India (Uttarakhand)
40 BHU IND
species :
Athyrium vidalii
(Franch. & Sav.) Nakai; Bot. Mag. (Tokyo) 39: 110 (1925)
Distribution: Russian Far East (Kuril Isl.); China (Anhui, Chongqing, Fujian, Gansu, Guangxi, Guizhou, S-Henan, Hubei, Hunan, W-Jiangxi, Shaanxi, Sichuan, Yunnan, Zhejiang); South Korea (incl. Cheju Isl.); Japan; Taiwan; Vietnam
31 RFE 36 CHI 38 JAP KOR TAI 41 VIE
Remarks: not in India
Synonyms: hidden
= Asplenium vidalii Franch. & Sav.; Enum. Pl. Jap. 2: 229, 624 (1877)
= Athyrium commixtum Koidz.; Bot. Mag. (Tokyo) 39: 13 (1925)
= Athyrium glabrescens Ching; Acta Bot. Boreal.-Occid. Sin. 6(3): 156 (1987)
= Athyrium glabrum Ching; Sporae Pterid. Sin. 205, t. 41, f. 31-32 (1976), nom. nud.
= Athyrium kanghsienense Ching & Y.P.Hsu; Fl. Tsinling. 2: 215 (1974)
= Athyrium mabianense Ching & Y.T.Hsieh; Acta Bot. Boreal.-Occid. Sin. 6(3): 157 (1986)
= Athyrium neowardii Ching; Acta Bot. Boreal.-Occid. Sin. 6(3): 155 (1986)
= Athyrium regulare Koidz.; Bot. Mag. (Tokyo) 88: 111 (1924)
= Athyrium taipaishanense Ching; Fl. Tsinling. 2: 217 (1974)
= Athyrium vidalii f. yamadae (Miyabe & Kudô) Sa.Kurata; Enum. Jap. Pterid.: 281, 338 (1961)
= Athyrium vidalii var. confusum Miyabe & Kudô; Fl. Hokkaido 1: 31 (1930)
= Athyrium vidalii var. yamadae (Miyabe & Kudô) Miyabe & Tatew.; Trans. Sapporo Nat. Hist. Soc. 17: 77 (1957)
= Athyrium wugongshanense Ching & Y.T.Hsieh; Acta Bot. Boreal.-Occid. Sin. 6(3): 155 (1986)
= Athyrium yamadae Miyabe & Kudô; Trans. Sapporo Nat. Hist. Soc. 8: 61 (1921)
variant :
var. amabile
(Ching) Z.R.Wang; Fl. Reipubl. Popularis Sin. [Fl. China Ed. Committee] 3(2): 199 (1999)
Distribution: China (Zhejiang)
36 CHI
Synonyms: hidden
= Athyrium amabile Ching; Acta Bot. Boreal.-Occid. Sin. 6(1): 16 (1986)
species :
Athyrium viridescentipes
Sa.Kurata; J. Geobot. 13: 75 (1965)
Distribution: Japan
38 JAP
Remarks: sect. Strigoathyrium, non-setose group
Conservation Status: IUCN R
species :
Athyrium viviparum
Christ; Bull. Acad. Int. Géogr. Bot. 13 (1910)
Distribution: China (Jiangxi, Hunan, Guangdong, Guangxi, Sichuan, Chongqing, Guizhou, Yunnan)
36 CHI
Remarks: sect. Strigoathyrium, strigillosum agg.
Synonyms: hidden
= Athyrium spinosissimum Ching; Acta Bot. Boreal.-Occid. Sin. 6(3): 155 (1986)
= Athyrium tianzishanense S.F.Wu & L.F.Zhang; Acta Phytotax. Sin. 28(2): 166 (1990)
= Athyrium yindeense Ching; Acta Bot. Boreal.-Occid. Sin. 6(3): 152 (1986)
species :
Athyrium wallichianum
Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 497 (1938), nom. nov.
