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Ver. 24.11
November 5th, 2024

046.0000 Family Athyriaceae Alston; Taxon 5: 25 (1956)
6-7 genera, ca. 760 spp. Circumscription following Rothfels et al. (2012). According to Fraser-Jenkins et al. (2018) it should be included in Woodsiaceae s. lat.
Species listed in database: 758; hybrids: 102
= Athyriinae Milde; Bot. Zeitung (Berlin) 28: 332 (1870)
= Athyrioideae B.K.Nayar; Taxon 19: 235 (1970)
= Physematieae J.Sm.; (1875)
= Deparioideae M.L.Wang & Y.T.Hsieh; Acta Phytotax. Sin. 42: 526 (2004)
= Diplazioideae Shmakov; Turczaninowia 4(1-2): 59 (2001), nom. superfl.
= Diplazioideae Ching & Z.R.Wang; Acta Phytotax. Sin. 35: 35 (1997)
= Diplazieae Pfeiff.; Nomencl. Bot. [Pfeiffer] 1(1): 628 (1873)
= Diplaziinae Payer; Bot. Crypt.: 198 (1850)
046.0002 Genus Anisocampium C.Presl; Abh. Königl. Böhm. Ges. Wiss., ser. 5, 6 [Epim. Bot.]: 58 (1851)
Distribution: E-Asia, S-Himalaya and Indochina, 1 to Sri Lanka and Malesia. Revision see Liu et al. (2011), systematics Wei et al. (2018). Fraser-Jenkins (2018) includes the genus in Athyrium s. lat.
Species listed in database: 4; hybrids: 1
= Kuniwatsukia Pic.Serm.; Webbia 28: 455 (1975)
= Microchlaena Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 322 (1938), non Wight & Arn. (1833)
species :
Anisocampium cumingianum C.Presl; Abh. Königl. Böhm. Ges. Wiss., ser. 5, 6 [Epim. Bot.]: 59 (1851)
Distribution: China (S-Yunnan); S-Taiwan; India (Karnataka, Kerala, ?Odisha, Tamil Nadu); Sri Lanka; Myanmar (Chin, etc.); Thailand; S-Laos; N-Cambodia; S-Sulawesi; Philippines (Luzon); E-Java; Lesser Sunda Isl. (Bali, Sumbawa, Timor)
36 CHI 38 TAI 40 IND SRL 41 CBD LAO MYA THA 42 JAW LSI PHI SUL
Conservation Status: very rare in Taiwan
Synonyms: hidden
= Anisocampium paucijugum (Alderw.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 151 (1947)
= Aspidium otaria Kunze ex Mett.; Abh. Senckenberg. Naturf. Ges. 2(2): [Pheg. u. Asp. 34, no. 73] (1858)
= Aspidium subpectinatum (Wall.) Christ; Bull. Acad. Int. Géogr. Bot. 248 (1902)
= Athyrium cumingianum (C.Presl) Ching; Index Filic., Suppl. Tert.: 40 (1934)
= Athyrium cumingianum (C.Presl) Milde; Bot. Zeitung (Berlin) 353 (1870), nomen
= Athyrium otaria (Kunze ex Mett.) Posth.; Proc. Fourth Pacif. Congress Java 137 (1929)
= Cyclodium cumingianum (C.Presl) T.Moore; Index Fil. [T. Moore] 83 (1857)
= Dryopteris otaria (Kunze ex Mett.) Kuntze; Revis. Gen. Pl. 2: 813 (1891)
= Dryopteris paucijuga (Alderw.) Rosenst.; Bull. Jard. Bot. Buitenzorg, sér. 2, 20: 17, t. 2 (1915)
= Goniopteris aristata Fée; Mém. Foug. 5 (Gen. Filic.): 253 (1852), nom. superfl.
= Goniopteris crenatodentata Fée; Mém. Foug. 5 (Gen. Filic.): 252 (1852)
= Goniopteris subpectinata (Wall.) J.Sm.; Hist. Fil. 192 (1875)
= Nephrodium aristatum (Fée) Hook.; Sp. Fil. [W. J. Hooker] 4: 62, t. 238 (1862)
= Nephrodium otaria (Kunze ex Mett.) Baker; Syn. Fil. [W. J. Hooker & Baker] 288 (1867)
= Nephrodium subpectinatum (Wall.) Bedd.; Suppl. Ferns Brit. Ind. 66 (1892)
= Oligocampia microcarpa Trevis.; Atti Reale Ist. Veneto Sci. Lett. Arti, ser. 2, 2: 165 (1851)
= Phegopteris paucijuga Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 16: 26 (1914)
= Pleocnemia aristata (Fée) Bedd.; Ferns S. India t. 83 (1863)
= Polypodium subpectinatum Wall.; Numer. List. [Wallich] no. 311 (1829), nomen
= Thelypteris aristata (Fée) C.F.Reed; Phytologia 17: 261 (1968)
species :
Anisocampium cuspidatum (Mett. ex Bedd.) Yea C.Liu, W.L.Chiou & M.Kato; Taxon 60(3): 829 (2011)