Distribution: China (Sichuan, NW-Yunnan); Tibet; Pakistan (Swat, Hazara); Nepal; ?Bhutan; India (Jammu & Kashmir, Arunachal Pradesh, Himachal Pradesh, Sikkim, Uttarakhand, West Bengal); Myanmar; Laos
36 CHI TIB 40 IND NEP PAK 41 LAO MYA
Synonyms: hidden
= Aspidium brunonianum Wall.; Numer. List. [Wallich] no. 344 (1829), nomen
= Aspidium brunonianum Wall. ex Mett.; Abh. Senckenberg. Naturf. Ges. 2(2) [Pheg. u. Asp.]: 338 (seors. 54) (1858)
= Dryopteris brunoniana (Wall. ex Mett.) Kuntze; Revis. Gen. Pl. 2: 812 (1891)
= Lastrea brunoniana (Wall. ex Mett.) C.Presl; Tent. Pterid. 76 (1836) (nomen) ex Bedd., Ferns Brit. India t. 37 (1865)
= Nephrodium brunonianum (Wall. ex Mett.) Hook.; Sp. Fil. [W. J. Hooker] 4: 113, t. 251 (1862)
species :
Athyrium wangii
Ching; Bull. Fan Mem. Inst. Biol. Bot., ser. 2, 1: 279 (1949)
Distribution: China (S-Yunnan, Hainan)
36 CHI
species :
Athyrium wardii
(Hook.) Makino; Bot. Mag. (Tokyo) 18: 15 (1899)
Distribution: Russian Far East (Kuril Isl.); China (Anhui, Chongqing, Fujian, Guangxi, Guizhou, Hubei, Hunan, Jiangxi, Sichuan, Yunnan, Zhejiang); South Korea (Cheju Isl.); Japan
31 RFE 36 CHI 38 JAP KOR
Remarks: sect. Strigoathyrium, non-setose group
Synonyms: hidden
= Asplenium wardii Hook.; Sp. Fil. [W. J. Hooker] 3: 189 (1860)
= Athyrium tsussimense Koidz.; Fl. Symb. Orient.-Asiat. 41 (1930)
= Athyrium wardii f. inadae (Tagawa) Ebihara; Bull. Nat. Mus. Nat. Sci., Ser. B, 43(1): 20 (2017)
= Athyrium wardii var. inadae Tagawa; Acta Phytotax. Geobot. 16: 178 (1956)
variant :
var. densipinnum
Z.R.Wang & Li Bing Zhang; Acta Bot. Yunnan. 17(3): 303 (1995)
Distribution: China (Sichuan)
36 CHI
variant :
var. glabratum
Y.T.Hsieh & Z.R.Wang; Fl. Reipubl. Popularis Sin. [Fl. China Ed. Committee] 3(2): 508 (1999)
Distribution: China (Zhejiang, Fujian, Hunan)
36 CHI
species :
Athyrium xichouense
Y.T.Hsieh & Z.R.Wang; Acta Phytotax. Sin. 27(2): 155 (1989)
Distribution: China (Sichuan, Yunnan); Tibet; Nepal; NE-India (Sikkim, Arunachal Pradesh, Meghalaya, Meghalaya, Mizoram, West Bengal)
36 CHI TIB 40 IND NEP
Remarks: kept separate (again) from A. silvicola by Fraser-Jenkins (2018)
species :
Athyrium yakusimense
Tagawa; Acta Phytotax. Geobot. 6: 94 (1937)
Distribution: Japan (Yakushima Isl.)