Distribution: China (SE-Yunnan, Guizhou, Guangxi); Tibet; Nepal; India (Arunachal Pradesh, Manipur, Meghalaya, Mizoram, Nagaland, Sikkim, Uttarakhand); E-Myanmar; N-Thailand
36 CHI TIB 40 IND NEP 41 MYA THA
Synonyms: hidden
= Aspidium cuspidatum Mett.; Abh. Senckenberg. Naturf. Ges. 2(2) [Pheg. u. Asp.]: 376 (1858), non Desv. 1827, nomen
= Aspidium yunnanense Christ; Bull. Herb. Boissier, sér. 1, 6: 965 (1898) (fide Fraser-Jenkins 1997)
= Athyrium cuspidatum (Mett. ex Bedd.) M.Kato; Bot. Mag. (Tokyo) 90(1017): 27 (1977)
= Christella cuspidata (Mett. ex Bedd.) H.Lév.; Fl. Kouy-Tcheou 474-476 (1915)
= Dryopteris cuspidata (Mett. ex Bedd.) Christ; Philipp. J. Sci. 2: 205 (1907)
= Dryopteris elongata (Wall. ex Hook.) Kuntze; Revis. Gen. Pl. 2: 811 (1891)
= Dryopteris khasiana C.Chr.; Index Filic. 272 (1905), nom. nov. for Polypodium elongatum Wall. ex Hook., nom. superfl.
= Dryopteris trinidadensis (Jenman) C.Chr.; Index Filic. 298 (1905) (type mislabeled)
= Dryopteris yunnanensis (Christ) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 122 (1947) (fide Fraser-Jenkins 1997)
= Kuniwatsukia cuspidata (Bedd.) Pic.Serm.; Webbia 28: 455 (1973)
= Lastraea cuspidata Mett. ex Bedd.; Ferns Brit. India 2: 111, IV, et correct. 348 (1870)
= Lastrea elongata (Wall. ex Hook.) Bedd. ex C.B.Clarke; Trans. Linn. Soc. London, Bot., ser. 2, 1: 513 (1880) non C. Presl
= Microchlaena cuspidata (Mett. ex Bedd.) Ching; Acta Phytotax. Sin. 9: 99 (1964)
= Nephrodium cuspidatum (Mett. ex Bedd.) Baker; Syn. Fil. [W. J. Hooker & Baker] 260 (1867), based on Aspidium cuspidatum Mett., non C. Presl 1825
= Phegopteris elongata (Wall. ex Hook.) J.Sm.; Hist. Fil. 233 (1875), non E. Fourn. (1872)
= Polypodium acuminatum Roxb. ex D.Don; Prodr. Fl. Nepal. 2 (1824), non Houtt. 1783, nec Roxb. 1844 et al.
= Polypodium elongatum Wall.; Numer. List. [Wallich] no. 309 (1829), nomen
= Polypodium elongatum Wall. ex Hook.; Sp. Fil. [W. J. Hooker] 4: 234 (1862), non Aiton (1789)
= Polypodium trinidadense Jenman; Gard. Chron., ser. 3, 18: 235 (1895) (type mislabeled)
species :
Anisocampium niponicum (Mett.) Yea C.Liu, W.L.Chiou & M.Kato; Taxon 60(3): 828 (2011)
Distribution: China (Anhui, Chongqing, Gansu, Guangdong, Guangxi, Guizhou, Hebei, Henan, Hubei, Hunan, Jiangsu, Jiangxi, Liaoning, Ningxia, Shaanxi, Shandong, Shanxi, Sichuan, Yunnan, Zhejiang); North Korea; South Korea (incl. Cheju Isl.); Japan (widespread); Taiwan; NE-India (Arunachal Pradesh, Manipur, Tripura, Mizoram, Nagaland); Myanmar; N-Vietnam; USA [I] (Alabama [I])
36 CHI 38 JAP KOR NKO TAI 40 IND 41 MYA VIE 78 usa
Conservation Status: IUCN VU for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Asplenium brevisorum Wall.; Numer. List. [Wallich] no. 220 (1829), nomen
= Asplenium brevisorum Wall. ex Hook.; Sp. Fil. [W. J. Hooker] 3: 229 (1860), nom. illeg. (fide Fraser-Jenkins et al. 2015)
= Asplenium niponicum Mett.; Ann. Mus. Bot. Lugduno-Batavi 2: 240 (1865)
= Asplenium niponicum var. longipes Franch. & Sav.
= Asplenium niponicum var. minus Franch. & Sav.
= Asplenium niponicum var. uropteron (Miq.) Franch. & Sav.; Enum. Pl. Jap. 2(1): 224 (1877)
= Asplenium uropteron Miq.; Ann. Mus. Bot. Lugduno-Batavi 8: 174 (1867)
= Athyrium anhweiense Ching & P.C.Chiu; Obs. Fl. Hwangshan. 63 (1965), nom. nud.
= Athyrium apiciflorum Ching; Obs. Fl. Hwangshan. 63 (1965), nom. nud.
= Athyrium biondii Christ; Nuovo Giorn. Bot. Ital., n. s. 4: 91 (1897)
= Athyrium brevisorum (Wall. ex Hook.) T.Moore; Index Fil. [T. Moore] 117 (1859) (fide Fraser-Jenkins et al. 2015)
= Athyrium chekiangense Ching; Y. L. Chang, Sporae Pterid. Sin. (1976), nom. nud.
= Athyrium concinnum Nakai; Bot. Mag. (Tokyo) 45: 92 (1931)
= Athyrium fissum Christ; Notul. Syst. (Paris) 1: 47 (1909)
= Athyrium goeringianum var. pictum Maxwell; Gard. Chron. 1862, II: 25 (1862)
= Athyrium matsumurae Christ ex Matsum.; Bot. Mag. (Tokyo) 24: 241 (1910)
= Athyrium niponicum (Mett) Hance; J. Linn. Soc., Bot. 13: 92 (1873)
= Athyrium niponicum f. cristato-flabellatum (Makino) Nameg. & Sa.Kurata; Enum. Jap. Pterid. 279, 338 (1961)
= Athyrium niponicum f. majus C.Chr.; Index Filic., Suppl. Tert.: 45 (1934), nom. nud.
= Athyrium niponicum var. cristato-flabellatum Makino; (1916)
= Athyrium niponicum var. pachyphlebium (C.Chr.) Kitag.; Neo-Lineam. Fl. Manshur.: 31 (1979)
= Athyrium niponicum var. pictum (Maxwell) Fraser-Jenk.; Fern Gaz. 12(4): 247 (1982)
= Athyrium pachyphlebium C.Chr.; Dansk Bot. Ark. 9(3): 55 (1937)
= Athyrium silvestrii Christ; Nuovo Giorn. Bot. Ital., n. s. 17: 226 (1910)
= Athyrium uropteron (Miq.) C.Chr.; Index Filic. 147 (1905)
= Athyrium yunnanense Christ; Bull. Acad. Int. Géogr. Bot. 134 (1907)
= Microchlaena yunnanensis (Christ) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 325 (1938)
species :
Anisocampium sheareri (Baker) Ching; Ill. Handb. Chin. Pl. Pterid. 111, f. 147 (1957)
Distribution: China (Anhui, Chongqing, Fujian, Gansu, Guangdong, Guangxi, Guizhou, Hubei, Hunan, Jiangsu, Jiangxi, Sichuan, Yunnan, Zhejiang); South Korea (incl. Cheju Isl.); Japan; Taiwan (very rare)
36 CHI 38 JAP KOR TAI
Remarks: very rare in Taiwan and not seen since 2001 (Knapp 2011 ff.)
Synonyms: hidden
= Aspidium otarioides Christ; Bull. Acad. Int. Géogr. Bot. 247 (1902)
= Aspidium polypodiforme (Makino) Makino; Bot. Mag. (Tokyo) 6(60): 46 (1892)
= Athyrium sheareri (Baker) Ching; C. Chr., Index Filic., Suppl. Tert.: 44 (1934)
= Dryopteris otarioides (Christ) C.Chr.; Index Filic. 282 (1905)
= Dryopteris polypodiformis (Makino) C.Chr.; Index Filic. 285 (1905)
= Dryopteris sheareri (Baker) C.Chr.; Index Filic. 292 (1905)
= Dryopteris subsagenioides Christ; Bull. Acad. Int. Géogr. Bot. 8 (1910)
= Nephrodium isolatum Baker; Gard. Chron., ser. 2, 14: 494 (1880)
= Nephrodium polypodiforme Makino; Bot. Mag. (Tokyo) 13: 58 (1891)
= Nephrodium sheareri Baker; J. Bot. 13: 200 (1875)
species :
Anisocampium ×saitoanum (Sugim.) M.Kato; Taxon 60(3): 829 (2011)
Distribution: Japan
38 JAP
Remarks: Athyrium niponicum x sheareri
Synonyms: hidden
= Athyrium sheareri f. saitoanum Sugim.; Keys Herb. Pl. Jap. Pterid. 404 (1966)
= Athyrium ×saitoanum (Sugim.) Seriz.; J. Phytogeogr. Taxon. 34(2): 68 (1986)
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A complete, synonymic checklist of the Higher Plants of the World


Version 24.11 (November 5th, 2024)

Covering 355,576 accepted species,
8,715 hybrids, 47,863 accepted infraspecific taxa and 1,000,581 synonyms,
in total 1,412,785* names

[*about 25,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus" are deliberately omitted]


With complete, detailed distribution data (countries and regions) and literature references

Arranged in linear phylogenetic sequence reflecting true relationships

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With 38,209 pictures of 15,787 species

Assembled through 40 years independently from other databases

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