38 JAP NNS
Conservation Status: CR in Japan
species :
Athyrium yokoscense
(Franch. & Sav.) Christ; Bull. Herb. Boissier, sér. 1, 4: 668 (1896)
Distribution: Russian Far East (Kuril Isl., Primorye); China (Anhui, Chongqing, NE-Guizhou, Heilongjiang, Henan, Hunan, Jiangsu, Jiangxi, Jilin, Liaoning, Shandong, Zhejiang); North Korea; South Korea (incl. Cheju Isl.); Japan
31 RFE 36 CHI 38 JAP KOR NKO
Synonyms: hidden
= Aspidium subspinulosum Christ; Bull. Herb. Boissier, sér. 2, 2: 829 (1902)
= Asplenium yokoscense Franch. & Sav.; Enum. Pl. Jap. 2: 225, 622 (1877)
= Athyrium demissum Christ; Repert. Spec. Nov. Regni Veg. 6: 284 (1908)
= Athyrium flaccidum Christ; Repert. Spec. Nov. Regni Veg. 6: 11 (1908)
= Athyrium pachysorum Christ; Notul. Syst. (Paris) 1: 48 (1909)
= Athyrium subspinulosum (Christ) Ching ex Vorosch.
= Dryopteris subspinulosa (Christ) C.Chr.; Index Filic. 296 (1905)
variant :
var. kirisimaense
(Tagawa) Li & J.Z.Wang; Clav. Pl. Chin. Bor.-Or., ed. 2 (ed. P. Y. Fu): 36 (1995)
Distribution: ?NE-China; Japan
38 JAP
Synonyms: hidden
= Athyrium kirisimaense Tagawa; Acta Phytotax. Geobot. 2: 22 (1933)
species :
Athyrium yuanyangense
Y.T.Hsieh & W.M.Chu; Acta Bot. Boreal.-Occid. Sin. 7(1): 54 (1987)
Distribution: China (S-Yunnan)
36 CHI
species :
Athyrium zayuense
Z.R.Wang; Acta Phytotax. Sin. 32(1): 84 (1994)
Distribution: Tibet (Zayu)
36 TIB
Synonyms: hidden
= Athyrium chayuense Z.R.Wang; Vasc. Pl. Hengduan Mount. 1: 84 (1993) (without latin descr. or type)
species :
Athyrium zhenfengense
Ching; Acta Bot. Boreal.-Occid. Sin. 6(3): 151 (1986)
Distribution: China (SW-Guizhou)
36 CHI
species :
Athyrium ×akiense
Sa.Kurata; J. Geobot. 15: 5 (1966)
Distribution: Japan
38 JAP
Remarks: Athyrium eremicola x otophorum
species :
Athyrium ×amagi-pedis
Sa.Kurata; Sci. Rep. Yokosuka City Mus. 6: 23 (1961)
Distribution: Japan
38 JAP
Remarks: Athyrium otophorum x wardii
species :
Athyrium ×anceps
Sa.Kurata; J. Geobot. 13: 76 (1965)
Distribution: Japan
38 JAP
Remarks: Athyrium arisanense x kenzo-satakei
species :
Athyrium ×austrojaponense
Sa.Kurata; J. Geobot. 16: 99 (1968)
Distribution: Japan
38 JAP
Remarks: Athyrium kuratae x otophorum
species :
Athyrium ×awatae
Seriz.; J. Phytogeogr. Taxon. 28(1): 34 (1980)
Distribution: Japan
38 JAP
Remarks: Athyrium oblitescens x otophorum
species :
Athyrium ×bicolor
Seriz.; J. Jap. Bot. 55(8): 252 (1980)
Distribution: Japan
38 JAP
Remarks: Athyrium neglectum x vidalii
species :
Athyrium ×boreo-occidentali-indobharaticola-birianum
Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 220 (2008)
Distribution: India (Uttarakhand)
40 IND
Remarks: Athyrium pectinatum x schimperi ssp. biserrulatum
species :
Athyrium ×calophyllum
Sa.Kurata ex Seriz.; J. Jap. Bot. 55(8): 254 (1980)
Distribution: Japan
38 JAP
Remarks: Athyrium palustre x vidalii
species :
Athyrium ×decurrentiserratum
K.Hori; Fern Gaz. 21(6): 262 (2021)
Distribution: Japan (Honshu)
38 JAP
species :
Athyrium ×dixitii
Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 225 (2008)
Distribution: India (Uttarakhand)
40 IND
Remarks: Athyrium anisopterum (kumaonicum) x fimbriatum
species :
Athyrium ×drepanopteroides
Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 214 (2008)
Distribution: Nepal; India (Meghalaya, West Bengal)
40 IND NEP
Remarks: Athyrium drepanopterum x eburneum (woodsioides)
species :
Athyrium ×flavosorum
Seriz.; J. Phytogeogr. Taxon. 33(2): 77 (1985)
Distribution: Japan
38 JAP
Remarks: Athyrium arisanense x delavayi
species :
Athyrium ×fuscopaleaceum
Sa.Kurata; Sci. Rep. Yokosuka City Mus. 6: 24 (1961)
Distribution: Japan
38 JAP
Remarks: Athyrium kuratae x clivicola
species :
Athyrium ×gurungae
Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 229 (2008)
Distribution: Nepal
40 NEP
Remarks: Athyrium pectinatum x foliolosum
species :
Athyrium ×hisatsuanum
Sa.Kurata; J. Geobot. 15: 6 (1966)
Distribution: Japan
38 JAP
Remarks: Athyrium clivicola x iseanum
species :
Athyrium ×ikutae
Sa.Kurata; J. Geobot. 16: 33 (1968)
Distribution: Japan
38 JAP
Remarks: Athyrium tozanense x yakusimense
species :
Athyrium ×inabaense
Seriz.; J. Phytogeogr. Taxon. 28(1): 34 (1980)
Distribution: Japan
38 JAP
Remarks: Athyrium iseanum x oblitescens
species :
Athyrium ×inouei
Sa.Kurata; J. Geobot. 17: 89 (1969)
Distribution: Japan
38 JAP
Remarks: Athyrium iseanum x yokoscense
species :
Athyrium ×kathmanduense
Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 211 (2008)
Distribution: China; Nepal; India (Darjeeling, Arunachal Pradesh, West Bengal, Meghalaya)
36 CHI 40 IND NEP
Remarks: Athyrium foliolosum x drepanopterum
Synonyms: hidden
= Athyrium darjeelingensis Fraser-Jenk. ex Subh.Chandra; Ferns India (Enum., Syn. & Dist.) 125 (2000) nom. nud.
species :
Athyrium ×kawabatae
Sa.Kurata; Sci. Rep. Yokosuka City Mus. 6: 22 (1961)
Distribution: Japan
38 JAP
Remarks: Athyrium iseanum x subrigescens
species :
Athyrium ×kawabataeoides
Sa.Kurata; J. Geobot. 16: 33 (1968)
Distribution: Japan
38 JAP
Remarks: Athyrium subrigescens x tozanense
species :
Athyrium ×kidoanum
Sa.Kurata; J. Geobot. 10: 10 (1961)
Distribution: Japan
38 JAP
Remarks: Athyrium iseanum x wardii
species :
Athyrium ×langtangense
Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 222 (2008), nom. inval., sine herbarium, et Annot. Checkl. Ind. Pterid. 2: 60 (2018)
Distribution: Nepal
40 NEP
Remarks: Athyrium anisopterum x micropterum
species :
Athyrium ×lobulosoimpolitum
Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 212 (2008)
Distribution: Nepal; Bhutan; India (Sikkim, Meghalaya, West Bengal)
40 BHU IND NEP
Remarks: Athyrium eburneum (woodsioides) x foliolosum
species :
Athyrium ×manickamii
Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 226 (2008)
Distribution: India (West Bengal)
40 IND
Remarks: Athyrium foliolosum x xichouense
species :
Athyrium ×mantoniae
Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 231 (2008)
Distribution: Sri Lanka
40 SRL
Remarks: Athyrium praetermissum x solenopteris
species :
Athyrium ×masayukianum
Sa.Kurata; J. Geobot. 13: 76 (1965)
Distribution: Japan
38 JAP
Remarks: Athyrium kuratae x iseanum
species :
Athyrium ×megayakusimense
Sa.Kurata; J. Geobot. 14: 5 (1965)
Distribution: Japan
38 JAP
Remarks: Athyrium kuratae x yakusimense
species :
Athyrium ×meghalaicum
Fraser-Jenk.; Annot. Checkl. Ind. Pterid. 2: 60 (2018)
Distribution: India (Meghalaya)
40 IND
Remarks: Athyrium foliolosum x puncticaule
species :
Athyrium ×mentiens
Sa.Kurata; J. Geobot. 16: 34 (1968)
Distribution: Japan
38 JAP
Remarks: Athyrium vidalii x yokoscense
species :
Athyrium ×minakuchii
Sa.Kurata; J. Geobot. 14: 85 (1966)
Distribution: Japan
38 JAP
Remarks: Athyrium otophorum x yokoscense
species :
Athyrium ×multifidum
Rosenst.; Repert. Spec. Nov. Regni Veg. 13: 125 (1914)
Distribution: China
36 CHI
Remarks: Athyrium deltoidofrons x vidalii
Synonyms: hidden
= Athyrium deltoidofrons var. latisectum (Rosenst.) Koidz.; Bot. Mag. (Tokyo) 38(449): 109 (1924)
= Athyrium deltoidofrons var. multifidum (Rosenst.) Koidz.; Bot. Mag. (Tokyo) 38(449): 109 (1924)
= Athyrium ×multifidum f. sakuraii (Rosenst.) Sa.Kurata; J. Geobot. 9: 100 (1961)
= Athyrium ×multifidum var. latisectum Rosenst.; Repert. Spec. Nov. Regni Veg. 13: 126 (1914)
= Athyrium ×multifidum var. sakuraii Rosenst.; Repert. Spec. Nov. Regni Veg. 13: 126 (1914)
species :
Athyrium ×neoelegans
Sa.Kurata; J. Geobot. 14: 4 (1965)
Distribution: Japan
38 JAP
Remarks: Athyrium subrigescens x yakusimense
species :
Athyrium ×nepalense
Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 212 (2008)
Distribution: Nepal; India (West Bengal)
40 IND NEP
Remarks: Athyrium pectinatum x drepanopterum
species :
Athyrium ×paludicola
Sa.Kurata ex Seriz.; J. Jap. Bot. 55: 254 (1980)
Distribution: Japan
38 JAP
Remarks: Athyrium deltoidofrons x palustre
species :
Athyrium ×pichisermollianum
Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 224 (2008)
Distribution: India (Uttarakhand)
40 IND
Remarks: Athyrium anisopterum (kumaonicum) x foliolosum
Synonyms: hidden
= Athyrium ×pangteyi Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes: 222 (2008)
species :
Athyrium ×pokharense
Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 214 (2008)
Distribution: Nepal
40 NEP
Remarks: Athyrium pectinatum x eburneum (woodsioides)
species :
Athyrium ×pseudobipinnatum
K.Hori; Fern Gaz. 21(6): 264 (2021)
Distribution: Japan (Honshu, Kyushu, Shikoku)
38 JAP
species :
Athyrium ×pseudoiseanum
Sa.Kurata; J. Geobot. 8: 84 (1960)
Distribution: Japan (Kyushu)
38 JAP
Remarks: Athyrium iseanum x otophorum
species :
Athyrium ×pseudopinetorum
Seriz.; J. Jap. Bot. 55(8): 253 (1980)
Distribution: Japan
38 JAP
Remarks: Athyrium pinetorum x vidalii
species :
Athyrium ×pseudospinescens
Seriz.; J. Phytogeogr. Taxon. 28(1): 35 (1980)
Distribution: Japan
38 JAP
Remarks: Athyrium iseanum x vidalii
species :
Athyrium ×pseudowardii
Seriz.; J. Phytogeogr. Taxon. 28(1): 35 (1980)
Distribution: Japan
38 JAP
Remarks: Athyrium vidalii x wardii
species :
Athyrium ×purpurascens
(Tagawa) Sa.Kurata; J. Geobot. 14: 4 (1965)
Distribution: Japan
38 JAP
Remarks: Athyrium kuratae (arisanense) x subrigescens
Synonyms: hidden
= Athyrium elegans var. purpurascens Tagawa; Acta Phytotax. Geobot. 3: 33 (1934)
species :
Athyrium ×purpureipes
Sa.Kurata; Sci. Rep. Yokosuka City Mus. 6: 23 (1961)
Distribution: Japan
38 JAP
Remarks: Athyrium clivicola x otophorum
species :
Athyrium ×pygmaei-silvae
Sa.Kurata; Sci. Rep. Yokosuka City Mus. 6: 20 (1961)
Distribution: Japan
38 JAP
Remarks: Athyrium kuratae x tozanense
species :
Athyrium ×quaesitum
Sa.Kurata; J. Geobot. 15: 5 (1966)
Distribution: Japan
38 JAP
Remarks: Athyrium otophorum x vidalii
species :
Athyrium ×sanjappae
Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 227 (2008)
Distribution: India (West Bengal)
40 IND
Remarks: Athyrium distans x foliolosum
species :
Athyrium ×satsumense
Seriz.; J. Phytogeogr. Taxon. 28(1): 34 (1980)
Distribution: Japan
38 JAP
Remarks: Athyrium kuratae x viridescentipes
species :
Athyrium ×sefuricola
Sa.Kurata; J. Geobot. 15: 6 (1966)
Distribution: Japan
38 JAP
Remarks: Athyrium kuratae (arisanense) x wardii
species :
Athyrium ×shiibaense
(Seriz.) Nakaike; New Fl. Japan, Pterid. 839 (1992)
Distribution: Japan
38 JAP
Remarks: Athyrium neglectum ssp. australe x vidalii
Synonyms: hidden
= Athyrium ×bicolor nothosubsp. shiibaense Seriz.; J. Phytogeogr. Taxon. 33(2): 77 (1985)
species :
Athyrium ×shikoku-montanum
Seriz.; J. Jap. Bot. 55(8): 255 (1980)
Distribution: Japan
38 JAP
Remarks: Athyrium reflexipinnum x vidalii
species :
Athyrium ×simlense
Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 231 (2008)
Distribution: India (Himachal Pradesh)
40 IND
Remarks: Athyrium setiferum x strigillosum
species :
Athyrium ×sledgei
Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 230 (2008)
Distribution: Sri Lanka
40 SRL
Remarks: Athyrium puncticaule x praetermissum
species :
Athyrium ×subchristensenianum
K.Hori; Fern Gaz. 21(6): 265 (2021)
Distribution: Japan (Honshu, Kyushu)
38 JAP
species :
Athyrium ×subcrassipes
Seriz.; J. Phytogeogr. Taxon. 33(2): 77 (1985)
Distribution: Japan
38 JAP
Remarks: Athyrium masamunei x tozanense (nigripes)
species :
Athyrium ×tokashikii
Sa.Kurata; J. Geobot. 12: 15 (1963)
Distribution: Japan
38 JAP
Remarks: Athyrium clivicola x wardii
species :
Athyrium ×tsurutanum
Sa.Kurata; J. Geobot. 13: 77 (1965)
Distribution: Japan
38 JAP
Remarks: Athyrium kenzo-satakei x otophorum
species :
Athyrium ×watanabei
Seriz.; J. Jap. Bot. 55(8): 252 (1980)
Distribution: Japan
38 JAP
Remarks: Athyrium deltoidofrons f. ohmurae x neglectum
species :
Athyrium ×yakuinsulare
Sa.Kurata; J. Geobot. 13: 77 (1965)
Distribution: Japan
38 JAP
Remarks: Athyrium kenzo-satakei x yakusimense
species :
Athyrium ×yakumonticola
Sa.Kurata; J. Geobot. 12: 15 (1963)
Distribution: Japan
38 JAP
Remarks: Athyrium iseanum x tozanense
